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<document id="BD0C4312E0074F4EBAA2675364A424EC" ID-CLB-Dataset="8062" ID-DOI="10.5852/ejt.2021.735.1247" ID-GBIF-Dataset="ab1c2bbe-9a7c-4796-9318-103a96c50a9c" ID-ISSN="2118-9773" ID-Zenodo-Dep="4558749" ID-ZooBank="F1CE3697-319D-4D02-A99F-11A0E16A8743" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1614156286742" checkinUser="plazi" docAuthor="Wittmann, Karl J. &amp; Abed-Navandi, Daniel" docDate="2021" docId="03F487FEFF8FA6071725FE93FBDAC161" docLanguage="en" docName="ejt-735_133-175.pdf" docOrigin="European Journal of Taxonomy 735" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Heteromysis (Olivemysis) sixi Wittmann &amp; Abed-Navandi 2021, sp. nov" docType="treatment" docUuid="A5C4DF45-0137-4A0F-AAB5-4C71E081755C" docUuidSource="ZooBank" docVersion="9" lastPageNumber="170" masterDocId="FFCDFF86FFAEA622151CFF9AFFEAC47A" masterDocTitle="Four new species of Heteromysis (Crustacea: Mysida) from public aquaria in Hawaii, Florida, and Western to Central Europe" masterLastPageNumber="175" masterPageNumber="133" pageNumber="166" updateTime="1698931019546" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FD52333E9C049D1AE969F5D911CB2E31">Four new species of Heteromysis (Crustacea: Mysida) from public aquaria in Hawaii, Florida, and Western to Central Europe</mods:title>
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<mods:namePart id="1577F0802757BB88A49C274612A4C634">Wittmann, Karl J.</mods:namePart>
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<mods:affiliation id="CE14B728D0284BF360DA6FD87D70E0F7">Medical University of Vienna, Department of Environmental Health, Kinderspitalgasse 15, 1090 Vienna, Austria.</mods:affiliation>
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<mods:namePart id="CAF24BFCE2273E9F4D7DD67043C08E68">Abed-Navandi, Daniel</mods:namePart>
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<mods:affiliation id="E15577ACBE1D3BA102856473F2F4FD3F">Haus des Meeres - Aqua Terra Zoo, Fritz Grünbaum Platz 1, 1060 Vienna, Austria.</mods:affiliation>
<mods:nameIdentifier id="FA7F93E784933CF2986A3BD638D8505D" type="email">daniel.abed@haus-des-meeres.at</mods:nameIdentifier>
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<mods:date id="EDF84805F68A41AE07A8A738DE961DBC">2021</mods:date>
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<treatment id="03F487FEFF8FA6071725FE93FBDAC161" ID-DOI="http://doi.org/10.5281/zenodo.4559766" ID-GBIF-Taxon="179906474" ID-Zenodo-Dep="4559766" LSID="urn:lsid:zoobank.org:act:A5C4DF45-0137-4A0F-AAB5-4C71E081755C" httpUri="http://treatment.plazi.org/id/03F487FEFF8FA6071725FE93FBDAC161" lastPageId="37" lastPageNumber="170" pageId="33" pageNumber="166">
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<paragraph id="8BE236E8FF8FA6031725FE93FC10C559" blockId="33.[363,1224,264,363]" box="[569,1018,264,291]" pageId="33" pageNumber="166">
<heading id="D0AA8184FF8FA6031725FE93FC10C559" box="[569,1018,264,291]" centered="true" fontSize="11" level="2" pageId="33" pageNumber="166" reason="2">
<taxonomicName id="4C5D4D6BFF8FA6031725FE93FC74C559" authority="Wittmann &amp; Abed-Navandi, 2021" authorityName="Wittmann &amp; Abed-Navandi" authorityYear="2021" box="[569,926,264,291]" class="Malacostraca" family="Mysidae" genus="Heteromysis" kingdom="Animalia" order="Mysida" pageId="33" pageNumber="166" phylum="Arthropoda" rank="species" species="sixi" status="sp. nov." subGenus="Olivemysis">
<emphasis id="B929EAFAFF8FA6031725FE93FC74C559" bold="true" box="[569,926,264,291]" pageId="33" pageNumber="166">
<emphasis id="B929EAFAFF8FA6031725FE93FD25C559" bold="true" box="[569,719,265,291]" italics="true" pageId="33" pageNumber="166">Heteromysis</emphasis>
(
<emphasis id="B929EAFAFF8FA60317FCFE92FC88C559" bold="true" box="[736,866,264,291]" italics="true" pageId="33" pageNumber="166">Olivemysis</emphasis>
)
<emphasis id="B929EAFAFF8FA6031668FE93FC74C559" bold="true" box="[884,926,265,291]" italics="true" pageId="33" pageNumber="166">sixi</emphasis>
</emphasis>
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<taxonomicNameLabel id="A21A5781FF8FA60316B9FE93FC10C559" box="[933,1018,265,291]" pageId="33" pageNumber="166" rank="species">sp. nov</taxonomicNameLabel>
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<subSubSection id="C3476563FF8FA6031477FEB7FCB5C511" pageId="33" pageNumber="166" type="description">
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urn:lsid:zoobank.org:act:
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<paragraph id="8BE236E8FF8FA60317C0FECBFCB5C511" blockId="33.[363,1224,264,363]" box="[732,863,337,363]" pageId="33" pageNumber="166">
<figureCitation id="13662A6DFF8FA60317C0FECBFCB5C511" box="[732,863,337,363]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="34.[189,232,1717,1743]" captionStartId-1="36.[189,232,1928,1954]" captionTargetBox-0="[280,1307,267,1680]" captionTargetBox-1="[190,1385,276,1875]" captionTargetId-0="figure-178@34.[279,1308,266,1681]" captionTargetId-1="figure-76@36.[189,1386,275,1881]" captionTargetPageId-0="34" captionTargetPageId-1="36" captionText-0="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." captionText-1="Fig. 13. Heteromysis sixisp. nov., holotype, ♂with body length 3.0 mm (NHMW26964). A. Thoracopod 8 with penis, caudal. BF. Series of male pleopods 15, caudal. G. Detail of male pleopod 2 (C) showing flagellate spine. H. Uropods, ventral; with setae and ventral wall of statocyst omitted. I. Telson." figureDoi-0="http://doi.org/10.5281/zenodo.4558775" figureDoi-1="http://doi.org/10.5281/zenodo.4558777" httpUri-0="https://zenodo.org/record/4558775/files/figure.png" httpUri-1="https://zenodo.org/record/4558777/files/figure.png" pageId="33" pageNumber="166">Figs 1213</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3476563FF8FA60315A1FE09FCF4C0B4" pageId="33" pageNumber="166" type="diagnosis">
<paragraph id="8BE236E8FF8FA60315A1FE09FEDCC5D7" blockId="33.[189,310,403,429]" box="[189,310,403,429]" pageId="33" pageNumber="166">
<heading id="D0AA8184FF8FA60315A1FE09FEDCC5D7" bold="true" box="[189,310,403,429]" fontSize="11" level="3" pageId="33" pageNumber="166" reason="3">
<emphasis id="B929EAFAFF8FA60315A1FE09FEDCC5D7" bold="true" box="[189,310,403,429]" pageId="33" pageNumber="166">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE236E8FF8FA60315A1FE58FCF4C0B4" blockId="33.[187,1401,450,1230]" pageId="33" pageNumber="166">
Only male
<typeStatus id="54E6884AFF8FA6031427FE58FE48C5A6" box="[315,418,450,476]" pageId="33" pageNumber="166" type="holotype">holotype</typeStatus>
known (ranges of meristic characters are here indicating differences between left and right side of the body). Rostrum (
<figureCitation id="13662A6DFF8FA6031754FE7CFD5FC67A" box="[584,693,486,512]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="33" pageNumber="166">Fig. 12A</figureCitation>
) triangular with narrowly rounded tip, ⅔ length of terminal segment of antennular trunk. Cornea (
<figureCitation id="13662A6DFF8FA6031763FD90FD39C65E" box="[639,723,522,548]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="33" pageNumber="166">Fig. 12</figureCitation>
AB) occupies distal third of eye surface. Eyestalks with smooth surface except for prominent tooth on disto-mesial edge. Antennular trunk (
<figureCitation id="13662A6DFF8FA60311BAFDB4FB11C632" box="[1190,1275,558,584]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="33" pageNumber="166">Fig. 12</figureCitation>
CE) with blade-like, subapically flagellate spine in addition to setae on disto-mesial edge. Distal 1624% of this spine with rugged, obliquely truncate margin. Antennal scale (
<figureCitation id="13662A6DFF8FA60316B9FDEFFBFBC6F5" box="[933,1041,629,655]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="33" pageNumber="166">Fig. 12A</figureCitation>
) undivided, extending shortly beyond antennal peduncle; scale length 2.43.0 times maximum width. Thoracic sternites (
<figureCitation id="13662A6DFF8FA603101FFD03FA9AC6C9" box="[1283,1392,665,691]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="33" pageNumber="166">Fig. 12H</figureCitation>
) with short median processes. Flagellum of thoracic exopod 1 with 8 segments, exopods 28 with nine segments (
<figureCitation id="13662A6DFF8FA6031420FD7BFE40C681" box="[316,426,737,763]" captionStart="Fig" captionStartId="36.[189,232,1928,1954]" captionTargetBox="[190,1385,276,1875]" captionTargetId="figure-76@36.[189,1386,275,1881]" captionTargetPageId="36" captionText="Fig. 13. Heteromysis sixisp. nov., holotype, ♂with body length 3.0 mm (NHMW26964). A. Thoracopod 8 with penis, caudal. BF. Series of male pleopods 15, caudal. G. Detail of male pleopod 2 (C) showing flagellate spine. H. Uropods, ventral; with setae and ventral wall of statocyst omitted. I. Telson." figureDoi="http://doi.org/10.5281/zenodo.4558777" httpUri="https://zenodo.org/record/4558777/files/figure.png" pageId="33" pageNumber="166">Fig. 13A</figureCitation>
). Carpopropodus of thoracic endopods 18 with 2, 2, 2, 4, 6, 6, 6, and 6 segments, respectively. Thoracic endopod 3 (
<figureCitation id="13662A6DFF8FA603177CFC9FFD5CC765" box="[608,694,773,799]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="33" pageNumber="166">Fig. 12</figureCitation>
IM) without tooth-like extension on disto-mesial edge of merus; carpopropodus length 2.5 times maximum width. Distal 4464% of male carpus 3 with total of six flagellate spines along inner margin; propodus without paradactylary setae. Penes (
<figureCitation id="13662A6DFF8FA603101CFCD7FA84C71D" box="[1280,1390,845,871]" captionStart="Fig" captionStartId="36.[189,232,1928,1954]" captionTargetBox="[190,1385,276,1875]" captionTargetId="figure-76@36.[189,1386,275,1881]" captionTargetPageId="36" captionText="Fig. 13. Heteromysis sixisp. nov., holotype, ♂with body length 3.0 mm (NHMW26964). A. Thoracopod 8 with penis, caudal. BF. Series of male pleopods 15, caudal. G. Detail of male pleopod 2 (C) showing flagellate spine. H. Uropods, ventral; with setae and ventral wall of statocyst omitted. I. Telson." figureDoi="http://doi.org/10.5281/zenodo.4558777" httpUri="https://zenodo.org/record/4558777/files/figure.png" pageId="33" pageNumber="166">Fig. 13A</figureCitation>
) cylindrical, apically blunt, ending in five blunt lobes; size ¾ length of ischium 8; small setae present at half length of penis. Pleopods (
<figureCitation id="13662A6DFF8FA6031730FC0EFD94C7D4" box="[556,638,916,942]" captionStart="Fig" captionStartId="36.[189,232,1928,1954]" captionTargetBox="[190,1385,276,1875]" captionTargetId="figure-76@36.[189,1386,275,1881]" captionTargetPageId="36" captionText="Fig. 13. Heteromysis sixisp. nov., holotype, ♂with body length 3.0 mm (NHMW26964). A. Thoracopod 8 with penis, caudal. BF. Series of male pleopods 15, caudal. G. Detail of male pleopod 2 (C) showing flagellate spine. H. Uropods, ventral; with setae and ventral wall of statocyst omitted. I. Telson." figureDoi="http://doi.org/10.5281/zenodo.4558777" httpUri="https://zenodo.org/record/4558777/files/figure.png" pageId="33" pageNumber="166">Fig. 13</figureCitation>
BG) rudimentary, unsegmented, with residual differentiation of endopod (pseudobranchial lobe). Male pleopods 1, 35 normal, without spines. Pleopod 2 with large smooth spine (stylet) on apex and three minute flagellate spines in series along distal half of outer margin of elongate distal portion. Exopod of uropods (
<figureCitation id="13662A6DFF8FA6031643FB9AFC27C060" box="[863,973,1024,1050]" captionStart="Fig" captionStartId="36.[189,232,1928,1954]" captionTargetBox="[190,1385,276,1875]" captionTargetId="figure-76@36.[189,1386,275,1881]" captionTargetPageId="36" captionText="Fig. 13. Heteromysis sixisp. nov., holotype, ♂with body length 3.0 mm (NHMW26964). A. Thoracopod 8 with penis, caudal. BF. Series of male pleopods 15, caudal. G. Detail of male pleopod 2 (C) showing flagellate spine. H. Uropods, ventral; with setae and ventral wall of statocyst omitted. I. Telson." figureDoi="http://doi.org/10.5281/zenodo.4558777" httpUri="https://zenodo.org/record/4558777/files/figure.png" pageId="33" pageNumber="166">Fig. 13H</figureCitation>
) 1.21.3 times length of endopod; endopod with only one small spine near statocyst; distal spine-free portion ⅔ length of endopod. Each lateral margin of telson (
<figureCitation id="13662A6DFF8FA60314F5FBD2FDA0C018" box="[489,586,1096,1122]" captionStart="Fig" captionStartId="36.[189,232,1928,1954]" captionTargetBox="[190,1385,276,1875]" captionTargetId="figure-76@36.[189,1386,275,1881]" captionTargetPageId="36" captionText="Fig. 13. Heteromysis sixisp. nov., holotype, ♂with body length 3.0 mm (NHMW26964). A. Thoracopod 8 with penis, caudal. BF. Series of male pleopods 15, caudal. G. Detail of male pleopod 2 (C) showing flagellate spine. H. Uropods, ventral; with setae and ventral wall of statocyst omitted. I. Telson." figureDoi="http://doi.org/10.5281/zenodo.4558777" httpUri="https://zenodo.org/record/4558777/files/figure.png" pageId="33" pageNumber="166">Fig. 13I</figureCitation>
) with six spines on distal half, proximal portion smooth. U-shaped apical cleft penetrates ¼ telson length; proximal 5/6 of cleft with 26 laminae. Disto-lateral lobes each with two spines on narrowly truncate apex. The outer apical spines are 1618% telson length; inner apical spines are 0.20.3 times length of outer ones.
</paragraph>
</subSubSection>
<subSubSection id="C3476563FF8FA60315A1FB6CFE46C1FC" pageId="33" pageNumber="166" type="etymology">
<paragraph id="8BE236E8FF8FA60315A1FB6CFEAEC16A" blockId="33.[189,324,1270,1296]" box="[189,324,1270,1296]" pageId="33" pageNumber="166">
<heading id="D0AA8184FF8FA60315A1FB6CFEAEC16A" bold="true" box="[189,324,1270,1296]" fontSize="11" level="3" pageId="33" pageNumber="166" reason="3">
<emphasis id="B929EAFAFF8FA60315A1FB6CFEAEC16A" bold="true" box="[189,324,1270,1296]" pageId="33" pageNumber="166">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BE236E8FF8FA60315A1FABFFE46C1FC" blockId="33.[189,1399,1317,1414]" pageId="33" pageNumber="166">
The species name is a masculine noun in genitive singular, dedicated on the occasion of his retirement to Franz Six, citizen scientist and 55 year-long promoter of the Haus des Meeres, a public marine aquarium in
<collectingRegion id="4999F80AFF8FA603144DFAF6FE42C1FC" box="[337,424,1388,1414]" country="Austria" name="Wien" pageId="33" pageNumber="166">Vienna</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3476563FF8FA60315A1FA34FDEEC23A" pageId="33" pageNumber="166" type="materials_examined">
<paragraph id="8BE236E8FF8FA60315A1FA34FE40C1B2" blockId="33.[189,426,1454,1480]" box="[189,426,1454,1480]" pageId="33" pageNumber="166">
<heading id="D0AA8184FF8FA60315A1FA34FE40C1B2" bold="true" box="[189,426,1454,1480]" fontSize="11" level="3" pageId="33" pageNumber="166" reason="3">
<emphasis id="B929EAFAFF8FA60315A1FA34FE40C1B2" bold="true" box="[189,426,1454,1480]" pageId="33" pageNumber="166">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BE236E8FF8FA60315F6FA44FDEEC23A" blockId="33.[189,1398,1502,1600]" pageId="33" pageNumber="166">
<materialsCitation id="3B353CB5FF8FA60315F6FA44FDEEC23A" ID-GBIF-Occurrence="3042073320" collectingDate="2020-01" collectionCode="NHMW" collectorName="Gwen Lentes" location="Waikiki Beach" municipality="Honolulu" pageId="33" pageNumber="166" specimenCode="NHMW 26964" specimenCount="1" stateProvince="HAWAII" typeStatus="holotype">
<emphasis id="B929EAFAFF8FA60315F6FA44FEB6C182" bold="true" box="[234,348,1502,1528]" pageId="33" pageNumber="166">
<typeStatus id="54E6884AFF8FA60315F6FA44FEB6C182" box="[234,348,1502,1528]" pageId="33" pageNumber="166" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingRegion id="4999F80AFF8FA60315A1F998FECCC266" box="[189,294,1538,1564]" pageId="33" pageNumber="166">HAWAII</collectingRegion>
<specimenCount id="9D5BFD61FF8FA6031420F998FEB8C264" box="[316,338,1538,1566]" pageId="33" pageNumber="166" type="generic"></specimenCount>
ad., BL 3.0 mm (on slides);
<collectingMunicipality id="6B86AC92FF8FA6031785F99BFCE1C261" box="[665,779,1537,1563]" pageId="33" pageNumber="166">Honolulu</collectingMunicipality>
,
<location id="8E826033FF8FA6031609F99BFC29C261" LSID="urn:lsid:plazi:treatment:03F487FEFF8FA6071725FE93FBDAC161:8E826033FF8FA6031609F99BFC29C261" box="[789,963,1537,1563]" municipality="Honolulu" name="Waikiki Beach" pageId="33" pageNumber="166" stateProvince="HAWAII">Waikiki Beach</location>
,
<location id="8E826033FF8FA60316D1F99BFB43C261" LSID="urn:lsid:plazi:treatment:03F487FEFF8FA6071725FE93FBDAC161:8E826033FF8FA60316D1F99BFB43C261" box="[973,1193,1537,1563]" municipality="Honolulu" name="Waikiki Aquarium" pageId="33" pageNumber="166" stateProvince="HAWAII">Waikiki Aquarium</location>
;
<date id="FFE31028FF8FA60311AFF998FACCC266" box="[1203,1318,1538,1564]" pageId="33" pageNumber="166" value="2020-01">
<collectingDate id="EFA7E9C0FF8FA60311AFF998FACCC266" box="[1203,1318,1538,1564]" pageId="33" pageNumber="166" value="2020-01">Jan. 2020</collectingDate>
</date>
;
<collectorName id="26A8533EFF8FA603102CF998FEE1C23A" pageId="33" pageNumber="166">Gwen Lentes</collectorName>
leg.;
<specimenCode id="DBFB9E93FF8FA6031451F9BCFE17C245" box="[333,509,1573,1600]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="33" pageNumber="166" type="Museum">NHMW 26964</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3476563FF8FA60715A1F9FDFB6EC0F9" lastPageId="37" lastPageNumber="170" pageId="33" pageNumber="166" type="description">
<paragraph id="8BE236E8FF8FA60315A1F9FDFEA4C2FB" blockId="33.[189,334,1639,1665]" box="[189,334,1639,1665]" pageId="33" pageNumber="166">
<heading id="D0AA8184FF8FA60315A1F9FDFEA4C2FB" bold="true" box="[189,334,1639,1665]" fontSize="11" level="3" pageId="33" pageNumber="166" reason="3">
<emphasis id="B929EAFAFF8FA60315A1F9FDFEA4C2FB" bold="true" box="[189,334,1639,1665]" pageId="33" pageNumber="166">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BE236E8FF8FA60315A1F90DFD8AC282" blockId="33.[188,1399,1687,1784]" pageId="33" pageNumber="166">All features of the diagnosis. Cephalothorax comprises 30% of body length, pleon (without telson) 55%, and carapace (without rostrum) 32%. Abdominal somites 15 measure 1.0, 0.9, 0.8, 0.9, and 0.9 times the length of somite 6, respectively.</paragraph>
<paragraph id="8BE236E8FF8FA60315A1F8BBFBBFC3F8" blockId="33.[189,1399,1824,1922]" pageId="33" pageNumber="166">
CARAPACE (
<figureCitation id="13662A6DFF8FA603145FF8BBFE58C341" box="[323,434,1825,1851]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="33" pageNumber="166">Fig. 12A</figureCitation>
). Carapace covers 75% of cephalothorax dorsally. Rostrum reaches to proximal third of artificially straight forward-oriented eyestalks (without cornea). Cervical sulcus short but strong. Posterior margin leaving 12 posterior thoracic somites mid-dorsally exposed.
</paragraph>
<paragraph id="8BE236E8FF8FA60315A1F831FA9CC392" blockId="33.[189,1399,1962,2025]" pageId="33" pageNumber="166">
EYES (
<figureCitation id="13662A6DFF8FA6031417F830FE88C3BF" box="[267,354,1962,1989]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="33" pageNumber="166">Fig. 12</figureCitation>
AB). Eyestalks and cornea dorsoventrally compressed (
<figureCitation id="13662A6DFF8FA6031104F830FB6DC3BF" box="[1048,1159,1962,1989]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="33" pageNumber="166">Fig. 12B</figureCitation>
). Eyestalks without scales. In dorsal view (
<figureCitation id="13662A6DFF8FA60314C9F854FDA8C392" box="[469,578,1998,2024]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="33" pageNumber="166">Fig. 12A</figureCitation>
) the cornea appears calotte-shaped, measuring 4/5 length of eyestalk
</paragraph>
<caption id="DF226660FF8CA60015A1F92FFB62C393" ID-DOI="http://doi.org/10.5281/zenodo.4558775" ID-Zenodo-Dep="4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="34" pageNumber="167" startId="34.[189,232,1717,1743]" targetBox="[280,1307,267,1680]" targetPageId="34">
<paragraph id="8BE236E8FF8CA60015A1F92FFB62C393" blockId="34.[188,1400,1717,2026]" pageId="34" pageNumber="167">
<emphasis id="B929EAFAFF8CA60015A1F92FFEF0C2B5" bold="true" box="[189,282,1717,1743]" pageId="34" pageNumber="167">Fig. 12.</emphasis>
<taxonomicName id="4C5D4D6BFF8CA600143EF92CFE0EC2B5" authority="Wittmann &amp; Abed-Navandi, 2021" authorityName="Wittmann &amp; Abed-Navandi" authorityYear="2021" box="[290,484,1718,1743]" class="Malacostraca" family="Mysidae" genus="Heteromysis" kingdom="Animalia" order="Mysida" pageId="34" pageNumber="167" phylum="Arthropoda" rank="species" species="sixi" status="sp. nov.">
<emphasis id="B929EAFAFF8CA600143EF92CFE0EC2B5" box="[290,484,1718,1743]" italics="true" pageId="34" pageNumber="167">Heteromysis sixi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A5781FF8CA60014F0F92CFDA0C2AA" box="[492,586,1718,1744]" pageId="34" pageNumber="167" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ with body length 3.0 mm (NHMW 26964).
<emphasis id="B929EAFAFF8CA60011ECF92FFAEFC2B5" bold="true" box="[1264,1285,1717,1743]" pageId="34" pageNumber="167">A</emphasis>
. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted.
<emphasis id="B929EAFAFF8CA60011F1F943FAEBC289" bold="true" box="[1261,1281,1753,1779]" pageId="34" pageNumber="167">B</emphasis>
. Detail of panel (A) showing right eye in lateral view.
<emphasis id="B929EAFAFF8CA60017D7F966FD35C36C" bold="true" box="[715,735,1788,1814]" pageId="34" pageNumber="167">C</emphasis>
. Right antennula, dorsal.
<emphasis id="B929EAFAFF8CA600110FF966FBA3C36C" bold="true" box="[1043,1097,1788,1814]" pageId="34" pageNumber="167">DE</emphasis>
. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages.
<emphasis id="B929EAFAFF8CA60015A1F8D8FF25C326" bold="true" box="[189,207,1858,1884]" pageId="34" pageNumber="167">F</emphasis>
. Palp of left mandible, caudal.
<emphasis id="B929EAFAFF8CA600174BF8D8FD84C326" bold="true" box="[599,622,1858,1884]" pageId="34" pageNumber="167">G</emphasis>
. Detail of panel (F) showing barbed seta from outer margin of median segment.
<emphasis id="B929EAFAFF8CA6001497F8FCFE48C3FA" bold="true" box="[395,418,1894,1920]" pageId="34" pageNumber="167">H</emphasis>
. Thoracic sternites 18 with median processes.
<emphasis id="B929EAFAFF8CA60016CFF8FCFC34C3FA" bold="true" box="[979,990,1894,1920]" pageId="34" pageNumber="167">I</emphasis>
. Right thoracic endopod 3, rostral.
<emphasis id="B929EAFAFF8CA60015A1F813FF1CC3D9" bold="true" box="[189,246,1929,1955]" pageId="34" pageNumber="167">JM</emphasis>
. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium.
<emphasis id="B929EAFAFF8CA600170CF836FDAEC3BC" bold="true" box="[528,580,1964,1990]" pageId="34" pageNumber="167">NS</emphasis>
. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted.
<emphasis id="B929EAFAFF8CA60015A0F855FF19C393" bold="true" box="[188,243,1999,2025]" pageId="34" pageNumber="167">TU</emphasis>
. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut.
</paragraph>
</caption>
<paragraph id="8BE236E8FF8DA60115A1FE93FD27C53D" blockId="35.[188,1398,265,328]" pageId="35" pageNumber="168">
(cornea not included). In lateral view (
<figureCitation id="13662A6DFF8DA6011764FE93FD35C559" box="[632,735,265,291]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12B</figureCitation>
) the cornea appears roughly trapezoid with well-rounded edges, length 1.41.5 times maximum width.
</paragraph>
<paragraph id="8BE236E8FF8DA60115A1FEF5FE0BC6B4" blockId="35.[188,1399,367,718]" pageId="35" pageNumber="168">
ANTENNULAE (
<figureCitation id="13662A6DFF8DA601147EFEF5FE52C5F3" box="[354,440,367,394]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12</figureCitation>
CE). Trunk extends 5662% its length beyond eyes, 1214% beyond antennal scale. Measured without apophysis along dorsal midline, the basal segment is 4748% trunk length, median segment 1018%, and terminal segment 3443%. Basal segment on basal half of its outer face with two small, stout, barbed setae as in
<figureCitation id="13662A6DFF8DA601178EFE41FD04C58F" box="[658,750,475,501]" captionStart="Fig" captionStartId="14.[189,232,1644,1670]" captionTargetBox="[305,1283,266,1610]" captionTargetId="figure-209@14.[304,1283,265,1611]" captionTargetPageId="14" captionText="Fig. 2. Heteromysis smithsoniana sp. nov., paratypes, ♂♂ with body length 4.8 mm (AD, IK, RS: NHMW 26971), 4.2 mm (EH, LN: NHMW 26972), and ♀ 4.2 mm (OQ: NHMW 26970). A. Male antennula, dorsal. BD. Details of antennula (A) showing modified spines (BC) and setae (D). E. Antenna with antennal gland, dorsal. F. Carapace expanded on slide. GH. Details of carapace (F) showing pore groups in front of cervical sulcus (G) and behind cardial sulcus (H). I. Palp of left mandible, rostral. JK. Details of mandibular palp (I) showing three scale groups (J) and one modified seta (K) from inner margin of median segment; most setae of terminal segment are below drawing plane. L. Right male thoracic sympod and endopod 3, rostral. MN. Details of male endopod 3 (L) showing modified spine (M) of carpus and modified seta (N) of merus. O. Left female thoracic endopod 3, rostral. PQ. Details of female endopod 3 (O) showing modified spine (P) of carpus and modified seta (Q) of merus. RS. Dactyli 1 and 3, with claw, setae omitted." figureDoi="http://doi.org/10.5281/zenodo.4558753" httpUri="https://zenodo.org/record/4558753/files/figure.png" pageId="35" pageNumber="168">Fig. 2D</figureCitation>
. Dorsal apophysis with three barbed setae and one whip seta (
<figureCitation id="13662A6DFF8DA60115E5FE65FE89C660" box="[249,355,511,538]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12E</figureCitation>
). Lateral apophysis with two barbed setae and one smooth seta. Median segment dorsally with large apophysis bearing three barbed setae and one whip seta. Terminal segment 1.31.4 times as long as wide. Its mid-dorsal apophysis with four small, barbed setae. Outer antennular flagellum is thicker than inner one by factor of 1.51.7 when measured near basis of flagella. Male lobe setose, inserts ventrally close to terminal margin of antennular trunk, length is 2231% width of terminal segment of trunk, its width 2225%.
</paragraph>
<paragraph id="8BE236E8FF8DA60115A1FD6CFDD8C722" blockId="35.[189,1398,758,856]" pageId="35" pageNumber="168">
ANTENNAE (
<figureCitation id="13662A6DFF8DA6011458FD6CFE5AC76A" box="[324,432,758,784]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12A</figureCitation>
). Three-segmented antennal peduncle with basal segment 20% length of peduncle, second segment 4748%, and third segment 33%. Sympod with tongue-like, terminally broad process on ventral face (as in
<figureCitation id="13662A6DFF8DA60114A0FCA4FDCFC722" box="[444,549,830,856]" captionStart="Fig" captionStartId="30.[189,232,1750,1776]" captionTargetBox="[266,1317,269,1716]" captionTargetId="figure-161@30.[265,1322,265,1718]" captionTargetPageId="30" captionText="Fig. 10. Heteromysis waikikensis sp. nov., holotype, ♂ with body length 3.9 mm (N: NHMW 26965), paratypes, ♂ 4.2 mm (AE, LM, OP: NHMW 26967) and ♀ 4.2 mm (FK: NHMW 26966). A. Male antennula, dorsal.BD. Details of male antennula (A) showing modified spines. E.Antenna with antennal gland, dorsal. F. Palp of left mandible, caudal. GH. Setae from outer margin of median segment of mandibular palp in ♀ (G) versus ♂ (H). I. Left female thoracic endopod 3, rostral. JK. Details of female endopod (I) 3 showing modified spines of carpus. L. Left male thoracic sympod and endopod 3, rostral. M. Detail of male endopod 3 (L) showing modified seta of merus. N. Scutellum paracaudale, lateral. OP. Modified spines from dorsolateral infoldings (O) and lateralia (P) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558771" httpUri="https://zenodo.org/record/4558771/files/figure.png" pageId="35" pageNumber="168">Fig. 10E</figureCitation>
).
</paragraph>
<paragraph id="8BE236E8FF8DA60115A1FC1BFC72C179" blockId="35.[188,1400,896,1283]" pageId="35" pageNumber="168">
MOUTHPARTS (
<figureCitation id="13662A6DFF8DA6011443FC1AFE5BC7E0" box="[351,433,896,923]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12</figureCitation>
FG). Mandibular palp (
<figureCitation id="13662A6DFF8DA60117C8FC1AFCCCC7E0" box="[724,806,896,922]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12</figureCitation>
FG) three-segmented. Proximal segment without setae, 1015% length of palp. Length of median segment 1.62.3 times its maximum width and 6168% length of palp. Inner margin of median segment with three sparsely barbed setae in subapical to subbasal position, and with 46 short setae on proximal third. Basal half of these setae thickened; with unilateral series of stiff barbs; distal half smooth (
<figureCitation id="13662A6DFF8DA6011794FB8AFD1EC050" box="[648,756,1040,1066]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12G</figureCitation>
). Outer margin of median segment all along with setae; most setae barbed on basal third. No scales present on median segment. Terminal segment strongly setose (most setae below drawing plane in
<figureCitation id="13662A6DFF8DA60117DEFBC2FCC1C009" box="[706,811,1112,1139]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12F</figureCitation>
), this segment 2728% palp length. Pars molaris with well-developed grinding surface in both mandibles. Pars incisiva with four teeth, digitus mobilis with three teeth, and pars centralis with 45 spiny teeth. Labium as described above for
<taxonomicName id="4C5D4D6BFF8DA60111AAFB3BFA9FC0C0" authority="Wittmann &amp; Abed-Navandi, 2021" authorityName="Wittmann &amp; Abed-Navandi" authorityYear="2021" box="[1206,1397,1184,1210]" class="Malacostraca" family="Mysidae" genus="Heteromysis" kingdom="Animalia" order="Mysida" pageId="35" pageNumber="168" phylum="Arthropoda" rank="species" species="smithsoniana" status="sp. nov.">
<emphasis id="B929EAFAFF8DA60111AAFB3BFA9FC0C0" box="[1206,1397,1184,1210]" italics="true" pageId="35" pageNumber="168">H. smithsoniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A5781FF8DA60115A1FB5FFEFDC0A5" box="[189,279,1221,1247]" pageId="35" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
Maxillula and maxilla as normal in this genus. Outer margin of exopod of maxilla all along with 16 plumose setae, two apical setae larger than remaining ones.
</paragraph>
<paragraph id="8BE236E8FF8DA60115A1FAB1FCEBC113" blockId="35.[189,1399,1323,1385]" pageId="35" pageNumber="168">
THORACIC STERNITES (
<figureCitation id="13662A6DFF8DA60114AFFAB1FDF4C13F" box="[435,542,1323,1349]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12H</figureCitation>
) with short median processes, in sternites 13, 8 terminally rounded, in sternites 47 ending in small triangular denticles.
</paragraph>
<paragraph id="8BE236E8FF8DA60115A1FA08FD8FC324" blockId="35.[188,1399,1425,1886]" pageId="35" pageNumber="168">
THORACOPODS (general;
<figureCitation id="13662A6DFF8DA60114CBFA0BFDDAC1D1" box="[471,560,1425,1451]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Figs 12</figureCitation>
IS, 13A). Length of flagella as well as of basal plates increase from exopod 1 to 56, and decrease from exopod 6‾8. Basal plates (
<figureCitation id="13662A6DFF8DA601168FFA2FFC17C1B5" box="[915,1021,1461,1487]" captionStart="Fig" captionStartId="36.[189,232,1928,1954]" captionTargetBox="[190,1385,276,1875]" captionTargetId="figure-76@36.[189,1386,275,1881]" captionTargetPageId="36" captionText="Fig. 13. Heteromysis sixisp. nov., holotype, ♂with body length 3.0 mm (NHMW26964). A. Thoracopod 8 with penis, caudal. BF. Series of male pleopods 15, caudal. G. Detail of male pleopod 2 (C) showing flagellate spine. H. Uropods, ventral; with setae and ventral wall of statocyst omitted. I. Telson." figureDoi="http://doi.org/10.5281/zenodo.4558777" httpUri="https://zenodo.org/record/4558777/files/figure.png" pageId="35" pageNumber="168">Fig. 13A</figureCitation>
) expanded, length 1.72.1 times width. Outer edge of plates angular, tip narrowly rounded in exopods 12, and pointed in remaining exopods. The first thoracopods with large, leaf-like, smooth epipod. Plumose seta on intersegmental joint between sympod of thoracopod 2 and the corresponding sternite. Basis of endopods 48 with a small, lappet-like apophysis on rostral face below endopod (
<figureCitation id="13662A6DFF8DA60116DCF9DFFBD8C225" box="[960,1074,1605,1631]" captionStart="Fig" captionStartId="36.[189,232,1928,1954]" captionTargetBox="[190,1385,276,1875]" captionTargetId="figure-76@36.[189,1386,275,1881]" captionTargetPageId="36" captionText="Fig. 13. Heteromysis sixisp. nov., holotype, ♂with body length 3.0 mm (NHMW26964). A. Thoracopod 8 with penis, caudal. BF. Series of male pleopods 15, caudal. G. Detail of male pleopod 2 (C) showing flagellate spine. H. Uropods, ventral; with setae and ventral wall of statocyst omitted. I. Telson." figureDoi="http://doi.org/10.5281/zenodo.4558777" httpUri="https://zenodo.org/record/4558777/files/figure.png" pageId="35" pageNumber="168">Fig. 13A</figureCitation>
). Total length increases in endopods 1‾6 and decreases in 6‾8. Ischium becomes longer and more slender from endopods 1 to 4, while both features remain subequal among endopods 48. Ischium shorter than merus in endopods 13 (
<figureCitation id="13662A6DFF8DA60115D9F92EFECFC2B4" box="[197,293,1716,1742]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12I</figureCitation>
) but longer than merus in endopods 48 (
<figureCitation id="13662A6DFF8DA6011605F92EFC6DC2B4" box="[793,903,1716,1742]" captionStart="Fig" captionStartId="36.[189,232,1928,1954]" captionTargetBox="[190,1385,276,1875]" captionTargetId="figure-76@36.[189,1386,275,1881]" captionTargetPageId="36" captionText="Fig. 13. Heteromysis sixisp. nov., holotype, ♂with body length 3.0 mm (NHMW26964). A. Thoracopod 8 with penis, caudal. BF. Series of male pleopods 15, caudal. G. Detail of male pleopod 2 (C) showing flagellate spine. H. Uropods, ventral; with setae and ventral wall of statocyst omitted. I. Telson." figureDoi="http://doi.org/10.5281/zenodo.4558777" httpUri="https://zenodo.org/record/4558777/files/figure.png" pageId="35" pageNumber="168">Fig. 13A</figureCitation>
). Thoracic endopods with claw 3 longest; claws 1, 48 about half length of claw 3; claw 2 miniaturized. Claws 14 smooth (
<figureCitation id="13662A6DFF8DA6011192F942FB0AC288" box="[1166,1248,1752,1778]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12</figureCitation>
NQ), claws 58 subapically unilaterally serrated (
<figureCitation id="13662A6DFF8DA601176AF966FD20C36C" box="[630,714,1788,1814]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12</figureCitation>
RS). Claws 1, 4 straight (
<figureCitation id="13662A6DFF8DA601111CF966FB65C36C" box="[1024,1167,1788,1814]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12N, Q</figureCitation>
), claws 23 weakly bent (
<figureCitation id="13662A6DFF8DA601141CF8BAFEBEC340" box="[256,340,1824,1850]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12</figureCitation>
OP), claws 58 strongly bent (
<figureCitation id="13662A6DFF8DA60117D0F8BAFCCBC340" box="[716,801,1824,1850]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="35" pageNumber="168">Fig. 12</figureCitation>
RS). Penes with one barbed plus one smooth seta in about central position (
<figureCitation id="13662A6DFF8DA60114F6F8DEFDBDC324" box="[490,599,1860,1886]" captionStart="Fig" captionStartId="36.[189,232,1928,1954]" captionTargetBox="[190,1385,276,1875]" captionTargetId="figure-76@36.[189,1386,275,1881]" captionTargetPageId="36" captionText="Fig. 13. Heteromysis sixisp. nov., holotype, ♂with body length 3.0 mm (NHMW26964). A. Thoracopod 8 with penis, caudal. BF. Series of male pleopods 15, caudal. G. Detail of male pleopod 2 (C) showing flagellate spine. H. Uropods, ventral; with setae and ventral wall of statocyst omitted. I. Telson." figureDoi="http://doi.org/10.5281/zenodo.4558777" httpUri="https://zenodo.org/record/4558777/files/figure.png" pageId="35" pageNumber="168">Fig. 13A</figureCitation>
).
</paragraph>
<paragraph id="8BE236E8FF8DA60115A1F81DFA92C392" blockId="35.[187,1400,1926,2025]" pageId="35" pageNumber="168">
MAXILLIPEDS. First maxilliped as described above for
<taxonomicName id="4C5D4D6BFF8DA6011600F81DFC30C3DA" authority="Wittmann &amp; Abed-Navandi, 2021" authorityName="Wittmann &amp; Abed-Navandi" authorityYear="2021" box="[796,986,1926,1952]" class="Malacostraca" family="Mysidae" genus="Heteromysis" kingdom="Animalia" order="Mysida" pageId="35" pageNumber="168" phylum="Arthropoda" rank="species" species="smithsoniana" status="sp. nov.">
<emphasis id="B929EAFAFF8DA6011600F81DFC30C3DA" box="[796,986,1926,1952]" italics="true" pageId="35" pageNumber="168">H. smithsoniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A5781FF8DA60116FEF81DFBD1C3DB" box="[994,1083,1927,1953]" pageId="35" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
Basis of second maxilliped with large, distinctly medially projecting endite. Combined praeischium plus ischium 0.9 times length of merus, carpopropodus plus dactylus 1.1 times merus. Dactylus very large, with dense brush formed
</paragraph>
<caption id="DF226660FF8AA60615A1F812FAF4C392" ID-DOI="http://doi.org/10.5281/zenodo.4558777" ID-Zenodo-Dep="4558777" httpUri="https://zenodo.org/record/4558777/files/figure.png" pageId="36" pageNumber="169" startId="36.[189,232,1928,1954]" targetBox="[190,1385,276,1875]" targetPageId="36">
<paragraph id="8BE236E8FF8AA60615A1F812FAF4C392" blockId="36.[185,1399,1928,2025]" pageId="36" pageNumber="169">
<emphasis id="B929EAFAFF8AA60615A1F812FEFCC3D8" bold="true" box="[189,278,1928,1954]" pageId="36" pageNumber="169">Fig. 13.</emphasis>
<taxonomicName id="4C5D4D6BFF8AA6061406F813FE33C3D8" authority="Wittmann &amp; Abed-Navandi, 2021" authorityName="Wittmann &amp; Abed-Navandi" authorityYear="2021" box="[282,473,1929,1954]" class="Malacostraca" family="Mysidae" genus="Heteromysis" kingdom="Animalia" order="Mysida" pageId="36" pageNumber="169" phylum="Arthropoda" rank="species" species="sixi" status="sp. nov.">
<emphasis id="B929EAFAFF8AA6061406F813FE33C3D8" box="[282,473,1929,1954]" italics="true" pageId="36" pageNumber="169">Heteromysis sixi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A5781FF8AA60614C0F812FDDCC3D8" box="[476,566,1928,1954]" pageId="36" pageNumber="169" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ with body length 3.0 mm (NHMW26964).
<emphasis id="B929EAFAFF8AA60611AAF812FB21C3D8" bold="true" box="[1206,1227,1928,1954]" pageId="36" pageNumber="169">A</emphasis>
. Thoracopod 8 with penis, caudal.
<emphasis id="B929EAFAFF8AA6061483F831FE39C3BF" bold="true" box="[415,467,1963,1989]" pageId="36" pageNumber="169">BF</emphasis>
. Series of male pleopods 15, caudal.
<emphasis id="B929EAFAFF8AA606168AF831FC47C3BF" bold="true" box="[918,941,1963,1989]" pageId="36" pageNumber="169">G</emphasis>
. Detail of male pleopod 2 (C) showing flagellate spine.
<emphasis id="B929EAFAFF8AA6061460F854FE79C392" bold="true" box="[380,403,1998,2024]" pageId="36" pageNumber="169">H</emphasis>
. Uropods, ventral; with setae and ventral wall of statocyst omitted.
<emphasis id="B929EAFAFF8AA60611B2F854FB53C392" bold="true" box="[1198,1209,1998,2024]" pageId="36" pageNumber="169">I</emphasis>
. Telson.
</paragraph>
</caption>
<paragraph id="8BE236E8FF8BA60715A1FE93FB2CC532" blockId="37.[189,1398,265,329]" pageId="37" pageNumber="170">by large numbers of normal setae and about ten modified setae, the latter apically bent, bearing two symmetrical series of denticles (stiff barbs) on each side in subbasal to median portions.</paragraph>
<paragraph id="8BE236E8FF8BA60715A1FEE8FBC0C6CE" blockId="37.[188,1399,369,693]" pageId="37" pageNumber="170">
GNATHOPODS (thoracic endopods 3;
<figureCitation id="13662A6DFF8BA6071746FEEBFD44C5F6" box="[602,686,369,396]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="37" pageNumber="170">Fig. 12</figureCitation>
IM). Ischium 1.7‾2.0 times as long as wide; merus 2.8‾3.3 as long as wide and 1.7‾2.3 length of ischium. Carpus 0.60.7 times length of merus, 0.91.1 times ischium. Claw 2.63.6 times length of dactylus, and 4452% carpopropodus. Distal half of ischium with 34 small whip setae (
<figureCitation id="13662A6DFF8BA60714E0FE7BFD81C581" box="[508,619,481,507]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="37" pageNumber="170">Fig. 12M</figureCitation>
) on inner margin; these setae inserted on short, terminally rounded projections. Six to eight such setae intermixed in series of larger smooth setae along 6080% of inner margin of merus, these setae not inserted on projections. Series of five unilaterally barbed setae (
<figureCitation id="13662A6DFF8BA6071016FDB1FA84C63F" box="[1290,1390,555,581]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="37" pageNumber="170">Fig.12L</figureCitation>
) near outer margin on distal ¾ of merus. These setae with series of normal barbs (cilia) along median to subapical portions, basal portions thickened. Carpus with proximal two spines unpaired, distal spines arranged in two pairs; only distal 24 spines with rugged anterior margins.
</paragraph>
<paragraph id="8BE236E8FF8BA60715A1FD44FB13C767" blockId="37.[189,1398,733,798]" pageId="37" pageNumber="170">
PLEOPODS (
<figureCitation id="13662A6DFF8BA6071427FD44FE66C682" box="[315,396,734,760]" captionStart="Fig" captionStartId="36.[189,232,1928,1954]" captionTargetBox="[190,1385,276,1875]" captionTargetId="figure-76@36.[189,1386,275,1881]" captionTargetPageId="36" captionText="Fig. 13. Heteromysis sixisp. nov., holotype, ♂with body length 3.0 mm (NHMW26964). A. Thoracopod 8 with penis, caudal. BF. Series of male pleopods 15, caudal. G. Detail of male pleopod 2 (C) showing flagellate spine. H. Uropods, ventral; with setae and ventral wall of statocyst omitted. I. Telson." figureDoi="http://doi.org/10.5281/zenodo.4558777" httpUri="https://zenodo.org/record/4558777/files/figure.png" pageId="37" pageNumber="170">Fig. 13</figureCitation>
BG). Length increases in series of pleopods 1, 3 = 4, 5, 2. For presence and numbers of spines on pleopod 2, see Diagnosis above. All setae of pleopods 15 plumose or barbed.
</paragraph>
<paragraph id="8BE236E8FF8BA60715A1FCDDFD64C78E" blockId="37.[188,1399,838,1013]" pageId="37" pageNumber="170">
TAIL FAN (
<figureCitation id="13662A6DFF8BA6071433FCDCFE95C71A" box="[303,383,838,865]" captionStart="Fig" captionStartId="36.[189,232,1928,1954]" captionTargetBox="[190,1385,276,1875]" captionTargetId="figure-76@36.[189,1386,275,1881]" captionTargetPageId="36" captionText="Fig. 13. Heteromysis sixisp. nov., holotype, ♂with body length 3.0 mm (NHMW26964). A. Thoracopod 8 with penis, caudal. BF. Series of male pleopods 15, caudal. G. Detail of male pleopod 2 (C) showing flagellate spine. H. Uropods, ventral; with setae and ventral wall of statocyst omitted. I. Telson." figureDoi="http://doi.org/10.5281/zenodo.4558777" httpUri="https://zenodo.org/record/4558777/files/figure.png" pageId="37" pageNumber="170">Fig. 13</figureCitation>
HI). Exopod of uropods reaches with 1419% its length beyond endopod and 4649% beyond telson. Endopod reaches with 2326% its length beyond telson. Statoliths mineralised with fluorite. Statoliths discoidal, diameter
<quantity id="4CA59B0DFF8BA607179DFC0AFD11C7D1" box="[641,763,912,939]" metricMagnitude="-2" metricUnit="m" metricValue="4.1" metricValueMax="4.2" metricValueMin="4.0" pageId="37" pageNumber="170" unit="mm" value="41.0" valueMax="42.0" valueMin="40.0">4042 µm</quantity>
(n = 2). Telson length 1.4 times maximum width, 0.9 times endopod of uropod, 0.7 times exopod of uropod, and 1.3 times last abdominal somite. For further details of telson, see Diagnosis above.
</paragraph>
<paragraph id="8BE236E8FF8BA60715A1FB84FB6EC0F9" blockId="37.[187,1398,1054,1155]" pageId="37" pageNumber="170">
FOREGUT (
<figureCitation id="13662A6DFF8BA6071433FB84FE95C042" box="[303,383,1054,1080]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="37" pageNumber="170">Fig. 12</figureCitation>
TU) essentially as described above for
<taxonomicName id="4C5D4D6BFF8BA6071654FB85FBEFC042" authority="Wittmann &amp; Abed-Navandi, 2021" authorityName="Wittmann &amp; Abed-Navandi" authorityYear="2021" box="[840,1029,1054,1080]" class="Malacostraca" family="Mysidae" genus="Heteromysis" kingdom="Animalia" order="Mysida" pageId="37" pageNumber="170" phylum="Arthropoda" rank="species" species="smithsoniana" status="sp. nov.">
<emphasis id="B929EAFAFF8BA6071654FB85FBEFC042" box="[840,1029,1054,1080]" italics="true" pageId="37" pageNumber="170">H. smithsoniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A5781FF8BA6071111FB84FB8FC042" box="[1037,1125,1054,1080]" pageId="37" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
Dorsolateral infoldings with two comparatively large, centrally serrated spines (
<figureCitation id="13662A6DFF8BA6071644FBD9FC29C027" box="[856,963,1091,1117]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="37" pageNumber="170">Fig. 12T</figureCitation>
); posterior part of lateralia with 23 smaller, centrally serrated spines (
<figureCitation id="13662A6DFF8BA6071752FBF2FD51C0F8" box="[590,699,1128,1154]" captionStart="Fig" captionStartId="34.[189,232,1717,1743]" captionTargetBox="[280,1307,267,1680]" captionTargetId="figure-178@34.[279,1308,266,1681]" captionTargetPageId="34" captionText="Fig. 12. Heteromysis sixi sp. nov., holotype, ♂ with body length 3.0 mm (NHMW 26964). A. Anterior body region, dorsal view; right antennula, left antenna, and setae of antennal scale omitted. B. Detail of panel (A) showing right eye in lateral view. C. Right antennula, dorsal. DE. Details of antennula (C) showing flagellate spine (D) and an example for whip setae (E) found here and on other appendages. F. Palp of left mandible, caudal. G. Detail of panel (F) showing barbed seta from outer margin of median segment. H. Thoracic sternites 18 with median processes. I. Right thoracic endopod 3, rostral. JM. Details of endopod 3 (I) showing modified spines (JK) of carpus, barbed seta (L) of merus, and whip seta (M) from ischium. NS. Series of dactyli with nail in thoracic endopods 15, 8; setae omitted. TU. Modified spines from dorsolateral infoldings (T) and lateralia (U) of foregut." figureDoi="http://doi.org/10.5281/zenodo.4558775" httpUri="https://zenodo.org/record/4558775/files/figure.png" pageId="37" pageNumber="170">Fig. 12U</figureCitation>
). The examined gut was almost empty.
</paragraph>
</subSubSection>
<subSubSection id="C3476563FF8BA60715A1FB31FBDAC161" pageId="37" pageNumber="170" type="distribution">
<paragraph id="8BE236E8FF8BA60715A1FB31FEBCC0BF" blockId="37.[189,342,1195,1221]" box="[189,342,1195,1221]" pageId="37" pageNumber="170">
<heading id="D0AA8184FF8BA60715A1FB31FEBCC0BF" bold="true" box="[189,342,1195,1221]" fontSize="11" level="3" pageId="37" pageNumber="170" reason="3">
<emphasis id="B929EAFAFF8BA60715A1FB31FEBCC0BF" bold="true" box="[189,342,1195,1221]" pageId="37" pageNumber="170">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BE236E8FF8BA60715A1FB46FBDAC161" blockId="37.[189,1399,1244,1308]" pageId="37" pageNumber="170">
The species is so far known only from a service tank of the Waikiki Aquarium, Honolulu, Hawaii. Origin most likely in coastal marine waters of the Central Pacific (
<tableCitation id="C6DF0353FF8BA60716DBFA9BFBF5C161" box="[967,1055,1281,1307]" captionStart="Table 1" captionStartId="2.[189,258,265,291]" captionTargetPageId="2" captionText="Table 1. Samples and data of Heteromysis S.I. Smith, 1873 obtained from aquarium institutions." httpUri="http://table.plazi.org/id/DF226660FFACA62015A1FE93FAC2C559" pageId="37" pageNumber="170" tableUuid="DF226660FFACA62015A1FE93FAC2C559">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>