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<mods:title id="73AEFCCBE4ED6725EBE9CCF07785602C">Revision of the genus Tmesiphorus LeConte, 1849 (Coleoptera: Staphylinidae: Pselaphinae) from Japan</mods:title>
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<taxonomicName id="4C224D17B3232B7462FAFE2BFDEA55ED" ID-CoL="579XY" authority="Weise, 1877" authorityName="Weise" authorityYear="1877" box="[151,568,463,490]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="costatus">
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<emphasis id="B956EA86B3232B7462FAFE2BFE4D55EE" bold="true" box="[151,415,463,490]" italics="true" pageId="2" pageNumber="69">Tmesiphorus costatus</emphasis>
<bibRefCitation id="EFB34B65B3232B7463C8FE2BFDEA55ED" author="Weise, J." box="[421,568,463,489]" pageId="2" pageNumber="69" pagination="88 - 100" refId="ref10783" refString="Weise, J. (1877) Japanische Staphilinidae und Pselaphidae. Deutsche entomologische Zeitschrift, 21, 88 - 100. https: // doi. org / 10.1002 / mmnd. 4800210118" type="journal article" year="1877">Weise, 1877</bibRefCitation>
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<paragraph id="8B9D3694B3232B7462FAFE10FD63560A" blockId="2.[151,689,463,562]" box="[151,689,499,526]" pageId="2" pageNumber="69">[Japanese name: Misuji-higekata-arizukamushi]</paragraph>
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(
<figureCitation id="13192A11B3232B7462F2FDF3FED05636" box="[159,258,535,562]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1841,1866]" captionStartId-1="4.[151,250,1939,1964]" captionStartId-2="5.[151,250,802,827]" captionTargetBox-0="[187,1401,477,1777]" captionTargetBox-1="[189,1398,506,1867]" captionTargetBox-2="[247,1335,193,773]" captionTargetId-0="figure@3.[186,1401,462,1777]" captionTargetId-1="figure@4.[189,1398,506,1867]" captionTargetId-2="figure@5.[227,1359,181,781]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. A, B: Habitus of Tmesiphorus costatus, C, D: left antenna. A, male; B, female; C, antenna in dorsal view; D, antennal club in ventral view by SEM." captionText-1="FIGURE 2. Tmesiphorus costatus, male. A, head in ventral view; B, head in dorsal view; C, pronotum; D, elytra; E, abdomen; F, left protarsus." captionText-2="FIGURE 3. Male genitalia of Tmesiphorus costatus. A, ventral view; B, lateral view; C, dorsal view." httpUri-0="https://zenodo.org/record/3346341/files/figure.png" httpUri-1="https://zenodo.org/record/3346343/files/figure.png" httpUri-2="https://zenodo.org/record/3346345/files/figure.png" pageId="2" pageNumber="69">Figs 13</figureCitation>
)
</paragraph>
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<taxonomicName id="4C224D17B3232B7462FAFDB8FDF05670" ID-CoL="579XY" authority="Weise, 1877: 99" authorityName="Weise" authorityPageNumber="99" authorityYear="1877" box="[151,546,603,628]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="costatus">
<emphasis id="B956EA86B3232B7462FAFDB8FEA75670" box="[151,373,604,628]" italics="true" pageId="2" pageNumber="69">Tmesiphorus costatus</emphasis>
<treatmentCitation id="0A831085B3232B746311FDBFFDF05670" author="Weise, J." box="[380,546,603,628]" page="99" pageId="2" pageNumber="69" year="1877">
<bibRefCitation id="EFB34B65B3232B746311FDBFFDF05670" author="Weise, J." box="[380,546,603,628]" pageId="2" pageNumber="69" pagination="88 - 100" refId="ref10783" refString="Weise, J. (1877) Japanische Staphilinidae und Pselaphidae. Deutsche entomologische Zeitschrift, 21, 88 - 100. https: // doi. org / 10.1002 / mmnd. 4800210118" type="journal article" year="1877">Weise, 1877: 99</bibRefCitation>
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(original description);
<treatmentCitation id="0A831085B3232B74617FFDBFFC015670" author="Raffray, A." box="[786,979,603,628]" page="374" pageId="2" pageNumber="69" year="1908">
<bibRefCitation id="EFB34B65B3232B74617FFDBFFC015670" author="Raffray, A." box="[786,979,603,628]" pageId="2" pageNumber="69" pagination="1 - 487" refId="ref10494" refString="Raffray, A. (1908) Coleoptera. Fam. Pselaphidae. Genera Insectorum, 64, 1 - 487, 9 pls." type="book chapter" year="1908">Raffray, 1908: 374</bibRefCitation>
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(catalogue);
<treatmentCitation id="0A831085B3232B746636FDBFFAC95670" author="Raffray, A." box="[1115,1307,603,628]" page="157" pageId="2" pageNumber="69" year="1911">
<bibRefCitation id="EFB34B65B3232B746636FDBFFAC95670" author="Raffray, A." box="[1115,1307,603,628]" pageId="2" pageNumber="69" pagination="1 - 222" refId="ref10519" refString="Raffray, A. (1911) Pselaphidae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 27. W. Junk, Berlin, pp. 1 - 222. https: // doi. org / 10.1007 / 978 - 94 - 011 - 9697 - 0 _ 1" type="book chapter" year="1911">Raffray, 1911: 157</bibRefCitation>
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(catalogue);
<treatmentCitation id="0A831085B3232B7462AAFD9EFE565696" author="Jeannel, R." box="[199,388,634,659]" page="118" pageId="2" pageNumber="69" year="1958">
<bibRefCitation id="EFB34B65B3232B7462AAFD9EFE565696" author="Jeannel, R." box="[199,388,634,659]" pageId="2" pageNumber="69" pagination="1 - 138" refId="ref9702" refString="Jeannel, R. (1958) Revision des Pselaphides du Japon. Memoires du Museum national d'histoire naturelle, Paris, Serie A, Zoologie, 18, 1 - 138." type="journal article" year="1958">Jeannel, 1958: 118</bibRefCitation>
</treatmentCitation>
(diagnosis);
<treatmentCitation id="0A831085B3232B74606AFD9FFD1E5697" author="Nomura, S." box="[519,716,634,659]" page="158" pageId="2" pageNumber="69" year="2001">
<bibRefCitation id="EFB34B65B3232B74606AFD9FFD1E5697" author="Nomura, S." box="[519,716,634,659]" pageId="2" pageNumber="69" pagination="141 - 161" refId="ref10278" refString="Nomura, S. (2001) Taxonomical review and a list of the pselaphine Species (Staphylinidae, Pselaphinae) known from Japan. Elytra, Tokyo, 29 (1), 141 - 161." type="journal article" year="2001">Nomura, 2001: 158</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0A831085B3232B74613DFD9EFBAD5696" author="Lobl, I. &amp; Besuchet, C." box="[848,1151,634,659]" page="327" pageId="2" pageNumber="69" year="2004">
<bibRefCitation id="EFB34B65B3232B74613DFD9EFBAD5696" author="Lobl, I. &amp; Besuchet, C." box="[848,1151,634,659]" pageId="2" pageNumber="69" pagination="272 - 329" refId="ref9960" refString="Lobl, I. &amp; Besuchet, C. (2004) Subfamily Pselaphinae Latreille, 1802. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 2. Hydrophiloidea-Histeroidea-Staphylinoidea. Apollo Books, Stenstrup, pp. 272 - 329." type="book chapter" year="2004">Löbl and Besuchet, 2004: 327</bibRefCitation>
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(catalogue);
<treatmentCitation id="0A831085B3232B74676EFD9FFF3F56B5" author="Nomura, S." page="101" pageId="2" pageNumber="69" year="2006">
<bibRefCitation id="EFB34B65B3232B74676EFD9FFF3F56B5" author="Nomura, S." pageId="2" pageNumber="69" pagination="99 - 102" refId="ref10316" refString="Nomura, S. (2006) Taxonomical notes on three pselaphine Species (Staphylinidae, Pselaphidae) described by Weise, 1877 from Japan. Elytra, Tokyo, 34 (1), 99 - 102." type="journal article" year="2006">Nomura, 2006: 101</bibRefCitation>
</treatmentCitation>
(taxonomic note);
<treatmentCitation id="0A831085B3232B7463C2FD7CFD4B56B5" author="Shibata, Y. &amp; Maruyama, M. &amp; Hoshina, H. &amp; Kishimoto, T. &amp; Naomi, S. &amp; Nomura, S. &amp; Puthz, V. &amp; Shimada, T. &amp; Watanabe, Y. &amp; Yamamoto, S." box="[431,665,664,689]" page="97" pageId="2" pageNumber="69" year="2013">
<bibRefCitation id="EFB34B65B3232B7463C2FD7CFD4B56B5" author="Shibata, Y. &amp; Maruyama, M. &amp; Hoshina, H. &amp; Kishimoto, T. &amp; Naomi, S. &amp; Nomura, S. &amp; Puthz, V. &amp; Shimada, T. &amp; Watanabe, Y. &amp; Yamamoto, S." box="[431,665,664,689]" pageId="2" pageNumber="69" pagination="69 - 218" refId="ref10679" refString="Shibata, Y., Maruyama, M., Hoshina, H., Kishimoto, T., Naomi, S., Nomura, S., Puthz, V., Shimada, T., Watanabe, Y. &amp; Yamamoto, S. (2013) Catalogue of Japanese Staphylinidae (Insecta: Coleoptera). Bulletin of the Kyushu University Museum, 11, 69 - 218." type="journal article" year="2013">
Shibata
<emphasis id="B956EA86B3232B74606CFD7EFDE656B5" box="[513,564,665,689]" italics="true" pageId="2" pageNumber="69">et al.</emphasis>
, 2013: 97
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(catalogue).
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</paragraph>
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<typeStatus id="54998836B3232B7462FAFD04FF0056FE" box="[151,210,736,762]" pageId="2" pageNumber="69">Type</typeStatus>
locality.
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<collectingCountry id="F3357604B3232B746351FD04FEAD56FE" box="[316,383,736,762]" name="Japan" pageId="2" pageNumber="69">Japan</collectingCountry>
,
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,
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.
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<emphasis id="B956EA86B3232B7462AAFCE0FEA7571A" bold="true" box="[199,373,771,798]" pageId="2" pageNumber="69">Type material.</emphasis>
<materialsCitation id="3B4A3CC9B3232B746311FCE0FBAF571A" ID-GBIF-Occurrence="2293867385" box="[380,1149,771,798]" collectionCode="MNHU" country="Japan" location="Hiller" pageId="2" pageNumber="69" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B956EA86B3232B746311FCE0FE3B571A" bold="true" box="[380,489,772,798]" pageId="2" pageNumber="69">Holotype</emphasis>
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:
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, “
<collectingCountry id="F3357604B3232B74605FFCE0FDA5571A" box="[562,631,772,798]" name="Japan" pageId="2" pageNumber="69">Japan</collectingCountry>
,
<location id="8EFD604FB3232B7460EEFCE7FD165719" LSID="urn:lsid:plazi:treatment:038B8782B3232B7262FAFE2BFB0055AB:8EFD604FB3232B7460EEFCE7FD165719" box="[643,708,771,797]" country="Japan" name="Hiller" pageId="2" pageNumber="69">Hiller</location>
//
<taxonomicName id="4C224D17B3232B74608CFCE7FC06571A" box="[737,980,771,798]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="costatus">Tmesiphorus costatus</taxonomicName>
W.” (
<collectionCode id="ED33AE51B3232B746678FCE0FBA3571A" box="[1045,1137,772,798]" pageId="2" pageNumber="69">MNHU</collectionCode>
).
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<emphasis id="B956EA86B3232B7462AAFCC3FE605746" bold="true" box="[199,434,807,834]" pageId="2" pageNumber="69">Material examined.</emphasis>
(
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,
<specimenCount id="9D24FD1DB3232B74639BFCCCFDC95746" box="[502,539,808,834]" pageId="2" pageNumber="69" type="female">4♀</specimenCount>
).
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<collectingCountry id="F3357604B3232B74605CFCCCFDA75746" box="[561,629,808,834]" name="Japan" pageId="2" pageNumber="69">Japan</collectingCountry>
: [
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]:
<specimenCount id="9D24FD1DB3232B746096FCCCFCCD5746" box="[763,799,808,834]" pageId="2" pageNumber="69" type="male">1♂</specimenCount>
,
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,
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,
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,
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9
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</date>
, Y. Hi- rano leg. (
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);
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,
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,
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,
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,
<location id="8EFD604FB3232B746105FCAFFC0C5761" LSID="urn:lsid:plazi:treatment:038B8782B3232B7262FAFE2BFB0055AB:8EFD604FB3232B746105FCAFFC0C5761" box="[872,990,843,869]" country="Japan" county="Ishikawa-ken" municipality="Suzu-shi" name="Suzu-jinja" pageId="2" pageNumber="69">Suzu-jinja</location>
,
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leg. (
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,
<collectingMunicipality id="6BF9ACEEB3232B7463C3FC94FDFB578E" box="[430,553,880,906]" pageId="2" pageNumber="69">Kamo-gun</collectingMunicipality>
,
<location id="8EFD604FB3232B74605EFC8BFD6B578D" LSID="urn:lsid:plazi:treatment:038B8782B3232B7262FAFE2BFB0055AB:8EFD604FB3232B74605EFC8BFD6B578D" box="[563,697,879,905]" country="Japan" county="Hiroshima-ken" municipality="Kamo-gun" name="Kurose-cho" pageId="2" pageNumber="69">Kurose-chô</location>
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,
<materialsCitation id="3B4A3CC9B3232B7460A9FC94FAA4578D" ID-GBIF-Occurrence="2293867384" box="[708,1398,879,906]" collectingDate="1988-05-02" collectionCode="NSMT" collectorName="I. Okamoto" country="Japan" location="Kure-shi" pageId="2" pageNumber="69" specimenCount="1" specimenCount-female="1" stateProvince="Hiroshima">
<collectingDate id="EFD8E9BCB3232B7460A9FC94FCFF578E" box="[708,813,880,906]" pageId="2" pageNumber="69" value="1988-05-02">
<date id="FF9C1054B3232B7460A9FC94FCFF578E" box="[708,813,880,906]" pageId="2" pageNumber="69" value="1988-05-02">2 V 1988</date>
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,
<collectorName id="26D75342B3232B746155FC94FC65578D" box="[824,951,879,906]" pageId="2" pageNumber="69">I. Okamoto</collectorName>
leg. (
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);
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,
<collectingRegion id="49E6F876B3232B7466E0FC8BFAD4578D" box="[1165,1286,879,905]" country="Japan" name="Hiroshima" pageId="2" pageNumber="69">Hiroshima</collectingRegion>
,
<location id="8EFD604FB3232B74677CFC8BFAA4578D" LSID="urn:lsid:plazi:treatment:038B8782B3232B7262FAFE2BFB0055AB:8EFD604FB3232B74677CFC8BFAA4578D" box="[1297,1398,879,905]" country="Japan" name="Kure-shi" pageId="2" pageNumber="69" stateProvince="Hiroshima">Kure-shi</location>
</materialsCitation>
,
<materialsCitation id="3B4A3CC9B3232B7467EDFC94FD2C57A9" ID-GBIF-Occurrence="2293867391" collectingDate="1994-09-10" collectionCode="NSMT" collectorName="I. Okamoto" country="Japan" location="Kure-shi" pageId="2" pageNumber="69" specimenCount="1" specimenCount-female="1" stateProvince="Hiroshima">
<collectingDate id="EFD8E9BCB3232B7467EDFC94FF2557AA" pageId="2" pageNumber="69" value="1994-09-10">
<date id="FF9C1054B3232B7467EDFC94FF2557AA" pageId="2" pageNumber="69" value="1994-09-10">10 IX 1994</date>
</collectingDate>
,
<collectorName id="26D75342B3232B74636EFC70FE5057A9" box="[259,386,915,942]" pageId="2" pageNumber="69">I. Okamoto</collectorName>
leg. (
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);
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, same data, but
</materialsCitation>
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<collectingDate id="EFD8E9BCB3232B746168FC70FCBF57AA" box="[773,877,916,942]" pageId="2" pageNumber="69" value="1988-05-02">
<date id="FF9C1054B3232B746168FC70FCBF57AA" box="[773,877,916,942]" pageId="2" pageNumber="69" value="1988-05-02">2 V 1988</date>
</collectingDate>
(
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);
<specimenCount id="9D24FD1DB3232B746189FC70FBDA57AA" box="[996,1032,916,942]" pageId="2" pageNumber="69" type="male">1♂</specimenCount>
,
<collectingCounty id="62FC4E18B3232B746678FC77FB1257A9" box="[1045,1216,915,941]" pageId="2" pageNumber="69">Hiroshima-ken</collectingCounty>
,
<collectingMunicipality id="6BF9ACEEB3232B7466A1FC77FF6857D5" pageId="2" pageNumber="69">Higashihiroshimashi</collectingMunicipality>
,
<location id="8EFD604FB3232B7462A5FC53FEE457D5" LSID="urn:lsid:plazi:treatment:038B8782B3232B7262FAFE2BFB0055AB:8EFD604FB3232B7462A5FC53FEE457D5" box="[200,310,951,977]" country="Japan" county="Hiroshima-ken" municipality="Higashihiroshimashi" name="Saijo-cho" pageId="2" pageNumber="69">Saijô-chô</location>
,
<location id="8EFD604FB3232B74632EFC53FE4F57D5" LSID="urn:lsid:plazi:treatment:038B8782B3232B7262FAFE2BFB0055AB:8EFD604FB3232B74632EFC53FE4F57D5" box="[323,413,951,977]" country="Japan" county="Hiroshima-ken" municipality="Higashihiroshimashi" name="Shitami" pageId="2" pageNumber="69">Shitami</location>
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,
<date id="FF9C1054B3232B7463C4FC5CFDFD57D6" box="[425,559,952,978]" pageId="2" pageNumber="69" value="2014-04-17">
17
<collectionCode id="ED33AE51B3232B7463A3FC5CFE3957D6" box="[462,491,952,978]" pageId="2" pageNumber="69">IV</collectionCode>
<materialsCitation id="3B4A3CC9B3232B746399FC5CFDFD57D6" ID-GBIF-Occurrence="2293867386" box="[500,559,952,978]" collectingDate="1988-05-02" collectionCode="KUM" collectorName="K. Maehara" country="Japan" county="Hiroshima-ken" location="Saijo-cho" municipality="Higashihiroshimashi" pageId="2" pageNumber="69" specimenCount="1" specimenCount-male="1">2014</materialsCitation>
</date>
,
<collectorName id="26D75342B3232B746051FC5CFD1357D5" box="[572,705,951,978]" pageId="2" pageNumber="69">K. Maehara</collectorName>
leg. (
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);
<specimenCount id="9D24FD1DB3232B74610CFC5CFC5757D6" box="[865,901,952,978]" pageId="2" pageNumber="69" type="male">1♂</specimenCount>
, same data, but
<date id="FF9C1054B3232B74662DFC5CFB1157D6" box="[1088,1219,952,978]" pageId="2" pageNumber="69" value="2014-04-24">
24
<collectionCode id="ED33AE51B3232B746608FC5CFB5057D6" box="[1125,1154,952,978]" pageId="2" pageNumber="69">IV</collectionCode>
<materialsCitation id="3B4A3CC9B3232B7466E6FC5CFB1157D6" ID-GBIF-Occurrence="2293867389" box="[1163,1219,952,978]" collectingDate="1988-05-02" collectionCode="KUM" collectorName="K. Maehara" country="Japan" county="Hiroshima-ken" location="Saijo-cho" municipality="Higashihiroshimashi" pageId="2" pageNumber="69" specimenCount="1" specimenCount-female="1">2014</materialsCitation>
</date>
(
<collectionCode id="ED33AE51B3232B7466B9FC5CFAC957D6" box="[1236,1307,952,978]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="69" type="Herbarium">KUM</collectionCode>
);
<specimenCount id="9D24FD1DB3232B74675DFC5CFA8157D6" box="[1328,1363,952,978]" pageId="2" pageNumber="69" type="female">1♀</specimenCount>
, same data, but
<date id="FF9C1054B3232B746292FC38FEAC57F2" box="[255,382,988,1014]" pageId="2" pageNumber="69" value="2017-04-18">
18
<collectionCode id="ED33AE51B3232B74634FFC38FEED57F2" box="[290,319,988,1014]" pageId="2" pageNumber="69">IV</collectionCode>
2017
</date>
(
<collectionCode id="ED33AE51B3232B7463E0FC38FE0657F2" box="[397,468,988,1014]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="2" pageNumber="69" type="Herbarium">KUM</collectionCode>
).
</paragraph>
<paragraph id="8B9D3694B3232B7462AAFC1BFE7B5082" blockId="2.[151,1437,735,2022]" pageId="2" pageNumber="69">
<emphasis id="B956EA86B3232B7462AAFC1BFE90501E" bold="true" box="[199,322,1023,1050]" pageId="2" pageNumber="69">Diagnosis.</emphasis>
<taxonomicName id="4C224D17B3232B74632AFC1BFDEB501D" box="[327,569,1023,1049]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="costatus">
<emphasis id="B956EA86B3232B74632AFC1BFDEB501D" box="[327,569,1023,1049]" italics="true" pageId="2" pageNumber="69">Tmesiphorus costatus</emphasis>
</taxonomicName>
is readily distinguishable from the other Japanese congeners by a combination of strong sulcus on each elytron that extends from the median fovea to the posterior 2/3 of the elytra, and tergites IVVI strongly carinate along their midlines. This species is smaller than the other Japanese
<taxonomicName id="4C224D17B3232B7466DEFBA3FA965065" authorityName="LeConte" authorityYear="1849" box="[1203,1348,1095,1121]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="genus">
<emphasis id="B956EA86B3232B7466DEFBA3FA965065" box="[1203,1348,1095,1121]" italics="true" pageId="2" pageNumber="69">Tmesiphorus</emphasis>
</taxonomicName>
species found in Honshû,
<collectingCountry id="F3357604B3232B74630CFB88FE765082" box="[353,420,1132,1158]" name="Japan" pageId="2" pageNumber="69">Japan</collectingCountry>
.
</paragraph>
<paragraph id="8B9D3694B3232B7462AAFB6BFF0D50CA" blockId="2.[151,1437,735,2022]" pageId="2" pageNumber="69">
<emphasis id="B956EA86B3232B7462AAFB6BFEA750AE" bold="true" box="[199,373,1167,1194]" pageId="2" pageNumber="69">Redescription.</emphasis>
Male (
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). Body length
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. Body dark brown to reddish-brown, elytra testaceous.
</paragraph>
<paragraph id="8B9D3694B3232B7462AAFB33FC3752C1" blockId="2.[151,1437,735,2022]" pageId="2" pageNumber="69">
Head (
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) about as long as wide, HL
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, HW
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, almost ovoid in dorsal view, coarsely punctate with setae; frons narrow, with longitudinal groove including fovea narrowed posteriorly to vertex; vertex with two foveae, with short longitudinal carina between vertexal foveae, shallowly depressed around vertexal foveae; eyes large, prominent; occiput rounded, with bundles of yellow setae; antennae (
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) slightly thick, antennomeres I thick, elongate, each 2.5 times longer than II; II as long as wide, each slightly wider than III; IIIVI each longer than wide, narrowed at base, thickened apically; VIIVIII each transverse; VII each slightly longer than VIII; VIII each slightly wider than VII; IXXI enlarged to form club, slightly curved inward in outer margin; IX each longer than wide, 1.3 times longer than I, 1.2 times wider than I, narrowed at base, thickened apically, longitudinally depressed with dense short setae on ventral surface; with a depression on ventral surface from basal 1/4 to apex, broadened apically, 1/3 as wide as segmental width on apical margin; X each longer than wide on inner margin, as long as wide on outer margin, longitudinally depressed with sparse setae on ventral side; XI slightly wider than IX at middle, produced outward, narrowed apically from middle, shallowly depressed with sparse setae on ventral side; maxillary palpi asymmetrical; segments II and III each with penicillate spine on lateral margin; segments
<collectionCode id="ED33AE51B3232B74628CF948FF2C52C2" box="[225,254,1708,1734]" pageId="2" pageNumber="69">IV</collectionCode>
thick, with rounded lobe on lateral margin, apical portion conical.
</paragraph>
<paragraph id="8B9D3694B3232B7562AAF934FA4654D3" blockId="2.[151,1437,735,2022]" lastBlockId="3.[151,1437,153,395]" lastPageId="3" lastPageNumber="70" pageId="2" pageNumber="69">
Pronotum (
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) about as long as wide,
<collectionCode id="ED33AE51B3232B7460C3F934FD1C52EE" box="[686,718,1744,1770]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="2" pageNumber="69" type="Museum">PL</collectionCode>
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,
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; dorsum broadly round, widest anterior to middle, narrowed posteriorly, sparsely covered with setae, with distinct median and lateral foveae at base, with prominent convexity extending from anterior margin to median fovea; elytra (
<figureCitation id="13192A11B3232B7466CEF8F3FAD45336" box="[1187,1286,1815,1842]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1964]" captionTargetBox="[189,1398,506,1867]" captionTargetId="figure@4.[189,1398,506,1867]" captionTargetPageId="4" captionText="FIGURE 2. Tmesiphorus costatus, male. A, head in ventral view; B, head in dorsal view; C, pronotum; D, elytra; E, abdomen; F, left protarsus." httpUri="https://zenodo.org/record/3346343/files/figure.png" pageId="2" pageNumber="69">Fig. 2-D</figureCitation>
) much wider than long, EL
<quantity id="4CDA9B71B3232B74635BF8DFFE095352" box="[310,475,1851,1878]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" metricValueMax="6.5" metricValueMin="6.3" pageId="2" pageNumber="69" unit="mm" value="0.64" valueMax="0.65" valueMin="0.63">0.630.65 mm</quantity>
, EW
<quantity id="4CDA9B71B3232B746075F8DFFD6F5352" box="[536,701,1851,1878]" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="2" pageNumber="69" unit="mm" value="0.925" valueMax="0.95" valueMin="0.9">0.900.95 mm</quantity>
, coarsely punctate with setae; each elytron with basal median and inner foveae pubescent, with inner and lateral longitudinal carinae, strongly sulcate from median fovea to posterior 3/4 of elytra; inner carinae extending from between two basal foveae to posterior 3/4 of elytra; lateral carinae extending from anterior to posterior margin; posterior margin with thick brush-like yellow setae; profemora thickest, with white setae on anterior face; mesofemora as thick as metafemora; protibiae thickest, arcuate and thickened at middle, as long as mesotibiae; metatibiae longest; tibiae all with dense yellow setae at apices; tarsi (
<figureCitation id="13192A11B3222B75669AFF7DFA8154B7" box="[1271,1363,153,179]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1964]" captionTargetBox="[189,1398,506,1867]" captionTargetId="figure@4.[189,1398,506,1867]" captionTargetPageId="4" captionText="FIGURE 2. Tmesiphorus costatus, male. A, head in ventral view; B, head in dorsal view; C, pronotum; D, elytra; E, abdomen; F, left protarsus." httpUri="https://zenodo.org/record/3346343/files/figure.png" pageId="3" pageNumber="70">Fig. 2-F</figureCitation>
) elongate; tarsomeres
<date id="FF9C1054B3222B756338FF59FEAF54D3" box="[341,381,189,215]" pageId="3" pageNumber="70">II 2</date>
/3 as long as III; protarsal claws asymmetrical, anterior claws each thicker than posterior one.
</paragraph>
<paragraph id="8B9D3694B3222B7562AAFF05FD35558F" blockId="3.[151,1437,153,395]" pageId="3" pageNumber="70">
Abdomen (
<figureCitation id="13192A11B3222B75632BFF05FE7754FF" box="[326,421,225,251]" captionStart="FIGURE 2" captionStartId="4.[151,250,1939,1964]" captionTargetBox="[189,1398,506,1867]" captionTargetId="figure@4.[189,1398,506,1867]" captionTargetPageId="4" captionText="FIGURE 2. Tmesiphorus costatus, male. A, head in ventral view; B, head in dorsal view; C, pronotum; D, elytra; E, abdomen; F, left protarsus." httpUri="https://zenodo.org/record/3346343/files/figure.png" pageId="3" pageNumber="70">Fig. 2-E</figureCitation>
) large, AL
<quantity id="4CDA9B71B3222B75604AFF05FD5C54FF" box="[551,654,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="3" pageNumber="70" unit="mm" value="0.78">0.78 mm</quantity>
, AW
<quantity id="4CDA9B71B3222B7560A0FF05FCA154FF" box="[717,883,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="9.6" metricValueMax="9.8" metricValueMin="9.399999999999999" pageId="3" pageNumber="70" unit="mm" value="0.96" valueMax="0.98" valueMin="0.94">0.940.98 mm</quantity>
, wider than long, with sparse setae; tergite
<collectionCode id="ED33AE51B3222B75670DFF05FAAF54FF" box="[1376,1405,225,251]" pageId="3" pageNumber="70">IV</collectionCode>
as long as
<collectionCode id="ED33AE51B3222B756282FEE1FEC2551B" box="[239,272,261,287]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="70">VI</collectionCode>
; tergite V longest; tergite VII small; tergite VIII smallest, semicircular; tergites IVVI each remarkably convex along midline; tergites
<collectionCode id="ED33AE51B3222B75639BFECDFDC15547" box="[502,531,297,323]" pageId="3" pageNumber="70">IV</collectionCode>
and V each with lateral longitudinal carinae, those of tergite
<collectionCode id="ED33AE51B3222B7566A7FECDFB355547" box="[1226,1255,297,323]" pageId="3" pageNumber="70">IV</collectionCode>
extending from anterior margin to posterior margin, those of tergite V extending from anterior to posterior margin; tergite
<collectionCode id="ED33AE51B3222B75672BFEA9FAB15563" box="[1350,1379,333,359]" pageId="3" pageNumber="70">IV</collectionCode>
with thick brush-like yellow setae along posterior margin.
</paragraph>
<caption id="DF5D661CB3222B7562FAF8D5FE3A536A" httpUri="https://zenodo.org/record/3346341/files/figure.png" pageId="3" pageNumber="70" startId="3.[151,250,1841,1866]" targetBox="[187,1401,477,1777]" targetPageId="3">
<paragraph id="8B9D3694B3222B7562FAF8D5FE3A536A" blockId="3.[151,1436,1841,1902]" pageId="3" pageNumber="70">
<emphasis id="B956EA86B3222B7562FAF8D5FE9D534E" bold="true" box="[151,335,1841,1866]" pageId="3" pageNumber="70">FIGURE 1. A, B:</emphasis>
Habitus of
<taxonomicName id="4C224D17B3222B7563AEF8D6FD4D534E" box="[451,671,1842,1866]" class="Insecta" family="Staphylinidae" genus="Tmesiphorus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="costatus">
<emphasis id="B956EA86B3222B7563AEF8D6FD4D534E" box="[451,671,1842,1866]" italics="true" pageId="3" pageNumber="70">Tmesiphorus costatus</emphasis>
</taxonomicName>
, C, D: left antenna. A, male; B, female; C, antenna in dorsal view; D, antennal club in ventral view by SEM.
</paragraph>
</caption>
<paragraph id="8B9D3694B3222B7262AAF878FE6754D3" blockId="3.[151,1437,1948,2011]" lastBlockId="4.[151,1437,153,431]" lastPageId="4" lastPageNumber="71" pageId="3" pageNumber="70">
Male genitalia (
<figureCitation id="13192A11B3222B756315F878FE6953B3" box="[376,443,1948,1975]" captionStart="FIGURE 3" captionStartId="5.[151,250,802,827]" captionTargetBox="[247,1335,193,773]" captionTargetId="figure@5.[227,1359,181,781]" captionTargetPageId="5" captionText="FIGURE 3. Male genitalia of Tmesiphorus costatus. A, ventral view; B, lateral view; C, dorsal view." httpUri="https://zenodo.org/record/3346345/files/figure.png" pageId="3" pageNumber="70">Fig. 3</figureCitation>
)
<quantity id="4CDA9B71B3222B7563A6F879FDFD53B3" box="[459,559,1949,1975]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="70" unit="mm" value="0.31">0.31 mm</quantity>
in length, well-sclerotized, small, oval; parameres symmetric, projecting ventrally; each paramere with three setae; median lobe nearly symmetric, strongly projecting ventrally, slender; basal bulb broad, weakly narrowed apically; apical part slightly asymmetric; secondary gonopore small; endophallus elongate, circular at base.
</paragraph>
<paragraph id="8B9D3694B3252B7262AAFF05FD7D5563" blockId="4.[151,1437,153,431]" pageId="4" pageNumber="71">
Female (
<figureCitation id="13192A11B3252B72634AFF05FE5454FF" box="[295,390,225,251]" captionStart="FIGURE 1" captionStartId="3.[151,250,1841,1866]" captionTargetBox="[187,1401,477,1777]" captionTargetId="figure@3.[186,1401,462,1777]" captionTargetPageId="3" captionText="FIGURE 1. A, B: Habitus of Tmesiphorus costatus, C, D: left antenna. A, male; B, female; C, antenna in dorsal view; D, antennal club in ventral view by SEM." httpUri="https://zenodo.org/record/3346341/files/figure.png" pageId="4" pageNumber="71">Fig. 1-B</figureCitation>
). BL
<quantity id="4CDA9B71B3252B7263AFFF05FDB454FF" box="[450,614,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="2.305" metricValueMax="2.39" metricValueMin="2.22" pageId="4" pageNumber="71" unit="mm" value="2.305" valueMax="2.39" valueMin="2.22">2.222.39 mm</quantity>
; HL
<quantity id="4CDA9B71B3252B7260F7FF05FCEC54FF" box="[666,830,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="4.55" metricValueMax="4.699999999999999" metricValueMin="4.4" pageId="4" pageNumber="71" unit="mm" value="0.45499999999999996" valueMax="0.47" valueMin="0.44">0.440.47 mm</quantity>
; HW
<quantity id="4CDA9B71B3252B726111FF05FBF254FF" box="[892,1056,225,251]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.6" metricValueMin="4.1" pageId="4" pageNumber="71" unit="mm" value="0.435" valueMax="0.46" valueMin="0.41">0.410.46 mm</quantity>
;
<collectionCode id="ED33AE51B3252B726647FF05FB9854FF" box="[1066,1098,225,251]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="4" pageNumber="71" type="Museum">PL</collectionCode>
<quantity id="4CDA9B71B3252B726622FF04FB2654FF" box="[1103,1268,224,251]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.0" metricValueMin="4.699999999999999" pageId="4" pageNumber="71" unit="mm" value="0.485" valueMax="0.5" valueMin="0.47">0.470.50 mm</quantity>
;
<collectionCode id="ED33AE51B3252B726690FF05FAF454FF" box="[1277,1318,225,251]" country="Thailand" name="Paleontological Collections" pageId="4" pageNumber="71">PW</collectionCode>
<quantity id="4CDA9B71B3252B726741FF04FF14551B" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.0" metricValueMin="4.4" pageId="4" pageNumber="71" unit="mm" value="0.47" valueMax="0.5" valueMin="0.44">0.440.50 mm</quantity>
; EL
<quantity id="4CDA9B71B3252B726296FEE0FE70551B" box="[251,418,260,287]" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="5.8999999999999995" metricValueMin="5.6" pageId="4" pageNumber="71" unit="mm" value="0.575" valueMax="0.59" valueMin="0.56">0.560.59 mm</quantity>
; EW
<quantity id="4CDA9B71B3252B72638CFEE1FD5A551B" box="[481,648,261,287]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" metricValueMax="8.8" metricValueMin="8.0" pageId="4" pageNumber="71" unit="mm" value="0.84" valueMax="0.88" valueMin="0.8">0.800.88 mm</quantity>
; AL
<quantity id="4CDA9B71B3252B7260D3FEE0FCB7551B" box="[702,869,260,287]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" metricValueMax="8.299999999999999" metricValueMin="7.5" pageId="4" pageNumber="71" unit="mm" value="0.79" valueMax="0.83" valueMin="0.75">0.750.83 mm</quantity>
; AW
<quantity id="4CDA9B71B3252B7261CEFEE1FB98551B" box="[931,1098,261,287]" metricMagnitude="-4" metricUnit="m" metricValue="9.1" metricValueMax="9.399999999999999" metricValueMin="8.8" pageId="4" pageNumber="71" unit="mm" value="0.9099999999999999" valueMax="0.94" valueMin="0.88">0.880.94 mm</quantity>
. Antennae with antennomeres IIVIII more slender than in male, successively broadening towards apices; antennomeres IXXI not modified, smaller than in male, antennal club unmodified.
</paragraph>
<paragraph id="8B9D3694B3252B7262AAFE94FB645588" blockId="4.[151,1437,153,431]" box="[199,1206,368,396]" pageId="4" pageNumber="71">
<emphasis id="B956EA86B3252B7262AAFE94FEB3558F" bold="true" box="[199,353,368,395]" pageId="4" pageNumber="71">Distribution.</emphasis>
<collectingCountry id="F3357604B3252B72630AFE95FE79558F" box="[359,427,369,395]" name="Japan" pageId="4" pageNumber="71">Japan</collectingCountry>
: Honshû (Iwate-ken, Ishikawa-ken, Hiroshima-ken, Yamaguchi-ken).
</paragraph>
<paragraph id="8B9D3694B3252B7262AAFE70FB0055AB" blockId="4.[151,1437,153,431]" box="[199,1234,404,431]" pageId="4" pageNumber="71">
<emphasis id="B956EA86B3252B7262AAFE70FEF555AB" bold="true" box="[199,295,404,431]" pageId="4" pageNumber="71">Biology.</emphasis>
The specimens were collected by flight intercept traps (
<collectionCode id="ED33AE51B3252B7261F7FE71FC1555AB" box="[922,967,405,431]" pageId="4" pageNumber="71">FIT</collectionCode>
) or by sifting leaf litter.
</paragraph>
</subSubSection>
</treatment>
</document>