treatments-xml/data/0F/FF/40/0FFF4071E9E95F51B2FC38B2C002F7F3.xml

<document ID-DOI="http://dx.doi.org/10.3897/zse.98.78202" ID-GBIF-Dataset="c24e9174-953e-457c-96b0-819cced80599" ID-GBIF-Taxon="193383545" ID-Pensoft-Pub="1860-0743-1-55" ID-Pensoft-UUID="A7ED6822DF245CC385E84C2DA8697421" ID-ZooBank="F6DA891882B849C5BD64FE152C5D71BE" LSID="urn:lsid:zoobank.org:act:42360820-446E-4054-B0DB-CF52D6DE4317" ModsDocID="1860-0743-98-1-55" checkinTime="1645053248707" checkinUser="pensoft" docAuthor="Garcia, Andres F., Vargas, Alex Gonzalez &amp; Estrada, Miguel Gutierrez" docDate="2022" docId="0FFF4071E9E95F51B2FC38B2C002F7F3" docLanguage="en" docName="ZoosystEvol 98(1): 55-63" docOrigin="Zoosystematics and Evolution 98 (1)" docPubDate="2022-02-16" docSource="http://dx.doi.org/10.3897/zse.98.78202" docTitle="Avima wayuunaiki García &amp; Vargas &amp; Estrada 2022, sp. nov." docType="treatment" docUuid="42360820-446E-4054-B0DB-CF52D6DE4317" docUuidSource="ZooBank" docVersion="3" id="A7ED6822DF245CC385E84C2DA8697421" lastPageNumber="55" masterDocId="A7ED6822DF245CC385E84C2DA8697421" masterDocTitle="New records and a new cave-dwelling species of Agoristenidae (Arachnida, Opiliones) from Colombia" masterLastPageNumber="63" masterPageNumber="55" pageId="0" pageNumber="55" updateTime="1645057482806" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>New records and a new cave-dwelling species of Agoristenidae (Arachnida, Opiliones) from Colombia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Garcia, Andres F.</mods:namePart>
<mods:affiliation>Departamento de Invertebrados, Museu Nacional / Universidade Federal do Rio de Janeiro (UFRJ), Quinta da Boa Vista, Sao Cristovao, 20.940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">agarciarinc@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vargas, Alex Gonzalez</mods:namePart>
<mods:affiliation>Grupo de Investigacion Ecologia y Biodiversidad en Ecosistemas Tropicales (EBET), Semillero de Investigacion Bioespeleologia Neotropical, Facultad de Ciencias Basicas, Universidad de La Guajira, Riohacha, Colombia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Estrada, Miguel Gutierrez</mods:namePart>
<mods:affiliation>Grupo de Investigacion Ecologia y Biodiversidad en Ecosistemas Tropicales (EBET), Semillero de Investigacion Bioespeleologia Neotropical, Facultad de Ciencias Basicas, Universidad de La Guajira, Riohacha, Colombia</mods:affiliation>
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<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-02-16</mods:number>
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<mods:number>98</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zse.98.78202</mods:url>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.98.78202</mods:identifier>
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<treatment ID-GBIF-Taxon="193383545" LSID="urn:lsid:zoobank.org:act:42360820-446E-4054-B0DB-CF52D6DE4317" httpUri="http://treatment.plazi.org/id/0FFF4071E9E95F51B2FC38B2C002F7F3" lastPageNumber="55" pageId="0" pageNumber="55">
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<paragraph pageId="0" pageNumber="55">
<taxonomicName LSID="http://zoobank.org/42360820-446E-4054-B0DB-CF52D6DE4317" authority="García &amp; Vargas &amp; Estrada, 2022" authorityName="García &amp; Vargas &amp; Estrada" authorityYear="2022" class="Arachnida" family="Agoristenidae" genus="Avima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Avima wayuunaiki" order="Opiliones" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="wayuunaiki" status="sp. nov.">Avima wayuunaiki</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="55">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="description">
<paragraph pageId="0" pageNumber="55">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in panoramic (A), dorsal (B), ventral (C), lateral (D), and frontal (E) views. Scale bars: 2 mm (A); 1 mm (B-E)." figureDoi="10.3897/zse.98.78202.figure1" httpUri="https://binary.pensoft.net/fig/647183" pageId="0" pageNumber="55">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Drawings of Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in dorsal (A) and lateral (B) views. Left coxa I in ventral view (C). Right chelicera in frontal view (D). Left pedipalp in ventral view (E). Scale bars: 1 mm (A, B, D, E); 0.5 mm (C)." figureDoi="10.3897/zse.98.78202.figure2" httpUri="https://binary.pensoft.net/fig/647184" pageId="0" pageNumber="55">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM of the male genitalia of Avima wayuunaiki sp. nov. (ICN-Ao- 1718), paratype. Distal portion of the penis in lateroapical (A), dorsoapical (B), lateral (C), ventral (D), and oblique (E) views; detail of stylus in lateral view (F). Abbreviations: MS = Macrosetae. Scale bars: 50 µm (A-E); 10 µm (F)." figureDoi="10.3897/zse.98.78202.figure3" httpUri="https://binary.pensoft.net/fig/647185" pageId="0" pageNumber="55">, 3</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Geographic distribution of Avima venezuelica, Avima troglobia and Avima wayuunaiki sp. nov. in Northern South America. A. Map with previous and new records of the species in Colombia and Venezuela (colored areas represent the WWF ecoregions (Olson et al. 2001 )); B. Banaderos cave in La Guajira (Colombia), habitat of A. wayuunaiki sp. nov.; C. Living specimen of A. wayuunaiki sp. nov. Photographs by Miguel Gutierrez Estrada (B), and Alex Gonzalez Vargas (C)." figureDoi="10.3897/zse.98.78202.figure6" httpUri="https://binary.pensoft.net/fig/647188" pageId="0" pageNumber="55">, 6A, C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="type data">
<paragraph pageId="0" pageNumber="55">Type data.</paragraph>
<paragraph pageId="0" pageNumber="55">
COLOMBIA• ♂ holotype: La Guajira, Hatonuevo, Cerro 
<normalizedToken originalValue="Bañaderos">Banaderos</normalizedToken>
, cueva [Luis Pablo Ojeda]; 
<geoCoordinate degrees="11" direction="north" minutes="7" orientation="latitude" precision="1" seconds="51.5" value="11.130972">11°7'51.5&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="72" direction="west" minutes="47" orientation="longitude" precision="1" seconds="23.9" value="-72.78997">72°47'23.9&quot;W</geoCoordinate>
[11.130972°, -72.789972°]; 978 m a.s.l.; 14 July 2015; CarBio Team 17 leg.; ICN-Ao-1976 • 7 ♂ 10 ♀: same data as holotype; ICN-Ao-1718 • 1 ♂ 1 ♀: same data as holotype; MUSENUV-Ar 2102 • 4 ♂ 2 ♀: same locality as holotype, vereda 
<normalizedToken originalValue="Bañaderos">Banaderos</normalizedToken>
; cueva [Luis Pablo Ojeda]; 
<geoCoordinate degrees="11" direction="north" minutes="7" orientation="latitude" precision="1" seconds="33.3" value="11.1259165">11°7'33.3&quot;N</geoCoordinate>
, 
<geoCoordinate degrees="72" direction="west" minutes="47" orientation="longitude" precision="1" seconds="06.9" value="-72.78525">72°47'06.9&quot;W</geoCoordinate>
[11.125899°, -72.785241°]; 785 m a.s.l.; 19 May 2018; Miguel 
<normalizedToken originalValue="Gutiérrez">Gutierrez</normalizedToken>
leg.; MNRJ 283 • 2 ♀: same locality as previous; CBUDC-ARA 335.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="description">
<paragraph pageId="0" pageNumber="55">Description.</paragraph>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">
Male 
<emphasis italics="true" pageId="0" pageNumber="55">Holotype</emphasis>
</emphasis>
(ICN-Ao-1976). Measurements. TL (ChL+DSL): 4.45, DSL: 2.66, CL: 1.07, CW: 1.79, AL: 1.59, AW: 2.21, BaCh: 0.55. Pedipalp: Tr: 0.60, Fe: 1.23, Pa: 0.72, Ti: 0.88, Ta: 0.70, Claw: 0.79. Leg I: Tr: 0.42, Fe: 4.10, Pa: 0.73, Ti: 3.19, Mt: 5.86, Ta: 1.31, TL: 15.61; Leg II: Tr: 0.57, Fe: 9.21, Pa: 0.98, Ti: 7.67, Mt: 12.40, Ta: 5.14, TL: 35.97. Leg III: Tr: 0.62, Fe: 5.92, Pa: 0.97, Ti: 3.65, Mt: 7.52, Ta: 1.58, TL: 20.26. Leg IV: Tr: 0.70, Fe: 8.65, Pa: 1.00, Ti: 4.85, Mt: 11.14, Ta: 1.82, TL: 28.16.
</paragraph>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Dorsum.</emphasis>
DS Epsilon type 2. Anterior and lateral margins of DS smooth. Ocularium low, smooth, and without median concavity (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in panoramic (A), dorsal (B), ventral (C), lateral (D), and frontal (E) views. Scale bars: 2 mm (A); 1 mm (B-E)." figureDoi="10.3897/zse.98.78202.figure1" httpUri="https://binary.pensoft.net/fig/647183" pageId="0" pageNumber="55">1A, B, E</figureCitation>
). Mesotergum slightly delimited, divided into four smooth areas: area I divided into two halves; areas II-IV undivided (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Drawings of Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in dorsal (A) and lateral (B) views. Left coxa I in ventral view (C). Right chelicera in frontal view (D). Left pedipalp in ventral view (E). Scale bars: 1 mm (A, B, D, E); 0.5 mm (C)." figureDoi="10.3897/zse.98.78202.figure2" httpUri="https://binary.pensoft.net/fig/647184" pageId="0" pageNumber="55">2A</figureCitation>
). Posterior margin of scutum substraight and with few granules (Figs 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in panoramic (A), dorsal (B), ventral (C), lateral (D), and frontal (E) views. Scale bars: 2 mm (A); 1 mm (B-E)." figureDoi="10.3897/zse.98.78202.figure1" httpUri="https://binary.pensoft.net/fig/647183" pageId="0" pageNumber="55">1D</figureCitation>
, 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Drawings of Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in dorsal (A) and lateral (B) views. Left coxa I in ventral view (C). Right chelicera in frontal view (D). Left pedipalp in ventral view (E). Scale bars: 1 mm (A, B, D, E); 0.5 mm (C)." figureDoi="10.3897/zse.98.78202.figure2" httpUri="https://binary.pensoft.net/fig/647184" pageId="0" pageNumber="55">2B</figureCitation>
). Free tergites I-III with some granules (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in panoramic (A), dorsal (B), ventral (C), lateral (D), and frontal (E) views. Scale bars: 2 mm (A); 1 mm (B-E)." figureDoi="10.3897/zse.98.78202.figure1" httpUri="https://binary.pensoft.net/fig/647183" pageId="0" pageNumber="55">1B, D, E</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.98.78202.figure1" httpUri="https://binary.pensoft.net/fig/647183" pageId="0" pageNumber="55" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">Figure 1.</emphasis>
<taxonomicName authorityName="García &amp; Vargas &amp; Estrada" authorityYear="2022" class="Arachnida" family="Agoristenidae" genus="Avima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Avima wayuunaiki" order="Opiliones" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="wayuunaiki">
<emphasis italics="true" pageId="0" pageNumber="55">Avima wayuunaiki</emphasis>
</taxonomicName>
sp. nov. (ICN-Ao-1976), male holotype. Habitus in panoramic (
<emphasis bold="true" pageId="0" pageNumber="55">A</emphasis>
), dorsal (
<emphasis bold="true" pageId="0" pageNumber="55">B</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="55">C</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="55">D</emphasis>
), and frontal (
<emphasis bold="true" pageId="0" pageNumber="55">E</emphasis>
) views. Scale bars: 2 mm (
<emphasis bold="true" pageId="0" pageNumber="55">A</emphasis>
); 1 mm (
<emphasis bold="true" pageId="0" pageNumber="55">B-E</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">
<emphasis italics="true" pageId="0" pageNumber="55">Venter</emphasis>
.
</emphasis>
Coxa I with a longitudinal row of tubercles and one large bicapitate tubercle on the anteroproximal margin (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Drawings of Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in dorsal (A) and lateral (B) views. Left coxa I in ventral view (C). Right chelicera in frontal view (D). Left pedipalp in ventral view (E). Scale bars: 1 mm (A, B, D, E); 0.5 mm (C)." figureDoi="10.3897/zse.98.78202.figure2" httpUri="https://binary.pensoft.net/fig/647184" pageId="0" pageNumber="55">2C</figureCitation>
); coxae II-IV with some granules; coxa II longer than coxa I; coxa III longer than coxae I and II; coxa IV backward projected. Stigmata oval, small and transverse (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in panoramic (A), dorsal (B), ventral (C), lateral (D), and frontal (E) views. Scale bars: 2 mm (A); 1 mm (B-E)." figureDoi="10.3897/zse.98.78202.figure1" httpUri="https://binary.pensoft.net/fig/647183" pageId="0" pageNumber="55">1C</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.98.78202.figure2" httpUri="https://binary.pensoft.net/fig/647184" pageId="0" pageNumber="55" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">Figure 2.</emphasis>
Drawings of 
<taxonomicName authorityName="García &amp; Vargas &amp; Estrada" authorityYear="2022" class="Arachnida" family="Agoristenidae" genus="Avima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Avima wayuunaiki" order="Opiliones" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="wayuunaiki">
<emphasis italics="true" pageId="0" pageNumber="55">Avima wayuunaiki</emphasis>
</taxonomicName>
sp. nov. (ICN-Ao-1976), male holotype. Habitus in dorsal (
<emphasis bold="true" pageId="0" pageNumber="55">A</emphasis>
) and lateral (
<emphasis bold="true" pageId="0" pageNumber="55">B</emphasis>
) views. Left coxa I in ventral view (
<emphasis bold="true" pageId="0" pageNumber="55">C</emphasis>
). Right chelicera in frontal view (
<emphasis bold="true" pageId="0" pageNumber="55">D</emphasis>
). Left pedipalp in ventral view (
<emphasis bold="true" pageId="0" pageNumber="55">E</emphasis>
). Scale bars: 1 mm (
<emphasis bold="true" pageId="0" pageNumber="55">A, B, D, E</emphasis>
); 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="55">C</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">
<emphasis italics="true" pageId="0" pageNumber="55">Chelicera</emphasis>
.
</emphasis>
Chelicera swollen. BaCh quadrate in dorsal view, with well-marked bulla, three mesal, five anterior, and three ectal tubercles. Hand with setiferous tubercles of different sizes reaching the medial and posterior region. Fixed finger with the inner surface finely grooved. Movable finger with one trapezoid, small, sub-basal tooth, and with a dentate distal inner surface (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Drawings of Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in dorsal (A) and lateral (B) views. Left coxa I in ventral view (C). Right chelicera in frontal view (D). Left pedipalp in ventral view (E). Scale bars: 1 mm (A, B, D, E); 0.5 mm (C)." figureDoi="10.3897/zse.98.78202.figure2" httpUri="https://binary.pensoft.net/fig/647184" pageId="0" pageNumber="55">2D</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">
<emphasis italics="true" pageId="0" pageNumber="55">Pedipalps</emphasis>
.
</emphasis>
All segments more slender than usual (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in panoramic (A), dorsal (B), ventral (C), lateral (D), and frontal (E) views. Scale bars: 2 mm (A); 1 mm (B-E)." figureDoi="10.3897/zse.98.78202.figure1" httpUri="https://binary.pensoft.net/fig/647183" pageId="0" pageNumber="55">1B</figureCitation>
). Trochanter with one subapical tubercle on the ventral face. Femur with a ventroectal row of four setiferous tubercles (the two basalmost largest and the two distalmost medium-sized), and one large ventromesal setiferous tubercle in the apical portion (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Drawings of Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in dorsal (A) and lateral (B) views. Left coxa I in ventral view (C). Right chelicera in frontal view (D). Left pedipalp in ventral view (E). Scale bars: 1 mm (A, B, D, E); 0.5 mm (C)." figureDoi="10.3897/zse.98.78202.figure2" httpUri="https://binary.pensoft.net/fig/647184" pageId="0" pageNumber="55">2E</figureCitation>
). Patella with one large mesal setiferous tubercle. Tibia ectal III, mesal IIi. Tarsus ectal IIi, mesal IIi.
</paragraph>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">
<emphasis italics="true" pageId="0" pageNumber="55">Legs</emphasis>
.
</emphasis>
Increasing in thickness from leg I to leg IV, all smooth (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in panoramic (A), dorsal (B), ventral (C), lateral (D), and frontal (E) views. Scale bars: 2 mm (A); 1 mm (B-E)." figureDoi="10.3897/zse.98.78202.figure1" httpUri="https://binary.pensoft.net/fig/647183" pageId="0" pageNumber="55">1A</figureCitation>
). Leg I filiform; coxae I-III with one dorsoanterior and one dorso-posterior tubercle; Coxa IV with some prolaterodistal tubercles; Fe III-IV darker than I-II. Fe IV length three and a half times DS length (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in panoramic (A), dorsal (B), ventral (C), lateral (D), and frontal (E) views. Scale bars: 2 mm (A); 1 mm (B-E)." figureDoi="10.3897/zse.98.78202.figure1" httpUri="https://binary.pensoft.net/fig/647183" pageId="0" pageNumber="55">1A</figureCitation>
); Ta I-II each with one smooth claw; Ta III-IV with two subparallel smooth claws and without tarsal process. Tarsal counts: 9(3)-9(3)/19(3)-18(3)/7-6/7-7.
</paragraph>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">
<emphasis italics="true" pageId="0" pageNumber="55">Penis</emphasis>
.
</emphasis>
LP small (width twice the height) and apically depressed, with anterolateral acute corners (crescent-shaped) apically pointed (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM of the male genitalia of Avima wayuunaiki sp. nov. (ICN-Ao- 1718), paratype. Distal portion of the penis in lateroapical (A), dorsoapical (B), lateral (C), ventral (D), and oblique (E) views; detail of stylus in lateral view (F). Abbreviations: MS = Macrosetae. Scale bars: 50 µm (A-E); 10 µm (F)." figureDoi="10.3897/zse.98.78202.figure3" httpUri="https://binary.pensoft.net/fig/647185" pageId="0" pageNumber="55">3A-E</figureCitation>
). Malleus with two pairs of branched MS-A, being MS-A2 far from the base of the stylus; one pair of branched MS-B; MS-C absent; two pairs of medium-sized MS-D located in a keel between the dorsal region of the LP and the base of the stylus (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM of the male genitalia of Avima wayuunaiki sp. nov. (ICN-Ao- 1718), paratype. Distal portion of the penis in lateroapical (A), dorsoapical (B), lateral (C), ventral (D), and oblique (E) views; detail of stylus in lateral view (F). Abbreviations: MS = Macrosetae. Scale bars: 50 µm (A-E); 10 µm (F)." figureDoi="10.3897/zse.98.78202.figure3" httpUri="https://binary.pensoft.net/fig/647185" pageId="0" pageNumber="55">3A-C, E</figureCitation>
); MS-E2 large and branched, MS-E1 short and conical, located slightly distal to MS-E2 on the ventral side of the LP (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM of the male genitalia of Avima wayuunaiki sp. nov. (ICN-Ao- 1718), paratype. Distal portion of the penis in lateroapical (A), dorsoapical (B), lateral (C), ventral (D), and oblique (E) views; detail of stylus in lateral view (F). Abbreviations: MS = Macrosetae. Scale bars: 50 µm (A-E); 10 µm (F)." figureDoi="10.3897/zse.98.78202.figure3" httpUri="https://binary.pensoft.net/fig/647185" pageId="0" pageNumber="55">3C, D</figureCitation>
). Stylus elongated, mostly straight (concave at the apex), and surpassing the LP; dorsal keel present, smooth, elevated at the medial region, with a dorsoapical sharp projection; tip dorsally projected (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. SEM of the male genitalia of Avima wayuunaiki sp. nov. (ICN-Ao- 1718), paratype. Distal portion of the penis in lateroapical (A), dorsoapical (B), lateral (C), ventral (D), and oblique (E) views; detail of stylus in lateral view (F). Abbreviations: MS = Macrosetae. Scale bars: 50 µm (A-E); 10 µm (F)." figureDoi="10.3897/zse.98.78202.figure3" httpUri="https://binary.pensoft.net/fig/647185" pageId="0" pageNumber="55">3E, F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zse.98.78202.figure3" httpUri="https://binary.pensoft.net/fig/647185" pageId="0" pageNumber="55" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">Figure 3.</emphasis>
SEM of the male genitalia of 
<taxonomicName authorityName="García &amp; Vargas &amp; Estrada" authorityYear="2022" class="Arachnida" family="Agoristenidae" genus="Avima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Avima wayuunaiki" order="Opiliones" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="wayuunaiki">
<emphasis italics="true" pageId="0" pageNumber="55">Avima wayuunaiki</emphasis>
</taxonomicName>
sp. nov. (ICN-Ao-1718), paratype. Distal portion of the penis in lateroapical (
<emphasis bold="true" pageId="0" pageNumber="55">A</emphasis>
), dorsoapical (
<emphasis bold="true" pageId="0" pageNumber="55">B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="55">C</emphasis>
), ventral (
<emphasis bold="true" pageId="0" pageNumber="55">D</emphasis>
), and oblique (
<emphasis bold="true" pageId="0" pageNumber="55">E</emphasis>
) views; detail of stylus in lateral view (
<emphasis bold="true" pageId="0" pageNumber="55">F</emphasis>
). Abbreviations: MS = Macrosetae. Scale bars: 50 
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="55">A-E</emphasis>
); 10 
<normalizedToken originalValue="µm">µm</normalizedToken>
(
<emphasis bold="true" pageId="0" pageNumber="55">F</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" italics="true" pageId="0" pageNumber="55">Color (in alcohol).</emphasis>
DS and chelicerae Light Yellow (86). Pedipalps, coxae, and trochanters I-IV Pale Yellow (89). Femora to tarsi I-IV Dark Yellowish Brown (78) (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Avima wayuunaiki sp. nov. (ICN-Ao- 1976), male holotype. Habitus in panoramic (A), dorsal (B), ventral (C), lateral (D), and frontal (E) views. Scale bars: 2 mm (A); 1 mm (B-E)." figureDoi="10.3897/zse.98.78202.figure1" httpUri="https://binary.pensoft.net/fig/647183" pageId="0" pageNumber="55">1</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="55">
<emphasis bold="true" pageId="0" pageNumber="55">Female.</emphasis>
Paratype (ICN-Ao-1718). Similar to male, except for abdomen wider at areas II-III, chelicerae not swollen, and leg IV slender. Measurements: TL (Ch+DSL): 3.50, DSL: 2.35, CL: 0.96, CW: 1.61, AL: 1.39, AW: 1.96, BaCh: 0.42. Pedipalp: Tr: 0.41, Fe: 1.12, Pa: 0.65, Ti: 0.84, Ta: 0.79, Claw: 0.78. Leg I: Tr: 0.32, Fe: 4.27, Pa: 0.69, Ti: 3.19, Mt: 6.04, Ta: 1.29, TL: 15.8; Leg II: Tr: 0.39, Fe: 9.53, Pa: 0.98, Ti: 8.70, Mt: 12.14, Ta: 5.34, TL: 37.08. Leg III: Tr: 0.39, Fe: 9.36, Pa: 1.04, Ti: 5.77, Mt: 13.25, Ta: 1.51, TL: 31.32. Leg IV: Tr: 0.68, Fe: 8.70, Pa: 0.94, Ti: 4.83, Mt: 11.08, Ta: 1.80, TL: 28.03. Tarsal counts:?-6(3)/? -18(3)/6-6/7-6.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="etymology">
<paragraph pageId="0" pageNumber="55">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="55">Wayuunaiki is the language spoken by the indigenous Wayuu people in northwestern Venezuela and northeastern Colombia on the Guajira Peninsula, where the species was collected. Noun in apposition.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="natural history">
<paragraph pageId="0" pageNumber="55">Natural history.</paragraph>
<paragraph pageId="0" pageNumber="55">
Specimens of 
<taxonomicName genus="A." lsidName="A. wayuunaiki" pageId="0" pageNumber="55" rank="species" species="wayuunaiki">
<emphasis italics="true" pageId="0" pageNumber="55">A. wayuunaiki</emphasis>
</taxonomicName>
sp. nov. were found inside a cave (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Geographic distribution of Avima venezuelica, Avima troglobia and Avima wayuunaiki sp. nov. in Northern South America. A. Map with previous and new records of the species in Colombia and Venezuela (colored areas represent the WWF ecoregions (Olson et al. 2001 )); B. Banaderos cave in La Guajira (Colombia), habitat of A. wayuunaiki sp. nov.; C. Living specimen of A. wayuunaiki sp. nov. Photographs by Miguel Gutierrez Estrada (B), and Alex Gonzalez Vargas (C)." figureDoi="10.3897/zse.98.78202.figure6" httpUri="https://binary.pensoft.net/fig/647188" pageId="0" pageNumber="55">6B, C</figureCitation>
) in the northeastern slopes of Sierra Nevada de Santa Marta. This species lives sympatrically with 
<taxonomicName authorityName="Cala-Riquelme, Gutierrez-Estrada &amp; Florez" authorityYear="2015" class="Arachnida" family="Sicariidae" genus="Loxosceles" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Loxosceles guajira" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="guajira">
<emphasis italics="true" pageId="0" pageNumber="55">Loxosceles guajira</emphasis>
</taxonomicName>
Cala-Riquelme, 
<normalizedToken originalValue="Gutiérrez-Estrada">Gutierrez-Estrada</normalizedToken>
&amp; 
<normalizedToken originalValue="Flórez">Florez</normalizedToken>
, 2015 (
<taxonomicName authorityName="Keyserling" authorityYear="1880" class="Arachnida" family="Sicariidae" kingdom="Animalia" lsidName="" order="Araneae" pageId="0" pageNumber="55" phylum="Arthropoda" rank="family">Sicariidae</taxonomicName>
) and 
<taxonomicName authorityName="DoNascimiento &amp; Prada-Pedreros" authorityYear="2020" class="Actinopterygii" family="Trichomycteridae" genus="Trichomycterus" kingdom="Animalia" lsidName="Trichomycterus spectrum" order="Siluriformes" pageId="0" pageNumber="55" phylum="Chordata" rank="species" species="spectrum">
<emphasis italics="true" pageId="0" pageNumber="55">Trichomycterus spectrum</emphasis>
</taxonomicName>
DoNascimiento &amp; Prada-Pedreros, 2020 (
<taxonomicName authorityName="Bleeker" authorityYear="1858" class="Actinopterygii" family="Trichomycteridae" kingdom="Animalia" lsidName="" order="Siluriformes" pageId="0" pageNumber="55" phylum="Chordata" rank="family">Trichomycteridae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>