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<document id="9A7DFC1C1FBBBA8F79F19DF86E86ED51" ID-DOI="10.24199/j.mmv.2011.68.01" ID-ISSN="1447-2554" ID-Zenodo-Dep="10665999" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708005216384" checkinUser="felipe" docAuthor="Darragh, Thomas A." docDate="2011" docId="242187E6FFFD7E6819EE21A3C3C9AF16" docLanguage="en" docName="MemMusVictoria.68.1-28.pdf" docOrigin="Memoirs of Museum Victoria 68" docSource="https://museumsvictoria.com.au/collections-research/journals/memoirs-of-museum-victoria/volume-68-2011/pages-1-28/" docStyle="DocumentStyle:5B3B5C052A93BFEEE334AD6097A94E45.2:MemMusVictoria.2003-.journal_article.type1" docStyleId="5B3B5C052A93BFEEE334AD6097A94E45" docStyleName="MemMusVictoria.2003-.journal_article.type1" docStyleVersion="2" docTitle="Zoila didymorhyncha Darragh 2011, sp. nov." docType="treatment" docVersion="2" lastPageNumber="9" masterDocId="D818FF9EFFFB7E601962277BC16FA93E" masterDocTitle="A revision of the Australian fossil species of Zoila (Gastropoda: Cypraeidae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="7" updateTime="1708005626492" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="860B6E80203BF1046F97F9AD2B83D6C2">A revision of the Australian fossil species of Zoila (Gastropoda: Cypraeidae)</mods:title>
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<mods:namePart id="1AF2E524C97261462239405C1948B642">Darragh, Thomas A.</mods:namePart>
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<mods:title id="17101C43EBE79DA438A50B7BAD849F3A">Memoirs of Museum Victoria</mods:title>
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<mods:identifier id="6A3EDEA24C92B6B212967BC081CCA482" type="Zenodo-Dep">10665999</mods:identifier>
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<treatment id="242187E6FFFD7E6819EE21A3C3C9AF16" LSID="urn:lsid:plazi:treatment:242187E6FFFD7E6819EE21A3C3C9AF16" httpUri="http://treatment.plazi.org/id/242187E6FFFD7E6819EE21A3C3C9AF16" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="E492657BFFFD7E6619EE21A3C0AFAFD0" box="[140,448,1752,1774]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="AC3736F0FFFD7E6619EE21A3C0AFAFD0" blockId="6.[140,448,1752,1774]" box="[140,448,1752,1774]" pageId="6" pageNumber="7">
<heading id="F77F819CFFFD7E6619EE21A3C0AFAFD0" box="[140,448,1752,1774]" fontSize="9" level="2" pageId="6" pageNumber="7" reason="5">
<taxonomicName id="6B884D73FFFD7E6619EE21A3C000AFD0" ID-CoL="5D6LF" authority="Darragh, 2011" authorityName="Darragh" authorityYear="2011" box="[140,367,1752,1774]" class="Gastropoda" family="Cypraeidae" genus="Zoila" kingdom="Animalia" order="Littorinimorpha" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="didymorhyncha" status="sp. nov.">
<emphasis id="9EFCEAE2FFFD7E6619EE21A3C000AFD0" bold="true" box="[140,367,1752,1774]" italics="true" pageId="6" pageNumber="7">Zoila didymorhyncha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="85CF5799FFFD7E66181721A2C0AFAFD0" box="[373,448,1753,1774]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E492657BFFFD7E6819EE207EC3C9AF16" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="AC3736F0FFFD7E6619EE207EC03AAE24" blockId="6.[140,341,1797,1819]" box="[140,341,1797,1819]" pageId="6" pageNumber="7">
<figureCitation id="34B32A75FFFD7E6619EE207EC194AE24" box="[140,251,1797,1819]" captionStart="Figure 1" captionStartId="5.[147,206,1767,1786]" captionTargetBox="[133,1468,233,1722]" captionTargetId="figure-7@5.[127,1474,227,1727]" captionTargetPageId="5" captionText="Figure 1. A, Cypraeorbis medius (Deshayes,1835), × 5;B, Cypraeorbis sphaeroides (Conrad,1848),× 5; C, Cypraeorbis ventripotens (Cossmann, 1903), × 5; D, Cypraeorbis viathomsoni sp. nov. WAM 72.296, × 5; E, Zoila didymorhyncha sp. nov. P302687, × 2.6; F, Zoila campestris sp. nov. WAM 89.437, × 2.6; G, Zoila fodinata sp. nov. WAM 89.637, × 3; H, Zoila chathamensis (Cernohorsky, 1971) GNSTM 87922, × 2.3. Arrows indicate the location of the notch." figureDoi="http://doi.org/10.5281/zenodo.10666001" httpUri="https://zenodo.org/record/10666001/files/figure.png" pageId="6" pageNumber="7">Figures 1E</figureCitation>
,
<figureCitation id="34B32A75FFFD7E661864207EC03AAE24" box="[262,341,1797,1819]" captionStart="Figure 6" captionStartId="10.[140,199,1535,1554]" captionTargetBox="[146,1420,267,1493]" captionTargetId="figure-137@10.[140,1425,224,1493]" captionTargetPageId="10" captionText="Figure 6. AE, I, Zoila didymorhyncha; AC, paratype, P302685, PL3022, × 1; DE, I, holotype, P302687, PL 3022, × 1; FH, Z. glomerabilis, paratype, P315526, Half Moon Bay, Torquay, × 1." figureDoi="http://doi.org/10.5281/zenodo.10666015" httpUri="https://zenodo.org/record/10666015/files/figure.png" pageId="6" pageNumber="7">6AE, I</figureCitation>
</paragraph>
<paragraph id="AC3736F0FFFD7E6619EE204AC5B2ADE6" blockId="6.[140,771,1841,1891]" lastBlockId="6.[809,1441,959,1240]" pageId="6" pageNumber="7">
<emphasis id="9EFCEAE2FFFD7E6619EE204AC06BAE78" box="[140,260,1841,1862]" italics="true" pageId="6" pageNumber="7">Description</emphasis>
. Shell solid, small for genus, elongate, subfusiform; base relatively flat, but rounded on either side of aperture. Spire not visible. Posterior canal long, with rounded sides, sunk into massive rounded posterior rostrum formed by extensions of inner and outer lips. Anterior canal long, with rounded sides, sunk into a thick, rather flat rostrum. Aperture somewhat sinuous; outer lip with 1024 well-developed teeth extending along entire lip; teeth completely obsolete on
<specimenCount id="BA8EFD79FFFD7E661C6E2334C4F2AD5B" box="[1292,1437,1103,1125]" count="1" pageId="6" pageNumber="7" type="generic">one specimen</specimenCount>
. Columella lip with 326 well-developed teeth extending along entire lip on most specimens; teeth obsolete on
<specimenCount id="BA8EFD79FFFD7E661C6D23F2C4F2ADA0" box="[1295,1437,1160,1182]" count="1" pageId="6" pageNumber="7" type="generic">one specimen</specimenCount>
. Fossula moderately developed, very shallow, bounded anteriorly by a small, weak, terminal ridge.
</paragraph>
<paragraph id="AC3736F0FFFD7E661A4B2392C52CAF4F" blockId="6.[809,942,1257,1280]" lastBlockId="6.[809,1440,1540,1649]" pageId="6" pageNumber="7">
<emphasis id="9EFCEAE2FFFD7E661A4B2392C2C8AC3E" box="[809,935,1257,1280]" italics="true" pageId="6" pageNumber="7">Dimensions</emphasis>
.
<materialsCitation id="1CE03CADFFFD7E661A4B217EC550AF4F" collectionCode="AMG" collectorName="Torquay BT &amp; Jan Juc Formation. Early Janjukian" county="Addiscot Beach" location="Demons Bluff Formation" municipality="Bl" pageId="6" pageNumber="7" specimenCode="PL3022" specimenCount="1" stateProvince="Victoria" typeStatus="holotype">
<emphasis id="9EFCEAE2FFFD7E661A4B217EC2C0AF27" box="[809,943,1540,1562]" italics="true" pageId="6" pageNumber="7">
<typeStatus id="73338852FFFD7E661A4B217EC237AF24" box="[809,856,1541,1562]" pageId="6" pageNumber="7">Type</typeStatus>
locality
</emphasis>
.
<specimenCode id="FC2E9E8BFFFD7E661AA2217FC57FAF27" box="[960,1040,1540,1561]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="6" pageNumber="7" type="Museum">PL3022</specimenCode>
cliff section at
<collectingCounty id="45564E7CFFFD7E661DDF217FC40CAF24" box="[1213,1379,1540,1562]" pageId="6" pageNumber="7">Addiscot Beach</collectingCounty>
, beds
<collectingMunicipality id="4C53AC8AFFFD7E661A4B215AC22EAF09" box="[809,833,1569,1591]" pageId="6" pageNumber="7">Bl</collectingMunicipality>
09l07, southwest of small gully, clay overlying
<location id="A957602BFFFD7E661C2E2159C2BDAF6A" LSID="urn:lsid:plazi:treatment:242187E6FFFD7E6819EE21A3C3C9AF16:A957602BFFFD7E661C2E2159C2BDAF6A" county="Addiscot Beach" municipality="Bl" name="Demons Bluff Formation" pageId="6" pageNumber="7" stateProvince="Victoria">Demons Bluff Formation</location>
,
<collectingRegion id="6E4CF812FFFD7E661ABD2145C55BAF6A" box="[991,1076,1598,1620]" country="Australia" name="Victoria" pageId="6" pageNumber="7">Victoria</collectingRegion>
.
<collectionCode id="CA99AE35FFFD7E661D232145C513AF6D" box="[1089,1148,1598,1619]" country="South Africa" httpUri="http://grbio.org/cool/b9qz-7eka" name="Albany Museum" pageId="6" pageNumber="7">AMG</collectionCode>
<collectorName id="017D5326FFFD7E661DE72145C46DAF6A" box="[1157,1282,1598,1620]" pageId="6" pageNumber="7">Torquay BT</collectorName>
6l9490.
<collectorName id="017D5326FFFD7E661C362145C550AF4F" pageId="6" pageNumber="7">Jan Juc Formation. Early Janjukian</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="AC3736F0FFFD7E661A4B21FCC2FAAFEB" blockId="6.[809,1440,1671,1750]" pageId="6" pageNumber="7">
<emphasis id="9EFCEAE2FFFD7E661A4B21FCC2D7AFA2" box="[809,952,1671,1692]" italics="true" pageId="6" pageNumber="7">Type material</emphasis>
.
<materialsCitation id="1CE03CADFFFD7E661AAA21FCC2FAAFEB" collectingDate="1985-12-04" collectingDateMax="1990-05-08" collectingDateMin="1985-12-04" collectionCode="T" collectorName="A. Darragh" pageId="6" pageNumber="7" specimenCount="2" typeStatus="holotype">
<typeStatus id="73338852FFFD7E661AAA21FCC549AFA3" box="[968,1062,1671,1693]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
P302687, collected
<collectionCode id="CA99AE35FFFD7E661D9921FCC463AFA2" box="[1275,1292,1671,1692]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="7">T</collectionCode>
.
<collectorName id="017D5326FFFD7E661C6D21FCC4E9AFA3" box="[1295,1414,1671,1693]" pageId="6" pageNumber="7">A. Darragh</collectorName>
,
<date id="D8361030FFFD7E661CF121FCC2A2AF87" pageId="6" pageNumber="7" value="1985-12-04">
<collectingDate id="C872E9D8FFFD7E661CF121FCC2A2AF87" pageId="6" pageNumber="7" value="1985-12-04">4 December 1985</collectingDate>
</date>
;
<typeStatus id="73338852FFFD7E661ABA21DFC55FAF87" box="[984,1072,1700,1721]" pageId="6" pageNumber="7" type="paratype">paratype</typeStatus>
P302685, collected
<collectionCode id="CA99AE35FFFD7E661C6121DFC47BAF87" box="[1283,1300,1700,1721]" name="Tavera, Department of Geology and Geophysics" pageId="6" pageNumber="7">T</collectionCode>
.
<collectorName id="017D5326FFFD7E661C7521DFC4E4AF87" box="[1303,1419,1699,1721]" pageId="6" pageNumber="7">A. Darragh</collectorName>
<date id="D8361030FFFD7E661CF121DFC2FEAFEB" pageId="6" pageNumber="7" value="1990-05-08">
<collectingDate id="C872E9D8FFFD7E661CF121DFC2FEAFEB" pageId="6" pageNumber="7" value="1990-05-08">8 May 1990</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="AC3736F0FFFD7E661A4B2196C59CAE3C" blockId="6.[809,1267,1772,1794]" box="[809,1267,1772,1794]" pageId="6" pageNumber="7">
<emphasis id="9EFCEAE2FFFD7E661A4B2196C2F4AE3C" box="[809,923,1773,1794]" italics="true" pageId="6" pageNumber="7">Time range</emphasis>
. Early Janjukian, Late Oligocene.
</paragraph>
<paragraph id="AC3736F0FFFD7E661A4B2063C2CFAE75" blockId="6.[809,1440,1816,1867]" pageId="6" pageNumber="7">
<emphasis id="9EFCEAE2FFFD7E661A4B2063C556AE13" box="[809,1081,1816,1837]" italics="true" pageId="6" pageNumber="7">Occurrence and material</emphasis>
.
<materialsCitation id="1CE03CADFFFD7E661D2F2063C2F3AE75" collectionCode="PL" pageId="6" pageNumber="7" specimenCode="PL3022" specimenCount="3">
<specimenCode id="FC2E9E8BFFFD7E661D2F2063C5F3AE13" box="[1101,1180,1816,1837]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="6" pageNumber="7" type="Museum">PL3022</specimenCode>
Addiscott Beach (
<specimenCount id="BA8EFD79FFFD7E661C0E2063C2FAAE75" count="3" pageId="6" pageNumber="7" type="generic">three specimens</specimenCount>
)
</materialsCitation>
.
</paragraph>
<caption id="F8F76678FFFC7E6719F121A6C0D6AE67" ID-DOI="http://doi.org/10.5281/zenodo.10666005" ID-Zenodo-Dep="10666005" httpUri="https://zenodo.org/record/10666005/files/figure.png" pageId="7" pageNumber="8" startId="7.[147,207,1757,1776]" targetBox="[155,1428,267,1736]" targetPageId="7" targetType="figure">
<paragraph id="AC3736F0FFFC7E6719F121A6C0D6AE67" blockId="7.[147,1448,1757,1881]" pageId="7" pageNumber="8">
Figure 3. AG,
<taxonomicName id="6B884D73FFFC7E67184321A6C26AAFCE" authority="Moodys Branch Formation" authorityName="Moodys Branch Formation" box="[289,773,1757,1776]" class="Gastropoda" family="Cypraeidae" genus="Cypraeorbis" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="ventripotens">
<emphasis id="9EFCEAE2FFFC7E67184321A6C366AFCE" box="[289,521,1757,1776]" italics="true" pageId="7" pageNumber="8">Cypraeorbis ventripotens</emphasis>
Moodys Branch Formation
</taxonomicName>
, late Eocene, Town Creek, Jackson, Mississippi, United States of America; AC, × 2; F, × 4; DE, G, × 2; HJ, N,
<taxonomicName id="6B884D73FFFC7E671B632183C550AE35" authority="MNHN, Chipola Formation, Burdigalian" authorityName="MNHN, Chipola Formation, Burdigalian" box="[513,1087,1784,1803]" class="Gastropoda" family="Cypraeidae" genus="Cypraeorbis" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="arlettae">
<emphasis id="9EFCEAE2FFFC7E671B632183C3AFAE35" box="[513,704,1784,1803]" italics="true" pageId="7" pageNumber="8">Cypraeorbis arlettae</emphasis>
MNHN, Chipola Formation, Burdigalian
</taxonomicName>
, early Miocene, Farley Creek, Calhoun County, Florida, USA; K, O,
<taxonomicName id="6B884D73FFFC7E6718C52069C287AE1B" authority="MNHN, Chipola Formation, Burdigalian" authorityName="MNHN, Chipola Formation, Burdigalian" box="[423,1000,1810,1829]" class="Gastropoda" family="Cypraeidae" genus="Cypraeorbis" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="wilcoxi">
<emphasis id="9EFCEAE2FFFC7E6718C52069C30DAE1B" box="[423,610,1810,1829]" italics="true" pageId="7" pageNumber="8">Cypraeorbis wilcoxi</emphasis>
MNHN, Chipola Formation, Burdigalian
</taxonomicName>
, early Miocene, Farley Creek, Calhoun County, Florida, United States of America; LM,
<taxonomicName id="6B884D73FFFC7E671B782056C299AE01" authority="Liltved and Le Roux, 1988" authorityName="Liltved and Le Roux" authorityYear="1988" box="[538,1014,1836,1855]" class="Gastropoda" family="Cypraeidae" genus="Barycypraea" kingdom="Animalia" order="Littorinimorpha" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="zietsmani">
<emphasis id="9EFCEAE2FFFC7E671B782056C380AE00" box="[538,751,1836,1855]" italics="true" pageId="7" pageNumber="8">Barycypraea zietsmani</emphasis>
Liltved and Le Roux, 1988
</taxonomicName>
, P31664, Alexandria Formation, late Neogene, Eastern Cape, South Africa, × 1.
</paragraph>
</caption>
<caption id="F8F76678FFF37E6819EE226EC450AC16" ID-DOI="http://doi.org/10.5281/zenodo.10666007" ID-Zenodo-Dep="10666007" box="[140,1343,1301,1320]" httpUri="https://zenodo.org/record/10666007/files/figure.png" pageId="8" pageNumber="9" startId="8.[140,199,1301,1320]" targetBox="[148,1417,230,1251]" targetPageId="8" targetType="figure">
<paragraph id="AC3736F0FFF37E6819EE226EC450AC16" blockId="8.[140,1343,1301,1320]" box="[140,1343,1301,1320]" pageId="8" pageNumber="9">
Figure 4. AF,
<taxonomicName id="6B884D73FFF37E681877226EC3EFAC16" authority="MNHN, Bartonian" authorityName="MNHN, Bartonian" box="[277,640,1301,1320]" class="Gastropoda" family="Cypraeidae" genus="Cypraeorbis" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="medius">
<emphasis id="9EFCEAE2FFF37E681877226EC0A2AC16" box="[277,461,1301,1320]" italics="true" pageId="8" pageNumber="9">Cypraeorbis medius</emphasis>
MNHN, Bartonian
</taxonomicName>
, late Eocene, Bois du Rois, Auvers sur Oise, France; AC, DE, × 1.5; F, × 2.
</paragraph>
</caption>
<paragraph id="AC3736F0FFF37E6819EE2208C381ACC2" blockId="8.[140,771,1395,1532]" pageId="8" pageNumber="9">
<emphasis id="9EFCEAE2FFF37E6819EE2208C18CACB6" box="[140,227,1395,1416]" italics="true" pageId="8" pageNumber="9">Remarks</emphasis>
. The small size and the massive development of the anterior and posterior rostra separate this species from all others in the genus. In morphology, it comes closest to
<taxonomicName id="6B884D73FFF37E681B1422D6C1A7ACDE" authority="(Tate)" baseAuthorityName="Tate" baseAuthorityYear="1892" class="Gastropoda" family="Cypraeidae" genus="Zoila" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="mulderi">
<emphasis id="9EFCEAE2FFF37E681B1422D6C26CACFC" box="[630,771,1453,1474]" italics="true" pageId="8" pageNumber="9">Zoila mulderi</emphasis>
(Tate)
</taxonomicName>
, but it has a massive anterior rostrum lacking in
<taxonomicName id="6B884D73FFF37E681BF822B1C26CACE1" baseAuthorityName="Tate" baseAuthorityYear="1892" box="[666,771,1482,1503]" class="Gastropoda" family="Cypraeidae" genus="Zoila" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="mulderi">
<emphasis id="9EFCEAE2FFF37E681BF822B1C26CACE1" box="[666,771,1482,1503]" italics="true" pageId="8" pageNumber="9">Z. mulderi</emphasis>
</taxonomicName>
and the posterior rostrum is not notched as it is in
<taxonomicName id="6B884D73FFF37E681B1D229CC387ACC5" baseAuthorityName="Tate" baseAuthorityYear="1892" box="[639,744,1510,1532]" class="Gastropoda" family="Cypraeidae" genus="Zoila" kingdom="Animalia" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="mulderi">
<emphasis id="9EFCEAE2FFF37E681B1D229CC387ACC5" box="[639,744,1510,1532]" italics="true" pageId="8" pageNumber="9">Z. mulderi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="AC3736F0FFF37E6819EE2169C3C9AF16" blockId="8.[140,678,1554,1576]" box="[140,678,1554,1576]" pageId="8" pageNumber="9">
<emphasis id="9EFCEAE2FFF37E6819EE2169C06FAF19" box="[140,256,1554,1575]" italics="true" pageId="8" pageNumber="9">Etymology.</emphasis>
Greek, didymos, double; rhynchos, snout.
</paragraph>
</subSubSection>
</treatment>
</document>