treatments-xml/data/03/8B/87/038B87F6A5751F6D0AEDB318BC7E50DA.xml

<document id="4B07BB9849C915EE105AC544824CA538" ID-CLB-Dataset="21269" ID-DOI="10.11646/zootaxa.4786.1.3" ID-GBIF-Dataset="70d743db-654b-4050-a05f-a5c71b54e14e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3990784" ID-ZooBank="625F197D-9D26-4B00-AF3E-3045586ECB07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597795225921" checkinUser="felipe" docAuthor="Salles, F. F., Nieto, C. &amp; Cruz, P. V." docDate="2020" docId="038B87F6A5751F6D0AEDB318BC7E50DA" docLanguage="en" docName="zootaxa.4786.1.3.pdf.imf" docOrigin="Zootaxa 4786 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rivudiva inma Salles &amp; Nieto &amp; Cruz 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="42" masterDocId="FFB2FF8EA5771F680A7AB129B91D5554" masterDocTitle="New species of Rivudiva Lugo-Ortiz &amp; McCafferty (Ephemeroptera: Baetidae) with comments on R. minantenna Lugo-Ortiz &amp; McCafferty and R. trichobasis Lugo-Ortiz &amp; McCafferty" masterLastPageNumber="52" masterPageNumber="37" pageNumber="39" updateTime="1698844779479" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E8A8092F028D32D80B1C491054CC30BD">New species of Rivudiva Lugo-Ortiz &amp; McCafferty (Ephemeroptera: Baetidae) with comments on R. minantenna Lugo-Ortiz &amp; McCafferty and R. trichobasis Lugo-Ortiz &amp; McCafferty</mods:title>
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<mods:namePart id="4A44935DE7E5DBB06AE0E3021A27BC1D">Salles, F. F.</mods:namePart>
<mods:affiliation id="54E2D2B4F18255E5D888F17DDC84BC15">Museu de Entomologia, Depto. de Entomologia, Universidade Federal de Viçosa, CEP 36.570 - 900, Viçosa, MG, Brazil. frederico. salles @ ufv. br; https: // orcid. org / 0000 - 0001 - 8331 - 5929</mods:affiliation>
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<mods:namePart id="E948E9048E02D06AF4866C4EC641C88D">Nieto, C.</mods:namePart>
<mods:affiliation id="02271BF0AD790083FD6A32AA85D3E676">Instituto de Biodiversidad Neotropical (IBN), CONICET-U. N. T. &amp; Facultad de Ciencias Naturales, Universidad Nacional de Tucumán, San Miguel de Tucumán, 4.000 Tucumán, Argentina.</mods:affiliation>
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<mods:namePart id="6798C5177D3670DD0CE004887F32EB6E">Cruz, P. V.</mods:namePart>
<mods:affiliation id="4B688DD7791F485CF53C2CF72CF14506">Universidade Federal de Rondônia - UNIR, Programa de Pós-Graduação em Ciências Ambientais, Laboratório de Biodiversidade e Conservação - LABICON, CEP: 76940 - 000, Rolim de Moura, Rondônia, Brazil.</mods:affiliation>
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<taxonomicName id="4C224D63A5751F6A0AEDB318B8E0571F" authority="Salles &amp; Nieto" authorityName="Salles &amp; Nieto &amp; Cruz" authorityYear="2020" box="[151,509,561,587]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="inma" status="sp. nov.">
<emphasis id="B956EAF2A5751F6A0AEDB318B85F571F" box="[151,322,561,587]" italics="true" pageId="2" pageNumber="39">Rivudiva inma</emphasis>
<emphasis id="B956EAF2A5751F6A0B33B318B8E0571F" bold="true" box="[329,509,561,587]" pageId="2" pageNumber="39">Salles &amp; Nieto</emphasis>
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, 
<taxonomicNameLabel id="A2655789A5751F6A0873B31BBB7A5718" box="[521,615,562,588]" pageId="2" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<paragraph id="8B9D36E0A5751F6A0AEDB35FBD8757DA" blockId="2.[151,1178,629,654]" box="[151,1178,629,654]" pageId="2" pageNumber="39">
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<taxonomicName id="4C224D63A5751F6A0AEDB35FB86D57D9" ID-CoL="6X948" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1998" box="[151,368,630,654]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="39" phylum="Arthropoda" rank="species" species="minantenna">
<emphasis id="B956EAF2A5751F6A0AEDB35FB86D57D9" box="[151,368,630,654]" italics="true" pageId="2" pageNumber="39">Rivudiva minantenna</emphasis>
</taxonomicName>
(misidentification): 
<treatmentCitation id="0A8310F1A5751F6A0845B35CBA8B57DA" author="Salles, F. F. &amp; Nascimento, J. M. C." box="[575,918,629,654]" page="233" pageId="2" pageNumber="39" year="2009">
<bibRefCitation id="EFB34B11A5751F6A0845B35CBA8B57DA" author="Salles, F. F. &amp; Nascimento, J. M. C." box="[575,918,629,654]" pageId="2" pageNumber="39" pagination="231 - 235" refId="ref7915" refString="Salles, F. F. &amp; Nascimento, J. M. C. (2009) The genus Rivudiva Lugo-Ortiz and McCafferty (Ephemeroptera: Baetidae): First generic description of adults, new combinations, and notes on the nymphs. Annales de Limnologie-International Journal of Limnology, 45 (4), 231 - 235. https: // doi. org / 10.1051 / limn / 2009023" type="journal article" year="2009">Salles &amp; Nascimento (2009: 233)</bibRefCitation>
</treatmentCitation>
; 
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<bibRefCitation id="EFB34B11A5751F6A09DAB35CBD8B57DA" author="Salles, F. F. &amp; Nascimento, J. M. C. D. &amp; Massariol, F. C. &amp; Angeli, K. B. &amp; Silva, P. B. &amp; Rudio, J. A. &amp; Boldrini, R." box="[928,1174,629,654]" pageId="2" pageNumber="39" pagination="293 - 307" refId="ref7988" refString="Salles, F. F., Nascimento, J. M. C. D., Massariol, F. C., Angeli, K. B., Silva, P. B., Rudio, J. A. &amp; Boldrini, R. (2010) Primeiro levantamento da fauna de Ephemeroptera (Insecta) do Espirito Santo, sudeste do Brasil. Biota Neotropica, 10 (1), 293 - 307. https: // doi. org / 10.1590 / S 1676 - 06032010000100025" type="journal article" year="2010">
Salles 
<emphasis id="B956EAF2A5751F6A0999B35FBD0B57DA" box="[995,1046,630,654]" italics="true" pageId="2" pageNumber="39">et al.</emphasis>
(2010: 303)
</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="C338656BA5751F6A0AEDB392B8365632" pageId="2" pageNumber="39" type="materials_examined">
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<emphasis id="B956EAF2A5751F6A0AEDB392B89B5782" bold="true" box="[151,390,699,726]" pageId="2" pageNumber="39">Material examined.</emphasis>
<materialsCitation id="3B4A3CBDA5751F6A0BE9B395B83A564A" ID-GBIF-Occurrence="2832053308" collectingDate="2009-02-19" collectorName="FF Salles" country="Brazil" elevation="720" latitude="-19.875834" location="Capitel de Santo Antonio" longLatPrecision="21" longitude="-40.530556" municipality="Santa Teresa" pageId="2" pageNumber="39" specimenCount="720" stateProvince="Espirito Santo" typeStatus="holotype">
<typeStatus id="54998842A5751F6A0BE9B395B8E65782" box="[403,507,700,726]" pageId="2" pageNumber="39" type="holotype">Holotype</typeStatus>
(male adult with corresponding exuviae): 
<collectingCountry id="F3357670A5751F6A0982B395BD7D5782" box="[1016,1120,700,726]" name="Brazil" pageId="2" pageNumber="39">BRAZIL</collectingCountry>
, 
<collectingRegion id="49E6F802A5751F6A0E0BB395BC455783" box="[1137,1368,700,727]" country="Brazil" name="Espirito Santo" pageId="2" pageNumber="39">Espírito Santo state</collectingRegion>
, 
<collectingMunicipality id="6BF9AC9AA5751F6A0F13B395B81C57AF" pageId="2" pageNumber="39">Santa Teresa</collectingMunicipality>
, 
<location id="8EFD603BA5751F6A0B74B3C9BB2E57AE" LSID="urn:lsid:plazi:treatment:038B87F6A5751F6D0AEDB318BC7E50DA:8EFD603BA5751F6A0B74B3C9BB2E57AE" box="[270,563,736,762]" country="Brazil" latitude="-19.875834" longLatPrecision="21" longitude="-40.530556" municipality="Santa Teresa" name="Capitel de Santo Antonio" pageId="2" pageNumber="39" stateProvince="Espirito Santo">Capitel de Santo Antônio</location>
, 
<geoCoordinate id="EE165027A5751F6A083BB3C9BBD257AE" box="[577,719,735,762]" degrees="19" direction="south" minutes="52" orientation="latitude" pageId="2" pageNumber="39" precision="15" seconds="33" value="-19.875834">S 19°52’33”</geoCoordinate>
<geoCoordinate id="EE165027A5751F6A08A2B3C8BA6E57AE" box="[728,883,735,763]" degrees="40" direction="west" minutes="31" orientation="longitude" pageId="2" pageNumber="39" precision="15" seconds="50" value="-40.530556">W 40°31’50”</geoCoordinate>
, 
<specimenCount id="9D24FD69A5751F6A09FBB3C9BACB57AE" box="[897,982,736,762]" pageId="2" pageNumber="39" type="generic">
<quantity id="4CDA9B05A5751F6A09FBB3C9BACF57AE" box="[897,978,736,762]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="2" pageNumber="39" unit="m" value="720.0">
<elevation id="000FD1D3A5751F6A09FBB3C9BACF57AE" box="[897,978,736,762]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="2" pageNumber="39" unit="m" value="720.0">720 m</elevation>
</quantity>
.
</specimenCount>
a.s.l., 
<date id="FF9C1020A5751F6A0E6DB3C9BD9657AE" box="[1047,1163,736,762]" pageId="2" pageNumber="39" value="2009-02-19">
<collectingDate id="EFD8E9C8A5751F6A0E6DB3C9BD9657AE" box="[1047,1163,736,762]" pageId="2" pageNumber="39" value="2009-02-19">19.ii.2009</collectingDate>
</date>
, 
<collectorName id="26D75336A5751F6A0EE3B3C8BC1857AE" box="[1177,1285,736,763]" pageId="2" pageNumber="39">FF Salles</collectorName>
leg. (
<collectionCode id="ED33AE25A5751F6A0F33B3C9BC8557AE" box="[1353,1432,736,762]" country="Brazil" httpUri="http://grbio.org/cool/3i20-rx1w" name="Vicosa, Universidade Federal de Vicosa, Museum of Entomology" pageId="2" pageNumber="39">UFVB</collectionCode>
, #EP000150)
</materialsCitation>
. 
<materialsCitation id="3B4A3CBDA5751F6A0B48B22CBA6F564A" ID-GBIF-Occurrence="2832053301" box="[306,882,772,799]" collectingDate="2009-02-19" collectorName="FF Salles" country="Brazil" elevation="720" latitude="-19.875834" location="Capitel de Santo Antonio" longLatPrecision="21" longitude="-40.530556" municipality="Santa Teresa" pageId="2" pageNumber="39" specimenCount="1" stateProvince="Espirito Santo" typeStatus="paratype">
<typeStatus id="54998842A5751F6A0B48B22CB8BE564B" box="[306,419,773,799]" pageId="2" pageNumber="39" type="paratype">Paratypes</typeStatus>
: same data as 
<typeStatus id="54998842A5751F6A083FB22DBBB7564A" box="[581,682,772,798]" pageId="2" pageNumber="39" type="holotype">holotype</typeStatus>
, 2 nymphs (
<collectionCode id="ED33AE25A5751F6A094FB22DBA76564A" box="[821,875,772,798]" pageId="2" pageNumber="39">IBN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B4A3CBDA5751F6A0905B22CBBEA5616" ID-GBIF-Occurrence="2832053302" collectingDate="2009-02-20" collectorName="FF Salles" country="Brazil" elevation="720" latitude="-19.875834" location="Capitel de Santo Antonio" longLatPrecision="21" longitude="-40.530556" municipality="Santa Teresa" pageId="2" pageNumber="39" specimenCount="1" stateProvince="Espirito Santo" typeStatus="holotype">
same data as 
<typeStatus id="54998842A5751F6A0E6CB22DBD66564A" box="[1046,1147,772,798]" pageId="2" pageNumber="39" type="holotype">holotype</typeStatus>
, except for 
<date id="FF9C1020A5751F6A0F7AB22DBC69564A" box="[1280,1396,772,798]" pageId="2" pageNumber="39" value="2009-02-20">
<collectingDate id="EFD8E9C8A5751F6A0F7AB22DBC69564A" box="[1280,1396,772,798]" pageId="2" pageNumber="39" value="2009-02-20">20.ii.2009</collectingDate>
</date>
, 14 nymphs (
<collectionCode id="ED33AE25A5751F6A0A84B201B8505616" box="[254,333,808,834]" country="Brazil" httpUri="http://grbio.org/cool/3i20-rx1w" name="Vicosa, Universidade Federal de Vicosa, Museum of Entomology" pageId="2" pageNumber="39">UFVB</collectionCode>
, 1 used for SEM images, #EP000151)
</materialsCitation>
; 
<materialsCitation id="3B4A3CBDA5751F6A097BB200B83A5632" ID-GBIF-Occurrence="2832053306" collectingDate="2008-10-26" collectorName="FF Salles" country="Brazil" elevation="720" latitude="-19.875834" location="Capitel de Santo Antonio" longLatPrecision="21" longitude="-40.530556" municipality="Santa Teresa" pageId="2" pageNumber="39" specimenCount="1" stateProvince="Espirito Santo" typeStatus="holotype">
same data as 
<typeStatus id="54998842A5751F6A09E9B201BAE55616" box="[915,1016,808,834]" pageId="2" pageNumber="39" type="holotype">holotype</typeStatus>
, except 
<date id="FF9C1020A5751F6A0E2BB201BDD95616" box="[1105,1220,808,834]" pageId="2" pageNumber="39" value="2008-10-26">
<collectingDate id="EFD8E9C8A5751F6A0E2BB201BDD95616" box="[1105,1220,808,834]" pageId="2" pageNumber="39" value="2008-10-26">26.x.2008</collectingDate>
</date>
, 4 nymphs (
<collectionCode id="ED33AE25A5751F6A0F33B201BC855616" box="[1353,1432,808,834]" country="Brazil" httpUri="http://grbio.org/cool/3i20-rx1w" name="Vicosa, Universidade Federal de Vicosa, Museum of Entomology" pageId="2" pageNumber="39">UFVB</collectionCode>
, #EP000152)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C338656BA5751F6A0AEDB291BAAD50F2" pageId="2" pageNumber="39" type="diagnosis">
<paragraph id="8B9D36E0A5751F6A0AEDB291B80D5686" blockId="2.[151,272,952,978]" box="[151,272,952,978]" pageId="2" pageNumber="39">
<heading id="D0D5818CA5751F6A0AEDB291B80D5686" bold="true" box="[151,272,952,978]" fontSize="11" level="3" pageId="2" pageNumber="39">
<emphasis id="B956EAF2A5751F6A0AEDB291B80D5686" bold="true" box="[151,272,952,978]" pageId="2" pageNumber="39">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36E0A5751F6A0AEDB529BD185042" blockId="2.[151,1437,1024,1447]" pageId="2" pageNumber="39">
<emphasis id="B956EAF2A5751F6A0AEDB529B9FB514E" box="[151,230,1024,1050]" italics="true" pageId="2" pageNumber="39">Nymph</emphasis>
. 1) antenna without spine-like setae on scape and pedicel; 2) distal margin of labrum with deep, wide, medial emargination (
<figureCitation id="13192A65A5751F6A0B13B50DB8B2516A" box="[361,431,1060,1086]" captionStart="FIGURES 1−6" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[167,1415,197,1905]" captionTargetId="figure@3.[153,1434,181,1920]" captionTargetPageId="3" captionText="FIGURES 1−6. Rivudiva inma, sp. nov., nymph: (1) Labrum; (2) Left mandible; (3) Right mandible; (4) Hypopharynx; (5) Maxilla; (6) Labium (left = ventral surface, right = dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.3990786" httpUri="https://zenodo.org/record/3990786/files/figure.png" pageId="2" pageNumber="39">Figs 1</figureCitation>
, 
<figureCitation id="13192A65A5751F6A0BC7B50DB8C0516A" box="[445,477,1060,1086]" captionStart="FIGURES 22−29" captionStartId="9.[151,264,1860,1885]" captionTargetBox="[241,1346,183,1834]" captionTargetId="figure@9.[241,1346,181,1834]" captionTargetPageId="9" captionText="FIGURES 22−29. Rivudiva spp. SEM photographs. (22) Labrum of R. inma, sp. nov.; (23) Labrum of R. trichobasis; (24) Maxilla of R. inma, sp. nov.; (25) Detail of crown of maxilla of R. inma, sp. nov., dorsal view; (26) Detail of crown of maxilla of R. trichobasis, lateral view; (27) Detail of crown of maxilla of R. minantenna, lateral view; (28) Scape and pedicel of R. trichobasis; (29) Outer and inner incisors of R. trichobasis. (ds, dentiseta; dc, dorsal canine; ii, inner incisor; oc, outer canine; oi, outer incisor; pe, pedicel; vc, ventral canine)." figureDoi="http://doi.org/10.5281/zenodo.3990794" httpUri="https://zenodo.org/record/3990794/files/figure.png" pageId="2" pageNumber="39">22</figureCitation>
); 3) dorsal surface of labrum with regular row of setae, plus few additional setae (
<figureCitation id="13192A65A5751F6A0F10B50DB9B85136" captionStart="FIGURES 1−6" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[167,1415,197,1905]" captionTargetId="figure@3.[153,1434,181,1920]" captionTargetPageId="3" captionText="FIGURES 1−6. Rivudiva inma, sp. nov., nymph: (1) Labrum; (2) Left mandible; (3) Right mandible; (4) Hypopharynx; (5) Maxilla; (6) Labium (left = ventral surface, right = dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.3990786" httpUri="https://zenodo.org/record/3990786/files/figure.png" pageId="2" pageNumber="39">Figs 1</figureCitation>
, 
<figureCitation id="13192A65A5751F6A0AC9B561B9CE5136" box="[179,211,1096,1122]" captionStart="FIGURES 22−29" captionStartId="9.[151,264,1860,1885]" captionTargetBox="[241,1346,183,1834]" captionTargetId="figure@9.[241,1346,181,1834]" captionTargetPageId="9" captionText="FIGURES 22−29. Rivudiva spp. SEM photographs. (22) Labrum of R. inma, sp. nov.; (23) Labrum of R. trichobasis; (24) Maxilla of R. inma, sp. nov.; (25) Detail of crown of maxilla of R. inma, sp. nov., dorsal view; (26) Detail of crown of maxilla of R. trichobasis, lateral view; (27) Detail of crown of maxilla of R. minantenna, lateral view; (28) Scape and pedicel of R. trichobasis; (29) Outer and inner incisors of R. trichobasis. (ds, dentiseta; dc, dorsal canine; ii, inner incisor; oc, outer canine; oi, outer incisor; pe, pedicel; vc, ventral canine)." figureDoi="http://doi.org/10.5281/zenodo.3990794" httpUri="https://zenodo.org/record/3990794/files/figure.png" pageId="2" pageNumber="39">22</figureCitation>
); 4) tuft on apex of lingua formed by long simple setae (
<figureCitation id="13192A65A5751F6A093EB561BA945136" box="[836,905,1096,1122]" captionStart="FIGURES 1−6" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[167,1415,197,1905]" captionTargetId="figure@3.[153,1434,181,1920]" captionTargetPageId="3" captionText="FIGURES 1−6. Rivudiva inma, sp. nov., nymph: (1) Labrum; (2) Left mandible; (3) Right mandible; (4) Hypopharynx; (5) Maxilla; (6) Labium (left = ventral surface, right = dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.3990786" httpUri="https://zenodo.org/record/3990786/files/figure.png" pageId="2" pageNumber="39">Fig. 4</figureCitation>
); 5) maxillary palp segment II apically rounded and with a nipple-like projection (
<figureCitation id="13192A65A5751F6A086CB545BB4251D2" box="[534,607,1131,1158]" captionStart="FIGURES 1−6" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[167,1415,197,1905]" captionTargetId="figure@3.[153,1434,181,1920]" captionTargetPageId="3" captionText="FIGURES 1−6. Rivudiva inma, sp. nov., nymph: (1) Labrum; (2) Left mandible; (3) Right mandible; (4) Hypopharynx; (5) Maxilla; (6) Labium (left = ventral surface, right = dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.3990786" httpUri="https://zenodo.org/record/3990786/files/figure.png" pageId="2" pageNumber="39">Figs 5</figureCitation>
, 
<figureCitation id="13192A65A5751F6A0816B545BB9151D2" box="[620,652,1132,1158]" captionStart="FIGURES 22−29" captionStartId="9.[151,264,1860,1885]" captionTargetBox="[241,1346,183,1834]" captionTargetId="figure@9.[241,1346,181,1834]" captionTargetPageId="9" captionText="FIGURES 22−29. Rivudiva spp. SEM photographs. (22) Labrum of R. inma, sp. nov.; (23) Labrum of R. trichobasis; (24) Maxilla of R. inma, sp. nov.; (25) Detail of crown of maxilla of R. inma, sp. nov., dorsal view; (26) Detail of crown of maxilla of R. trichobasis, lateral view; (27) Detail of crown of maxilla of R. minantenna, lateral view; (28) Scape and pedicel of R. trichobasis; (29) Outer and inner incisors of R. trichobasis. (ds, dentiseta; dc, dorsal canine; ii, inner incisor; oc, outer canine; oi, outer incisor; pe, pedicel; vc, ventral canine)." figureDoi="http://doi.org/10.5281/zenodo.3990794" httpUri="https://zenodo.org/record/3990794/files/figure.png" pageId="2" pageNumber="39">24</figureCitation>
); 6) glossa with base ventrally expanded, apex obliquely truncate and with dorsal thick setae distomedially (
<figureCitation id="13192A65A5751F6A083CB5B9BB9151FF" box="[582,652,1168,1195]" captionStart="FIGURES 1−6" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[167,1415,197,1905]" captionTargetId="figure@3.[153,1434,181,1920]" captionTargetPageId="3" captionText="FIGURES 1−6. Rivudiva inma, sp. nov., nymph: (1) Labrum; (2) Left mandible; (3) Right mandible; (4) Hypopharynx; (5) Maxilla; (6) Labium (left = ventral surface, right = dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.3990786" httpUri="https://zenodo.org/record/3990786/files/figure.png" pageId="2" pageNumber="39">Fig. 6</figureCitation>
); 7) hind wing pads present; 8) trochanter with robust, short and apically pointed setae (
<figureCitation id="13192A65A5751F6A0B0FB59DB8A4519B" box="[373,441,1204,1231]" captionStart="FIGURES 7−10" captionStartId="4.[151,264,1898,1923]" captionTargetBox="[182,1395,193,1828]" captionTargetId="figure@4.[151,1436,181,1873]" captionTargetPageId="4" captionText="FIGURES 7−10. Rivudiva inma, sp. nov., nymph. (7) Fore leg; (8) Tarsal claw; (9) Posterior margin of tergum IV; (10) Paraproct." figureDoi="http://doi.org/10.5281/zenodo.3990788" httpUri="https://zenodo.org/record/3990788/files/figure.png" pageId="2" pageNumber="39">Fig. 7</figureCitation>
); 9) outer dorsal row, medial dorsal row and inner dorsal row of femora with robust blunt setae, those of outer row at most 1/3 maximum width of femur (
<figureCitation id="13192A65A5751F6A0923B5F1BABC51A7" box="[857,929,1240,1267]" captionStart="FIGURES 7−10" captionStartId="4.[151,264,1898,1923]" captionTargetBox="[182,1395,193,1828]" captionTargetId="figure@4.[151,1436,181,1873]" captionTargetPageId="4" captionText="FIGURES 7−10. Rivudiva inma, sp. nov., nymph. (7) Fore leg; (8) Tarsal claw; (9) Posterior margin of tergum IV; (10) Paraproct." figureDoi="http://doi.org/10.5281/zenodo.3990788" httpUri="https://zenodo.org/record/3990788/files/figure.png" pageId="2" pageNumber="39">Figs 7</figureCitation>
, 
<figureCitation id="13192A65A5751F6A09D6B5F1BAEA51A6" box="[940,1015,1240,1266]" captionStart="FIGURES 30−37" captionStartId="10.[151,264,1910,1935]" captionTargetBox="[222,1364,181,1885]" captionTargetId="figure@10.[222,1364,181,1885]" captionTargetPageId="10" captionText="FIGURES 30−37. Rivudiva spp. SEM photographs. (30) Fore femur of R. inma, sp. nov.; (31) Fore femur of R. inma, sp. nov., detail of outer margin; (32) Femur of R. minantenna; (33) Femur of R. trichobasis; (34) Tarsal claw of R. inma, sp. nov., inner view; (35) Tarsal claw of R. minantenna, lateral view; (36) Abdominal sternum of R. trichobasis; (37) Abdominal sternum of R. trichobasis, detail of setae." figureDoi="http://doi.org/10.5281/zenodo.3990796" httpUri="https://zenodo.org/record/3990796/files/figure.png" pageId="2" pageNumber="39">30–31</figureCitation>
); 10) abdominal sterna without robust apically pointed setae; 11) paraproct with around 15 marginal spines (
<figureCitation id="13192A65A5751F6A09DEB5D5BAE55042" box="[932,1016,1276,1302]" captionStart="FIGURES 7−10" captionStartId="4.[151,264,1898,1923]" captionTargetBox="[182,1395,193,1828]" captionTargetId="figure@4.[151,1436,181,1873]" captionTargetPageId="4" captionText="FIGURES 7−10. Rivudiva inma, sp. nov., nymph. (7) Fore leg; (8) Tarsal claw; (9) Posterior margin of tergum IV; (10) Paraproct." figureDoi="http://doi.org/10.5281/zenodo.3990788" httpUri="https://zenodo.org/record/3990788/files/figure.png" pageId="2" pageNumber="39">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8B9D36E0A5751F6A0ABDB408BAAD50F2" blockId="2.[151,1437,1024,1447]" pageId="2" pageNumber="39">
<emphasis id="B956EAF2A5751F6A0ABDB408B80B506E" box="[199,278,1313,1338]" italics="true" pageId="2" pageNumber="39">Imago.</emphasis>
1) dorsal portion of turbinate eyes with inner margins parallel (
<figureCitation id="13192A65A5751F6A09A1B409BD00506E" box="[987,1053,1312,1338]" captionStart="FIGURES 1−6" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[167,1415,197,1905]" captionTargetId="figure@3.[153,1434,181,1920]" captionTargetPageId="3" captionText="FIGURES 1−6. Rivudiva inma, sp. nov., nymph: (1) Labrum; (2) Left mandible; (3) Right mandible; (4) Hypopharynx; (5) Maxilla; (6) Labium (left = ventral surface, right = dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.3990786" httpUri="https://zenodo.org/record/3990786/files/figure.png" pageId="2" pageNumber="39">Fig. 1</figureCitation>
of 
<bibRefCitation id="EFB34B11A5751F6A0E38B409BC6F506E" author="Salles, F. F. &amp; Nascimento, J. M. C." box="[1090,1394,1312,1338]" pageId="2" pageNumber="39" pagination="231 - 235" refId="ref7915" refString="Salles, F. F. &amp; Nascimento, J. M. C. (2009) The genus Rivudiva Lugo-Ortiz and McCafferty (Ephemeroptera: Baetidae): First generic description of adults, new combinations, and notes on the nymphs. Annales de Limnologie-International Journal of Limnology, 45 (4), 231 - 235. https: // doi. org / 10.1051 / limn / 2009023" type="journal article" year="2009">Salles &amp; Nascimento 2009</bibRefCitation>
); 2) length of forewing about 2.7× width; 3) hind wings present (fig. 5 of 
<bibRefCitation id="EFB34B11A5751F6A09D6B46DBDFD500A" author="Salles, F. F. &amp; Nascimento, J. M. C." box="[940,1248,1348,1374]" pageId="2" pageNumber="39" pagination="231 - 235" refId="ref7915" refString="Salles, F. F. &amp; Nascimento, J. M. C. (2009) The genus Rivudiva Lugo-Ortiz and McCafferty (Ephemeroptera: Baetidae): First generic description of adults, new combinations, and notes on the nymphs. Annales de Limnologie-International Journal of Limnology, 45 (4), 231 - 235. https: // doi. org / 10.1051 / limn / 2009023" type="journal article" year="2009">Salles &amp; Nascimento 2009</bibRefCitation>
); 4) forceps segment III oblong, 2× as long as wide (fig. 6 of 
<bibRefCitation id="EFB34B11A5751F6A08D8B441BAC850D6" author="Salles, F. F. &amp; Nascimento, J. M. C." box="[674,981,1384,1410]" pageId="2" pageNumber="39" pagination="231 - 235" refId="ref7915" refString="Salles, F. F. &amp; Nascimento, J. M. C. (2009) The genus Rivudiva Lugo-Ortiz and McCafferty (Ephemeroptera: Baetidae): First generic description of adults, new combinations, and notes on the nymphs. Annales de Limnologie-International Journal of Limnology, 45 (4), 231 - 235. https: // doi. org / 10.1051 / limn / 2009023" type="journal article" year="2009">Salles &amp; Nascimento 2009</bibRefCitation>
); 6) posterior margin of subgenital plate with a medial pointed projection (fig. 6 of 
<bibRefCitation id="EFB34B11A5751F6A080EB4A5BAB950F2" author="Salles, F. F. &amp; Nascimento, J. M. C." box="[628,932,1420,1446]" pageId="2" pageNumber="39" pagination="231 - 235" refId="ref7915" refString="Salles, F. F. &amp; Nascimento, J. M. C. (2009) The genus Rivudiva Lugo-Ortiz and McCafferty (Ephemeroptera: Baetidae): First generic description of adults, new combinations, and notes on the nymphs. Annales de Limnologie-International Journal of Limnology, 45 (4), 231 - 235. https: // doi. org / 10.1051 / limn / 2009023" type="journal article" year="2009">Salles &amp; Nascimento 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C338656BA5751F6D0AEDB4D1BC7E50DA" lastPageId="5" lastPageNumber="42" pageId="2" pageNumber="39" type="description">
<paragraph id="8B9D36E0A5751F6A0AEDB4D1B8355346" blockId="2.[151,296,1528,1554]" box="[151,296,1528,1554]" pageId="2" pageNumber="39">
<heading id="D0D5818CA5751F6A0AEDB4D1B8355346" bold="true" box="[151,296,1528,1554]" fontSize="11" level="3" pageId="2" pageNumber="39">
<emphasis id="B956EAF2A5751F6A0AEDB4D1B8355346" bold="true" box="[151,296,1528,1554]" pageId="2" pageNumber="39">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D36E0A5751F6A0AEDB769BBC8530F" blockId="2.[151,1437,1599,2023]" box="[151,725,1599,1627]" pageId="2" pageNumber="39">
<emphasis id="B956EAF2A5751F6A0AEDB769B9F3530E" box="[151,238,1600,1626]" italics="true" pageId="2" pageNumber="39">Nymph.</emphasis>
Lengths: body, 
<quantity id="4CDA9B05A5751F6A0BD8B716B8E6530F" box="[418,507,1599,1627]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="39" unit="mm" value="5.2">5.2 mm</quantity>
; antennae, 
<quantity id="4CDA9B05A5751F6A0802B769BBCC530F" box="[632,721,1600,1627]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="39" unit="mm" value="1.8">1.8 mm</quantity>
.
</paragraph>
<paragraph id="8B9D36E0A5751F6D0ABDB74DBA5E54A7" blockId="2.[151,1437,1599,2023]" lastBlockId="5.[151,1437,152,1422]" lastPageId="5" lastPageNumber="42" pageId="2" pageNumber="39">
Head. Coloration: pale yellow. Antenna pale yellow. Compound eyes pale yellow. Antenna. Scape and pedicel without spine-like setae. Labrum (
<figureCitation id="13192A65A5751F6A0863B7A1BB7C53F6" box="[537,609,1672,1698]" captionStart="FIGURES 1−6" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[167,1415,197,1905]" captionTargetId="figure@3.[153,1434,181,1920]" captionTargetPageId="3" captionText="FIGURES 1−6. Rivudiva inma, sp. nov., nymph: (1) Labrum; (2) Left mandible; (3) Right mandible; (4) Hypopharynx; (5) Maxilla; (6) Labium (left = ventral surface, right = dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.3990786" httpUri="https://zenodo.org/record/3990786/files/figure.png" pageId="2" pageNumber="39">Figs 1</figureCitation>
, 
<figureCitation id="13192A65A5751F6A080AB7A1BB8D53F6" box="[624,656,1672,1698]" captionStart="FIGURES 22−29" captionStartId="9.[151,264,1860,1885]" captionTargetBox="[241,1346,183,1834]" captionTargetId="figure@9.[241,1346,181,1834]" captionTargetPageId="9" captionText="FIGURES 22−29. Rivudiva spp. SEM photographs. (22) Labrum of R. inma, sp. nov.; (23) Labrum of R. trichobasis; (24) Maxilla of R. inma, sp. nov.; (25) Detail of crown of maxilla of R. inma, sp. nov., dorsal view; (26) Detail of crown of maxilla of R. trichobasis, lateral view; (27) Detail of crown of maxilla of R. minantenna, lateral view; (28) Scape and pedicel of R. trichobasis; (29) Outer and inner incisors of R. trichobasis. (ds, dentiseta; dc, dorsal canine; ii, inner incisor; oc, outer canine; oi, outer incisor; pe, pedicel; vc, ventral canine)." figureDoi="http://doi.org/10.5281/zenodo.3990794" httpUri="https://zenodo.org/record/3990794/files/figure.png" pageId="2" pageNumber="39">22</figureCitation>
). Rectangular, broader than long; length about 0.6x maximum width; distal margin with wide medial emargination; anterolateral and distal margins with bifid and pectinate setae; dorsal surface near distal margin with somewhat regular row of several simple setae. Left mandible (
<figureCitation id="13192A65A5751F6A0E92B7F9BC3353BE" box="[1256,1326,1744,1770]" captionStart="FIGURES 1−6" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[167,1415,197,1905]" captionTargetId="figure@3.[153,1434,181,1920]" captionTargetPageId="3" captionText="FIGURES 1−6. Rivudiva inma, sp. nov., nymph: (1) Labrum; (2) Left mandible; (3) Right mandible; (4) Hypopharynx; (5) Maxilla; (6) Labium (left = ventral surface, right = dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.3990786" httpUri="https://zenodo.org/record/3990786/files/figure.png" pageId="2" pageNumber="39">Fig. 2</figureCitation>
). Incisors cleft in two sets; outer and inner set of incisors respectively with 4 + 2 denticles; prostheca robust; margin between prostheca and mola straight; tuft of spine-like setae at base of mola present; subtriangular process wide; denticles of mola constricted; lateral margins convex. Right mandible (
<figureCitation id="13192A65A5751F6A092DB615BA835202" box="[855,926,1852,1878]" captionStart="FIGURES 1−6" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[167,1415,197,1905]" captionTargetId="figure@3.[153,1434,181,1920]" captionTargetPageId="3" captionText="FIGURES 1−6. Rivudiva inma, sp. nov., nymph: (1) Labrum; (2) Left mandible; (3) Right mandible; (4) Hypopharynx; (5) Maxilla; (6) Labium (left = ventral surface, right = dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.3990786" httpUri="https://zenodo.org/record/3990786/files/figure.png" pageId="2" pageNumber="39">Fig. 3</figureCitation>
). Incisors deeply cleft in two sets; outer and inner set of incisors respectively with 3 + 3 denticles; prostheca bifurcated at middle and pectinate; margin between prostheca and mola straight; tuft of spine-like setae at base of mola present; denticles of mola not constricted; apex of mola with two simple setae; lateral margins convex. Hypopharynx (
<figureCitation id="13192A65A5751F6A09DDB681BAF15296" box="[935,1004,1960,1986]" captionStart="FIGURES 1−6" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[167,1415,197,1905]" captionTargetId="figure@3.[153,1434,181,1920]" captionTargetPageId="3" captionText="FIGURES 1−6. Rivudiva inma, sp. nov., nymph: (1) Labrum; (2) Left mandible; (3) Right mandible; (4) Hypopharynx; (5) Maxilla; (6) Labium (left = ventral surface, right = dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.3990786" httpUri="https://zenodo.org/record/3990786/files/figure.png" pageId="2" pageNumber="39">Fig. 4</figureCitation>
). Lingua subequal to superlingua, subquadrangular with a distomedial pointed projection covered by tuft of long simple setae; superlingua with angulate outer margin; short, thin, simple setae scattered over distal margin of lingua and superlingua. Maxilla (
<figureCitation id="13192A65A5721F6D0F66B1B0BC7B55E7" box="[1308,1382,152,179]" captionStart="FIGURES 1−6" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[167,1415,197,1905]" captionTargetId="figure@3.[153,1434,181,1920]" captionTargetPageId="3" captionText="FIGURES 1−6. Rivudiva inma, sp. nov., nymph: (1) Labrum; (2) Left mandible; (3) Right mandible; (4) Hypopharynx; (5) Maxilla; (6) Labium (left = ventral surface, right = dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.3990786" httpUri="https://zenodo.org/record/3990786/files/figure.png" pageId="5" pageNumber="42">Figs 5</figureCitation>
, 
<figureCitation id="13192A65A5721F6D0F08B1B0B9AA5583" captionStart="FIGURES 22−29" captionStartId="9.[151,264,1860,1885]" captionTargetBox="[241,1346,183,1834]" captionTargetId="figure@9.[241,1346,181,1834]" captionTargetPageId="9" captionText="FIGURES 22−29. Rivudiva spp. SEM photographs. (22) Labrum of R. inma, sp. nov.; (23) Labrum of R. trichobasis; (24) Maxilla of R. inma, sp. nov.; (25) Detail of crown of maxilla of R. inma, sp. nov., dorsal view; (26) Detail of crown of maxilla of R. trichobasis, lateral view; (27) Detail of crown of maxilla of R. minantenna, lateral view; (28) Scape and pedicel of R. trichobasis; (29) Outer and inner incisors of R. trichobasis. (ds, dentiseta; dc, dorsal canine; ii, inner incisor; oc, outer canine; oi, outer incisor; pe, pedicel; vc, ventral canine)." figureDoi="http://doi.org/10.5281/zenodo.3990794" httpUri="https://zenodo.org/record/3990794/files/figure.png" pageId="5" pageNumber="42">24– 25</figureCitation>
). Maxillary palp 1.5× longer than galea-lacinia; segment II and III combined 0.8× the length of segment I, apex with inner nipple-like projection; maxillary palp with fine and simple setae scattered over surface. Labium (
<figureCitation id="13192A65A5721F6D0F30B1C9BC9355AF" box="[1354,1422,224,251]" captionStart="FIGURES 1−6" captionStartId="3.[151,264,1945,1970]" captionTargetBox="[167,1415,197,1905]" captionTargetId="figure@3.[153,1434,181,1920]" captionTargetPageId="3" captionText="FIGURES 1−6. Rivudiva inma, sp. nov., nymph: (1) Labrum; (2) Left mandible; (3) Right mandible; (4) Hypopharynx; (5) Maxilla; (6) Labium (left = ventral surface, right = dorsal surface)." figureDoi="http://doi.org/10.5281/zenodo.3990786" httpUri="https://zenodo.org/record/3990786/files/figure.png" pageId="5" pageNumber="42">Fig. 6</figureCitation>
). Glossa oblong, basally expanded, narrowing apically and subequal to paraglossa; apex with eight short spine-like setae; outer margin bare; inner margin with row of thick setae on apical 2/3; ventral surface covered by thin long setae. Paraglossa curved inward, but with distinct apical margin; apex with three rows of robust long spine-like setae; outer margin with long thin setae on distal 1/2. Labial palp with segment I subequal in length to segments II and III combined; segment II with rounded medial protuberance, outer margin and distomedial protuberance covered with thin, long and simple setae; segment III conical, length 0.7× width, covered by thin simple setae on outer margin, dorsal surface with robust spine-like setae near outer margin.
</paragraph>
<caption id="DF5D6668A5741F6B0AEDB6B0BA215282" ID-DOI="http://doi.org/10.5281/zenodo.3990786" ID-Zenodo-Dep="3990786" httpUri="https://zenodo.org/record/3990786/files/figure.png" pageId="3" pageNumber="40" startId="3.[151,264,1945,1970]" targetBox="[167,1415,197,1905]" targetPageId="3">
<paragraph id="8B9D36E0A5741F6B0AEDB6B0BA215282" blockId="3.[151,1436,1945,2006]" pageId="3" pageNumber="40">
<emphasis id="B956EAF2A5741F6B0AEDB6B0B82352E6" bold="true" box="[151,318,1945,1970]" pageId="3" pageNumber="40">FIGURES 1−6.</emphasis>
<taxonomicName id="4C224D63A5741F6B0B3DB6B3B8C152E5" authority="Salles &amp; Nieto &amp; Cruz, 2020" authorityName="Salles &amp; Nieto &amp; Cruz" authorityYear="2020" box="[327,476,1946,1970]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="40" phylum="Arthropoda" rank="species" species="inma" status="sp. nov.">
<emphasis id="B956EAF2A5741F6B0B3DB6B3B8C152E5" box="[327,476,1946,1970]" italics="true" pageId="3" pageNumber="40">Rivudiva inma</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A2655789A5741F6B0B90B6B3BB2052E5" box="[490,573,1946,1969]" pageId="3" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
, nymph: (1) Labrum; (2) Left mandible; (3) Right mandible; (4) Hypopharynx; (5) Maxilla; (6) Labium (left = ventral surface, right = dorsal surface).
</paragraph>
</caption>
<caption id="DF5D6668A5731F6C0AEDB643B9CE52F3" ID-DOI="http://doi.org/10.5281/zenodo.3990788" ID-Zenodo-Dep="3990788" httpUri="https://zenodo.org/record/3990788/files/figure.png" pageId="4" pageNumber="41" startId="4.[151,264,1898,1923]" targetBox="[182,1395,193,1828]" targetPageId="4">
<paragraph id="8B9D36E0A5731F6C0AEDB643B9CE52F3" blockId="4.[151,1436,1898,1959]" pageId="4" pageNumber="41">
<emphasis id="B956EAF2A5731F6C0AEDB643B85E52D7" bold="true" box="[151,323,1898,1923]" pageId="4" pageNumber="41">FIGURES 7−10</emphasis>
. 
<taxonomicName id="4C224D63A5731F6C0B2BB642B8F852D7" authority="Salles &amp; Nieto &amp; Cruz, 2020" authorityName="Salles &amp; Nieto &amp; Cruz" authorityYear="2020" box="[337,485,1899,1923]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="41" phylum="Arthropoda" rank="species" species="inma" status="sp. nov.">
<emphasis id="B956EAF2A5731F6C0B2BB642B8F852D7" box="[337,485,1899,1923]" italics="true" pageId="4" pageNumber="41">Rivudiva inma</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A2655789A5731F6C0B88B645BB5E52D7" box="[498,579,1900,1923]" pageId="4" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
, nymph. (7) Fore leg; (8) Tarsal claw; (9) Posterior margin of tergum IV; (10) Paraproct.
</paragraph>
</caption>
<paragraph id="8B9D36E0A5721F6D0ABDB0D5BC5357BF" blockId="5.[151,1437,152,1422]" pageId="5" pageNumber="42">
Thorax. Pale yellow. Hind wing pads present. Legs pale yellow. Foreleg (
<figureCitation id="13192A65A5721F6D0E5CB0D5BD6E5743" box="[1062,1139,508,535]" captionStart="FIGURES 7−10" captionStartId="4.[151,264,1898,1923]" captionTargetBox="[182,1395,193,1828]" captionTargetId="figure@4.[151,1436,181,1873]" captionTargetPageId="4" captionText="FIGURES 7−10. Rivudiva inma, sp. nov., nymph. (7) Fore leg; (8) Tarsal claw; (9) Posterior margin of tergum IV; (10) Paraproct." figureDoi="http://doi.org/10.5281/zenodo.3990788" httpUri="https://zenodo.org/record/3990788/files/figure.png" pageId="5" pageNumber="42">Figs 7</figureCitation>
, 
<figureCitation id="13192A65A5721F6D0EF8B0D5BDD05742" box="[1154,1229,508,534]" captionStart="FIGURES 30−37" captionStartId="10.[151,264,1910,1935]" captionTargetBox="[222,1364,181,1885]" captionTargetId="figure@10.[222,1364,181,1885]" captionTargetPageId="10" captionText="FIGURES 30−37. Rivudiva spp. SEM photographs. (30) Fore femur of R. inma, sp. nov.; (31) Fore femur of R. inma, sp. nov., detail of outer margin; (32) Femur of R. minantenna; (33) Femur of R. trichobasis; (34) Tarsal claw of R. inma, sp. nov., inner view; (35) Tarsal claw of R. minantenna, lateral view; (36) Abdominal sternum of R. trichobasis; (37) Abdominal sternum of R. trichobasis, detail of setae." figureDoi="http://doi.org/10.5281/zenodo.3990796" httpUri="https://zenodo.org/record/3990796/files/figure.png" pageId="5" pageNumber="42">30–31</figureCitation>
). Ratio of foreleg 1.2:(1):0.9:0.3. Fore femur. Length about 2.7× maximum width; outer and inner margins with short to elongate clavate setae, anterior and posterior surface covered with longitudinal row of clavate robust setae; length of setae on outer margin at most 1/3 maximum width of femur. Tibia. Dorsally bare; inner margin with a row of spine-like setae increasing in length toward apex. Tarsus. Inner margin with a row of spine-like setae. Tarsal claws as in 
<figureCitation id="13192A65A5721F6D0F22B3A3BC8157F0" box="[1368,1436,650,676]" captionStart="FIGURES 7−10" captionStartId="4.[151,264,1898,1923]" captionTargetBox="[182,1395,193,1828]" captionTargetId="figure@4.[151,1436,181,1873]" captionTargetPageId="4" captionText="FIGURES 7−10. Rivudiva inma, sp. nov., nymph. (7) Fore leg; (8) Tarsal claw; (9) Posterior margin of tergum IV; (10) Paraproct." figureDoi="http://doi.org/10.5281/zenodo.3990788" httpUri="https://zenodo.org/record/3990788/files/figure.png" pageId="5" pageNumber="42">Figs 8</figureCitation>
and 
<figureCitation id="13192A65A5721F6D0ABCB387B9F8579C" box="[198,229,686,712]" captionStart="FIGURES 30−37" captionStartId="10.[151,264,1910,1935]" captionTargetBox="[222,1364,181,1885]" captionTargetId="figure@10.[222,1364,181,1885]" captionTargetPageId="10" captionText="FIGURES 30−37. Rivudiva spp. SEM photographs. (30) Fore femur of R. inma, sp. nov.; (31) Fore femur of R. inma, sp. nov., detail of outer margin; (32) Femur of R. minantenna; (33) Femur of R. trichobasis; (34) Tarsal claw of R. inma, sp. nov., inner view; (35) Tarsal claw of R. minantenna, lateral view; (36) Abdominal sternum of R. trichobasis; (37) Abdominal sternum of R. trichobasis, detail of setae." figureDoi="http://doi.org/10.5281/zenodo.3990796" httpUri="https://zenodo.org/record/3990796/files/figure.png" pageId="5" pageNumber="42">34</figureCitation>
. Mid and hind leg. Femora narrower than fore femora; setae on anterior surface of femora longer than in fore femur; setae on posterior surface longer and pointed. Tibiae and tarsi with one row of short spine-like setae.
</paragraph>
<paragraph id="8B9D36E0A5721F6D0ABDB3DDBB2C562D" blockId="5.[151,1437,152,1422]" pageId="5" pageNumber="42">
Abdomen. Pale yellow, terga II–III and VI yellowish brown, anterior margins of segments II-VI brownish (
<figureCitation id="13192A65A5721F6D0F10B3DDB9FC5666" captionStart="FIGURES 38−40" captionStartId="11.[151,264,1939,1964]" captionTargetBox="[153,1436,446,1914]" captionTargetId="figure@11.[151,1436,443,1914]" captionTargetPageId="11" captionText="FIGURES 38−40. Habitus of living nymphs of Rivudiva spp. (38) Male of Rivudiva inma, sp. nov.; (39) Female of Rivudiva inma, sp. nov.; (40) Female of R. trichobasis." figureDoi="http://doi.org/10.5281/zenodo.3990798" httpUri="https://zenodo.org/record/3990798/files/figure.png" pageId="5" pageNumber="42">Figs 38–39</figureCitation>
). Tergal surface covered by pores and short spine-like setae; posterior margin with contiguous spination (
<figureCitation id="13192A65A5721F6D0F15B231B9BA5601" captionStart="FIGURES 7−10" captionStartId="4.[151,264,1898,1923]" captionTargetBox="[182,1395,193,1828]" captionTargetId="figure@4.[151,1436,181,1873]" captionTargetPageId="4" captionText="FIGURES 7−10. Rivudiva inma, sp. nov., nymph. (7) Fore leg; (8) Tarsal claw; (9) Posterior margin of tergum IV; (10) Paraproct." figureDoi="http://doi.org/10.5281/zenodo.3990788" httpUri="https://zenodo.org/record/3990788/files/figure.png" pageId="5" pageNumber="42">Fig. 9</figureCitation>
). Sterna pale yellow, bare. Gills whitish. Paraproct (
<figureCitation id="13192A65A5721F6D0894B212BA225601" box="[750,831,827,853]" captionStart="FIGURES 7−10" captionStartId="4.[151,264,1898,1923]" captionTargetBox="[182,1395,193,1828]" captionTargetId="figure@4.[151,1436,181,1873]" captionTargetPageId="4" captionText="FIGURES 7−10. Rivudiva inma, sp. nov., nymph. (7) Fore leg; (8) Tarsal claw; (9) Posterior margin of tergum IV; (10) Paraproct." figureDoi="http://doi.org/10.5281/zenodo.3990788" httpUri="https://zenodo.org/record/3990788/files/figure.png" pageId="5" pageNumber="42">Fig. 10</figureCitation>
) with around 15 marginal spines. Cercus and terminal filament without large lateral spines.
</paragraph>
<paragraph id="8B9D36E0A5721F6D0ABDB2AAB9C15694" blockId="5.[151,1437,152,1422]" pageId="5" pageNumber="42">
<emphasis id="B956EAF2A5721F6D0ABDB2AAB81256C8" box="[199,271,899,924]" italics="true" pageId="5" pageNumber="42">Imago</emphasis>
. Adequately characterized by 
<bibRefCitation id="EFB34B11A5721F6D0818B2ABBABE56C8" author="Salles, F. F. &amp; Nascimento, J. M. C." box="[610,931,898,924]" pageId="5" pageNumber="42" pagination="231 - 235" refId="ref7915" refString="Salles, F. F. &amp; Nascimento, J. M. C. (2009) The genus Rivudiva Lugo-Ortiz and McCafferty (Ephemeroptera: Baetidae): First generic description of adults, new combinations, and notes on the nymphs. Annales de Limnologie-International Journal of Limnology, 45 (4), 231 - 235. https: // doi. org / 10.1051 / limn / 2009023" type="journal article" year="2009">Salles &amp; Nascimento (2009)</bibRefCitation>
, although incorrectly associated to 
<taxonomicName id="4C224D63A5721F6D0F54B2AAB9C85694" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1998" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="minantenna">
<emphasis id="B956EAF2A5721F6D0F54B2AAB9C85694" italics="true" pageId="5" pageNumber="42">R. minantenna</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9D36E0A5721F6D0ABDB2E0BDA8517E" blockId="5.[151,1437,152,1422]" pageId="5" pageNumber="42">
<emphasis id="B956EAF2A5721F6D0ABDB2E0B85556B7" bold="true" box="[199,328,969,995]" pageId="5" pageNumber="42">Etymology</emphasis>
. After the Instituto Nacional da Mata Atlântica (INMA), the Brazilian National Institute of the Atlantic Forest. INMA is located in the city of Santa Teresa, locality of the new species, and one of the institute’s most important mission is the conservation of the Atlantic Forest and, therefore, of the new species.
</paragraph>
<paragraph id="8B9D36E0A5721F6D0ABDB51ABDA85188" blockId="5.[151,1437,152,1422]" pageId="5" pageNumber="42">
<emphasis id="B956EAF2A5721F6D0ABDB51AB855511A" bold="true" box="[199,328,1075,1102]" pageId="5" pageNumber="42">Comments</emphasis>
. Specimens of this species have been incorrectly assigned to 
<taxonomicName id="4C224D63A5721F6D0E68B51DBDA55119" authorityName="Lugo-Ortiz &amp; McCafferty" authorityYear="1998" box="[1042,1208,1076,1101]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="minantenna">
<emphasis id="B956EAF2A5721F6D0E68B51DBDA55119" box="[1042,1208,1076,1101]" italics="true" pageId="5" pageNumber="42">R. minantenna</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFB34B11A5721F6D0E9DB51AB8325125" author="Salles, F. F. &amp; Nascimento, J. M. C." pageId="5" pageNumber="42" pagination="231 - 235" refId="ref7915" refString="Salles, F. F. &amp; Nascimento, J. M. C. (2009) The genus Rivudiva Lugo-Ortiz and McCafferty (Ephemeroptera: Baetidae): First generic description of adults, new combinations, and notes on the nymphs. Annales de Limnologie-International Journal of Limnology, 45 (4), 231 - 235. https: // doi. org / 10.1051 / limn / 2009023" type="journal article" year="2009">Salles &amp; Nascimento (2009)</bibRefCitation>
due to several characteristics shared by them (see discussion for more details). The adults of the new species differ from the adults of 
<taxonomicName id="4C224D63A5721F6D087AB552BB9951C0" baseAuthorityName="Traver" baseAuthorityYear="1971" box="[512,644,1146,1172]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="covaloae">
<emphasis id="B956EAF2A5721F6D087AB552BB9951C0" box="[512,644,1146,1172]" italics="true" pageId="5" pageNumber="42">R. covaloae</emphasis>
</taxonomicName>
, the other species with hind wings, by the shape of the distal margin of the subgenital plate. In 
<taxonomicName id="4C224D63A5721F6D0BDBB5B6BB3B51EC" baseAuthorityName="Traver" baseAuthorityYear="1971" box="[417,550,1182,1208]" class="Insecta" family="Baetidae" genus="Rivudiva" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="42" phylum="Arthropoda" rank="species" species="covaloae">
<emphasis id="B956EAF2A5721F6D0BDBB5B6BB3B51EC" box="[417,550,1182,1208]" italics="true" pageId="5" pageNumber="42">R. covaloae</emphasis>
</taxonomicName>
the medial spine lies in a concavity of this margin (see fig. 2 of 
<bibRefCitation id="EFB34B11A5721F6D0F78B5B6BC9251EC" author="Traver, J. R." box="[1282,1423,1182,1209]" pageId="5" pageNumber="42" pagination="58 - 63" refId="ref8109" refString="Traver, J. R. (1971) Four new species of Neotropical Baetis (Ephemeroptera: Baetidae). Proceedings of the Entomological Society of Washington, 73, 58 - 63." type="journal article" year="1971">Traver 1971</bibRefCitation>
), while in the new species the distal margin is convex (see fig. 6 of 
<bibRefCitation id="EFB34B11A5721F6D0903B5E8BDB45188" author="Salles, F. F. &amp; Nascimento, J. M. C." box="[889,1193,1217,1244]" pageId="5" pageNumber="42" pagination="231 - 235" refId="ref7915" refString="Salles, F. F. &amp; Nascimento, J. M. C. (2009) The genus Rivudiva Lugo-Ortiz and McCafferty (Ephemeroptera: Baetidae): First generic description of adults, new combinations, and notes on the nymphs. Annales de Limnologie-International Journal of Limnology, 45 (4), 231 - 235. https: // doi. org / 10.1051 / limn / 2009023" type="journal article" year="2009">Salles &amp; Nascimento 2009</bibRefCitation>
).
</paragraph>
<paragraph id="8B9D36E0A5721F6D0ABDB5CDBA7C51AB" blockId="5.[151,1437,152,1422]" box="[199,865,1252,1279]" pageId="5" pageNumber="42">
<emphasis id="B956EAF2A5721F6D0ABDB5CDB87C51AB" bold="true" box="[199,353,1252,1279]" pageId="5" pageNumber="42">Distribution.</emphasis>
Southeastern 
<collectingCountry id="F3357670A5721F6D0B85B5CCBB5B51AB" box="[511,582,1253,1279]" name="Brazil" pageId="5" pageNumber="42">Brazil</collectingCountry>
, 
<collectingRegion id="49E6F802A5721F6D0828B5CCBBEC51AB" box="[594,753,1253,1279]" country="Brazil" name="Espirito Santo" pageId="5" pageNumber="42">Espírito Santo</collectingRegion>
(
<figureCitation id="13192A65A5721F6D097AB5CCBA4951AB" box="[768,852,1253,1279]" captionStart="FIGURE 41" captionStartId="13.[151,250,2017,2042]" captionTargetBox="[165,1418,319,1979]" captionTargetId="figure@13.[158,1430,309,1992]" captionTargetPageId="13" captionText="FIGURE 41. Map of South America showing the distribution of the distinct species of Rivudiva." figureDoi="http://doi.org/10.5281/zenodo.3990800" httpUri="https://zenodo.org/record/3990800/files/figure.png" pageId="5" pageNumber="42">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="8B9D36E0A5721F6D0ABDB421BC7E50DA" blockId="5.[151,1437,152,1422]" pageId="5" pageNumber="42">
<emphasis id="B956EAF2A5721F6D0ABDB421B8025077" bold="true" box="[199,287,1288,1315]" pageId="5" pageNumber="42">Biology</emphasis>
. Nymphs of the new species were found inhabiting the sandy substrates of few streams in Augusto Ruschi Biological Reserve, 
<collectingRegion id="49E6F802A5721F6D0B02B405BB055012" box="[376,536,1324,1350]" country="Brazil" name="Espirito Santo" pageId="5" pageNumber="42">Espírito Santo</collectingRegion>
, or in adjacent areas. They are difficult to collect due to an apparently low abundance and/or to an apparently patchy distribution in the extensive sandy areas of these streams. Besides, the color pattern of the nymphs generates a remarkable camouflage with its habitat (see fig. 8 of 
<bibRefCitation id="EFB34B11A5721F6D0E54B45DBC4550DA" author="Salles, F. F. &amp; Nascimento, J. M. C." box="[1070,1368,1396,1422]" pageId="5" pageNumber="42" pagination="231 - 235" refId="ref7915" refString="Salles, F. F. &amp; Nascimento, J. M. C. (2009) The genus Rivudiva Lugo-Ortiz and McCafferty (Ephemeroptera: Baetidae): First generic description of adults, new combinations, and notes on the nymphs. Annales de Limnologie-International Journal of Limnology, 45 (4), 231 - 235. https: // doi. org / 10.1051 / limn / 2009023" type="journal article" year="2009">Salles &amp; Nascimento 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>