646 lines
106 KiB
XML
646 lines
106 KiB
XML
<document id="2EDCE43D827333FA39BF20F26FE1FE34" ID-DOI="10.5281/zenodo.13620799" ID-ISSN="1732-2421" ID-Zenodo-Dep="13620799" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1723143778560" checkinUser="felipe" docAuthor="Jouve, Stéphane, Iarochène, Mohamed, Bouya, Baâdi & Amaghzaz, Mbarek" docDate="2005" docId="711F661DFF90FF9FFF90FE61FEFDDE21" docLanguage="en" docName="ActaPalaeontolPol.50.3.581-594.pdf" docOrigin="Acta Palaeontologica Polonica 50 (3)" docStyle="DocumentStyle:93F5927C23D90727DE855278A6F17B37.1:ActaPalaeontolPol.2002-2009.journal_article" docStyleId="93F5927C23D90727DE855278A6F17B37" docStyleName="ActaPalaeontolPol.2002-2009.journal_article" docStyleVersion="1" docTitle="Arambourgisuchus khouribgaensis Jouve, Iarochène, Bouya & Amaghzaz, 2005, sp. nov." docType="treatment" docVersion="3" lastPageNumber="589" masterDocId="8D261E65FF92FF97FFF6FFBDFFE8DD30" masterDocTitle="A new dyrosaurid crocodyliform from the Palaeocene of Morocco and a phylogenetic analysis of Dyrosauridae" masterLastPageNumber="594" masterPageNumber="581" pageNumber="583" updateTime="1725043448023" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B659C2BB486B2B3BA0DBC193BC043177">A new dyrosaurid crocodyliform from the Palaeocene of Morocco and a phylogenetic analysis of Dyrosauridae</mods:title>
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<mods:namePart id="F38D3C1BBD429B70276D8B249C2CDAAB">Jouve, Stéphane</mods:namePart>
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<mods:namePart id="028696C438431E1425A241AE4C82584A">Iarochène, Mohamed</mods:namePart>
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<mods:namePart id="21846E3F1200EC0FD4CB162D69D19D82">Bouya, Baâdi</mods:namePart>
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<mods:roleTerm id="BFE04FDD736D64F7C733DAE9722E6540">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="076E8DE503DE9426E6D46FA1A1067EE2">Amaghzaz, Mbarek</mods:namePart>
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<mods:typeOfResource id="2F2D24BA3EA204B205691398DC62AAEB">text</mods:typeOfResource>
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<mods:title id="7500AF2DA997EB4FB89B03FA86D80E4F">Acta Palaeontologica Polonica</mods:title>
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<mods:part id="C7998438416A90F744C9373AA0D8ACA0">
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<mods:date id="97D7F97AE152B3D4ABEF51A2F862013C">2005</mods:date>
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<mods:number id="46FC24F5EBBD320F153AAE04E8667C03">50</mods:number>
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<mods:number id="EA2D3E0FC83FBDCF5090DF74A10F2E42">3</mods:number>
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<mods:classification id="9065317E916CDC742C0559CB8B9690B2">journal article</mods:classification>
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<mods:identifier id="5DF54A82E3ADF0528EB7337D0020B71A" type="DOI">10.5281/zenodo.13620799</mods:identifier>
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<mods:identifier id="0B95D73645DBFD464D9D950AA014339F" type="ISSN">1732-2421</mods:identifier>
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<mods:identifier id="D1B5E72D44F2FFC71E67B7F966A89DA6" type="Zenodo-Dep">13620799</mods:identifier>
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<treatment id="711F661DFF90FF9FFF90FE61FEFDDE21" LSID="urn:lsid:plazi:treatment:711F661DFF90FF9FFF90FE61FEFDDE21" httpUri="http://treatment.plazi.org/id/711F661DFF90FF9FFF90FE61FEFDDE21" lastPageId="8" lastPageNumber="589" pageId="2" pageNumber="583">
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<subSubSection id="B1AC8480FF90FF95FF90FE61FD79DCC7" box="[102,657,475,504]" pageId="2" pageNumber="583" type="nomenclature">
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<paragraph id="F909D70BFF90FF95FF90FE61FD79DCC7" blockId="2.[102,775,475,862]" box="[102,657,475,504]" pageId="2" pageNumber="583">
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<heading id="A2416067FF90FF95FF90FE61FD79DCC7" box="[102,657,475,504]" fontSize="12" level="2" pageId="2" pageNumber="583" reason="9">
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<taxonomicName id="3EB6AC88FF90FF95FF90FE61FDCCDCC8" authorityName="Jouve & Iarochène & Bouya & Amaghzaz" authorityYear="2005" box="[102,548,476,504]" class="Reptilia" family="Dyrosauridae" genus="Arambourgisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="2" pageNumber="583" phylum="Chordata" rank="species" species="khouribgaensis" status="sp. nov.">
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<emphasis id="CBC20B19FF90FF95FF90FE61FDCCDCC8" box="[102,548,476,504]" italics="true" pageId="2" pageNumber="583">Arambourgisuchus khouribgaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D0F1B662FF90FF95FDDBFE66FD79DCC7" box="[557,657,475,503]" pageId="2" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B1AC8480FF90FF95FF90FDB8FAB0D93A" pageId="2" pageNumber="583" type="etymology">
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<paragraph id="F909D70BFF90FF95FF90FDB8FF5DDF06" blockId="2.[102,775,475,862]" pageId="2" pageNumber="583">
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<emphasis id="CBC20B19FF90FF95FF90FDB8FF25DF2A" box="[102,205,517,538]" italics="true" pageId="2" pageNumber="583">Etymology</emphasis>
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: Khouribga (place name), referring to the city near the
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<typeStatus id="260D69A9FF90FF95FD2BFDB8FCEEDF2A" box="[733,774,517,538]" pageId="2" pageNumber="583">type</typeStatus>
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locality.
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</paragraph>
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<paragraph id="F909D70BFF90FF95FF90FDF8FF29DE32" blockId="2.[102,775,475,862]" pageId="2" pageNumber="583">
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<materialsCitation id="49DEDD56FF90FF95FF90FDF8FF29DE32" collectionCode="OCP" pageId="2" pageNumber="583" specimenCode="DEK-GE 300" specimenCount="2" typeStatus="holotype">
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<emphasis id="CBC20B19FF90FF95FF90FDF8FF56DF6A" box="[102,190,581,602]" italics="true" pageId="2" pageNumber="583">
|
||
<typeStatus id="260D69A9FF90FF95FF90FDF8FF56DF6A" box="[102,190,581,602]" pageId="2" pageNumber="583" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
:
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<collectionCode id="9FA74FCEFF90FF95FF3FFDF8FF10DF6A" box="[201,248,581,602]" pageId="2" pageNumber="583">OCP</collectionCode>
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<specimenCode id="A9107F70FF90FF95FF0BFDF8FE97DF6A" box="[253,383,581,602]" collectionCode="DEK-GE" pageId="2" pageNumber="583">DEK−GE 300</specimenCode>
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(
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<figureCitation id="618DCB8EFF90FF95FE7DFDF8FE27DF6A" box="[395,463,581,602]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="2" pageNumber="583">Figs. 2</figureCitation>
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,
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<figureCitation id="618DCB8EFF90FF95FE2EFDF8FE0CDF6A" box="[472,484,581,602]" captionStart="Fig" captionStartId="5.[102,135,548,568]" captionTargetBox="[108,1485,235,515]" captionTargetId="figure-793@5.[102,1382,226,519]" captionTargetPageId="5" captionText="Fig. 5. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, detail of the right posttemporal fenestra in posterior view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620807" httpUri="https://zenodo.org/record/13620807/files/figure.png" pageId="2" pageNumber="583">5</figureCitation>
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–7), a nearly complete crushed skull, lacking the anterior part of the rostrum. Originally preserved in a phosphatic block, it has been mechanically prepared on both sides. It is very crushed and dorsoventrally flattened from the anterior level of the palatine to its posterior end. Some sutures are hardly visible, but almost all the bones can be reconstituted. All the measurements are taken for the
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<typeStatus id="260D69A9FF90FF95FF90FD50FF55DE32" box="[102,189,749,770]" pageId="2" pageNumber="583" type="holotype">holotype</typeStatus>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="F909D70BFF90FF95FF90FCACFEFBDE6E" blockId="2.[102,775,475,862]" pageId="2" pageNumber="583">
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<emphasis id="CBC20B19FF90FF95FF90FCACFE8CDE16" box="[102,356,785,806]" italics="true" pageId="2" pageNumber="583">Type locality and horizon</emphasis>
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: The phosphate mine of Sidi Chenane, in the Ouled Abdoun Basin,
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<collectingCountry id="81A1979BFF90FF95FEAEFC90FE5ADE72" box="[344,434,813,834]" name="Morocco" pageId="2" pageNumber="583">Morocco</collectingCountry>
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; from the “couche (bed) 2a”, Thanetian (Palaeocene).
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</paragraph>
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<paragraph id="F909D70BFF90FF95FF90FCCEFAE7DE71" blockId="2.[102,774,881,1098]" lastBlockId="2.[812,1485,232,833]" pageId="2" pageNumber="583">
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<emphasis id="CBC20B19FF90FF95FF90FCCEFF30DEBA" box="[102,216,883,906]" italics="true" pageId="2" pageNumber="583">Diagnosis</emphasis>
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.—Dyrosaurid with about 20 to 21 robust but sharp teeth on the upper tooth row (on each ramus); teeth moderately elongated, with a posterior carina which ends before reaching the base of the teeth (not in the more posterior teeth), buccal and lingual surface smooth or lightly striated; interfenestral bar very narrow (sagittal crest); supraoccipital, with parietal, tapers posteriorly between the two occipital tuberosities; occipital tuberosities well developed, and dorsoventrally flattened; suture between basioccipital and exoccipital (posteriorly to basioccipital tuberosities) deeply within a groove; mandibular symphysis long, wider than high in its median part, and ending posteriorly at the level of the sixteenth tooth; splenials end dorsally between the level of the tenth and the eleventh tooth. Differs from
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<taxonomicName id="3EB6AC88FF90FF95FABEFE14FA24DCF0" authorityName="Pomel" authorityYear="1894" box="[1352,1484,425,448]" class="Reptilia" family="Dyrosauridae" genus="Dyrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="2" pageNumber="583" phylum="Chordata" rank="genus">
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<emphasis id="CBC20B19FF90FF95FABEFE14FA24DCF0" box="[1352,1484,425,448]" italics="true" pageId="2" pageNumber="583">Dyrosaurus</emphasis>
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</taxonomicName>
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in having less numerous, and more massive teeth, a narrower interfenestral bar, a posterior wall of the supratemporal fenestra very inclined dorsally, the occipital tuberosities dorsoventrally flattened, and the supraoccipital which tapers posteriorly; from
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<taxonomicName id="3EB6AC88FF90FF95FC0BFDF5FB7ADF50" authorityName="Dollo" authorityYear="1914" box="[1021,1170,584,608]" class="Reptilia" family="Dyrosauridae" genus="Congosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="2" pageNumber="583" phylum="Chordata" rank="genus">
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<emphasis id="CBC20B19FF90FF95FC0BFDF5FB7ADF50" box="[1021,1170,584,608]" italics="true" pageId="2" pageNumber="583">Congosaurus</emphasis>
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</taxonomicName>
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in having a longer snout, more massive and more widely separated teeth; from
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<taxonomicName id="3EB6AC88FF90FF95FA71FDD4FC9FDF90" baseAuthorityName="Jouve and Schwarz" baseAuthorityYear="2004" class="Reptilia" family="Dyrosauridae" genus="Hyposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="2" pageNumber="583" phylum="Chordata" rank="genus">
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<emphasis id="CBC20B19FF90FF95FA71FDD4FC9FDF90" italics="true" pageId="2" pageNumber="583">Hyposaurus</emphasis>
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</taxonomicName>
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in having a longer snout, and more massive teeth, more ventrally projected basioccipital tuberosities, and the supraoccipital which tapers posteriorly; from
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<taxonomicName id="3EB6AC88FF90FF95FAB4FD75FCB3DE30" authorityName="Swinton" authorityYear="1930" class="Reptilia" family="Dyrosauridae" genus="Rhabdognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="2" pageNumber="583" phylum="Chordata" rank="genus">
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<emphasis id="CBC20B19FF90FF95FAB4FD75FCB3DE30" italics="true" pageId="2" pageNumber="583">Rhabdognathus</emphasis>
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</taxonomicName>
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in having more ventrally projected basioccipital tuberosities, the occipital tuberosities dorsoventrally flattened, and the supraoccipital which tapers posteriorly.
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</paragraph>
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<paragraph id="F909D70BFF90FF95FCDAFCECFAA2DE59" blockId="2.[812,1485,849,1099]" box="[812,1354,849,874]" pageId="2" pageNumber="583">
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<emphasis id="CBC20B19FF90FF95FCDAFCECFC66DE59" box="[812,910,849,873]" italics="true" pageId="2" pageNumber="583">Material</emphasis>
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.—In addition to the
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<typeStatus id="260D69A9FF90FF95FB8EFCECFB32DE5A" box="[1144,1242,849,874]" pageId="2" pageNumber="583" type="holotype">holotype</typeStatus>
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there are:
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</paragraph>
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<paragraph id="F909D70BFF90FF95FCA4FCCCFAB0D93A" blockId="2.[812,1485,849,1099]" pageId="2" pageNumber="583">
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OCP DEK−GE 18 (
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<figureCitation id="618DCB8EFF90FF95FBCEFCCCFB68DEBA" box="[1080,1152,881,906]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="2" pageNumber="583">Fig. 3</figureCitation>
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). An almost complete skull, with mandible. The skull is crushed, and the occipital part is strongly damaged. In the left ramus of the mandible the posterior part is missing a little behind the end of the mandibular symphysis; the right mandible is badly preserved.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="B1AC8480FF90FF9FFCA4FBAFFEFDDE21" lastPageId="8" lastPageNumber="589" pageId="2" pageNumber="583" type="description">
|
||
<paragraph id="F909D70BFF90FF95FCA4FBAFFA68D97B" blockId="2.[812,1485,849,1099]" pageId="2" pageNumber="583">OCP DEK−GE 269. A posterior part of a mandibular symphysis with five or six tooth alveoli on each side.</paragraph>
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<paragraph id="F909D70BFF90FF95FD1DFAD4FCA8D84F" box="[747,832,1385,1407]" pageId="2" pageNumber="583">
|
||
<quantity id="3E4E7AEEFF90FF95FD1DFAD4FCA8D84F" box="[747,832,1385,1407]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="583" unit="mm" value="100.0">100 mm</quantity>
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||
</paragraph>
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||
<caption id="ADC98783FF90FF95FF90F805FF49DAD8" pageId="2" pageNumber="583" startId="2.[102,135,1976,1996]" targetType="figure">
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<paragraph id="F909D70BFF90FF95FF90F805FF49DAD8" blockId="2.[102,1486,1976,2024]" pageId="2" pageNumber="583">
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Fig. 2.
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<taxonomicName id="3EB6AC88FF90FF95FF52F805FE37DAFC" authorityName="Jouve & Iarochène & Bouya & Amaghzaz" authorityYear="2005" box="[164,479,1976,1996]" class="Reptilia" family="Dyrosauridae" genus="Arambourgisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="2" pageNumber="583" phylum="Chordata" rank="species" species="khouribgaensis" status="gen. et sp. nov.">
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||
<emphasis id="CBC20B19FF90FF95FF52F805FE37DAFC" box="[164,479,1976,1996]" italics="true" pageId="2" pageNumber="583">Arambourgisuchus khouribgaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D0F1B662FF90FF95FE12F804FD85DAFC" box="[484,621,1977,1996]" pageId="2" pageNumber="583" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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, OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (
|
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<emphasis id="CBC20B19FF90FF95FADCF805FAD2DAFC" bold="true" box="[1322,1338,1976,1996]" pageId="2" pageNumber="583">A</emphasis>
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||
) and ventral (
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||
<emphasis id="CBC20B19FF90FF95FA40F804FA2DDAFC" bold="true" box="[1462,1477,1977,1996]" pageId="2" pageNumber="583">B</emphasis>
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) views.
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</paragraph>
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</caption>
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<paragraph id="F909D70BFF91FF94FF2FFB82FBACD963" blockId="3.[217,663,1087,1108]" box="[217,1092,1087,1108]" lastBlockId="3.[875,1092,1087,1108]" pageId="3" pageNumber="584">quadratojugal postorbital jugal orbit tooth splenial</paragraph>
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<caption id="ADC98783FF91FF94FF90FBD3FD9AD9AE" ID-DOI="http://doi.org/10.5281/zenodo.13620803" ID-Zenodo-Dep="13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="3" pageNumber="584" startId="3.[102,135,1134,1154]" targetBox="[200,1388,233,1080]" targetPageId="3" targetType="figure">
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||
<paragraph id="F909D70BFF91FF94FF90FBD3FD9AD9AE" blockId="3.[102,1486,1134,1182]" pageId="3" pageNumber="584">
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Fig. 3.
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<taxonomicName id="3EB6AC88FF91FF94FF53FBD3FE08D9B2" authorityName="Jouve & Iarochène & Bouya & Amaghzaz" authorityYear="2005" box="[165,480,1134,1154]" class="Reptilia" family="Dyrosauridae" genus="Arambourgisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="3" pageNumber="584" phylum="Chordata" rank="species" species="khouribgaensis" status="gen. et sp. nov.">
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||
<emphasis id="CBC20B19FF91FF94FF53FBD3FE08D9B2" box="[165,480,1134,1154]" italics="true" pageId="3" pageNumber="584">Arambourgisuchus khouribgaensis</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="D0F1B662FF91FF94FE13FBD2FD87D9B2" box="[485,623,1135,1154]" pageId="3" pageNumber="584" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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, OCP DEK−GE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (
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<emphasis id="CBC20B19FF91FF94FF2DFB37FF03D9AE" bold="true" box="[219,235,1162,1182]" pageId="3" pageNumber="584">A</emphasis>
|
||
) and explanatory drawing of the same (
|
||
<emphasis id="CBC20B19FF91FF94FDA0FB36FD8DD9AE" bold="true" box="[598,613,1163,1182]" pageId="3" pageNumber="584">B</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F909D70BFF91FF94FF7AFB6DFDA3D839" blockId="3.[102,775,1232,1289]" pageId="3" pageNumber="584">OCP DEK−GE 1200. The anteriormost portion of a mandibular symphysis exposed in ventral view.</paragraph>
|
||
<paragraph id="F909D70BFF91FF94FF90FA9EFF18D80C" blockId="3.[102,240,1315,1340]" box="[102,240,1315,1340]" pageId="3" pageNumber="584">
|
||
<emphasis id="CBC20B19FF91FF94FF90FA9EFF18D80C" box="[102,240,1315,1340]" italics="true" pageId="3" pageNumber="584">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF91FF94FF90FAF2FEC2D898" blockId="3.[102,775,1359,2026]" pageId="3" pageNumber="584">
|
||
<emphasis id="CBC20B19FF91FF94FF90FAF2FEC7D857" box="[102,303,1359,1383]" italics="true" pageId="3" pageNumber="584">Cranial openings.</emphasis>
|
||
—The external nares are poorly preserved. They seem large, dorsally directed, and surrounded by the premaxillae only.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF91FF94FF7AFA0DFE7FDB39" blockId="3.[102,775,1359,2026]" pageId="3" pageNumber="584">The orbits are rounded, dorsolaterally oriented, bordered posterolaterally by a prominent anterolateral postorbital process typical of dyrosaurids.</paragraph>
|
||
<paragraph id="F909D70BFF91FF94FF7AF9ACFCECDBDA" blockId="3.[102,775,1359,2026]" pageId="3" pageNumber="584">
|
||
The supratemporal fenestrae are longer (about
|
||
<quantity id="3E4E7AEEFF91FF94FD73F9ACFD22DB1A" box="[645,714,1553,1578]" metricMagnitude="-1" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="584" unit="cm" value="17.0">17 cm</quantity>
|
||
) than wide (about
|
||
<quantity id="3E4E7AEEFF91FF94FF0AF98CFED4DB7A" box="[252,316,1585,1610]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="584" unit="cm" value="6.0">6 cm</quantity>
|
||
), separated medially by a very narrow interfenestral bar constituted by the frontal (for about one fifth) and in a major part by the parietal (
|
||
<figureCitation id="618DCB8EFF91FF94FDC4F9CCFD7DDBBA" box="[562,661,1649,1674]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="3" pageNumber="584">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF91FF94FD54F9CCFD20DBBA" box="[674,712,1649,1674]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="3" pageNumber="584">4A</figureCitation>
|
||
). The fenestra appears to be bordered laterally more by the postorbital than by the squamosal. The posterior margin is constituted for about the same proportion by squamosal and parietal.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF91FF94FF7AF94CFD78DADA" blockId="3.[102,775,1359,2026]" pageId="3" pageNumber="584">
|
||
<materialsCitation id="49DEDD56FF91FF94FF7AF94CFD64DADA" collectionCode="OCP" pageId="3" pageNumber="584" specimenCode="DEK-GE 300" specimenCount="1" typeStatus="holotype">
|
||
The infratemporal fenestra is not preserved, but can be reconstructed after the
|
||
<typeStatus id="260D69A9FF91FF94FEBBF8ACFE47DA1A" box="[333,431,1809,1834]" pageId="3" pageNumber="584" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="9FA74FCEFF91FF94FE4BF8ACFE1DDA1A" box="[445,501,1809,1834]" pageId="3" pageNumber="584">OCP</collectionCode>
|
||
<specimenCode id="A9107F70FF91FF94FE0DF8ACFD7EDA1A" box="[507,662,1809,1834]" collectionCode="DEK-GE" pageId="3" pageNumber="584">DEK−GE 300</specimenCode>
|
||
;
|
||
<figureCitation id="618DCB8EFF91FF94FD56F8ACFCE9DA1A" box="[672,769,1809,1834]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="3" pageNumber="584">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF91FF94FF90F88CFF64DA7A" box="[102,140,1841,1866]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="3" pageNumber="584">4A</figureCitation>
|
||
). It is anteriorly limited by the postorbital bar, constituted half by the postorbital and jugal, this latter constituting the major part of the ventral margin. The quadratojugal borders the posterior margin, constitutes the posterior part of the dorsal edge, excluding the quadrate from the dorsal margin. It participates for a small length to the ventral edge
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF91FF94FCA4FB6DFC05D81B" blockId="3.[812,1486,1232,2026]" pageId="3" pageNumber="584">The temporal canal, wider than high, is largely surrounded by the parietal (two third), with the squamosal contributing to its lateral margin.</paragraph>
|
||
<paragraph id="F909D70BFF91FF94FCA4FA88FAC0D8E2" blockId="3.[812,1486,1232,2026]" pageId="3" pageNumber="584">
|
||
The suborbital fenestra is formed medially by the palatine and the pterygoid, posterolaterally by the ectopterygoid, and anterolaterally by the maxilla (
|
||
<figureCitation id="618DCB8EFF91FF94FB89FAC5FB08D8A1" box="[1151,1248,1400,1425]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="3" pageNumber="584">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF91FF94FB1CFAC5FAE6D8A1" box="[1258,1294,1400,1425]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="3" pageNumber="584">4B</figureCitation>
|
||
). The medial and lateral edges are curved, and the most posterior part is acute and bears the ectopterygoid−pterygoid suture.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF91FF94FCABFA60FABADA30" blockId="3.[812,1486,1232,2026]" pageId="3" pageNumber="584">
|
||
The choanae are deep, ventrally oriented (slightly caudally), and separated by a pterygoidian septum (
|
||
<figureCitation id="618DCB8EFF91FF94FAC2FA43FA7ADB27" box="[1332,1426,1534,1559]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="3" pageNumber="584">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF91FF94FA6BFA43FA29DB27" box="[1437,1473,1534,1559]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="3" pageNumber="584">4B</figureCitation>
|
||
). They are largely surrounded by the pterygoid, only the most anterior border being formed by the palatine. The choanae are not very abruptly pierced within the pterygoids: the choanae lie within a depression on the ventral pterygoidal surface. The depression extends posterolaterally on to the lateral branch of the pterygoid, tapering before the pterygoid contacts the ectopterygoid. Thus, the choanal margin is not abrupt.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF91FF93FCA4F8B7FE27D9E7" blockId="3.[812,1486,1232,2026]" lastBlockId="4.[102,774,1182,1239]" lastPageId="4" lastPageNumber="585" pageId="3" pageNumber="584">
|
||
On the occipital face, the parietal forms the dorsomedial quarter of the posttemporal fenestra (
|
||
<figureCitation id="618DCB8EFF91FF94FB3DF896FAE7DA74" box="[1227,1295,1835,1860]" captionStart="Fig" captionStartId="5.[102,135,548,568]" captionTargetBox="[108,1485,235,515]" captionTargetId="figure-793@5.[102,1382,226,519]" captionTargetPageId="5" captionText="Fig. 5. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, detail of the right posttemporal fenestra in posterior view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620807" httpUri="https://zenodo.org/record/13620807/files/figure.png" pageId="3" pageNumber="584">Fig. 5</figureCitation>
|
||
). The squamosal contributes to it dorsolaterally (less than the lateral half), and very slightly to the ventrolateral margin. The supraoccipital constitutes a small part of the ventral margin of the posttemporal fenestra (the rest formed by exoccipital), but constitutes the major part of this same margin to contact the squamosal more deeply within the posttemporal fenestra (excluding the exoccipital) (
|
||
<figureCitation id="618DCB8EFF96FF93FE8BFB03FE29D9E7" box="[381,449,1214,1239]" captionStart="Fig" captionStartId="5.[102,135,548,568]" captionTargetBox="[108,1485,235,515]" captionTargetId="figure-793@5.[102,1382,226,519]" captionTargetPageId="5" captionText="Fig. 5. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, detail of the right posttemporal fenestra in posterior view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620807" httpUri="https://zenodo.org/record/13620807/files/figure.png" pageId="4" pageNumber="585">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="ADC98783FF96FF93FF90FBE0FB70D941" ID-DOI="http://doi.org/10.5281/zenodo.13620805" ID-Zenodo-Dep="13620805" box="[102,1176,1116,1137]" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="4" pageNumber="585" startId="4.[102,135,1117,1137]" targetBox="[171,1417,225,1090]" targetPageId="4" targetType="figure">
|
||
<paragraph id="F909D70BFF96FF93FF90FBE0FB70D941" blockId="4.[102,1176,1116,1137]" box="[102,1176,1116,1137]" pageId="4" pageNumber="585">
|
||
Fig. 4.
|
||
<taxonomicName id="3EB6AC88FF96FF93FF5AFBE0FE00D941" authorityName="Jouve & Iarochène & Bouya & Amaghzaz" authorityYear="2005" box="[172,488,1117,1137]" class="Reptilia" family="Dyrosauridae" genus="Arambourgisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="4" pageNumber="585" phylum="Chordata" rank="species" species="khouribgaensis" status="gen. et sp. nov.">
|
||
<emphasis id="CBC20B19FF96FF93FF5AFBE0FE00D941" box="[172,488,1117,1137]" italics="true" pageId="4" pageNumber="585">Arambourgisuchus khouribgaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D0F1B662FF96FF93FE18FBE3FD92D941" box="[494,634,1118,1137]" pageId="4" pageNumber="585" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, reconstruction of skull in dorsal (
|
||
<emphasis id="CBC20B19FF96FF93FC46FBE1FC28D940" bold="true" box="[944,960,1116,1136]" pageId="4" pageNumber="585">A</emphasis>
|
||
) and ventral (
|
||
<emphasis id="CBC20B19FF96FF93FBB6FBE0FBA7D940" bold="true" box="[1088,1103,1117,1136]" pageId="4" pageNumber="585">B</emphasis>
|
||
) views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F909D70BFF96FF93FF90FB5AFE33DBEF" blockId="4.[102,775,1255,1857]" pageId="4" pageNumber="585">
|
||
<emphasis id="CBC20B19FF96FF93FF90FB5AFF08D9CF" box="[102,224,1255,1279]" italics="true" pageId="4" pageNumber="585">Premaxilla</emphasis>
|
||
.—The premaxilla is only preserved on OCP DEK−GE 18 (
|
||
<figureCitation id="618DCB8EFF96FF93FF0DFABAFED4D810" box="[251,316,1287,1312]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="4" pageNumber="585">Fig. 3</figureCitation>
|
||
). It is crushed, but its general shape can be reconstructed. Only the three first right teeth are preserved, but four teeth seem to have existed on each premaxilla. The first tooth is smaller than the two other, and a deep concavity is present between the first and the second. The second tooth is larger, but less than the third tooth which seems to be the largest (apparently confirmed by the large space between the second and third mandibular tooth). The fourth premaxillary tooth is not preserved on the skull of OCP DEK−GE 18, but the space between its third and fourth mandibular teeth is short (as in OCP DEK−GE 1200), implying the fourth tooth would have been the smallest. The space between the fourth and the first maxillary tooth is large, dues to the large size of the fourth dentary tooth, and the premaxilla−maxilla suture is at this level on the lateral surface of the snout.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF96FF93FF7AF955FF28DA71" blockId="4.[102,775,1255,1857]" pageId="4" pageNumber="585">
|
||
The dorsal posterior process of the premaxilla is long and ends posteriorly at the level of the third maxillary tooth (
|
||
<figureCitation id="618DCB8EFF96FF93FF98F895FF5ADA71" box="[110,178,1832,1857]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="4" pageNumber="585">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF96FF93FF90F8EFFD11DADA" blockId="4.[102,775,1873,2026]" pageId="4" pageNumber="585">
|
||
<emphasis id="CBC20B19FF96FF93FF90F8EFFF53DA5A" box="[102,187,1874,1898]" italics="true" pageId="4" pageNumber="585">Maxilla</emphasis>
|
||
.—The number of teeth on each maxilla can be estimated to about 17 (
|
||
<figureCitation id="618DCB8EFF96FF93FEBFF8CCFE67DABA" box="[329,399,1905,1930]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="4" pageNumber="585">Fig. 3</figureCitation>
|
||
). The snout is relatively narrow, slender, and exhibits a longirostrine
|
||
<typeStatus id="260D69A9FF96FF93FE09F82FFDC6DA9A" box="[511,558,1938,1962]" pageId="4" pageNumber="585">type</typeStatus>
|
||
morphology (
|
||
<figureCitation id="618DCB8EFF96FF93FD26F82CFF7BDAFA" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[102,135,1976,1996]" captionStartId-1="3.[102,135,1134,1154]" captionStartId-2="4.[102,135,1117,1137]" captionTargetBox-1="[200,1388,233,1080]" captionTargetBox-2="[171,1417,225,1090]" captionTargetId-1="figure-433@3.[298,1372,225,1080]" captionTargetId-2="figure-520@4.[278,1288,249,1063]" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." captionText-1="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." captionText-2="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi-1="http://doi.org/10.5281/zenodo.13620803" figureDoi-2="http://doi.org/10.5281/zenodo.13620805" httpUri-1="https://zenodo.org/record/13620803/files/figure.png" httpUri-2="https://zenodo.org/record/13620805/files/figure.png" pageId="4" pageNumber="585">Figs. 2–4</figureCitation>
|
||
). The maxilla is relatively sculptured laterally compared to other skull area, with longitudinal deep ridges and furrows.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF96FF93FCA4FB23FA62D807" blockId="4.[812,1485,1182,1756]" pageId="4" pageNumber="585">
|
||
The alveoli are well developed, circular, with their base in relief and very marked ventrally (
|
||
<figureCitation id="618DCB8EFF96FF93FB51FB03FB17D9E7" box="[1191,1279,1214,1239]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="4" pageNumber="585">Fig. 2B</figureCitation>
|
||
). They are largely spaced (interalveolar space larger than the alveoli diameter for the teeth anterior to the thirteenth one), diameter and space decreasing posteriorly from the thirteenth tooth.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF96FF93FCACFAFDFB38DB09" blockId="4.[812,1485,1182,1756]" pageId="4" pageNumber="585">The maxilla is laterally straight, and the tooth row does not display a festooned outline in dorsal view. Occlusal pits are present from the space between the twelfth and thirteenth right maxillary alveoli, and between the thirteenth and fourteenth left alveoli. These pits, lined with the alveoli tooth row, increase in depth posteriorly. Anteriorly, the space between the right and left alveoli is large (twice the diameter of the alveoli) and increases posteriorly.</paragraph>
|
||
<paragraph id="F909D70BFF96FF93FCA4F9FEFB52DBEC" blockId="4.[812,1485,1182,1756]" pageId="4" pageNumber="585">
|
||
The contact with the palatines is expected between the level of the eleventh and twelfth maxillary alveoli (OCP DEK−GE 300;
|
||
<figureCitation id="618DCB8EFF96FF93FC23F93EFBC5DBAC" box="[981,1069,1667,1692]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="4" pageNumber="585">Fig. 4B</figureCitation>
|
||
). The maxilla contacts the ectopterygoid medially, and ends ventrally on the jugal. Dorsally, the maxillae are separated by the nasal.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF96FF92FCDAF94FFD13DF89" blockId="4.[812,1485,1777,2026]" lastBlockId="5.[102,774,640,697]" lastPageId="5" lastPageNumber="586" pageId="4" pageNumber="585">
|
||
<emphasis id="CBC20B19FF96FF93FCDAF94FFC84DA3A" box="[812,876,1778,1802]" italics="true" pageId="4" pageNumber="585">Nasal</emphasis>
|
||
.—The nasal is a single bone (the two nasals are fused), ornamented with only discrete and sparse furrows. It is narrow between the maxillae, and its anterior process penetrates deeply between the posterior premaxillae processes, but terminates
|
||
<quantity id="3E4E7AEEFF96FF93FC9DF8CFFC49DAB9" box="[875,929,1906,1929]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="585" unit="cm" value="7.0">7 cm</quantity>
|
||
(in OCP DEK−GE 18) posterior to the external nares, which it does not reach (
|
||
<figureCitation id="618DCB8EFF96FF93FBADF82CFB73DA9A" box="[1115,1179,1937,1962]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="4" pageNumber="585">Fig. 3</figureCitation>
|
||
). The nasal is narrow anteriorly between the maxillae, with a constant width from the posterior contact with the premaxillae to the anterior contact with the lacrimals. Then, it widens posteriorly and finally sends a long posterior process between frontal and prefrontal (
|
||
<figureCitation id="618DCB8EFF97FF92FD5AFD1DFD06DF89" box="[684,750,672,697]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="ADC98783FF97FF92FF90FD99FC8CDF64" ID-DOI="http://doi.org/10.5281/zenodo.13620807" ID-Zenodo-Dep="13620807" httpUri="https://zenodo.org/record/13620807/files/figure.png" pageId="5" pageNumber="586" startId="5.[102,135,548,568]" targetBox="[108,1485,235,515]" targetPageId="5" targetType="figure">
|
||
<paragraph id="F909D70BFF97FF92FF90FD99FC8CDF64" blockId="5.[102,1485,548,596]" pageId="5" pageNumber="586">
|
||
Fig. 5.
|
||
<taxonomicName id="3EB6AC88FF97FF92FF51FD99FE0BDF08" authorityName="Jouve & Iarochène & Bouya & Amaghzaz" authorityYear="2005" box="[167,483,548,568]" class="Reptilia" family="Dyrosauridae" genus="Arambourgisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="586" phylum="Chordata" rank="species" species="khouribgaensis" status="gen. et sp. nov.">
|
||
<emphasis id="CBC20B19FF97FF92FF51FD99FE0BDF08" box="[167,483,548,568]" italics="true" pageId="5" pageNumber="586">Arambourgisuchus khouribgaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D0F1B662FF97FF92FE1FFD98FD9EDF08" box="[489,630,549,568]" pageId="5" pageNumber="586" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, detail of the right posttemporal fenestra in posterior view. Photograph (
|
||
<emphasis id="CBC20B19FF97FF92FE38FDFDFE36DF64" bold="true" box="[462,478,576,596]" pageId="5" pageNumber="586">A</emphasis>
|
||
) and explanatory drawing of the same (
|
||
<emphasis id="CBC20B19FF97FF92FCBEFDFCFCBFDF64" bold="true" box="[840,855,577,596]" pageId="5" pageNumber="586">B</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F909D70BFF97FF92FF90FD71FE88DE55" blockId="5.[102,775,716,869]" pageId="5" pageNumber="586">
|
||
<emphasis id="CBC20B19FF97FF92FF90FD71FF33DFD4" box="[102,219,716,740]" italics="true" pageId="5" pageNumber="586">Prefrontal</emphasis>
|
||
.—Only the right prefrontal is preserved (on OCP DEK−GE 300;
|
||
<figureCitation id="618DCB8EFF97FF92FEF9FD51FE81DE35" box="[271,361,748,773]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Fig. 2A</figureCitation>
|
||
), almost complete (the most anterior part is absent). It is short, narrow and longer than wide (
|
||
<figureCitation id="618DCB8EFF97FF92FD38FCB1FF62DE75" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF97FF92FF62FC91FF53DE75" box="[148,187,812,837]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
|
||
). Dorsally, the contact with the frontal is as long as its contact with the nasal.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF97FF92FF90FCC5FDE7D920" blockId="5.[102,775,887,1040]" pageId="5" pageNumber="586">
|
||
<emphasis id="CBC20B19FF97FF92FF90FCC5FF25DEA0" box="[102,205,888,912]" italics="true" pageId="5" pageNumber="586">Lacrimal</emphasis>
|
||
.—The right lacrimal of specimen OCP DEK−GE 18 is well preserved (
|
||
<figureCitation id="618DCB8EFF97FF92FEB9FC2AFE79DE80" box="[335,401,919,944]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="5" pageNumber="586">Fig. 3</figureCitation>
|
||
), and is well anteriorly expanded, reaching the level of the tenth maxillary tooth. Its ornamentation is very light, with some shallow pits. It is large, and forms the anterior margin of the orbit.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF97FF92FF90FB9EFEB5D8AC" blockId="5.[102,775,1059,1501]" pageId="5" pageNumber="586">
|
||
<emphasis id="CBC20B19FF97FF92FF90FB9EFF54D90B" box="[102,188,1059,1083]" italics="true" pageId="5" pageNumber="586">Frontal</emphasis>
|
||
.—It extends anteriorly within the nasal as far as the prefrontal, and its width, at the orbital level appears moderate (
|
||
<figureCitation id="618DCB8EFF97FF92FF98FBDEFF39D94C" box="[110,209,1123,1148]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF97FF92FF2AFBDEFEEAD94C" box="[220,258,1123,1148]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
|
||
). Its lateral extension is short, and contacts the postorbital to constitute the postorbital bar. Posteromedially, at the angle between the postorbital bar and the interfenestral bar, the frontal forms a very light dorsal overhang within the supratemporal fenestra. Ventrally, and below this overhang, the frontal contacts an extremely laterally elongated laterosphenoid in a flat lateroventral extension. Posteromedially, the frontal takes part in the narrow interfenestral bar (
|
||
<quantity id="3E4E7AEEFF97FF92FD39FAFEFCEED86B" box="[719,774,1347,1372]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="586" unit="cm" value="1.0">1 cm</quantity>
|
||
width), since its total proportion is about one−fifth of the total length (
|
||
<figureCitation id="618DCB8EFF97FF92FF4DFA3EFEF7D8AC" box="[187,287,1411,1436]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF97FF92FEDCFA3EFEB9D8AC" box="[298,337,1411,1436]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF97FF92FF7AFA19FDC4D8ED" blockId="5.[102,775,1059,1501]" pageId="5" pageNumber="586">The frontal is smooth, without ornamentation, with only three medial grooves between the orbits.</paragraph>
|
||
<paragraph id="F909D70BFF97FF92FF90FA4DFE34DBD9" blockId="5.[102,775,1520,2026]" pageId="5" pageNumber="586">
|
||
<emphasis id="CBC20B19FF97FF92FF90FA4DFF2ADB38" box="[102,194,1520,1544]" italics="true" pageId="5" pageNumber="586">Parietal</emphasis>
|
||
.—The interfenestral bar, formed by the frontal and the parietal, decreases in width just posteriorly to the contact between these two bones: the width, which reached
|
||
<quantity id="3E4E7AEEFF97FF92FD50F98DFD34DB78" box="[678,732,1584,1609]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="586" unit="cm" value="1.0">1 cm</quantity>
|
||
anteriorly, is only 0.5 cm in width posterior to the suture (OCP DEK−GE 300;
|
||
<figureCitation id="618DCB8EFF97FF92FEE1F9CDFE96DBB9" box="[279,382,1648,1673]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF97FF92FE78F9CDFE5CDBB9" box="[398,436,1648,1673]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
|
||
). Anteroventrally, its contact with the laterosphenoid is visible, with a long anterior process between the frontal and the laterosphenoid, reducing the contact between these two bones.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF97FF92FF7AF94CFAD8D909" blockId="5.[102,775,1520,2026]" lastBlockId="5.[812,1485,640,1305]" pageId="5" pageNumber="586">
|
||
Ventrally, the parietal−laterosphenoid suture appears parallel to the skull roof. Posteriorly, the interfenestral bar is broken and separated from the occipital part of the skull. In the angle between the interfenestral bar and the posttemporal bar (formed by the parietal and squamosal), the parietal sends an anterodorsal overhang into the supratemporal fenestra. This overhang is more developed than the anterior one, beginning laterally about
|
||
<quantity id="3E4E7AEEFF97FF92FE93F86CFE72DAD9" box="[357,410,2001,2026]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="586" unit="cm" value="1.0">1 cm</quantity>
|
||
medial to the squamosal−parietal suture, and appears to end rapidly in the interfenestral bar. The posterior wall of the supratemporal fenestra is very inclined posteriorly, and is largely visible in dorsal view (
|
||
<figureCitation id="618DCB8EFF97FF92FA63FD7DFCB9DFC9" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF97FF92FCAAFD5DFC6BDFC9" box="[860,899,736,761]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
|
||
). The parietal−quadrate suture, visible in dorsal view on the posterior wall of the supratemporal fenestra, continues in the same direction as the parietal−laterosphenoid suture. It extends dorsally to join the squamosal−quadrate suture at the same level (in the right supratemporal fenestra) or slightly below the temporal canal (in the left supratemporal fenestra), about
|
||
<quantity id="3E4E7AEEFF97FF92FC87FC1DFC40DE88" box="[881,936,928,953]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="586" unit="cm" value="1.0">1 cm</quantity>
|
||
laterally to the temporal canal. The ventral squamosal−parietal suture joins the temporal canal in its most lateral part, continues dorsally to this one (crosses the temporal canal medially, in its one−third dorsomedial part), and continues in a vertical direction (with zigzag).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF97FF92FCA4FBFDFA9ED829" blockId="5.[812,1485,640,1305]" pageId="5" pageNumber="586">
|
||
The parietal is not ornamented on either the interfenestral bar or on the posterior skull roof. It contributes to half of the posterior wall of the supratemporal fenestra, and tapers posteromedially with a strong and acute process in the occipital face between the occipital tuberosities. The dorsal surface of this process is posteroventrally inclined, and the supraoccipital forms its ventrooccipital part (
|
||
<figureCitation id="618DCB8EFF97FF92FAD5FABDFA80D829" box="[1315,1384,1280,1305]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK− GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="5" pageNumber="586">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF97FF92FCDAFA94FBF7D8D2" blockId="5.[812,1486,1321,2026]" pageId="5" pageNumber="586">
|
||
<emphasis id="CBC20B19FF97FF92FCDAFA94FC42D871" box="[812,938,1321,1345]" italics="true" pageId="5" pageNumber="586">Postorbital</emphasis>
|
||
.—It is badly preserved on OCP DEK−GE 18 (
|
||
<figureCitation id="618DCB8EFF97FF92FCC3FAF4FC9FD852" box="[821,887,1353,1378]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="5" pageNumber="586">Fig. 3</figureCitation>
|
||
), and extremely fractured in OCP DEK−GE 300 (
|
||
<figureCitation id="618DCB8EFF97FF92FA56FAF4FCBAD8B2" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Fig. 2A</figureCitation>
|
||
). The postorbital bar, comprised of the postorbital and the jugal, seems to be gently concave medially. It was probably inclined ventrolaterally, not ornamented, mediolaterally flat and longer than wide.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF97FF92FCA4FA54FAAADA32" blockId="5.[812,1486,1321,2026]" pageId="5" pageNumber="586">
|
||
The postorbital is the most important part of the lateral arcade of the supratemporal fenestra (
|
||
<figureCitation id="618DCB8EFF97FF92FB33F9B4FAC2DB12" box="[1221,1322,1545,1570]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF97FF92FAC1F9B4FAB4DB12" box="[1335,1372,1545,1570]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
|
||
, 7). It appears dorsally and laterally ornamented with spaced pits, contacts the squamosal posteriorly, and the quadratojugal posteroventrally. It seems to participate largely to the dorsal margin of the infratemporal fenestra (Fig. 7). Anteriorly, it bears a robust lateral process, directed anteroventrally, and ornamented with deep furrows, which seems to contact the ventral margin of the orbit (jugal) (
|
||
<figureCitation id="618DCB8EFF97FF92FB42F954FAEADA32" box="[1204,1282,1769,1794]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="5" pageNumber="586">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF97FF92FAF9F954FADEDA32" box="[1295,1334,1769,1794]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="5" pageNumber="586">4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF97FF91FCDAF8AFFDC1D9D2" blockId="5.[812,1486,1321,2026]" lastBlockId="6.[102,775,1033,2026]" lastPageId="6" lastPageNumber="587" pageId="5" pageNumber="586">
|
||
<emphasis id="CBC20B19FF97FF92FCDAF8AFFC4FDA1A" box="[812,935,1810,1834]" italics="true" pageId="5" pageNumber="586">Squamosal</emphasis>
|
||
.—The squamosal forms the posterolateral part of the supratemporal fenestra. It is relatively narrow at the level of the posttemporal bar (1.5 cm in its minimum anteroposterior length) (
|
||
<figureCitation id="618DCB8EFF97FF92FC04F8CCFBA3DABA" box="[1010,1099,1905,1930]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="5" pageNumber="586">Fig. 2A</figureCitation>
|
||
). Weakly high in the posterior wall of the supratemporal fenestra, it constitutes the major part of the dorsal border of the temporal canal. It contacts the postorbital anterolaterally, but is separated from the quadratojugal by the quadrate. It takes part in the dorsal part of the external ear, and sends an important squamosal wing roofing the external otic aperture (Fig. 7). Posterior to the ear, the squamosal constitutes a long and high blade sinking deeply beneath the skull roof and forms the anterior wall of the paroccipital process. The paroccipital process extends more ventrally than the level of the skull roof.
|
||
</paragraph>
|
||
<caption id="ADC98783FF94FF91FF90FCC0FD28DEF9" ID-DOI="http://doi.org/10.5281/zenodo.13620809" ID-Zenodo-Dep="13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="6" pageNumber="587" startId="6.[102,136,893,913]" targetBox="[102,774,231,864]" targetPageId="6" targetType="figure">
|
||
<paragraph id="F909D70BFF94FF91FF90FCC0FD28DEF9" blockId="6.[102,775,893,969]" pageId="6" pageNumber="587">
|
||
Fig. 6.
|
||
<taxonomicName id="3EB6AC88FF94FF91FF5CFCC0FE1DDEA1" authorityName="Jouve & Iarochène & Bouya & Amaghzaz" authorityYear="2005" box="[170,501,893,913]" class="Reptilia" family="Dyrosauridae" genus="Arambourgisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="6" pageNumber="587" phylum="Chordata" rank="species" species="khouribgaensis" status="gen. et sp. nov.">
|
||
<emphasis id="CBC20B19FF94FF91FF5CFCC0FE1DDEA1" box="[170,501,893,913]" italics="true" pageId="6" pageNumber="587">Arambourgisuchus khouribgaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D0F1B662FF94FF91FE0AFCC3FD78DEA1" box="[508,656,894,913]" pageId="6" pageNumber="587" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, OCP DEKGE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (
|
||
<emphasis id="CBC20B19FF94FF91FEECFC08FEC2DEF9" bold="true" box="[282,298,949,969]" pageId="6" pageNumber="587">A</emphasis>
|
||
) and explanatory drawing of the same (
|
||
<emphasis id="CBC20B19FF94FF91FD55FC0BFD5ADEF9" bold="true" box="[675,690,950,969]" pageId="6" pageNumber="587">B</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F909D70BFF94FF91FF7AFB57FDB0D892" blockId="6.[102,775,1033,2026]" pageId="6" pageNumber="587">
|
||
In occipital view, the squamosal contributes dorsolaterally to the posttemporal fenestra (less than the lateral half), and to a small lateral part of the occipital tuberosity. It contributes to a small part to the occipital face, and more posteriorly, it participates in the dorsal edge of the paroccipital process, ending before the extremity one of the process (
|
||
<figureCitation id="618DCB8EFF94FF91FDF1FA34FDA3D892" box="[519,587,1417,1442]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK− GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="6" pageNumber="587">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF94FF91FF90FA0FFEA9DA7A" blockId="6.[102,775,1033,2026]" pageId="6" pageNumber="587">
|
||
<emphasis id="CBC20B19FF94FF91FF90FA0FFF4CD8FA" box="[102,164,1458,1482]" italics="true" pageId="6" pageNumber="587">Jugal</emphasis>
|
||
.—Very fragmentary (
|
||
<figureCitation id="618DCB8EFF94FF91FE5CFA0CFDE1D8FA" box="[426,521,1457,1482]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Fig. 2A</figureCitation>
|
||
), the jugal forms the lateroventral edge of the orbit, and the ventral part of the postorbital pillar (
|
||
<figureCitation id="618DCB8EFF94FF91FEC1FA4CFE7DDB3A" box="[311,405,1521,1546]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="6" pageNumber="587">Fig. 4A</figureCitation>
|
||
). There is no lateral jugal edge raised bordering this pillar, and the postorbital pillar is not laterally in continuity with the lateral edge of the jugal. The postorbital bar is displaced medially and is not laterally in alignment with the lateral jugal edge. The base of the postorbital pillar is completely pierced by a foramen, anteroposteriorly directed. Apparently, the posterior aperture of this foramen is situated on the external face of the postorbital pillar. Posteriorly, the internal part of the jugal is exposed and exhibits a deep and long groove, including two distinct foramina (
|
||
<figureCitation id="618DCB8EFF94FF91FF2CF88CFEDCDA7A" box="[218,308,1841,1866]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Fig. 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF94FF91FF7AF8ECFE0FDADA" blockId="6.[102,775,1033,2026]" pageId="6" pageNumber="587">
|
||
The jugal is laterally ornamented with spaced deep pits. Posteriorly, it is high, slightly convex dorsally, lateromedially narrow and ends just before the quadratojugal lateral notch (see below). Anteriorly, it reaches the level of the anterior process of the prefrontal (
|
||
<figureCitation id="618DCB8EFF94FF91FEADF86CFE40DADA" box="[347,424,2001,2026]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="6" pageNumber="587">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF94FF91FE43F86CFE33DADA" box="[437,475,2001,2026]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="6" pageNumber="587">4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF94FF91FCDAFF55FC95DCD1" blockId="6.[812,1485,232,611]" pageId="6" pageNumber="587">
|
||
<emphasis id="CBC20B19FF94FF91FCDAFF55FC3ADC30" box="[812,978,232,256]" italics="true" pageId="6" pageNumber="587">Quadratojugal</emphasis>
|
||
.—It is well developed, and contributes to the jaw joint for one quarter (
|
||
<figureCitation id="618DCB8EFF94FF91FBA6FEB5FB7EDC11" box="[1104,1174,264,289]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Fig. 2</figureCitation>
|
||
). It is laterally straight, and extends slightly ventrally in its posterior portion. A deep lateral notch just before the articulation marks off the jaw joint segment (preserved on the right quadratojugal). Medially, the contact with the jugal is long, and the space between the quadratojugal and the ectopterygoid is small on the jugal (
|
||
<quantity id="3E4E7AEEFF94FF91FCC3FE75FC9ADCD1" box="[821,882,456,481]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="587" unit="cm" value="2.0">2 cm</quantity>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF94FF91FCA4FE57FA6ADF53" blockId="6.[812,1485,232,611]" pageId="6" pageNumber="587">The quadratojugal constitutes the posterior edge of the infratemporal fenestra, and forms a part of its dorsal margin (Fig. 7). Dorsolaterally, it is separated from the squamosal by the quadrate (there is no contact with the squamosal).</paragraph>
|
||
<paragraph id="F909D70BFF94FF91FCDAFDCBFA5DDE7F" blockId="6.[812,1485,630,847]" pageId="6" pageNumber="587">
|
||
<emphasis id="CBC20B19FF94FF91FCDAFDCBFC39DFBE" box="[812,977,630,654]" italics="true" pageId="6" pageNumber="587">Supraoccipital</emphasis>
|
||
.—It is a small bone, “V” shaped in posterior view, which contributes to the posteromedial occipital process with the parietal (
|
||
<figureCitation id="618DCB8EFF94FF91FBDCFD0BFB66DFFF" box="[1066,1166,694,719]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF94FF91FB6DFD0BFB28DFFF" box="[1179,1216,694,719]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="6" pageNumber="587">4A</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF94FF91FB3AFD0BFB34DFFF" box="[1228,1244,694,719]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK− GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="6" pageNumber="587">6</figureCitation>
|
||
). It forms the medioventral mid part of the posttemporal fenestra, posteriorly on the occipital tuberosity, but contacts the squamosal one centimeter anteriorly, within the posttemporal fenestra (
|
||
<figureCitation id="618DCB8EFF94FF91FA8CFCABFA57DE1F" box="[1402,1471,790,815]" captionStart="Fig" captionStartId="5.[102,135,548,568]" captionTargetBox="[108,1485,235,515]" captionTargetId="figure-793@5.[102,1382,226,519]" captionTargetPageId="5" captionText="Fig. 5. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, detail of the right posttemporal fenestra in posterior view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620807" httpUri="https://zenodo.org/record/13620807/files/figure.png" pageId="6" pageNumber="587">Fig. 5</figureCitation>
|
||
). It does not seem to contribute to the occipital tuberosities.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF94FF91FCDAFCDFFBBBD9EB" blockId="6.[812,1485,866,1534]" pageId="6" pageNumber="587">
|
||
<emphasis id="CBC20B19FF94FF91FCDAFCDFFC43DE4A" box="[812,939,866,890]" italics="true" pageId="6" pageNumber="587">Exoccipital</emphasis>
|
||
.—They form the main part of the occipital face, contributing laterally to the occipital condyle (each exoccipital constitutes to one third of the half width), and almost completely surround the foramen magnum (three−quarter) (
|
||
<figureCitation id="618DCB8EFF94FF91FC93FC5FFC44DECB" box="[869,940,994,1019]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK− GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="6" pageNumber="587">Fig. 6</figureCitation>
|
||
). Dorsally, they form the occipital tuberosities, which are well developed, dorsoventrally flattened, and posteriorly directed under the posttemporal fenestra. Laterally, they constitute the main part of the robust paroccipital process, and surround dorsally, medially and ventrally to the cranio−quadrate canal. The paroccipital process is flat, composed of the exoccipital and squamosal, and its posterior extremity is quadrangular.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF94FF91FCA4FB59FAF1D84D" blockId="6.[812,1485,866,1534]" pageId="6" pageNumber="587">
|
||
Ventrally, the exoccipitals contribute posterolaterally, one−third to each basioccipital tuberosity in a very broad ventral process (
|
||
<figureCitation id="618DCB8EFF94FF91FC3EFA99FBD9D80D" box="[968,1073,1316,1341]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF94FF91FBB4FA99FB8DD80D" box="[1090,1125,1316,1341]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="6" pageNumber="587">4B</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF94FF91FB81FA99FB73D80D" box="[1143,1179,1316,1341]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK− GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="6" pageNumber="587">6B</figureCitation>
|
||
). The suture between the basioccipital and the exoccipital, posteriorly to the basioccipital tuberosities, lay deeply in a cavity.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF94FF91FCA4FA38FB37D8CE" blockId="6.[812,1485,866,1534]" pageId="6" pageNumber="587">
|
||
The foramen for nerve XII is small and laterally directed on the exoccipital. The vagus foramen and posterior carotid foramen are situated anterior to the foramen for nerve XII, and directed ventroposteriorly (
|
||
<figureCitation id="618DCB8EFF94FF91FB7BFA58FB39D8CE" box="[1165,1233,1509,1534]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK− GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="6" pageNumber="587">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF94FF91FCDAF9AFFB8EDADA" blockId="6.[812,1485,1553,2026]" pageId="6" pageNumber="587">
|
||
<emphasis id="CBC20B19FF94FF91FCDAF9AFFC28DB1A" box="[812,960,1554,1578]" italics="true" pageId="6" pageNumber="587">Basioccipital</emphasis>
|
||
.—It constitutes the main part of the occipital condyle, which is very wide. Its anterior part (basioccipital tuberosities) is not strongly projected ventrally and therefore does not proceed ventrally to the occipital condyle in occipital view (
|
||
<figureCitation id="618DCB8EFF94FF91FC60F92CFC32DB9A" box="[918,986,1681,1706]" captionStart="Fig" captionStartId="6.[102,136,893,913]" captionTargetBox="[102,774,231,864]" captionTargetId="figure-805@6.[159,677,230,816]" captionTargetPageId="6" captionText="Fig. 6. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK− GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in occipital view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620809" httpUri="https://zenodo.org/record/13620809/files/figure.png" pageId="6" pageNumber="587">Fig. 6</figureCitation>
|
||
). The area between the basioccipital tuberosities and the occipital condyle is gently curved dorsally, almost horizontally oriented. In ventral view, the posterior margin of the basioccipital tuberosities is concave anteriorly, relatively thin in its medial part (just behind the medial eustachian foramen), with a small medial ridge posteriorly to the medial eustachian foramen (
|
||
<figureCitation id="618DCB8EFF94FF91FB9CF8ECFB24DA5A" box="[1130,1228,1873,1898]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF94FF91FB2EF8ECFB14DA5A" box="[1240,1276,1873,1898]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="6" pageNumber="587">4B</figureCitation>
|
||
). The anteroposterior thinness of the medial part of the basioccipital, posterior to the medial eustachian foramen, separates the two basioccipital tuberosities, each one having a “drop” shape in ventral view (
|
||
<figureCitation id="618DCB8EFF94FF91FC31F86CFBC1DADA" box="[967,1065,2001,2026]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="6" pageNumber="587">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF94FF91FBC3F86CFBB1DADA" box="[1077,1113,2001,2026]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="6" pageNumber="587">4B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF95FF90FF90FDDDFF07DEF1" blockId="7.[102,241,608,628]" lastBlockId="7.[112,239,941,961]" pageId="7" pageNumber="588">laterosphenoid parietal frontal postorbital postorbital bar jugal ectopterygoid</paragraph>
|
||
<paragraph id="F909D70BFF95FF90FF90FBA5FEE5D984" blockId="7.[102,775,1048,1204]" pageId="7" pageNumber="588">
|
||
<emphasis id="CBC20B19FF95FF90FF90FBA5FF15D900" box="[102,253,1048,1072]" italics="true" pageId="7" pageNumber="588">Basisphenoid</emphasis>
|
||
.—It seems to enclose completely the medial eustachian foramen, but the limit between basioccipital and basisphenoid cannot be seen. The bone is narrow laterally to the medial eustachian foramen, with a lateral “pinching” (
|
||
<figureCitation id="618DCB8EFF95FF90FF98FB26FF38D984" box="[110,208,1179,1204]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Figs. 2B</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF95FF90FF2AFB26FEE8D984" box="[220,256,1179,1204]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">4B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF95FF90FF90FB77FD15D8F3" blockId="7.[102,775,1226,1475]" pageId="7" pageNumber="588">
|
||
<emphasis id="CBC20B19FF95FF90FF90FB77FF27D9D2" box="[102,207,1226,1250]" italics="true" pageId="7" pageNumber="588">Quadrate</emphasis>
|
||
.—It is long, posteroventrally directed, and forms the jaw joint with the quadratojugal (
|
||
<figureCitation id="618DCB8EFF95FF90FDE8FB57FD6ED833" box="[542,646,1258,1283]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF95FF90FD61FB57FD40D833" box="[663,680,1258,1283]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">4</figureCitation>
|
||
). In the supratemporal fenestra, it contacts the squamosal lateral to the temporal canal. Anterolaterally, it slips in between the squamosal and the quadratojugal by a thin anterior process that contacts the postorbital. Anteriorly, on its ventral face, the quadrate bears a small crest at its mid width, which probably corresponds to the “crest B” of
|
||
<bibRefCitation id="9D27AAFAFF95FF90FE42FA17FD82D8F3" author="Iordansky, N. N." box="[436,618,1450,1475]" pageId="7" pageNumber="588" pagination="256 - 280" refId="ref10559" refString="Iordansky, N. N. 1964. The jaw muscles of the crocodiles and some relating structures of the crocodilian skull. Anatomischer Anzeiger 115: 256 - 280." type="journal article" year="1964">Iordansky (1964</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="9D27AAFAFF95FF90FD84FA17FD44D8F3" author="Iordansky, N. N." box="[626,684,1450,1475]" pageId="7" pageNumber="588" pagination="567 - 575" refId="ref10590" refString="Iordansky, N. N. 1967. Peculiarities of the crocodilian skull related to the feeding function and origin of Gavialidae [in Russian with English abstract]. Zoologiceskij zurnal 46: 567 - 575." type="journal article" year="1967">1967</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="9D27AAFAFF95FF90FD43FA17FD18D8F3" author="Iordansky, N. N." box="[693,752,1450,1475]" pageId="7" pageNumber="588" pagination="201 - 284" refId="ref10628" refString="Iordansky, N. N. 1973. The skull of the Crocodilia. In: C. Gans and T. S. Parson (eds.), Biology of the Reptilia, Morphology D, Volume 4, 201 - 284. Academic Press, London." type="book chapter" year="1973">1973</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF95FF90FF90FA64FDE6DA28" blockId="7.[102,775,1496,1816]" pageId="7" pageNumber="588">
|
||
<emphasis id="CBC20B19FF95FF90FF90FA64FF2DD8C1" box="[102,197,1497,1521]" italics="true" pageId="7" pageNumber="588">Palatine</emphasis>
|
||
.—The palatines are very crushed (
|
||
<figureCitation id="618DCB8EFF95FF90FDADFA65FD5ED8C1" box="[603,694,1496,1521]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Fig. 2B</figureCitation>
|
||
). They seem enlarged and curved laterally before contacting the maxillae anteriorly. This contact, dues to the bad preservation, is not available. Its posterior border seems flattened, enlarged posteriorly and deviates laterally before joining the pterygoid (lateral border of the palatine not parallel, but laterally curved anteriorly and posteriorly, medial border of suborbital fenestra medially curved). The contact with the pterygoid is not clearly visible, but it seems to form only the anterior part of the choanae (
|
||
<figureCitation id="618DCB8EFF95FF90FE5EF942FDE9DA28" box="[424,513,1791,1816]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">Fig. 4B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF95FF90FF90F893FBFBDEFF" blockId="7.[102,774,1838,2026]" lastBlockId="7.[895,1486,834,991]" pageId="7" pageNumber="588">
|
||
<emphasis id="CBC20B19FF95FF90FF90F893FEECDA76" box="[102,260,1838,1862]" italics="true" pageId="7" pageNumber="588">Ectopterygoid</emphasis>
|
||
.—The left ectopterygoid is well preserved (
|
||
<figureCitation id="618DCB8EFF95FF90FF98F8F3FF2DDA57" box="[110,197,1870,1895]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Fig. 2B</figureCitation>
|
||
). It is wide and twisted between its contacts with the jugal and the pterygoid. It curves gently anteriorly to form the posterolateral part of the suborbital fenestra (
|
||
<figureCitation id="618DCB8EFF95FF90FD68F82DFD11DA99" box="[670,761,1936,1961]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">Fig. 4B</figureCitation>
|
||
). The posterior process is wide, decreasing posteriorly, and covers the pterygoid almost as far as its posterior extremity occipital tuberosities exoccipital squamosal quadrate quadratojugal Fig. 7.
|
||
<taxonomicName id="3EB6AC88FF95FF90FB67FCFFFA25DE66" authorityName="Jouve & Iarochène & Bouya & Amaghzaz" authorityYear="2005" box="[1169,1485,834,854]" class="Reptilia" family="Dyrosauridae" genus="Arambourgisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="7" pageNumber="588" phylum="Chordata" rank="species" species="khouribgaensis" status="gen. et sp. nov.">
|
||
<emphasis id="CBC20B19FF95FF90FB67FCFFFA25DE66" box="[1169,1485,834,854]" italics="true" pageId="7" pageNumber="588">Arambourgisuchus khouribgaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D0F1B662FF95FF90FBA4FCE3FB09DE41" box="[1106,1249,862,881]" pageId="7" pageNumber="588" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, OCP DEK−GE 300, Sidi pterygoid Chenane,
|
||
<collectingCountry id="81A1979BFF95FF90FB5DFCC7FB17DEBD" box="[1195,1279,890,909]" name="Morocco" pageId="7" pageNumber="588">Morocco</collectingCountry>
|
||
, late Palaeocene, detail of the left posterior part of the skull in left lateral view. Photograph (
|
||
<emphasis id="CBC20B19FF95FF90FACAFC0DFAA4DEF4" bold="true" box="[1340,1356,944,964]" pageId="7" pageNumber="588">A</emphasis>
|
||
) and explanatorus transiliens
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF95FF90FBA4FC71FAB6DEEF" blockId="7.[895,1486,834,991]" box="[1106,1374,971,991]" pageId="7" pageNumber="588">
|
||
tory drawing of the same (
|
||
<emphasis id="CBC20B19FF95FF90FAB7FC71FAB8DEEF" bold="true" box="[1345,1360,972,991]" pageId="7" pageNumber="588">B</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF95FF90FCDAFBA5FBB9D961" blockId="7.[812,1485,1048,2026]" pageId="7" pageNumber="588">
|
||
(
|
||
<figureCitation id="618DCB8EFF95FF90FCC3FBA5FC65D901" box="[821,909,1048,1073]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Fig. 2B</figureCitation>
|
||
). Its contact with the maxilla seems short, whereas that with the jugal is long.
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF95FF90FCDAFBDCFAA5D9E9" blockId="7.[812,1485,1048,2026]" pageId="7" pageNumber="588">
|
||
<emphasis id="CBC20B19FF95FF90FCDAFBDCFC72D949" box="[812,922,1121,1145]" italics="true" pageId="7" pageNumber="588">Pterygoid</emphasis>
|
||
.—The pterygoids are damaged (
|
||
<figureCitation id="618DCB8EFF95FF90FAF1FBDDFAB5D949" box="[1287,1373,1120,1145]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Fig. 2B</figureCitation>
|
||
), but their shape can be trace out. Anteriorly, they almost completely surround the choanae. A septum, formed by the pterygoids separates the choanae in two openings (
|
||
<figureCitation id="618DCB8EFF95FF90FB1EFB7DFAA8D9E9" box="[1256,1344,1216,1241]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">Fig. 4B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF95FF90FCA4FB5CFACBDB0A" blockId="7.[812,1485,1048,2026]" pageId="7" pageNumber="588">
|
||
The pterygoids diverge posterolateraly to form two lateral wings in contact with the ectopterygoid. In front and anterolaterally to the choanae, they seem flattened and enlarged laterally; they are anteroposteriorly narrow between the choanae and their contact with the ectopterygoids (
|
||
<figureCitation id="618DCB8EFF95FF90FA56FADCFCB8D8AA" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">Fig. 4B</figureCitation>
|
||
). Contact with the ectopterygoid is small, and increases rapidly lateroposteriorly with the increase of the anteroposterior length of the pterygoid wing. The lateral part of this wing is extremely thickened dorsoventrally to form a strong
|
||
<emphasis id="CBC20B19FF95FF90FCDAF9BFFC0CDB29" box="[812,996,1537,1561]" italics="true" pageId="7" pageNumber="588">torus transiliens</emphasis>
|
||
, with a very important anterior thickening, decreasing progressively posteriorly (Fig. 7).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF95FF90FCDAF9F4FA2BDA32" blockId="7.[812,1485,1048,2026]" pageId="7" pageNumber="588">
|
||
<emphasis id="CBC20B19FF95FF90FCDAF9F4FC30DB51" box="[812,984,1609,1633]" italics="true" pageId="7" pageNumber="588">Laterosphenoid</emphasis>
|
||
.—Anteriorly, it is laterally expanded ventrally to the frontal, and its anterior margin is lateromedially directed (
|
||
<figureCitation id="618DCB8EFF95FF90FC60F934FC12DB92" box="[918,1018,1673,1698]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Figs. 2A</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF95FF90FBF1F934FBC5DB92" box="[1031,1069,1673,1698]" captionStart="Fig" captionStartId="4.[102,135,1117,1137]" captionTargetBox="[171,1417,225,1090]" captionTargetId="figure-520@4.[278,1288,249,1063]" captionTargetPageId="4" captionText="Fig. 4. Arambourgisuchus khouribgaensis gen. et sp. nov., reconstruction of skull in dorsal (A) and ventral (B) views." figureDoi="http://doi.org/10.5281/zenodo.13620805" httpUri="https://zenodo.org/record/13620805/files/figure.png" pageId="7" pageNumber="588">4A</figureCitation>
|
||
). The posterior part is dorsoventrally crushed, and its shape is hardly distinguished (
|
||
<figureCitation id="618DCB8EFF95FF90FADBF914FA99DBF2" box="[1325,1393,1705,1730]" captionStart="Fig" captionStartId="2.[102,135,1976,1996]" captionText="Fig. 2. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 300, Sidi Chenane, Morocco, late Palaeocene, skull in dorsal (A) and ventral (B) views." pageId="7" pageNumber="588">Fig. 2</figureCitation>
|
||
). Suture with the frontal and parietal (in continuity) seems horizontal, parallel to the skull roof (dorsal limit of the interfenestral bar).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF95FF9FFCDAF8AFFD91DC91" blockId="7.[812,1485,1048,2026]" lastBlockId="8.[102,775,232,613]" lastPageId="8" lastPageNumber="589" pageId="7" pageNumber="588">
|
||
<emphasis id="CBC20B19FF95FF90FCDAF8AFFC7FDA1A" box="[812,919,1810,1834]" italics="true" pageId="7" pageNumber="588">Mandible</emphasis>
|
||
.—Two isolated mandibular fragments have been found, one anterior, preserved from the first to the twelfth tooth (OCP DEK−GE 1200;
|
||
<figureCitation id="618DCB8EFF95FF90FB82F8ECFB20DA5A" box="[1140,1224,1873,1898]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEK−GE 1200 in ventral view. B. OCP DEK−GE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." figureDoi="http://doi.org/10.5281/zenodo.13620813" httpUri="https://zenodo.org/record/13620813/files/figure.png" pageId="7" pageNumber="588">Fig 8A</figureCitation>
|
||
), and a more posterior part, including the eleventh to the sixteenth teeth (OCP DEK−GE 269;
|
||
<figureCitation id="618DCB8EFF95FF90FC20F82CFBA7DA9A" box="[982,1103,1937,1962]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEK−GE 1200 in ventral view. B. OCP DEK−GE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." figureDoi="http://doi.org/10.5281/zenodo.13620813" httpUri="https://zenodo.org/record/13620813/files/figure.png" pageId="7" pageNumber="588">Fig. 8B, C</figureCitation>
|
||
). Unfortunately, only a small part of its posterior branch is preserved on the specimen OCP DEK−GE 300, but too poorly preserved to be interpreted. The first tooth is relatively robust (
|
||
<figureCitation id="618DCB8EFF9AFF9FFE4DFF55FDFDDC31" box="[443,533,232,257]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEK−GE 1200 in ventral view. B. OCP DEK−GE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." figureDoi="http://doi.org/10.5281/zenodo.13620813" httpUri="https://zenodo.org/record/13620813/files/figure.png" pageId="8" pageNumber="589">Fig. 8A</figureCitation>
|
||
), the second alveolus smaller, and the space between it and the third one is greater than between the other. The third tooth is about the same size as the second, and it is near the very enlarged fourth. This one has its alveolar edge higher dorsally. The next alveoli seem relatively regular in their spacing and diameter.
|
||
</paragraph>
|
||
<caption id="ADC98783FF9AFF9FFCDAFF57FBBEDC09" pageId="8" pageNumber="589" startId="8.[812,867,234,255]" targetType="table">
|
||
<paragraph id="F909D70BFF9AFF9FFCDAFF57FBBEDC09" blockId="8.[812,1485,234,313]" pageId="8" pageNumber="589">Table 1. Measurements of the skull of different dyrosaurids. Abbreviations: DL, dorsal length of the skull; PreoL, preorbital length; R, ration, preorbital length/dorsal length.</paragraph>
|
||
</caption>
|
||
<paragraph id="F909D70BFF9AFF9FFBE0FEEBFA62DF1D" pageId="8" pageNumber="589">
|
||
<table id="8BB625ABFF9A0068FCC2FEEBFA5BDF1D" box="[820,1459,342,557]" gridcols="4" gridrows="7" pageId="8" pageNumber="589">
|
||
<tr id="4786D549FF9A0068FCC2FEEBFA5BDC5B" box="[820,1459,342,363]" gridrow="0" pageId="8" pageNumber="589" rowspan-0="1">
|
||
<th id="0457BC35FF9A0068FBE0FEEBFB81DC5B" box="[1046,1129,342,363]" gridcol="1" gridrow="0" pageId="8" pageNumber="589">DL (cm)</th>
|
||
<th id="0457BC35FF9A0068FB65FEEBFAE9DC5B" box="[1171,1281,342,363]" gridcol="2" gridrow="0" pageId="8" pageNumber="589">PreoL (cm)</th>
|
||
<th id="0457BC35FF9A0068FADCFEEBFA5BDC5B" box="[1322,1459,342,363]" gridcol="3" gridrow="0" pageId="8" pageNumber="589">R (PreSL/DL)</th>
|
||
</tr>
|
||
<tr id="4786D549FF9A0068FCC2FECAFA5BDCBC" box="[820,1459,375,396]" gridrow="1" pageId="8" pageNumber="589">
|
||
<th id="0457BC35FF9A0068FCC2FECAFC0ADCBC" box="[820,994,375,396]" gridcol="0" gridrow="1" pageId="8" pageNumber="589">
|
||
<taxonomicName id="3EB6AC88FF9AFF9FFCC2FECAFC6BDCBC" authorityName="Swinton" authorityYear="1930" box="[820,899,375,396]" class="Reptilia" family="Dyrosauridae" genus="Rhabdognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="rarus">
|
||
<emphasis id="CBC20B19FF9AFF9FFCC2FECAFC6BDCBC" box="[820,899,375,396]" italics="true" pageId="8" pageNumber="589">R. rarus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="0457BC35FF9A0068FBE0FECAFB81DCBC" box="[1046,1129,375,396]" gridcol="1" gridrow="1" pageId="8" pageNumber="589">73.1</td>
|
||
<td id="0457BC35FF9A0068FB65FECAFAE9DCBC" box="[1171,1281,375,396]" gridcol="2" gridrow="1" pageId="8" pageNumber="589">53.4</td>
|
||
<td id="0457BC35FF9A0068FADCFECAFA5BDCBC" box="[1322,1459,375,396]" gridcol="3" gridrow="1" pageId="8" pageNumber="589">73.05</td>
|
||
</tr>
|
||
<tr id="4786D549FF9A0068FCC2FE2AFA5BDC9C" box="[820,1459,407,428]" gridrow="2" pageId="8" pageNumber="589">
|
||
<th id="0457BC35FF9A0068FCC2FE2AFC0ADC9C" box="[820,994,407,428]" gridcol="0" gridrow="2" pageId="8" pageNumber="589">
|
||
<taxonomicName id="3EB6AC88FF9AFF9FFCC2FE2AFC39DC9C" baseAuthorityName="Thomas" baseAuthorityYear="1893" box="[820,977,407,428]" class="Reptilia" family="Crocodylidae" genus="Crocodilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="phosphaticus">
|
||
<emphasis id="CBC20B19FF9AFF9FFCC2FE2AFC39DC9C" box="[820,977,407,428]" italics="true" pageId="8" pageNumber="589">D. phosphaticus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="0457BC35FF9A0068FBE0FE2AFB81DC9C" box="[1046,1129,407,428]" gridcol="1" gridrow="2" pageId="8" pageNumber="589">104</td>
|
||
<td id="0457BC35FF9A0068FB65FE2AFAE9DC9C" box="[1171,1281,407,428]" gridcol="2" gridrow="2" pageId="8" pageNumber="589">75</td>
|
||
<td id="0457BC35FF9A0068FADCFE2AFA5BDC9C" box="[1322,1459,407,428]" gridcol="3" gridrow="2" pageId="8" pageNumber="589">72.12</td>
|
||
</tr>
|
||
<tr id="4786D549FF9A0068FCC2FE0AFA5BDCFC" box="[820,1459,439,460]" gridrow="3" pageId="8" pageNumber="589">
|
||
<th id="0457BC35FF9A0068FCC2FE0AFC0ADCFC" box="[820,994,439,460]" gridcol="0" gridrow="3" pageId="8" pageNumber="589">
|
||
<taxonomicName id="3EB6AC88FF9AFF9FFCC2FE0AFC0ADCFC" authorityName="Jouve & Iarochène & Bouya & Amaghzaz" authorityYear="2005" box="[820,994,439,460]" class="Reptilia" family="Dyrosauridae" genus="Arambourgisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="khouribgaensis">
|
||
<emphasis id="CBC20B19FF9AFF9FFCC2FE0AFC0ADCFC" box="[820,994,439,460]" italics="true" pageId="8" pageNumber="589">A. khouribgaensis</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="0457BC35FF9A0068FBE0FE0AFB81DCFC" box="[1046,1129,439,460]" gridcol="1" gridrow="3" pageId="8" pageNumber="589">100</td>
|
||
<td id="0457BC35FF9A0068FB65FE0AFAE9DCFC" box="[1171,1281,439,460]" gridcol="2" gridrow="3" pageId="8" pageNumber="589">71.5</td>
|
||
<td id="0457BC35FF9A0068FADCFE0AFA5BDCFC" box="[1322,1459,439,460]" gridcol="3" gridrow="3" pageId="8" pageNumber="589">71.50</td>
|
||
</tr>
|
||
<tr id="4786D549FF9A0068FCC2FE65FA5BDCDD" box="[820,1459,472,493]" gridrow="4" pageId="8" pageNumber="589">
|
||
<th id="0457BC35FF9A0068FCC2FE65FC0ADCDD" box="[820,994,472,493]" gridcol="0" gridrow="4" pageId="8" pageNumber="589">
|
||
<taxonomicName id="3EB6AC88FF9AFF9FFCC2FE65FC56DCDD" authorityName="Bergounioux" authorityYear="1955" box="[820,958,472,493]" class="Reptilia" family="Dyrosauridae" genus="Phosphatosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="gavialoides">
|
||
<emphasis id="CBC20B19FF9AFF9FFCC2FE65FC56DCDD" box="[820,958,472,493]" italics="true" pageId="8" pageNumber="589">P. gavialoides</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="0457BC35FF9A0068FBE0FE65FB81DCDD" box="[1046,1129,472,493]" gridcol="1" gridrow="4" pageId="8" pageNumber="589">107</td>
|
||
<td id="0457BC35FF9A0068FB65FE65FAE9DCDD" box="[1171,1281,472,493]" gridcol="2" gridrow="4" pageId="8" pageNumber="589">72</td>
|
||
<td id="0457BC35FF9A0068FADCFE65FA5BDCDD" box="[1322,1459,472,493]" gridcol="3" gridrow="4" pageId="8" pageNumber="589">67.29</td>
|
||
</tr>
|
||
<tr id="4786D549FF9A0068FCC2FE45FA5BDF3D" box="[820,1459,504,525]" gridrow="5" pageId="8" pageNumber="589">
|
||
<th id="0457BC35FF9A0068FCC2FE45FC0ADF3D" box="[820,994,504,525]" gridcol="0" gridrow="5" pageId="8" pageNumber="589">
|
||
<taxonomicName id="3EB6AC88FF9AFF9FFCC2FE45FC58DF3D" authorityName="Halstead" authorityYear="1975" box="[820,944,504,525]" class="Reptilia" family="Dyrosauridae" genus="Sokotosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="ianwilsoni">
|
||
<emphasis id="CBC20B19FF9AFF9FFCC2FE45FC58DF3D" box="[820,944,504,525]" italics="true" pageId="8" pageNumber="589">S. ianwilsoni</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="0457BC35FF9A0068FBE0FE45FB81DF3D" box="[1046,1129,504,525]" gridcol="1" gridrow="5" pageId="8" pageNumber="589">60.1</td>
|
||
<td id="0457BC35FF9A0068FB65FE45FAE9DF3D" box="[1171,1281,504,525]" gridcol="2" gridrow="5" pageId="8" pageNumber="589">39.7</td>
|
||
<td id="0457BC35FF9A0068FADCFE45FA5BDF3D" box="[1322,1459,504,525]" gridcol="3" gridrow="5" pageId="8" pageNumber="589">66.06</td>
|
||
</tr>
|
||
<tr id="4786D549FF9A0068FCC2FDA5FA5BDF1D" box="[820,1459,536,557]" gridrow="6" pageId="8" pageNumber="589">
|
||
<th id="0457BC35FF9A0068FCC2FDA5FC0ADF1D" box="[820,994,536,557]" gridcol="0" gridrow="6" pageId="8" pageNumber="589">
|
||
<taxonomicName id="3EB6AC88FF9AFF9FFCC2FDA5FC76DF1D" authorityName="Owen" authorityYear="1849" box="[820,926,536,557]" class="Reptilia" family="Dyrosauridae" genus="Hyposaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="8" pageNumber="589" phylum="Chordata" rank="species" species="rogersii">
|
||
<emphasis id="CBC20B19FF9AFF9FFCC2FDA5FC76DF1D" box="[820,926,536,557]" italics="true" pageId="8" pageNumber="589">H. rogersii</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="0457BC35FF9A0068FBE0FDA5FB81DF1D" box="[1046,1129,536,557]" gridcol="1" gridrow="6" pageId="8" pageNumber="589">42.9</td>
|
||
<td id="0457BC35FF9A0068FB65FDA5FAE9DF1D" box="[1171,1281,536,557]" gridcol="2" gridrow="6" pageId="8" pageNumber="589">28.2</td>
|
||
<td id="0457BC35FF9A0068FADCFDA5FA5BDF1D" box="[1322,1459,536,557]" gridcol="3" gridrow="6" pageId="8" pageNumber="589">65.73</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF9AFF9FFF7AFE17FDB4DF33" blockId="8.[102,775,232,613]" pageId="8" pageNumber="589">
|
||
The mandible is wider than high (
|
||
<figureCitation id="618DCB8EFF9AFF9FFDE7FE17FD89DCF3" box="[529,609,426,451]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="8" pageNumber="589">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="618DCB8EFF9AFF9FFD86FE17FD5EDCF3" box="[624,694,426,451]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEK−GE 1200 in ventral view. B. OCP DEK−GE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." figureDoi="http://doi.org/10.5281/zenodo.13620813" httpUri="https://zenodo.org/record/13620813/files/figure.png" pageId="8" pageNumber="589">8B, C</figureCitation>
|
||
), lacks festooned outlines, and there are occlusal pits posterior to the fifteenth teeth (OCP DEK−GE 269;
|
||
<figureCitation id="618DCB8EFF9AFF9FFE0EFE57FDA6DF33" box="[504,590,490,515]" captionStart="Fig" captionStartId="9.[102,135,926,946]" captionTargetBox="[246,1339,230,896]" captionTargetId="figure-560@9.[241,1347,225,899]" captionTargetPageId="9" captionText="Fig. 8. Arambourgisuchus khouribgaensis gen. et sp. nov., Sidi Chenane, Morocco, late Palaeocene.Elements of mandibles. A. OCP DEK−GE 1200 in ventral view. B. OCP DEK−GE 269 in dorsal (B1) and lateral (B2) views. The arrows indicate the teeth that have been added recently, and which do not belong to the same specimen." figureDoi="http://doi.org/10.5281/zenodo.13620813" httpUri="https://zenodo.org/record/13620813/files/figure.png" pageId="8" pageNumber="589">Fig. 8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF9AFF9FFF7AFDB1FE55DF55" blockId="8.[102,775,232,613]" pageId="8" pageNumber="589">
|
||
The symphysis ends posteriorly at the level of the sixteenth tooth, and the splenial ends anteriorly between the tenth and the eleventh (
|
||
<figureCitation id="618DCB8EFF9AFF9FFE9DFDF1FE58DF55" box="[363,432,588,613]" captionStart="Fig" captionStartId="3.[102,135,1134,1154]" captionTargetBox="[200,1388,233,1080]" captionTargetId="figure-433@3.[298,1372,225,1080]" captionTargetPageId="3" captionText="Fig. 3. Arambourgisuchus khouribgaensis gen. et sp. nov., OCP DEK−GE 18, Sidi Chenane, Morocco, late Palaeocene, skull and mandible in dorsal view. Photograph (A) and explanatory drawing of the same (B)." figureDoi="http://doi.org/10.5281/zenodo.13620803" httpUri="https://zenodo.org/record/13620803/files/figure.png" pageId="8" pageNumber="589">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F909D70BFF9AFF9FFF90FDC4FEFDDE21" blockId="8.[102,775,632,785]" pageId="8" pageNumber="589">
|
||
<emphasis id="CBC20B19FF9AFF9FFF90FDC4FF4CDFA1" box="[102,164,633,657]" italics="true" pageId="8" pageNumber="589">Teeth</emphasis>
|
||
.—They are robust, not very slender, but relatively sharp, with posterior carinae which ends before the base of the teeth, when the anterior one reaches the base. The superficial striae are variably present on some of the teeth, and are absent or weak.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |