treatments-xml/data/39/B1/97/39B19787089453358F3403B9F3CE2DD3.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.926.48745" ID-GBIF-Dataset="b64e4708-0cb5-407c-a5fe-68ff80068598" ID-PMC="PMC7170979" ID-Pensoft-Pub="1313-2970-926-81" ID-Pensoft-UUID="05C7D06F79EE57818F1FEC5600CC45FF" ID-PubMed="32336921" ID-ZooBank="5E83DB403A11442E85F36C8DCCF487E4" ModsDocID="1313-2970-926-81" checkinTime="1586879005274" checkinUser="pensoft" docAuthor="Song, Young Sun &amp; Kim, Jin-Koo" docDate="2020" docId="39B19787089453358F3403B9F3CE2DD3" docLanguage="en" docName="ZooKeys 926: 81-94" docOrigin="ZooKeys 926" docSource="http://dx.doi.org/10.3897/zookeys.926.48745" docTitle="Eptatretus wandoensis Song &amp; Kim 2020, sp. nov." docType="treatment" docUuid="9C6CA8CC-BC42-48E2-87EE-47702CC46D49" docUuidSource="ZooBank" docVersion="4" id="05C7D06F79EE57818F1FEC5600CC45FF" lastPageNumber="81" masterDocId="05C7D06F79EE57818F1FEC5600CC45FF" masterDocTitle="A new species of hagfish, Eptatretus wandoensis sp. nov. (Agnatha, Myxinidae), from the southwestern Sea of Korea" masterLastPageNumber="94" masterPageNumber="81" pageNumber="81" updateTime="1668168720297" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A new species of hagfish, Eptatretus wandoensis sp. nov. (Agnatha, Myxinidae), from the southwestern Sea of Korea</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Song, Young Sun</mods:namePart>
<mods:affiliation>Department of Marine Biology, Pukyong National University, Busan, 48513, Korea</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kim, Jin-Koo</mods:namePart>
<mods:affiliation>Department of Marine Biology, Pukyong National University, Busan, 48513, Korea</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-8499-406X</mods:nameIdentifier>
<mods:nameIdentifier type="email">taengko@hanmail.net</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>926</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>81</mods:start>
<mods:end>94</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.926.48745</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.926.48745</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-926-81</mods:identifier>
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<treatment ID-GBIF-Taxon="163626717" LSID="urn:lsid:zoobank.org:act:9C6CA8CC-BC42-48E2-87EE-47702CC46D49" httpUri="http://treatment.plazi.org/id/39B19787089453358F3403B9F3CE2DD3" lastPageNumber="81" pageId="0" pageNumber="81">
<subSubSection pageId="0" pageNumber="81" type="nomenclature">
<paragraph pageId="0" pageNumber="81">
<taxonomicName LSID="39B19787-0894-5335-8F34-03B9F3CE2DD3" authority="Song &amp; Kim, 2020" authorityName="Song &amp; Kim" authorityYear="2020" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus wandoensis" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="wandoensis" status="sp. nov.">Eptatretus wandoensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="81">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sampling location of Eptatretus wandoensis sp. nov. in Korea." figureDoi="10.3897/zookeys.926.48745.figure1" httpUri="https://binary.pensoft.net/fig/399904" pageId="0" pageNumber="81">Figures 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Overall view of Eptatretus wandoensis sp. nov., A holotype, PKU 62167, 292.0 mm in total length (TL) B paratype, PKU 62169, 202.0 mm TL C paratype, PKU 62171, 290.0 mm TL D paratype, PKU 62173, 275.0 mm TL, photographed prior to preservation. Scale bars: 1 mm." figureDoi="10.3897/zookeys.926.48745.figure2" httpUri="https://binary.pensoft.net/fig/399905" pageId="0" pageNumber="81">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eptatretus wandoensis sp. nov. PKU 62167, prior to preservation A head dorsal view, white arrow indicates a white mid-dorsal line B head ventral view C gill apertures (GA) and pharyngocutaneous duct (PCD), note the location of GA and PCD D ventral view of body, black arrow indicates the ventral fin-fold (VFF) E dorsal region of body with a white mid-dorsal line." figureDoi="10.3897/zookeys.926.48745.figure3" httpUri="https://binary.pensoft.net/fig/399906" pageId="0" pageNumber="81">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Anatomical morphology of Eptatretus wandoensis sp. nov. PKU 62167. A palatine tooth, anterior unicusps (AUC) and posterior unicusps (PUC) B branchial region dissected, gill pouches (GP), median ventral aorta (MVA), separated ventral aorta (SVA), efferent branchial duct (EBD), pharyngocutaneous duct (PCD), dental muscle (DM), liver (L), and heart (H)." figureDoi="10.3897/zookeys.926.48745.figure4" httpUri="https://binary.pensoft.net/fig/399907" pageId="0" pageNumber="81">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Phylogenetic tree of hagfishes based on mitochondrial DNA sequences, constructed with the Neighbor-joining (NJ) method using Kimura 2 - parameter distances A mitochondrial DNA 16 S rRNA sequences B mitochondrial DNA cytochrome c oxidase subunit I (COI) sequences. Numbers above tree branches are bootstrap values based on 1000 replicates. Scale bar represent nucleotide substitutions per site." figureDoi="10.3897/zookeys.926.48745.figure5" httpUri="https://binary.pensoft.net/fig/399908" pageId="0" pageNumber="81">, 5</figureCitation>
, 
<figureCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric and meristic measurements of Eptatretus wandoensis sp. nov., and congeneric species with five gill apertures (E. cheni, E. nelsoni and E. yangi) and white mid-dorsal line (E. burgeri and E. minor)." pageId="0" pageNumber="81">Table 1 New English name: Five-gilled white mid-dorsal line hagfish; new Korean name: Huin-jul-wae-meok-jang-eo</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="type locality">
<paragraph pageId="0" pageNumber="81">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="81">
The coast of Yeoseo-do (southwestern Sea of Korea): Yeoseo-ri, Cheongsan-myeon, Wando-gun, Jeollanam-do, Republic of Korea, 
<geoCoordinate degrees="33" direction="north" minutes="59" orientation="latitude" precision="1" seconds="56.5" value="33.999027">33°59'56.5&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="126" direction="east" minutes="53" orientation="longitude" precision="1" seconds="57.0" value="126.89916">126°53'57.0&quot;E</geoCoordinate>
, caught by fishing traps, 60-80 m (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sampling location of Eptatretus wandoensis sp. nov. in Korea." figureDoi="10.3897/zookeys.926.48745.figure1" httpUri="https://binary.pensoft.net/fig/399904" pageId="0" pageNumber="81">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="holotype">
<paragraph pageId="0" pageNumber="81">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="81">
PKU 62167, 292.0 mm TL, Yeoseo-ri, Cheongsan-myeon, Wando-gun, Jeollanam-do, Republic of Korea, 
<geoCoordinate degrees="33" direction="north" minutes="59" orientation="latitude" precision="1" seconds="56.5" value="33.999027">33°59'56.5&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="126" direction="east" minutes="53" orientation="longitude" precision="1" seconds="57.0" value="126.89916">126°53'57.0&quot;E</geoCoordinate>
, caught by fishing traps, 60-80 m, 26 Jun 2018.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="paratypes">
<paragraph pageId="0" pageNumber="81">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="81">
PKU 62169 (1 specimen), 202.0 mm TL, Yeoseo-ri, Cheongsan-myeon, Wando-gun, Jeollanam-do, Republic of Korea, 
<geoCoordinate degrees="33" direction="north" minutes="59" orientation="latitude" precision="1" seconds="56.5" value="33.999027">33°59'56.5&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="126" direction="east" minutes="53" orientation="longitude" precision="1" seconds="57.0" value="126.89916">126°53'57.0&quot;E</geoCoordinate>
, fishing trap, 60-80 m, 12 May 2018; PKU 62171, PKU 62173, 275.0-290.0 mm TL, Yeoseo-ri, Cheongsan-myeon, Wando-gun, Jeollanam-do, Republic of Korea, 
<geoCoordinate degrees="33" direction="north" minutes="59" orientation="latitude" precision="1" seconds="56.5" value="33.999027">33°59'56.5&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="126" direction="east" minutes="53" orientation="longitude" precision="1" seconds="57.0" value="126.89916">126°53'57.0&quot;E</geoCoordinate>
, fishing trap, 60-80 m, 26 Jun 2018.
</paragraph>
<caption doi="10.3897/zookeys.926.48745.figure1" httpUri="https://binary.pensoft.net/fig/399904" pageId="0" pageNumber="81" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figure 1.</emphasis>
Sampling location of 
<taxonomicName authorityName="Song &amp; Kim" authorityYear="2020" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus wandoensis" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus wandoensis</emphasis>
</taxonomicName>
sp. nov. in Korea.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="diagnosis">
<paragraph pageId="0" pageNumber="81">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="81">Gill apertures, 5; eyespots, conspicuous; fused cusps, 3/2; total cusps, 40-43; 1 GP at end of dental muscle; total slime pores, 74-82 (prebranchial, 14-18; branchial, 4; trunk, 46-49; tail, 9-11); branchial length, 5.2%-6.2% of TL; pharyngocutaneous duct confluent with last gill aperture; ventral artery splitting at approximately 3-4 GP; dorsal region with dark brown body color, ventral region with white body color; white mid-dorsal line, conspicuous.</paragraph>
<caption ID-Table-UUID="70A097ED6F5468D04B603B0FD927841F" httpUri="http://table.plazi.org/id/70A097ED6F5468D04B603B0FD927841F" pageId="0" pageNumber="81" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Table 1.</emphasis>
Morphometric and meristic measurements of 
<taxonomicName authorityName="Song &amp; Kim" authorityYear="2020" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus wandoensis" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus wandoensis</emphasis>
</taxonomicName>
sp. nov., and congeneric species with five gill apertures (
<taxonomicName lsidName="E. cheni" pageId="0" pageNumber="81" rank="species" species="cheni">
<emphasis italics="true" pageId="0" pageNumber="81">E. cheni</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="E. nelsoni" pageId="0" pageNumber="81" rank="species" species="nelsoni">
<emphasis italics="true" pageId="0" pageNumber="81">E. nelsoni</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="E. yangi" pageId="0" pageNumber="81" rank="species" species="yangi">
<emphasis italics="true" pageId="0" pageNumber="81">E. yangi</emphasis>
</taxonomicName>
) and white mid-dorsal line (
<taxonomicName lsidName="E. burgeri" pageId="0" pageNumber="81" rank="species" species="burgeri">
<emphasis italics="true" pageId="0" pageNumber="81">E. burgeri</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="E. minor" pageId="0" pageNumber="81" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="81">E. minor</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="81">
<table pageId="0" pageNumber="81">
<tr pageId="0" pageNumber="81">
<th colspan="1" isEmpty="true" pageId="0" pageNumber="81" rowspan="2">-</th>
<th colspan="2" pageId="0" pageNumber="81" rowspan="1">
<taxonomicName authorityName="Song &amp; Kim" authorityYear="2020" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus wandoensis" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus wandoensis</emphasis>
</taxonomicName>
sp. nov.
</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="2">
<taxonomicName lsidName="E. cheni" pageId="0" pageNumber="81" rank="species" species="cheni">
<emphasis italics="true" pageId="0" pageNumber="81">E. cheni</emphasis>
</taxonomicName>
*
</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="2">
<taxonomicName lsidName="E. nelsoni" pageId="0" pageNumber="81" rank="species" species="nelsoni">
<emphasis italics="true" pageId="0" pageNumber="81">E. nelsoni</emphasis>
</taxonomicName>
*
</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="2">
<taxonomicName lsidName="E. yangi" pageId="0" pageNumber="81" rank="species" species="yangi">
<emphasis italics="true" pageId="0" pageNumber="81">E. yangi</emphasis>
</taxonomicName>
*
</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="2">
<taxonomicName lsidName="E. burgeri" pageId="0" pageNumber="81" rank="species" species="burgeri">
<emphasis italics="true" pageId="0" pageNumber="81">E. burgeri</emphasis>
</taxonomicName>
*
</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="2">
<taxonomicName lsidName="E. minor" pageId="0" pageNumber="81" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="81">E. minor</emphasis>
</taxonomicName>
**
</th>
</tr>
<tr pageId="0" pageNumber="81">
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">Holotype</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">Paratypes (3)</th>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Gill aperture (GA)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6-7</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Gill pouch (GP)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">NSP</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">paired</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="8" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">Cusps</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">MUC (multi)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3/2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3/2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3/3</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3/2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3/2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3/2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3/3</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">AUC (outer)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7-8</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">9-11</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5-8</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5-8</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6-8</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">8-11</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">PUC (inner)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">8</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">8-9</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">9-10</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5-8</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6-9</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7-9</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">8-10</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Total Cusps</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">42</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">40-43</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">50-53</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">32-40</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">32-40</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">35-42</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">46-54</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="8" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">Slime pores</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Prebranchial</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">14</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">15-18</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">24-27</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">13-20</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">16-23</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">18-23</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">15-18</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Branchial</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">4-5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">4-6</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Trunk</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">47</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">46-49</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">41-47</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">33-39</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">39-47</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">45-51</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">41-48</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Tail</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">11</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">9-11</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7-10</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6-10</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7-12</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">11-14</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">11-14</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">Total pores</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">76</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">74-82</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">75-81</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">57-67</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">68-79</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">81-92</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">74-82</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="8" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">Length in % of TL</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Prebranchial</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">24.7</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">24.4-26.3 (25.3)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">33.3-35.5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">30.5-32.6</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">29.2-32.0</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">25.2-29.6</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">20.1-25.9</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Branchial</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5.9</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5.2-5.8 (5.5)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">2.2-3.4</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">1.1-2.8</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">1.1-1.7</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6.2-7.8</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5.1-7.2</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Trunk</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">56.5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">54.9-59.3 (56.9)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">45.9-50.8</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">49.5-52.6</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">53.2-54.9</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">47.6-55.0</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">50.6-55.9</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Tail</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">13.4</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">12.8-14.0 (13.5)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">13.2-16.7</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">15.0-18.0</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">12.2-15.6</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">13.2-17.0</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">13.9-18.3</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Nostril to mouth</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3.4</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3.7-3.8 (3.8)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Nostril width</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">1.7</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">0.8-1.5 (1.2)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Nostril length</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">0.6</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">0.7-1.6 (1.1)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Mouth width</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3.2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3.3-3.8 (3.6)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Pre-eyespot to nostril</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">4.9</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">4.4-5.2 (4.8)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="8" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">Depth in % of TL</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">w/VFF</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7.5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6.9-9.7 (8.3)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">8.1-9.0</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">15.0-15.5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6.9-10.4</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">4.7-8.5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7.1-11.4</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Branchial region</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6.3</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5.6-7.7 (6.7)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Over caudal</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7.8</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7.6-9.3 (8.6)</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7.6-10.2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">8.9-10.1</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6.5-10.0</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5.1-8.5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5.3-11.6</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="81">
<tableNote pageId="0" pageNumber="81">
* 
<bibRefCitation author="McMillan, CB" journalOrPublisher="Zoological Studies" pageId="0" pageNumber="81" pagination="51 - 73" refId="B20" refString="McMillan, CB, Wisner, RL, 2004. Review of the Hagfishes (Myxinidae: Myxiniformes) of the Northwestern Pacific Ocean, with Descriptions of Three New Species, Eptatretus fernholmi, Paramyxine moki, P. walkeri. Zoological Studies 43 (1): 51 - 73" title="Review of the Hagfishes (Myxinidae: Myxiniformes) of the Northwestern Pacific Ocean, with Descriptions of Three New Species, Eptatretus fernholmi, Paramyxine moki, P. walkeri." volume="43" year="2004">McMillan and Wisner (2004)</bibRefCitation>
, ** 
<bibRefCitation author="Fernholm, B" journalOrPublisher="Fishery Bulletin" pageId="0" pageNumber="81" pagination="69 - 83" refId="B4" refString="Fernholm, B, Hubbs, CL, 1981. Western Atlantic hagfishes of the genus Eptatretus (Myxinidae) with description of two new species. Fishery Bulletin 79: 69 - 83" title="Western Atlantic hagfishes of the genus Eptatretus (Myxinidae) with description of two new species." volume="79" year="1981">Fernholm and Hubbs (1981)</bibRefCitation>
; Abbreviation: NSP (nasal-sinus papillae), MUC (multicusps), AUC (anterior unicusps), PUC (posterior unicusps), VFF (ventral fin-fold).
</tableNote>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="description">
<paragraph pageId="0" pageNumber="81">Description.</paragraph>
<paragraph pageId="0" pageNumber="81">
Body elongated; laterally compressed at trunk and strongly compressed at tail (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Overall view of Eptatretus wandoensis sp. nov., A holotype, PKU 62167, 292.0 mm in total length (TL) B paratype, PKU 62169, 202.0 mm TL C paratype, PKU 62171, 290.0 mm TL D paratype, PKU 62173, 275.0 mm TL, photographed prior to preservation. Scale bars: 1 mm." figureDoi="10.3897/zookeys.926.48745.figure2" httpUri="https://binary.pensoft.net/fig/399905" pageId="0" pageNumber="81">2</figureCitation>
). Rostrum slightly blunt and round (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eptatretus wandoensis sp. nov. PKU 62167, prior to preservation A head dorsal view, white arrow indicates a white mid-dorsal line B head ventral view C gill apertures (GA) and pharyngocutaneous duct (PCD), note the location of GA and PCD D ventral view of body, black arrow indicates the ventral fin-fold (VFF) E dorsal region of body with a white mid-dorsal line." figureDoi="10.3897/zookeys.926.48745.figure3" httpUri="https://binary.pensoft.net/fig/399906" pageId="0" pageNumber="81">3A</figureCitation>
). Nasal-sinus papilla absent. Eyespots present (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eptatretus wandoensis sp. nov. PKU 62167, prior to preservation A head dorsal view, white arrow indicates a white mid-dorsal line B head ventral view C gill apertures (GA) and pharyngocutaneous duct (PCD), note the location of GA and PCD D ventral view of body, black arrow indicates the ventral fin-fold (VFF) E dorsal region of body with a white mid-dorsal line." figureDoi="10.3897/zookeys.926.48745.figure3" httpUri="https://binary.pensoft.net/fig/399906" pageId="0" pageNumber="81">3A</figureCitation>
). Pre-eyespots shorter than branchial region (4.4%-4.9% of TL). Three pairs of barbels on head: first (1.5%-1.6% of TL) and second barbels (1.6%-1.9% of TL) nearly equal in size; third barbel is longer (2.1%-2.3% of TL) and tips of third barbels extend at the mouth (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eptatretus wandoensis sp. nov. PKU 62167, prior to preservation A head dorsal view, white arrow indicates a white mid-dorsal line B head ventral view C gill apertures (GA) and pharyngocutaneous duct (PCD), note the location of GA and PCD D ventral view of body, black arrow indicates the ventral fin-fold (VFF) E dorsal region of body with a white mid-dorsal line." figureDoi="10.3897/zookeys.926.48745.figure3" httpUri="https://binary.pensoft.net/fig/399906" pageId="0" pageNumber="81">3B</figureCitation>
). Five pairs of GP and apertures; each gill aperture arranged regularly spaced in a straight line (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eptatretus wandoensis sp. nov. PKU 62167, prior to preservation A head dorsal view, white arrow indicates a white mid-dorsal line B head ventral view C gill apertures (GA) and pharyngocutaneous duct (PCD), note the location of GA and PCD D ventral view of body, black arrow indicates the ventral fin-fold (VFF) E dorsal region of body with a white mid-dorsal line." figureDoi="10.3897/zookeys.926.48745.figure3" httpUri="https://binary.pensoft.net/fig/399906" pageId="0" pageNumber="81">3C</figureCitation>
). Teeth row comb-like, consisting of two rows with tips sharp and curved rearward (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Anatomical morphology of Eptatretus wandoensis sp. nov. PKU 62167. A palatine tooth, anterior unicusps (AUC) and posterior unicusps (PUC) B branchial region dissected, gill pouches (GP), median ventral aorta (MVA), separated ventral aorta (SVA), efferent branchial duct (EBD), pharyngocutaneous duct (PCD), dental muscle (DM), liver (L), and heart (H)." figureDoi="10.3897/zookeys.926.48745.figure4" httpUri="https://binary.pensoft.net/fig/399907" pageId="0" pageNumber="81">4A</figureCitation>
); in the outer row, 3 multicusps and 7-8 unicusps; in the inner row, 2 multicusps and 8-9 unicusps; total number of cusps, 40-43. Dental muscle thick and long, posterior tip of dental muscle located in first GP (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Anatomical morphology of Eptatretus wandoensis sp. nov. PKU 62167. A palatine tooth, anterior unicusps (AUC) and posterior unicusps (PUC) B branchial region dissected, gill pouches (GP), median ventral aorta (MVA), separated ventral aorta (SVA), efferent branchial duct (EBD), pharyngocutaneous duct (PCD), dental muscle (DM), liver (L), and heart (H)." figureDoi="10.3897/zookeys.926.48745.figure4" httpUri="https://binary.pensoft.net/fig/399907" pageId="0" pageNumber="81">4B</figureCitation>
). Slime pores: prebranchial, 14-18; branchial, 4; trunk, 46-49; tail, 9-11; total, 74-82. Body proportions are as follows: prebranchial length, 24.4%-26.3% of TL; branchial length, 5.2%-6.2% of TL; trunk length, 54.9%-59.3% of TL; tail length, 12.8%-14.0% of TL; cutaneous duct, 7.6%-9.3% of TL; branchial duct (with ventral fin-fold), 6.9%-9.7% of TL; and branchial duct (alone), 5.6%-7.7% of TL (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric and meristic measurements of Eptatretus wandoensis sp. nov., and congeneric species with five gill apertures (E. cheni, E. nelsoni and E. yangi) and white mid-dorsal line (E. burgeri and E. minor)." httpUri="http://table.plazi.org/id/70A097ED6F5468D04B603B0FD927841F" pageId="0" pageNumber="81" tableUuid="70A097ED6F5468D04B603B0FD927841F">1</tableCitation>
). Posterior-most EBD confluent with pharyngocutaneous duct on left side, forming a larger aperture (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Anatomical morphology of Eptatretus wandoensis sp. nov. PKU 62167. A palatine tooth, anterior unicusps (AUC) and posterior unicusps (PUC) B branchial region dissected, gill pouches (GP), median ventral aorta (MVA), separated ventral aorta (SVA), efferent branchial duct (EBD), pharyngocutaneous duct (PCD), dental muscle (DM), liver (L), and heart (H)." figureDoi="10.3897/zookeys.926.48745.figure4" httpUri="https://binary.pensoft.net/fig/399907" pageId="0" pageNumber="81">4B</figureCitation>
). All efferent branchial ducts are equal in length. VA consists of two SBAs and one medial section, bifurcating at approximately the third or fourth GP. First through third pairs of ABAs, which cannot be regarded as branches of the VA, branch from SBAs; however, fourth and fifth ABAs on left and right branch from the medial section of the ventral artery (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Anatomical morphology of Eptatretus wandoensis sp. nov. PKU 62167. A palatine tooth, anterior unicusps (AUC) and posterior unicusps (PUC) B branchial region dissected, gill pouches (GP), median ventral aorta (MVA), separated ventral aorta (SVA), efferent branchial duct (EBD), pharyngocutaneous duct (PCD), dental muscle (DM), liver (L), and heart (H)." figureDoi="10.3897/zookeys.926.48745.figure4" httpUri="https://binary.pensoft.net/fig/399907" pageId="0" pageNumber="81">4B</figureCitation>
). Ventral fin-fold weakly developed or vestigial, beginning approximately at middle of body and extending to cloaca (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Eptatretus wandoensis sp. nov. PKU 62167, prior to preservation A head dorsal view, white arrow indicates a white mid-dorsal line B head ventral view C gill apertures (GA) and pharyngocutaneous duct (PCD), note the location of GA and PCD D ventral view of body, black arrow indicates the ventral fin-fold (VFF) E dorsal region of body with a white mid-dorsal line." figureDoi="10.3897/zookeys.926.48745.figure3" httpUri="https://binary.pensoft.net/fig/399906" pageId="0" pageNumber="81">3D</figureCitation>
). Caudal fin-fold weakly developed, beginning posterior cloaca and extending around tail to dorsal surface.
</paragraph>
<caption doi="10.3897/zookeys.926.48745.figure2" httpUri="https://binary.pensoft.net/fig/399905" pageId="0" pageNumber="81" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figure 2.</emphasis>
Overall view of 
<taxonomicName authorityName="Song &amp; Kim" authorityYear="2020" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus wandoensis" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus wandoensis</emphasis>
</taxonomicName>
sp. nov., 
<emphasis bold="true" pageId="0" pageNumber="81">A</emphasis>
holotype, PKU 62167, 292.0 mm in total length (TL) 
<emphasis bold="true" pageId="0" pageNumber="81">B</emphasis>
paratype, PKU 62169, 202.0 mm TL 
<emphasis bold="true" pageId="0" pageNumber="81">C</emphasis>
paratype, PKU 62171, 290.0 mm TL 
<emphasis bold="true" pageId="0" pageNumber="81">D</emphasis>
paratype, PKU 62173, 275.0 mm TL, photographed prior to preservation. Scale bars: 1 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.926.48745.figure3" httpUri="https://binary.pensoft.net/fig/399906" pageId="0" pageNumber="81" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figure 3.</emphasis>
<taxonomicName authorityName="Song &amp; Kim" authorityYear="2020" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus wandoensis" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus wandoensis</emphasis>
</taxonomicName>
sp. nov. PKU 62167, prior to preservation 
<emphasis bold="true" pageId="0" pageNumber="81">A</emphasis>
head dorsal view, white arrow indicates a white mid-dorsal line 
<emphasis bold="true" pageId="0" pageNumber="81">B</emphasis>
head ventral view 
<emphasis bold="true" pageId="0" pageNumber="81">C</emphasis>
gill apertures (GA) and pharyngocutaneous duct (PCD), note the location of GA and PCD 
<emphasis bold="true" pageId="0" pageNumber="81">D</emphasis>
ventral view of body, black arrow indicates the ventral fin-fold (VFF) 
<emphasis bold="true" pageId="0" pageNumber="81">E</emphasis>
dorsal region of body with a white mid-dorsal line.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.926.48745.figure4" httpUri="https://binary.pensoft.net/fig/399907" pageId="0" pageNumber="81" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figure 4.</emphasis>
Anatomical morphology of 
<taxonomicName authorityName="Song &amp; Kim" authorityYear="2020" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus wandoensis" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus wandoensis</emphasis>
</taxonomicName>
sp. nov. PKU 62167. 
<emphasis bold="true" pageId="0" pageNumber="81">A</emphasis>
palatine tooth, anterior unicusps (AUC) and posterior unicusps (PUC) 
<emphasis bold="true" pageId="0" pageNumber="81">B</emphasis>
branchial region dissected, gill pouches (GP), median ventral aorta (MVA), separated ventral aorta (SVA), efferent branchial duct (EBD), pharyngocutaneous duct (PCD), dental muscle (DM), liver (L), and heart (H).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="81">
Coloration when fresh: Body uniformly dark brown or purplish dorsally and white ventrally; white mid-dorsal line conspicuous, beginning from the upper region of the first prebranchial slime pore to around the tail. Eyespots conspicuous; whole barbels (rarely the tip) pale, and pale around mouth. Each gill aperture and pharyngocutaneous duct aperture with white margin; most slime pores blackish (except for tail region), tail slime pores same as surrounding color. White around cloaca; ventral fin-fold with a white line along the ventral midline; posterior margin of caudal fin pale (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Overall view of Eptatretus wandoensis sp. nov., A holotype, PKU 62167, 292.0 mm in total length (TL) B paratype, PKU 62169, 202.0 mm TL C paratype, PKU 62171, 290.0 mm TL D paratype, PKU 62173, 275.0 mm TL, photographed prior to preservation. Scale bars: 1 mm." figureDoi="10.3897/zookeys.926.48745.figure2" httpUri="https://binary.pensoft.net/fig/399905" pageId="0" pageNumber="81">2</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="81">Coloration when preserved: Body brown to dark brown dorsally and murky white ventrally (more conspicuous than fresh specimen). Eyespots conspicuous; all slime pores surrounded by conspicuous white ring. Each gill aperture and pharyngocutaneous duct aperture conspicuous; ventral fin-fold pale; white mid-dorsal line inconspicuous.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="distribution">
<paragraph pageId="0" pageNumber="81">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="81">Southwestern Sea of Korea.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="biology_ecology">
<paragraph pageId="0" pageNumber="81">Biology.</paragraph>
<paragraph pageId="0" pageNumber="81">Attains a maximum TL of 292.0 mm (fresh specimen); this specimen is female, without mature eggs in the body cavity. A female specimen of 290.0 mm TL carries approximately 20 developing eggs, which have no terminal anchor filaments or hooks; each egg approximately 4-7 mm in diameter and 10-12 mm in length.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="etymology">
<paragraph pageId="0" pageNumber="81">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="81">
The specific name, 
<taxonomicName authorityName="Song &amp; Kim" authorityYear="2020" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="wandoensis" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">wandoensis</emphasis>
</taxonomicName>
, refers to the type locality, in Korea.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="morphological comparisons">
<paragraph pageId="0" pageNumber="81">Morphological comparisons</paragraph>
<paragraph pageId="0" pageNumber="81">
<taxonomicName authorityName="Song &amp; Kim" authorityYear="2020" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus wandoensis" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus wandoensis</emphasis>
</taxonomicName>
sp. nov. is most similar to 
<taxonomicName genus="Eptatetus" lsidName="Eptatetus burgeri" pageId="0" pageNumber="81" rank="species" species="burgeri">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatetus burgeri</emphasis>
</taxonomicName>
(Girard, 1855) and 
<taxonomicName authorityName="Fernholm &amp; Hubbs" authorityYear="1981" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus minor" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus minor</emphasis>
</taxonomicName>
Fernholm &amp; Hubbs, 1981 due to the presence of a light mid-dorsal line, gill apertures regularly spaced in a straight line, and EBDs of equal length. These three species differ from each other in the number of gill apertures (5 for 
<taxonomicName lsidName="E. wandoensis" pageId="0" pageNumber="81" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">E. wandoensis</emphasis>
</taxonomicName>
, compared to 6 for 
<taxonomicName lsidName="E. burgeri" pageId="0" pageNumber="81" rank="species" species="burgeri">
<emphasis italics="true" pageId="0" pageNumber="81">E. burgeri</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="E. minor" pageId="0" pageNumber="81" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="81">E. minor</emphasis>
</taxonomicName>
), body color (dark brown or purplish dorsally and white ventrally for 
<taxonomicName lsidName="E. wandoensis" pageId="0" pageNumber="81" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">E. wandoensis</emphasis>
</taxonomicName>
, compared to brown for 
<taxonomicName lsidName="E. burgeri" pageId="0" pageNumber="81" rank="species" species="burgeri">
<emphasis italics="true" pageId="0" pageNumber="81">E. burgeri</emphasis>
</taxonomicName>
and gray/brown pale for 
<taxonomicName lsidName="E. minor" pageId="0" pageNumber="81" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="81">E. minor</emphasis>
</taxonomicName>
), prebranchial slime pores (14-18 for 
<taxonomicName lsidName="E. wandoensis" pageId="0" pageNumber="81" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">E. wandoensis</emphasis>
</taxonomicName>
, compared to 18-23 for 
<taxonomicName lsidName="E. burgeri" pageId="0" pageNumber="81" rank="species" species="burgeri">
<emphasis italics="true" pageId="0" pageNumber="81">E. burgeri</emphasis>
</taxonomicName>
), total slime pores (74-82 for 
<taxonomicName lsidName="E. wandoensis" pageId="0" pageNumber="81" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">E. wandoensis</emphasis>
</taxonomicName>
, compared to 81-92 for 
<taxonomicName lsidName="E. burgeri" pageId="0" pageNumber="81" rank="species" species="burgeri">
<emphasis italics="true" pageId="0" pageNumber="81">E. burgeri</emphasis>
</taxonomicName>
), ventral fin-fold (weakly developed for 
<taxonomicName lsidName="E. wandoensis" pageId="0" pageNumber="81" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">E. wandoensis</emphasis>
</taxonomicName>
, compared to well developed for 
<taxonomicName lsidName="E. burgeri" pageId="0" pageNumber="81" rank="species" species="burgeri">
<emphasis italics="true" pageId="0" pageNumber="81">E. burgeri</emphasis>
</taxonomicName>
), multicusps (3/2 for 
<taxonomicName lsidName="E. wandoensis" pageId="0" pageNumber="81" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">E. wandoensis</emphasis>
</taxonomicName>
, compared to 3/3 for 
<taxonomicName lsidName="E. minor" pageId="0" pageNumber="81" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="81">E. minor</emphasis>
</taxonomicName>
), total cusps (40-43 for 
<taxonomicName lsidName="E. wandoensis" pageId="0" pageNumber="81" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">E. wandoensis</emphasis>
</taxonomicName>
, compared to 46-52 for 
<taxonomicName lsidName="E. minor" pageId="0" pageNumber="81" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="81">E. minor</emphasis>
</taxonomicName>
) (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphometric and meristic measurements of Eptatretus wandoensis sp. nov., and congeneric species with five gill apertures (E. cheni, E. nelsoni and E. yangi) and white mid-dorsal line (E. burgeri and E. minor)." httpUri="http://table.plazi.org/id/70A097ED6F5468D04B603B0FD927841F" pageId="0" pageNumber="81" tableUuid="70A097ED6F5468D04B603B0FD927841F">1</tableCitation>
), nasal-sinus papillae (absent for 
<taxonomicName lsidName="E. wandoensis" pageId="0" pageNumber="81" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">E. wandoensis</emphasis>
</taxonomicName>
, compared to paired for 
<taxonomicName lsidName="E. minor" pageId="0" pageNumber="81" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="81">E. minor</emphasis>
</taxonomicName>
), and eyespots (present for 
<taxonomicName lsidName="E. wandoensis" pageId="0" pageNumber="81" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">E. wandoensis</emphasis>
</taxonomicName>
, compared to absent for 
<taxonomicName lsidName="E. minor" pageId="0" pageNumber="81" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="81">E. minor</emphasis>
</taxonomicName>
). 
<taxonomicName authorityName="Song &amp; Kim" authorityYear="2020" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus wandoensis" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus wandoensis</emphasis>
</taxonomicName>
sp. nov. can be distinguished from 
<taxonomicName baseAuthorityName="Shen &amp; Tao" baseAuthorityYear="1975" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus cheni" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="cheni">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus cheni</emphasis>
</taxonomicName>
(Shen &amp; Tao, 1975), 
<taxonomicName baseAuthorityName="Kuo, Huang &amp; Mok" baseAuthorityYear="1994" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus nelsoni" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="nelsoni">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus nelsoni</emphasis>
</taxonomicName>
(Kuo, Huang &amp; Mok, 1994), and 
<taxonomicName baseAuthorityName="Teng" baseAuthorityYear="1958" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus yangi" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="yangi">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus yangi</emphasis>
</taxonomicName>
(Teng, 1958) by the presence of regularly spaced gill apertures in a linear (vs. irregular and crowded for 
<taxonomicName lsidName="E. cheni" pageId="0" pageNumber="81" rank="species" species="cheni">
<emphasis italics="true" pageId="0" pageNumber="81">E. cheni</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="E. nelsoni" pageId="0" pageNumber="81" rank="species" species="nelsoni">
<emphasis italics="true" pageId="0" pageNumber="81">E. nelsoni</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="E. yangi" pageId="0" pageNumber="81" rank="species" species="yangi">
<emphasis italics="true" pageId="0" pageNumber="81">E. yangi</emphasis>
</taxonomicName>
) arrangement; equal length of all EBDs (vs. length of first efferent branchial duct notably longer than that of the most posterior efferent branchial duct); 40-43 total cusps (vs. 50-53 for 
<taxonomicName lsidName="E. cheni" pageId="0" pageNumber="81" rank="species" species="cheni">
<emphasis italics="true" pageId="0" pageNumber="81">E. cheni</emphasis>
</taxonomicName>
; 32-40 for 
<taxonomicName lsidName="E. nelsoni" pageId="0" pageNumber="81" rank="species" species="nelsoni">
<emphasis italics="true" pageId="0" pageNumber="81">E. nelsoni</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="E. yangi" pageId="0" pageNumber="81" rank="species" species="yangi">
<emphasis italics="true" pageId="0" pageNumber="81">E. yangi</emphasis>
</taxonomicName>
); 4 branchial slime pores (vs. no branchial slime pores); prebranchial length, 24.4%-26.3% of TL (vs. more than 29.0% of TL); branchial length, 5.2%-5.9% of TL (vs. less than 3.4% of TL); trunk length, 54.9-59.3% of TL (vs. less than 54.9%); eyespots conspicuous (vs. inconspicuous), and dorsal dark brown and ventral white body color (vs. brownish-grey). In comparison to 
<taxonomicName authorityName="Cloquet in Cloquet" authorityYear="1819" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus</emphasis>
</taxonomicName>
species occurring in Korean and Japanese waters, this new species is well distinguished from the three most common hagfishes, 
<taxonomicName baseAuthorityName="Dean" baseAuthorityYear="1904" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus atami" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="atami">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus atami</emphasis>
</taxonomicName>
(Dean, 1904), 
<taxonomicName baseAuthorityName="McMillan &amp; Wisner" baseAuthorityYear="2004" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus walkeri" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="walkeri">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus walkeri</emphasis>
</taxonomicName>
(McMillan &amp; Wisner, 2004), and 
<taxonomicName baseAuthorityName="Dean" baseAuthorityYear="1904" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus okinoseanus" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="okinoseanus">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus okinoseanus</emphasis>
</taxonomicName>
(Dean, 1904) based on the difference of gill apertures (5 in 
<taxonomicName lsidName="E. wandoensis" pageId="0" pageNumber="81" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">E. wandoensis</emphasis>
</taxonomicName>
sp. nov. vs. 6 in 
<taxonomicName lsidName="E. atami" pageId="0" pageNumber="81" rank="species" species="atami">
<emphasis italics="true" pageId="0" pageNumber="81">E. atami</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="E. walkeri" pageId="0" pageNumber="81" rank="species" species="walkeri">
<emphasis italics="true" pageId="0" pageNumber="81">E. walkeri</emphasis>
</taxonomicName>
vs. 8 in 
<taxonomicName lsidName="E. okinoseanus" pageId="0" pageNumber="81" rank="species" species="okinoseanus">
<emphasis italics="true" pageId="0" pageNumber="81">E. okinoseanus</emphasis>
</taxonomicName>
), branchial slime pores (4, 0-1, 0, and 6-8), and a white mid-dorsal line (present, absent, absent, and absent) (Table 
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparison of meristic and proportional measurements among Eptatretus species occurring in Korean and Japanese waters." httpUri="http://table.plazi.org/id/4E1D4A6635B1F7EAC9CD6307826FB677" pageId="0" pageNumber="81" tableUuid="4E1D4A6635B1F7EAC9CD6307826FB677">2</tableCitation>
).
</paragraph>
<caption ID-Table-UUID="4E1D4A6635B1F7EAC9CD6307826FB677" httpUri="http://table.plazi.org/id/4E1D4A6635B1F7EAC9CD6307826FB677" pageId="0" pageNumber="81" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Table 2.</emphasis>
Comparison of meristic and proportional measurements among 
<taxonomicName authorityName="Cloquet in Cloquet" authorityYear="1819" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus</emphasis>
</taxonomicName>
species occurring in Korean and Japanese waters.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="81">
<table pageId="0" pageNumber="81">
<tr pageId="0" pageNumber="81">
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">Characters</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">
<taxonomicName lsidName="E. wandoensis" pageId="0" pageNumber="81" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">E. wandoensis</emphasis>
</taxonomicName>
sp. nov.
</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">
<taxonomicName lsidName="E. atami" pageId="0" pageNumber="81" rank="species" species="atami">
<emphasis italics="true" pageId="0" pageNumber="81">E. atami</emphasis>
</taxonomicName>
*
</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">
<taxonomicName lsidName="E. walkeri" pageId="0" pageNumber="81" rank="species" species="walkeri">
<emphasis italics="true" pageId="0" pageNumber="81">E. walkeri</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">
<taxonomicName lsidName="E. okinoseanus" pageId="0" pageNumber="81" rank="species" species="okinoseanus">
<emphasis italics="true" pageId="0" pageNumber="81">E. okinoseanus</emphasis>
</taxonomicName>
*
</th>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Gill aperture</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">8</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Gill pouch</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">8</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">NSP</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">absent</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="5" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">Cusps</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">MUC</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3/2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3/3</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3/2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3/2</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">AUC</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7-8</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">9-10</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6-9</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7-10</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">PUC</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">8-9</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">8-10</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7-9</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7-10</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Total</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">40-43</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">47-52</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">36-44</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">40-49</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="5" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">Slime pores</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Prebranchial</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">14-18</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">12-19</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">15-22</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">13-17</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Branchial</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">0-1</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">0</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6-8</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Trunk</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">46-49</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">43-47</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">40-48</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">54-61</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Tail</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">9-11</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">9-12</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">8-13</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">10-14</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">Total pores</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">74-82</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">71-78</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">68-79</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">87-97</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="5" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">Length in % of TL</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Prebranchial</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">24.4-26.3</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">26.6-30.2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">24.2-39.1</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">19.2-22.6</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Branchial</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5.2-5.9</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">1.3-4.2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">2.0-3.8</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6.2-9.2</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Trunk</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">54.9-59.3</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">53.9-56.1</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">50.8-68.6</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">50.4-59.4</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Tail</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">12.8-14.0</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">11.1-14.2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">10.7-16.1</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">12.7-15.5</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="5" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">Depth (mm)</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">w/VFF</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6.9-9.7</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">8.1-9.0</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5.0-11.1</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5.7-8.1</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">Over caudal</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7.6-9.3</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">7.4-8.8</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6.3-11.4</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">6.2-9.0</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="81">
<tableNote pageId="0" pageNumber="81">
* 
<bibRefCitation author="McMillan, CB" journalOrPublisher="Zoological Studies" pageId="0" pageNumber="81" pagination="51 - 73" refId="B20" refString="McMillan, CB, Wisner, RL, 2004. Review of the Hagfishes (Myxinidae: Myxiniformes) of the Northwestern Pacific Ocean, with Descriptions of Three New Species, Eptatretus fernholmi, Paramyxine moki, P. walkeri. Zoological Studies 43 (1): 51 - 73" title="Review of the Hagfishes (Myxinidae: Myxiniformes) of the Northwestern Pacific Ocean, with Descriptions of Three New Species, Eptatretus fernholmi, Paramyxine moki, P. walkeri." volume="43" year="2004">McMillan and Wisner (2004)</bibRefCitation>
; Abbreviation: NSP (nasal-sinus papillae), MUC (multicusps), AUC (anterior unicusps), PUC (posterior unicusps), VFF (ventral fin-fold).
</tableNote>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="genetic comparisons">
<paragraph pageId="0" pageNumber="81">Genetic comparisons</paragraph>
<paragraph pageId="0" pageNumber="81">
Differences among mtDNA sequences obtained from the holotype and paratypes of 
<taxonomicName authorityName="Song &amp; Kim" authorityYear="2020" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus wandoensis" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus wandoensis</emphasis>
</taxonomicName>
sp. nov. were consistent with species-level divergences in other hagfish species (
<bibRefCitation DOI="https://doi.org/10.1111/jzs.12035" author="Fernholm, B" journalOrPublisher="Journal of Zoological Systematics and Evolutionary Research" pageId="0" pageNumber="81" pagination="296 - 307" refId="B5" refString="Fernholm, B, Noren, M, Kullander, SO, Quattrini, AM, Zintzen, V, Roberts, CD, Mok, HK, Kuo, CH, 2013. Hagfish phylogeny and taxonomy, with description of the new genus Rubicundus (Craniata, Myxinidae). Journal of Zoological Systematics and Evolutionary Research 51 (4): 296 - 307, DOI: https://doi.org/10.1111/jzs.12035" title="Hagfish phylogeny and taxonomy, with description of the new genus Rubicundus (Craniata, Myxinidae)." url="https://doi.org/10.1111/jzs.12035" volume="51" year="2013">Fernholm et al. 2013</bibRefCitation>
). The phylogenetic relationships of myxinid species, inferred from neighbor-joining trees, showed large genetic distances between similar hagfish species using mtDNA 16S rRNA (477 bp) and cytochrome c oxidase subunit I (COI) (466 bp) sequences. 
<taxonomicName authorityName="Song &amp; Kim" authorityYear="2020" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus wandoensis" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus wandoensis</emphasis>
</taxonomicName>
sp. nov. is separated from other congeneric species by high genetic divergences of 0.9%-7.5% in 16S rRNA sequences and 4.9%-13.9% in COI sequences (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Phylogenetic tree of hagfishes based on mitochondrial DNA sequences, constructed with the Neighbor-joining (NJ) method using Kimura 2 - parameter distances A mitochondrial DNA 16 S rRNA sequences B mitochondrial DNA cytochrome c oxidase subunit I (COI) sequences. Numbers above tree branches are bootstrap values based on 1000 replicates. Scale bar represent nucleotide substitutions per site." figureDoi="10.3897/zookeys.926.48745.figure5" httpUri="https://binary.pensoft.net/fig/399908" pageId="0" pageNumber="81">5</figureCitation>
). The respective genetic distances between this species and 
<taxonomicName lsidName="E. burgeri" pageId="0" pageNumber="81" rank="species" species="burgeri">
<emphasis italics="true" pageId="0" pageNumber="81">E. burgeri</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="E. minor" pageId="0" pageNumber="81" rank="species" species="minor">
<emphasis italics="true" pageId="0" pageNumber="81">E. minor</emphasis>
</taxonomicName>
were 0.9% and 4.5% in 16S rRNA sequences and 8.5% and 13.9% in COI sequences. In addition, phylogenetic analysis of 16S rRNA sequences showed that 
<taxonomicName lsidName="E. wandoensis" pageId="0" pageNumber="81" rank="species" species="wandoensis">
<emphasis italics="true" pageId="0" pageNumber="81">E. wandoensis</emphasis>
</taxonomicName>
sp. nov. is well separated from other five-gilled hagfishes (
<taxonomicName lsidName="E. cheni" pageId="0" pageNumber="81" rank="species" species="cheni">
<emphasis italics="true" pageId="0" pageNumber="81">E. cheni</emphasis>
</taxonomicName>
, 
<taxonomicName lsidName="E. nelsoni" pageId="0" pageNumber="81" rank="species" species="nelsoni">
<emphasis italics="true" pageId="0" pageNumber="81">E. nelsoni</emphasis>
</taxonomicName>
, and 
<taxonomicName lsidName="E. yangi" pageId="0" pageNumber="81" rank="species" species="yangi">
<emphasis italics="true" pageId="0" pageNumber="81">E. yangi</emphasis>
</taxonomicName>
), with genetic differences of 7.5%, 1.4%, and 1.6%, respectively. 
<taxonomicName baseAuthorityName="Shen &amp; Tao" baseAuthorityYear="1975" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus cheni" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="species" species="cheni">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus cheni</emphasis>
</taxonomicName>
is located at a basal position of hagfishes and well nested in the 
<taxonomicName authorityName="Cloquet in Cloquet" authorityYear="1819" class="Myxini" family="Myxinidae" genus="Eptatretus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eptatretus" order="Myxiniformes" pageId="0" pageNumber="81" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="81">Eptatretus</emphasis>
</taxonomicName>
clade.
</paragraph>
<caption doi="10.3897/zookeys.926.48745.figure5" httpUri="https://binary.pensoft.net/fig/399908" pageId="0" pageNumber="81" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figure 5.</emphasis>
Phylogenetic tree of hagfishes based on mitochondrial DNA sequences, constructed with the Neighbor-joining (NJ) method using Kimura 2-parameter distances 
<emphasis bold="true" pageId="0" pageNumber="81">A</emphasis>
mitochondrial DNA 16S rRNA sequences 
<emphasis bold="true" pageId="0" pageNumber="81">B</emphasis>
mitochondrial DNA cytochrome c oxidase subunit I (COI) sequences. Numbers above tree branches are bootstrap values based on 1000 replicates. Scale bar represent nucleotide substitutions per site.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>