256 lines
33 KiB
XML
256 lines
33 KiB
XML
<document id="7B977E7214401BE02B8562C6CEC7EB5A" ID-CLB-Dataset="295747" ID-DOI="10.1590/1982-0224-2022-0089" ID-GBIF-Dataset="9ec7a6a4-dd99-4d15-a180-0e3dbed303da" ID-ISSN="1982-0224" ID-Zenodo-Dep="11125059" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1712845780947" checkinUser="felipe" docAuthor="Domínguez-Cisneros, Sara E., Domínguez-Domínguez, Omar, Velázquez-Velázquez, Ernesto & Pérez-Rodríguez, Rodolfo" docDate="2023" docId="03F48798FFE4FFF98D45AC7AD8226EC4" docLanguage="en" docName="NeotropIchthyol.2023.21.1.e220089.pdf" docOrigin="Neotropical Ichthyology (e 220089) 21 (1)" docSource="http://dx.doi.org/10.1590/1982-0224-2022-0089" docStyle="DocumentStyle:FC0C3FD10650AF21F8DA93D98D275823.3:NeotropIchthyol.2020-.journal_article" docStyleId="FC0C3FD10650AF21F8DA93D98D275823" docStyleName="NeotropIchthyol.2020-.journal_article" docStyleVersion="3" docTitle="Profundulus Hubbs 1924" docType="treatment" docVersion="3" lastPageNumber="15" masterDocId="FFCDFFE0FFE8FFF78E7DAA14DB646F1F" masterDocTitle="Redescription and diagnoses of the genera Profundulus and Tlaloc (Cyprinodontiformes: Profundulidae), Mesoamerican endemic fishes" masterLastPageNumber="22" masterPageNumber="1" pageNumber="13" updateTime="1715071952175" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5992E5F5C9CE771FB7780FB13EFDCA3C">Redescription and diagnoses of the genera Profundulus and Tlaloc (Cyprinodontiformes: Profundulidae), Mesoamerican endemic fishes</mods:title>
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<mods:namePart id="EEF930D1205FCD3242E6CE03180A7136">Domínguez-Cisneros, Sara E.</mods:namePart>
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<mods:affiliation id="7B2E53E65A8557A200A4B952D1B9A8B8">Doctorado en Biodiversidad y Conservación de Ecosistemas Tropicales, Instituto de Ciencias Biológicas, Universidad de Ciencias y Artes de Chiapas, Libramiento Norte, 1150, Col. Lajas Maciel, 29029 Tuxtla Gutiérrez, CHIS, México. (SEDC) sara. dominguez @ unicach. mx. & Colección de Peces, Museo de ZoologÍa, Instituto de Ciencias Biológicas, Universidad de Ciencias y Artes de Chiapas, Libramiento Norte, 1150, Col. Lajas Maciel, 29029 Tuxtla Gutiérrez, CHIS, México. (EVV) ernesto. velazquez @ unicach. mx (corresponding author).</mods:affiliation>
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<mods:nameIdentifier id="0660D49D3E9854A66B067F20A454CD29" type="email">ernesto.velazquez@unicach.mx</mods:nameIdentifier>
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<mods:namePart id="91910B79FFA1ACFE3DB3FAEA296D3761">Domínguez-Domínguez, Omar</mods:namePart>
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<mods:affiliation id="818CD6265C65CD582CB1C5AEDBC5DAB7">Laboratorio de BiologÍa Acuática, Facultad de BiologÍa, Universidad Michoacana de San Nicolás de Hidalgo, Edificio “ R ” planta baja, Ciudad Universitaria, Francisco J. Mujica S / N, 58030 Morelia, MEX, México. (ODD) goodeido @ yahoo. com. mx, (RPR) rperezr 38 @ gmail. com.</mods:affiliation>
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<mods:nameIdentifier id="E1543F9BCF2E405DF115A2E86BE21258" type="email">goodeido@yahoo.com.mx</mods:nameIdentifier>
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<mods:namePart id="D95CD99AECD4ADBF302652A5FB09C19C">Velázquez-Velázquez, Ernesto</mods:namePart>
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<mods:affiliation id="62C11323B728FE67C9C5616CCE406C2C">Colección de Peces, Museo de ZoologÍa, Instituto de Ciencias Biológicas, Universidad de Ciencias y Artes de Chiapas, Libramiento Norte, 1150, Col. Lajas Maciel, 29029 Tuxtla Gutiérrez, CHIS, México. (EVV) ernesto. velazquez @ unicach. mx (corresponding author).</mods:affiliation>
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<mods:nameIdentifier id="4424F60C2A91C291169FF7B97C06DA08" type="email">ernesto.velazquez@unicach.mx</mods:nameIdentifier>
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<mods:namePart id="CABA47768D7180D0859756974460420A">Pérez-Rodríguez, Rodolfo</mods:namePart>
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<mods:affiliation id="1219164A5B59E31005F13137D29A832B">Laboratorio de BiologÍa Acuática, Facultad de BiologÍa, Universidad Michoacana de San Nicolás de Hidalgo, Edificio “ R ” planta baja, Ciudad Universitaria, Francisco J. Mujica S / N, 58030 Morelia, MEX, México. (ODD) goodeido @ yahoo. com. mx, (RPR) rperezr 38 @ gmail. com.</mods:affiliation>
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<mods:nameIdentifier id="A530E71C4BCDC6340B782B902C749C12" type="email">goodeido@yahoo.com.mx</mods:nameIdentifier>
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<mods:title id="0526C0A30B0655A05DFF32069A3E9474">Neotropical Ichthyology</mods:title>
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<mods:date id="8CC53338C9D48CB2B07A933F12D4B30F">2023</mods:date>
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<mods:title id="88699DE325DD936FC00D9A68E6C8746F">e 220089</mods:title>
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<mods:number id="77ECD7589B480D591B1DCF41BF0B4EC8">2023-02-20</mods:number>
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<mods:number id="B416B6E44A31EE1076A28FDBCB6B540B">21</mods:number>
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<mods:url id="E633CE147E2FCA90B2DBE26900F2A6D8">http://dx.doi.org/10.1590/1982-0224-2022-0089</mods:url>
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<mods:classification id="47B84680D2BA89497DE45713B5E7B3BF">journal article</mods:classification>
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<mods:identifier id="D84D678D58AB97FACDD89289269A296F" type="DOI">10.1590/1982-0224-2022-0089</mods:identifier>
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<treatment id="03F48798FFE4FFF98D45AC7AD8226EC4" ID-DOI="http://doi.org/10.5281/zenodo.11125077" ID-Zenodo-Dep="11125077" LSID="urn:lsid:plazi:treatment:03F48798FFE4FFF98D45AC7AD8226EC4" httpUri="http://treatment.plazi.org/id/03F48798FFE4FFF98D45AC7AD8226EC4" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13">
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<subSubSection id="C3476505FFE4FFFB8D45AC7ADF076999" box="[824,1123,1646,1670]" pageId="12" pageNumber="13" type="nomenclature">
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<paragraph id="8BE2368EFFE4FFFB8D45AC7ADF076999" blockId="12.[824,1123,1646,1670]" box="[824,1123,1646,1670]" pageId="12" pageNumber="13">
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<heading id="D0AA81E2FFE4FFFB8D45AC7ADF076999" box="[824,1123,1646,1670]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="13" reason="2">
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<taxonomicName id="4C5D4D0DFFE4FFFB8D45AC7ADF076999" ID-CoL="6WH8" ID-ENA="28769" authority="Hubbs, 1924" authorityName="Hubbs" authorityYear="1924" box="[824,1123,1646,1670]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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<emphasis id="B929EA9CFFE4FFFB8D45AC7AD8A96999" bold="true" box="[824,973,1646,1670]" pageId="12" pageNumber="13">Profundulus</emphasis>
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Hubbs, 1924
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3476505FFE4FFFB8F95ACA8D98469D0" box="[488,736,1724,1743]" pageId="12" pageNumber="13" type="reference_group">
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<paragraph id="8BE2368EFFE4FFFB8F95ACA8D98469D0" blockId="12.[488,1392,1724,1743]" box="[488,736,1724,1743]" pageId="12" pageNumber="13">
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<taxonomicName id="4C5D4D0DFFE4FFFB8F95ACA8D98469D0" authority="Hubbs, 1924: 12" authorityName="Hubbs" authorityPageNumber="12" authorityYear="1924" box="[488,736,1724,1743]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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<emphasis id="B929EA9CFFE4FFFB8F95ACA8D92969D0" box="[488,589,1724,1743]" italics="true" pageId="12" pageNumber="13">Profundulus</emphasis>
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Hubbs, 1924:12
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3476505FFE4FFFB8C98ACA8DE1469D0" box="[741,1392,1724,1743]" pageId="12" pageNumber="13" type="type_taxon">
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<paragraph id="8BE2368EFFE4FFFB8C98ACA8DE1469D0" blockId="12.[488,1392,1724,1743]" box="[741,1392,1724,1743]" pageId="12" pageNumber="13">
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(
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<typeStatus id="54E6882CFFE4FFFB8C96ACA8D87769D0" box="[747,787,1724,1743]" pageId="12" pageNumber="13">type</typeStatus>
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species
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<taxonomicName id="4C5D4D0DFFE4FFFB8D26ACA8DFED69D0" authority="Gunther, 1866" authorityName="Gunther" authorityYear="1866" box="[859,1161,1724,1743]" family="Fundulidae" genus="Fundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="punctatus">
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<emphasis id="B929EA9CFFE4FFFB8D26ACA8D89A69D0" box="[859,1022,1724,1743]" italics="true" pageId="12" pageNumber="13">Fundulus punctatus</emphasis>
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Günther, 1866
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</taxonomicName>
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, by original designation).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3476505FFE4FFFA8C7BAD3CDE176D6F" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="diagnosis">
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<paragraph id="8BE2368EFFE4FFFA8C7BAD3CDE176D6F" blockId="12.[488,1459,1832,1931]" lastBlockId="13.[488,1459,190,625]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
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<emphasis id="B929EA9CFFE4FFFB8C7BAD3CD9E2685F" bold="true" box="[518,646,1832,1856]" pageId="12" pageNumber="13">Diagnosis.</emphasis>
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<taxonomicName id="4C5D4D0DFFE4FFFB8CECAD3CD86B685F" authorityName="Hubbs" authorityYear="1924" box="[657,783,1832,1856]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="12" pageNumber="13" phylum="Chordata" rank="genus">
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<emphasis id="B929EA9CFFE4FFFB8CECAD3CD86B685F" box="[657,783,1832,1856]" italics="true" pageId="12" pageNumber="13">Profundulus</emphasis>
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</taxonomicName>
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is diagnosed here based on the following characters: The mesethmoid is small, crescent-shaped, and does not extend beyond the margin of the vomer (
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<figureCitation id="13662A0BFFE4FFFB8C3DAD67D9AC6894" box="[576,712,1907,1931]" captionStart="FIGURE 2" captionStartId="4.[488,561,1761,1781]" captionTargetBox="[502,1446,1202,1716]" captionTargetId="figure-298@4.[488,1459,1179,1744]" captionTargetPageId="4" captionText="FIGURE 2 | Ventral view of the neurocranium, in specimens cleared and stained. A. Tlaloc portillorum (MZ-UNICACH 7222, 70.5 mm SL); B. Vomer region of T. candalarius (MZ-UNICACH 3899, 61.7 mm SL); C. Vomer region of Profundulus kreiseri (MZ-UNICACH 7214, 59.6 mm SL); and D. Neurocranium of P. mixtlanensis (MZ-UNICACH 6716, 55.6 mm SL). Abbreviations: V, vomer (Y-shaped); E, lateral ethmoid; M, mesethmoid; P, parasphenoid; AF, autopterotic fossa." figureDoi="http://doi.org/10.5281/zenodo.11125063" httpUri="https://zenodo.org/record/11125063/files/figure.png" pageId="12" pageNumber="13">Figs. 2C–D</figureCitation>
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); the vomer is greatly broadened anteriorly, lacks lateral processes and is in contact with the lateral ethmoids; the anterior portion of the parasphenoid is just in contact with the mesethmoid and does not extend beyond the center of the mesethmoid; the autopterotic fossa is large (
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<figureCitation id="13662A0BFFE5FFFA8A60AB1CDF1E6E3E" box="[1053,1146,264,289]" captionStart="FIGURE 2" captionStartId="4.[488,561,1761,1781]" captionTargetBox="[502,1446,1202,1716]" captionTargetId="figure-298@4.[488,1459,1179,1744]" captionTargetPageId="4" captionText="FIGURE 2 | Ventral view of the neurocranium, in specimens cleared and stained. A. Tlaloc portillorum (MZ-UNICACH 7222, 70.5 mm SL); B. Vomer region of T. candalarius (MZ-UNICACH 3899, 61.7 mm SL); C. Vomer region of Profundulus kreiseri (MZ-UNICACH 7214, 59.6 mm SL); and D. Neurocranium of P. mixtlanensis (MZ-UNICACH 6716, 55.6 mm SL). Abbreviations: V, vomer (Y-shaped); E, lateral ethmoid; M, mesethmoid; P, parasphenoid; AF, autopterotic fossa." figureDoi="http://doi.org/10.5281/zenodo.11125063" httpUri="https://zenodo.org/record/11125063/files/figure.png" pageId="13" pageNumber="14">Fig. 2D</figureCitation>
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); the dorsal margin of the interoperculum, with a short extension, does not exceed the edge of the bone (
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<figureCitation id="13662A0BFFE5FFFA8B2CAB3ADEC16E59" box="[1361,1445,302,326]" captionStart="FIGURE 3" captionStartId="5.[488,561,1276,1296]" captionTargetBox="[503,1417,186,1201]" captionTargetId="figure-21@5.[488,1459,171,1226]" captionTargetPageId="5" captionText="FIGURE 3 | Lateral view of left jaws, suspensorium and opercular series in specimens cleared and stained. A. Tlaloc portillorum (MZ-UNICACH 7222, 70.5 mm SL); B. Profundulus mixtlanensis (MZ-UNICACH 6716, 55.6 mm SL). Arrow points to dorsal margin of the interoperculum, with a long or short extension. Abbreviations: de, dentary; pm, premaxilla; ra, retroarticular; mx, maxilla; pl, palatine; qu, quadrate; ar, articular; ms, mesopterygoid; io, interopercle; sy, sympletic; hy, hyomandibula; po, preopercle; op, opercle; so, subopercle." figureDoi="http://doi.org/10.5281/zenodo.11125065" httpUri="https://zenodo.org/record/11125065/files/figure.png" pageId="13" pageNumber="14">Fig. 3B</figureCitation>
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); the ventral margin of the lacrimal slightly concave (
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<figureCitation id="13662A0BFFE5FFFA8A50AB47DFD06E74" box="[1069,1204,339,363]" captionStart="FIGURE 4" captionStartId="8.[488,561,1201,1221]" captionTargetBox="[518,1367,189,1178]" captionTargetId="figure-25@8.[484,1462,168,1182]" captionTargetPageId="8" captionText="FIGURE 4 | Ventral view of left lacrimal. A. Tlaloc portillorum (MZ-UNICACH 7222, 70.5 mm SL); B. T. hildebrandi (MZ-UNICACH 2266, 75.1 mm SL); C. Profundulus mixtlanensis (MZ-UNICACH 6716, 55.6 mm SL); and D. P. punctatus (MZ-UNICACH 6632, 67.3 mm SL). Arrow point to ventral margin of the lacrimal." figureDoi="http://doi.org/10.5281/zenodo.11125069" httpUri="https://zenodo.org/record/11125069/files/figure.png" pageId="13" pageNumber="14">Figs. 4C–D</figureCitation>
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).
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<taxonomicName id="4C5D4D0DFFE5FFFA8AB5AB47DE226E74" authorityName="Hubbs" authorityYear="1924" box="[1224,1350,339,363]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
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<emphasis id="B929EA9CFFE5FFFA8AB5AB47DE226E74" box="[1224,1350,339,363]" italics="true" pageId="13" pageNumber="14">Profundulus</emphasis>
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</taxonomicName>
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is further distinguished from
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<taxonomicName id="4C5D4D0DFFE5FFFA8CB9AB6CD8626E8F" authorityName="Alvarez, Carranza" authorityYear="1951" box="[708,774,376,400]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
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<emphasis id="B929EA9CFFE5FFFA8CB9AB6CD8626E8F" box="[708,774,376,400]" italics="true" pageId="13" pageNumber="14">Tlaloc</emphasis>
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</taxonomicName>
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in having more than the basal half of the caudal fin densely scaled (except in
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<taxonomicName id="4C5D4D0DFFE5FFFA8CDAAB8BD86C6EA9" authorityName="Matamoros, Schaefer, Hernandez & Chakrabarty" authorityYear="2012" box="[679,776,414,438]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="kreiseri">
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<emphasis id="B929EA9CFFE5FFFA8CDAAB8BD86C6EA9" box="[679,776,414,438]" italics="true" pageId="13" pageNumber="14">P. kreiseri</emphasis>
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</taxonomicName>
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) (
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<emphasis id="B929EA9CFFE5FFFA8D5DAB8BD8516EA9" box="[800,821,415,438]" italics="true" pageId="13" pageNumber="14">vs</emphasis>
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. less than the basal half of caudal fin densely scaled) (
|
||
<figureCitation id="13662A0BFFE5FFFA8BFBAB8AD96D6EC4" captionStart="FIGURE 5" captionStartId="10.[488,561,977,997]" captionTargetBox="[488,1459,173,954]" captionTargetId="figure-24@10.[488,1459,173,954]" captionTargetPageId="10" captionText="FIGURE 5 | Male general morphology and caudal fin squamation, life colour patterns in: A. Tlaloc labialis; and B. Profundulus punctatus. Solid arrow points to the squamation of the caudal fin, dashed arrow points to the humeral spot." figureDoi="http://doi.org/10.5281/zenodo.11125071" httpUri="https://zenodo.org/record/11125071/files/figure.png" pageId="13" pageNumber="14">Fig. 5B</figureCitation>
|
||
); by a humeral spot (except in
|
||
<taxonomicName id="4C5D4D0DFFE5FFFA8D12ABD0D8A56EC4" authorityName="Dominguez-Cisneros, Velazquez-Velazquez, McMahan & Matamoros" authorityYear="2021" box="[879,961,451,475]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="adani">
|
||
<emphasis id="B929EA9CFFE5FFFA8D12ABD0D8A56EC4" box="[879,961,451,475]" italics="true" pageId="13" pageNumber="14">P. adani</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<emphasis id="B929EA9CFFE5FFFA8DA6ABD0D8946EC4" box="[987,1008,452,475]" italics="true" pageId="13" pageNumber="14">vs</emphasis>
|
||
. humeral spot absent) (
|
||
<figureCitation id="13662A0BFFE5FFFA8A81ABD7DE306EC4" box="[1276,1364,451,475]" captionStart="FIGURE 5" captionStartId="10.[488,561,977,997]" captionTargetBox="[488,1459,173,954]" captionTargetId="figure-24@10.[488,1459,173,954]" captionTargetPageId="10" captionText="FIGURE 5 | Male general morphology and caudal fin squamation, life colour patterns in: A. Tlaloc labialis; and B. Profundulus punctatus. Solid arrow points to the squamation of the caudal fin, dashed arrow points to the humeral spot." figureDoi="http://doi.org/10.5281/zenodo.11125071" httpUri="https://zenodo.org/record/11125071/files/figure.png" pageId="13" pageNumber="14">Fig. 5B</figureCitation>
|
||
); by the origin of the dorsal fin positioned at a vertical line slightly anterior to the origin of the anal fin (
|
||
<emphasis id="B929EA9CFFE5FFFA8C34A81BD93A6D39" box="[585,606,527,550]" italics="true" pageId="13" pageNumber="14">vs</emphasis>
|
||
. origin of the dorsal fin positioned at a vertical line posterior to the origin of the anal fin); by short epiotic processes, not extending beyond the first vertebra (except in
|
||
<taxonomicName id="4C5D4D0DFFE5FFFA8C78A84DD91D6D6F" authorityName="Matamoros, Dominguez-Cisneros, Velazquez-Velazquez & McMahan" authorityYear="2018" box="[517,633,601,624]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="parentiae">
|
||
<emphasis id="B929EA9CFFE5FFFA8C78A84DD91D6D6F" box="[517,633,601,624]" italics="true" pageId="13" pageNumber="14">P. parentiae</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<emphasis id="B929EA9CFFE5FFFA8CECA84DD9C26D6F" box="[657,678,601,624]" italics="true" pageId="13" pageNumber="14">vs</emphasis>
|
||
. long epiotic processes, extending beyond the second vertebra).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3476505FFE5FFF98C7BA8B7D9D26E74" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
|
||
<paragraph id="8BE2368EFFE5FFFA8C7BA8B7D99A6CDE" blockId="13.[488,1460,675,1296]" pageId="13" pageNumber="14">
|
||
<emphasis id="B929EA9CFFE5FFFA8C7BA8B7D9F86DA4" bold="true" box="[518,668,675,699]" pageId="13" pageNumber="14">Description.</emphasis>
|
||
Morphometric data appear in
|
||
<tableCitation id="C6DF0335FFE5FFFA8A7EA8B7DF366DA4" box="[1027,1106,675,699]" captionStart="TABLE 2" captionStartId="9.[134,196,193,213]" captionTargetBox="[151,1437,250,893]" captionTargetId="graphics-43@9.[136,1459,234,909]" captionTargetPageId="9" captionText="TABLE 2 | Meristic data of Tlaloc and Profundulus species. Asterisk mark the number of specimens analyzed, in parenthesis the modal values." httpUri="http://table.plazi.org/id/DF226606FFE1FFFE8EFBAAD5DED06FCA" pageId="13" pageNumber="14" tableUuid="DF226606FFE1FFFE8EFBAAD5DED06FCA">Tab. 2</tableCitation>
|
||
. Body rather robust, elongate; head compressed, large (24.1–35.3%); the interorbital broad (38.2–58.7%) and typically concave or nearly flat. Ascending premaxillary process short and broad and bluntly rounded at the tip. Mouth subterminal, lower jaw broad, heavy and protruding so that upper jaw is included. Dorsal fin rounded in both males and females with its basal length short (11.1–19.2%). Anal fins of males rounded, slightly elongated in females. Caudal peduncle relatively short (11.1–23.2%). Median hypural plate divided into subequal parts by an open groove.
|
||
</paragraph>
|
||
<paragraph id="8BE2368EFFE5FFFA8C7BA9DADF046B64" blockId="13.[488,1460,675,1296]" pageId="13" pageNumber="14">
|
||
Lateral-line system of sensory canals and pores on the head of
|
||
<taxonomicName id="4C5D4D0DFFE5FFFA8A89A9DADE166CF9" authorityName="Hubbs" authorityYear="1924" box="[1268,1394,974,998]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis id="B929EA9CFFE5FFFA8A89A9DADE166CF9" box="[1268,1394,974,998]" italics="true" pageId="13" pageNumber="14">Profundulus</emphasis>
|
||
</taxonomicName>
|
||
: The supraorbital series follows the
|
||
<typeStatus id="54E6882CFFE5FFFA8D2FA9E0D8EA6B13" box="[850,910,1012,1036]" pageId="13" pageNumber="14">Type</typeStatus>
|
||
II designation of Gosline (1949), with canals between pores 1–2a, 2b–4a, 4b–7; preopercular pores 6–8 (mode = 7); preorbital pores 3–5 (mode = 4); mandibular pores 4–5 (mode = 5); generally, there is a lack of rostral pores, exceptionally there may be
|
||
<quantity id="4CA59B6BFFE5FFFA8D1BAE77D8A06B64" box="[870,964,1123,1147]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="13" pageNumber="14" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">2 to 4 in</quantity>
|
||
some species.
|
||
</paragraph>
|
||
<paragraph id="8BE2368EFFE5FFFA8C7BAE9CDF126A0F" blockId="13.[488,1460,675,1296]" pageId="13" pageNumber="14">
|
||
Meristic traits are as follows (
|
||
<tableCitation id="C6DF0335FFE5FFFA8D39AE9CD8EA6BBE" box="[836,910,1160,1185]" captionStart="TABLE 2" captionStartId="9.[134,196,193,213]" captionTargetBox="[151,1437,250,893]" captionTargetId="graphics-43@9.[136,1459,234,909]" captionTargetPageId="9" captionText="TABLE 2 | Meristic data of Tlaloc and Profundulus species. Asterisk mark the number of specimens analyzed, in parenthesis the modal values." httpUri="http://table.plazi.org/id/DF226606FFE1FFFE8EFBAAD5DED06FCA" pageId="13" pageNumber="14" tableUuid="DF226606FFE1FFFE8EFBAAD5DED06FCA">Tab. 2</tableCitation>
|
||
): anal: 11–17; dorsal: 10–15; pelvic: 5–7; pectoral: 13–19; caudal: 17–24; vertebrae totals: 31–35; gill rakers on anterior arm of the first arch: 14–22; branchiostegal rays, 6; scales lateral series: 29–35; scales predorsal: 18–29; scales around body: 23–35; scales around peduncle: 16–25.
|
||
</paragraph>
|
||
<paragraph id="8BE2368EFFE5FFFA8C7BAF57D80B6AD4" blockId="13.[488,1459,1347,1483]" pageId="13" pageNumber="14">
|
||
<emphasis id="B929EA9CFFE5FFFA8C7BAF57D9F56A44" bold="true" box="[518,657,1347,1371]" pageId="13" pageNumber="14">Coloration.</emphasis>
|
||
The form and coloration of the nuptial adult male are shown in
|
||
<figureCitation id="13662A0BFFE5FFFA8BFBAF57D96E6A9F" captionStart="FIGURE 5" captionStartId="10.[488,561,977,997]" captionTargetBox="[488,1459,173,954]" captionTargetId="figure-24@10.[488,1459,173,954]" captionTargetPageId="10" captionText="FIGURE 5 | Male general morphology and caudal fin squamation, life colour patterns in: A. Tlaloc labialis; and B. Profundulus punctatus. Solid arrow points to the squamation of the caudal fin, dashed arrow points to the humeral spot." figureDoi="http://doi.org/10.5281/zenodo.11125071" httpUri="https://zenodo.org/record/11125071/files/figure.png" pageId="13" pageNumber="14">Fig. 5B</figureCitation>
|
||
. Most of the body dark, often with irregular brown-dark spots on the scales, on the sides of the body on to the caudal fin. A golden yellow blotch covers the operculum and reaches the base of the pectoral fin.
|
||
</paragraph>
|
||
<paragraph id="8BE2368EFFE5FFFA8C7BAFEAD9A4685F" blockId="13.[488,1460,1534,1856]" pageId="13" pageNumber="14">
|
||
<emphasis id="B929EA9CFFE5FFFA8C7BAFEAD8BF6909" bold="true" box="[518,987,1534,1558]" pageId="13" pageNumber="14">Sexual dimorphism and contact organs.</emphasis>
|
||
The sexual dimorphism is not very evident in species of the genus
|
||
<taxonomicName id="4C5D4D0DFFE5FFFA8C95AC37D8026924" authorityName="Hubbs" authorityYear="1924" box="[744,870,1571,1595]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis id="B929EA9CFFE5FFFA8C95AC37D8026924" box="[744,870,1571,1595]" italics="true" pageId="13" pageNumber="14">Profundulus</emphasis>
|
||
</taxonomicName>
|
||
. However, the males are slightly larger than females; the largest specimen recorded was a male from
|
||
<taxonomicName id="4C5D4D0DFFE5FFFA8D8AAC5DDFF9697F" baseAuthorityName="Gunther" baseAuthorityYear="1866" box="[1015,1181,1608,1632]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="guatemalensis">
|
||
<emphasis id="B929EA9CFFE5FFFA8D8AAC5DDFF9697F" box="[1015,1181,1608,1632]" italics="true" pageId="13" pageNumber="14">P. guatemalensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="4CA59B6BFFE5FFFA8AD6AC5DDE74697F" box="[1195,1296,1609,1633]" metricMagnitude="-2" metricUnit="m" metricValue="8.39" pageId="13" pageNumber="14" unit="mm" value="83.9">83.9 mm</quantity>
|
||
SL), while the maximum size recorded in a female was in
|
||
<taxonomicName id="4C5D4D0DFFE5FFFA8DB1AC7BDF786999" authorityName="Dominguez-Cisneros, Velazquez-Velazquez, McMahan & Matamoros" authorityYear="2021" box="[972,1052,1646,1670]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="adani">
|
||
<emphasis id="B929EA9CFFE5FFFA8DB1AC7BDF786999" box="[972,1052,1646,1670]" italics="true" pageId="13" pageNumber="14">P. adani</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="4CA59B6BFFE5FFFA8A56AC7ADFF4699A" box="[1067,1168,1646,1670]" metricMagnitude="-2" metricUnit="m" metricValue="7.79" pageId="13" pageNumber="14" unit="mm" value="77.9">77.9 mm</quantity>
|
||
SL). In males and females of all species of
|
||
<taxonomicName id="4C5D4D0DFFE5FFFA8CD9AC87D84669B4" authorityName="Hubbs" authorityYear="1924" box="[676,802,1683,1707]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis id="B929EA9CFFE5FFFA8CD9AC87D84669B4" box="[676,802,1683,1707]" italics="true" pageId="13" pageNumber="14">Profundulus</emphasis>
|
||
</taxonomicName>
|
||
, the dorsal and the anal fins have a similar morphology. Like
|
||
<taxonomicName id="4C5D4D0DFFE5FFFA8C5DACACD90669CF" authorityName="Alvarez, Carranza" authorityYear="1951" box="[544,610,1720,1744]" family="Profundulidae" genus="Tlaloc" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis id="B929EA9CFFE5FFFA8C5DACACD90669CF" box="[544,610,1720,1744]" italics="true" pageId="13" pageNumber="14">Tlaloc</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13662A0BFFE5FFFA8C0FACACD9D769CE" box="[626,691,1720,1745]" captionStart="FIGURE 7" captionStartId="11.[488,561,1836,1856]" captionTargetBox="[488,1459,1159,1814]" captionTargetId="figure-88@11.[488,1459,1159,1814]" captionTargetPageId="11" captionText="FIGURE 7 | Bony spinules in fin rays of male Tlaloc labialis, indicated by arrow (MZ-UNICACH 6740, 75.7 mm SL), articulated with the lateral surfaces of the anal fin rays." figureDoi="http://doi.org/10.5281/zenodo.11125079" httpUri="https://zenodo.org/record/11125079/files/figure.png" pageId="13" pageNumber="14">Fig. 7</figureCitation>
|
||
) the males of
|
||
<taxonomicName id="4C5D4D0DFFE5FFFA8D32ACACD8A969CF" authorityName="Hubbs" authorityYear="1924" box="[847,973,1720,1744]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis id="B929EA9CFFE5FFFA8D32ACACD8A969CF" box="[847,973,1720,1744]" italics="true" pageId="13" pageNumber="14">Profundulus</emphasis>
|
||
</taxonomicName>
|
||
have contact organs or spinules, articulated with the lateral surfaces of the anal fin rays with some of them conspicuously long; this is the most notable, though not striking difference between males and females of all
|
||
<taxonomicName id="4C5D4D0DFFE5FFFA8F95AD3CD9A4685F" box="[488,704,1832,1856]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B929EA9CFFE5FFFA8F95AD3CD902685F" box="[488,614,1832,1856]" italics="true" pageId="13" pageNumber="14">Profundulus</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BE2368EFFE6FFF98C7BAAAAD9D26E74" blockId="14.[488,1459,190,363]" pageId="14" pageNumber="15">
|
||
<emphasis id="B929EA9CFFE6FFF98C7BAAAAD82F6FC9" bold="true" box="[518,843,190,214]" pageId="14" pageNumber="15">Geographical distribution.</emphasis>
|
||
On the Pacific slope, from the río Malinaltepec, río Papagayo basin, of
|
||
<collectingRegion id="4999F86CFFE6FFF98CB3AAF7D85C6FE4" box="[718,824,227,251]" country="Mexico" name="Guerrero" pageId="14" pageNumber="15">Guerrero</collectingRegion>
|
||
, southeastern
|
||
<collectingCountry id="F34A761EFFE6FFF98DA3AAF7DF516FE4" box="[990,1077,227,251]" name="Mexico" pageId="14" pageNumber="15">Mexico</collectingCountry>
|
||
, southward to the río Lempa in
|
||
<collectingCountry id="F34A761EFFE6FFF98F95AB1CD90D6E3F" box="[488,617,264,288]" name="El Salvador" pageId="14" pageNumber="15">El Salvador</collectingCountry>
|
||
and
|
||
<collectingCountry id="F34A761EFFE6FFF98CDAAB1CD87C6E3F" box="[679,792,264,288]" name="Honduras" pageId="14" pageNumber="15">Honduras</collectingCountry>
|
||
; on the Atlantic slope, from the upper reaches of the río Papaloapan of
|
||
<collectingRegion id="4999F86CFFE6FFF98CE8AB3AD98F6E59" box="[661,747,302,326]" country="Mexico" name="Oaxaca" pageId="14" pageNumber="15">Oaxaca</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34A761EFFE6FFF98C86AB3AD8366E59" box="[763,850,302,326]" name="Mexico" pageId="14" pageNumber="15">Mexico</collectingCountry>
|
||
, to the ríos Chamelecón and Ulúa in northwestern
|
||
<collectingCountry id="F34A761EFFE6FFF98F95AB47D9336E74" box="[488,599,339,363]" name="Honduras" pageId="14" pageNumber="15">Honduras</collectingCountry>
|
||
(
|
||
<figureCitation id="13662A0BFFE6FFF98C1BAB47D9CD6E74" box="[614,681,339,363]" captionStart="FIGURE 8" captionStartId="12.[137,210,1201,1221]" captionTargetBox="[159,1431,180,1164]" captionTargetId="figure-157@12.[136,1459,156,1178]" captionTargetPageId="12" captionText="FIGURE 8 | Geographical distribution of species of the genera Tlaloc and Profundulus in southern Mexico and Central America." figureDoi="http://doi.org/10.5281/zenodo.11125081" httpUri="https://zenodo.org/record/11125081/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3476505FFE6FFF98C7BAB8AD8226EC4" pageId="14" pageNumber="15" type="etymology">
|
||
<paragraph id="8BE2368EFFE6FFF98C7BAB8AD8226EC4" blockId="14.[488,1459,414,476]" pageId="14" pageNumber="15">
|
||
<emphasis id="B929EA9CFFE6FFF98C7BAB8AD9F56EA9" bold="true" box="[518,657,414,438]" pageId="14" pageNumber="15">Etymology.</emphasis>
|
||
<taxonomicName id="4C5D4D0DFFE6FFF98CE6AB8AD87D6EA9" authorityName="Hubbs" authorityYear="1924" box="[667,793,414,438]" family="Profundulidae" genus="Profundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="B929EA9CFFE6FFF98CE6AB8AD87D6EA9" box="[667,793,414,438]" italics="true" pageId="14" pageNumber="15">Profundulus</emphasis>
|
||
</taxonomicName>
|
||
in reference to its presumed primitive relationship to the genus
|
||
<taxonomicName id="4C5D4D0DFFE6FFF98C4CABD7D8266EC4" authority="(Parenti, 1981)" baseAuthorityName="Parenti" baseAuthorityYear="1981" box="[561,834,451,475]" family="Fundulidae" genus="Fundulus" kingdom="Animalia" order="Cyprinodontiformes" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="B929EA9CFFE6FFF98C4CABD7D9F76EC4" box="[561,659,451,475]" italics="true" pageId="14" pageNumber="15">Fundulus</emphasis>
|
||
(Parenti, 1981)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |