173 lines
19 KiB
XML
173 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="39E4A79AABF551FDA178E60435341183" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina pallens Moellendorff 1894" docType="treatment" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
|
|
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
|
|
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
|
|
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
|
|
<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Panha, Somsak</mods:namePart>
|
|
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
|
|
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2022</mods:date>
|
|
<mods:detail type="pubDate">
|
|
<mods:number>2022-08-25</mods:number>
|
|
</mods:detail>
|
|
<mods:detail type="volume">
|
|
<mods:number>1119</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>1</mods:start>
|
|
<mods:end>115</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
|
|
<mods:identifier type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
|
|
<mods:identifier type="Pensoft-UUID">F158A7C5261D52B69288A62F3C777CAF</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-GBIF-Taxon="199838982" LSID="urn:lsid:plazi:treatment:39E4A79AABF551FDA178E60435341183" httpUri="http://treatment.plazi.org/id/39E4A79AABF551FDA178E60435341183" lastPageNumber="1" pageId="0" pageNumber="1">
|
|
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName LSID="39E4A79A-ABF5-51FD-A178-E60435341183" authority="Moellendorff, 1894" authorityName="Moellendorff" authorityYear="1894" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina pallens" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pallens">
|
|
Pupina pallens
|
|
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
|
|
, 1894
|
|
</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="description">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">Figs 21N, O</figureCitation>
|
|
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A Pupina artata, specimen CUMZ 12038 from Tham Suea Temple, Krabi B, C Pupina pallens B lectotype SMF 109951 and C specimen CUMZ 12042 from Suan Wiwek Bureau of Monks, Prachuap Khiri Khan D-F Pupina limitanea D, E syntypes NHMUK 1903.7.1.2967 from East of Burma & Siam and F specimen CUMZ 12044 from Mae Lana junction, Mae Hong Son G Pupina bensoni sp. nov., holotype CUMZ 12045 / 1. Photo: H. Taylor, NHM (D, E)." figureDoi="10.3897/zookeys.1119.85400.figure24" httpUri="https://binary.pensoft.net/fig/735385" pageId="0" pageNumber="1">, 24B, C</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="reference_group">
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Moellendorff" authorityYear="1894" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina pallens" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pallens">Pupina pallens</taxonomicName>
|
|
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
|
|
, 1894: 155, pl. 16, figs 27, 28. Type locality: Samui Islands, Gulf of Siam [Samui Island, Surat Thani Province, Thailand].
|
|
<bibRefCitation author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
|
|
: 431.
|
|
<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
|
|
: 92.
|
|
<bibRefCitation author="Sutcharit, C" journalOrPublisher="Zino Publishing, Bangkok" pageId="0" pageNumber="1" refId="B156" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2018. Land snails: The invaluable bio-resources for the Kingdom of Thailand. Zino Publishing, Bangkok" title="Land snails: The invaluable bio-resources for the Kingdom of Thailand." year="2018">Sutcharit et al. 2018</bibRefCitation>
|
|
: fig. 5-13i.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Moellendorff" authorityYear="1894" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) pallens" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pallens" subGenus="Tylotoechus">Pupina (Tylotoechus) pallens</taxonomicName>
|
|
-
|
|
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
|
|
: 318, 319.
|
|
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
|
|
: 34.
|
|
<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="41 - 56" refId="B198" refString="Zilch, A, 1957. Die Typen und Typoide des Natur-Museums Senckenberg, 19: Mollusca, Cyclophoridae, Pupininae. Archiv fuer Molluskenkunde 86: 41 - 56" title="Die Typen und Typoide des Natur-Museums Senckenberg, 19: Mollusca, Cyclophoridae, Pupininae." volume="86" year="1957">Zilch 1957</bibRefCitation>
|
|
: 47, pl. 2, fig. 16.
|
|
<bibRefCitation author="Hemmen, J" journalOrPublisher="Schriften zur Malakozoologie" pageId="0" pageNumber="1" pagination="35 - 70" refId="B66" refString="Hemmen, J, Hemmen, C, 2001. Aktualisierte Liste der terrestrischen Gastropoden Thailands. Schriften zur Malakozoologie 18: 35 - 70" title="Aktualisierte Liste der terrestrischen Gastropoden Thailands." volume="18" year="2001">Hemmen and Hemmen 2001</bibRefCitation>
|
|
: 39.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="type material">
|
|
<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
|
|
SMF 109951 (Figs
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21N</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A Pupina artata, specimen CUMZ 12038 from Tham Suea Temple, Krabi B, C Pupina pallens B lectotype SMF 109951 and C specimen CUMZ 12042 from Suan Wiwek Bureau of Monks, Prachuap Khiri Khan D-F Pupina limitanea D, E syntypes NHMUK 1903.7.1.2967 from East of Burma & Siam and F specimen CUMZ 12044 from Mae Lana junction, Mae Hong Son G Pupina bensoni sp. nov., holotype CUMZ 12045 / 1. Photo: H. Taylor, NHM (D, E)." figureDoi="10.3897/zookeys.1119.85400.figure24" httpUri="https://binary.pensoft.net/fig/735385" pageId="0" pageNumber="1">24B</figureCitation>
|
|
) and paralectotypes SMF 109952 (4 shells), SMF 109953 (2 shells) from Golf von Siam: Koh Samui.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
|
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
CUMZ 12041 (1 shell) from Bang Phu Temple, Sam Roi Yot District, Prachuap Khiri Khan Province,
|
|
<date value="2020-10-19">19 Oct. 2020</date>
|
|
. CUMZ 12042 (14 shells; Figs
|
|
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Shells of Pupina artata species group from mainland Southeast Asia A-M Pupina artata A syntype UMZC I. 102960. A and specimens B CUMZ 12011 C CUMZ 12023 D CUMZ 12038 E CUMZ 12026 F CUMZ 12015 G CUMZ 12009 H CUMZ 12001 I CUMZ 12003 J NHMUK 1906.4.4.28 K CUMZ 12037 L CUMZ 12031, and M CUMZ 12018 N, O Pupina pallens N lectotype SMF 109951 and O specimen CUMZ 12042 P-R Pupina limitanea P, Q syntypes NHMUK 1903.7.1.2967 and R specimen CUMZ 12044 S Pupina billeti, holotype MNHN-IM- 2000 - 35841 T Pupina verneaui, syntype MNHN-IM- 2000 - 35843 U, V Pupina hungerfordiana, specimens NHMUK 91.3.14.686 - 7 W, X Pupina bensoni sp. nov. W holotype CUMZ 12045 / 1 and X paratype CUMZ 12046 / 1. Photo: H. Taylor, NHM (A, P, Q), P. Maestrati, MNHN (S, T)." figureDoi="10.3897/zookeys.1119.85400.figure21" httpUri="https://binary.pensoft.net/fig/735382" pageId="0" pageNumber="1">21O</figureCitation>
|
|
,
|
|
<figureCitation captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. A Pupina artata, specimen CUMZ 12038 from Tham Suea Temple, Krabi B, C Pupina pallens B lectotype SMF 109951 and C specimen CUMZ 12042 from Suan Wiwek Bureau of Monks, Prachuap Khiri Khan D-F Pupina limitanea D, E syntypes NHMUK 1903.7.1.2967 from East of Burma & Siam and F specimen CUMZ 12044 from Mae Lana junction, Mae Hong Son G Pupina bensoni sp. nov., holotype CUMZ 12045 / 1. Photo: H. Taylor, NHM (D, E)." figureDoi="10.3897/zookeys.1119.85400.figure24" httpUri="https://binary.pensoft.net/fig/735385" pageId="0" pageNumber="1">24C</figureCitation>
|
|
) from Suan Wiwek Bureau of Monks, Sam Roi Yot District, Prachuap Khiri Khan Province,
|
|
<date value="2020-10-21">21 Oct. 2020</date>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">Shell ovate; last whorl ca. three quarters of shell height. Apertural lip slightly thickened, not expanded; basal position widened. Both parietal and columellar teeth fin-shaped and slightly thickened; parietal tooth covering posterior canal; columellar tooth next to slit-like anterior canal.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
|
|
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<taxonomicName authorityName="Moellendorff" authorityYear="1894" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina pallens" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pallens">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pupina pallens</emphasis>
|
|
</taxonomicName>
|
|
can be distinguished from all other species in the
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="artata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
|
|
</taxonomicName>
|
|
species group from mainland Southeast Asia by the widened basal position of the apertural lip.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
The type locality (
|
|
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
|
|
) and Prachuap Khiri Khan Province, western Thailand.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<bibRefCitation author="von Moellendorff, OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="146 - 156" refId="B192" refString="von Moellendorff, OF, 1894. On a collection of land-shells from the Samui Islands, Gulf of Siam. Proceedings of the Zoological Society of London 62: 146 - 156" title="On a collection of land-shells from the Samui Islands, Gulf of Siam." volume="62" year="1894">
|
|
von
|
|
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
|
|
(1894)
|
|
</bibRefCitation>
|
|
stated that this species is different from
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. arula" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="arula">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. arula</emphasis>
|
|
</taxonomicName>
|
|
in having "the more obtuse spire, the more distorted last whorl, and consequently the aperture placed more to the right and protracted at the base, the thinner outer peristome, the broader columella, the broad triangular parietal lamella, and the narrower lower incision". More sampling of this species, with both morphometric and molecular phylogenetic analyses, are needed to resolve the relationship between
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. pallens" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="pallens">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. pallens</emphasis>
|
|
</taxonomicName>
|
|
and other species in the
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. artata" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="artata">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. artata</emphasis>
|
|
</taxonomicName>
|
|
species group.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |