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<document id="6566DC408BDD92ADB7FA9F31BB686D96" ID-CLB-Dataset="3948" ID-DOI="10.5852/ejt.2021.775.1551" ID-GBIF-Dataset="cc10c087-bec3-4947-81d7-b3e5580a5c94" ID-ISSN="2118-9773" ID-Zenodo-Dep="5589689" ID-ZooBank="976D7020-5904-4951-97CE-B4FE58DA12A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634822039012" checkinUser="felipe" docAuthor="González-Alvarado, Arturo &amp; Vaz-De, Fernando Z." docDate="2021" docId="03F48795DC5AFFBAA5EEA6D7FC4AFDEB" docLanguage="en" docName="ejt-775_86-106.pdf" docOrigin="European Journal of Taxonomy 775" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Deltochilum gilli Gonzalez-Alvarado &amp; Vaz-de-Mello 2021" docType="treatment" docVersion="9" lastPageNumber="95" masterDocId="FFCDFFEDDC5DFFB3A45AA753FFCDFFC7" masterDocTitle="Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum)" masterLastPageNumber="106" masterPageNumber="86" pageNumber="93" updateTime="1699464201688" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="05C6FA692819B4266263B58497047AAE">Taxonomic revision of the Deltohyboma Lane, 1946 gilli species group (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilum)</mods:title>
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<mods:namePart id="49BCDF7F690BB2F7902C7A935E87000B">González-Alvarado, Arturo</mods:namePart>
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<mods:affiliation id="912FA0020A817A86F6BB157652A95A26">Programa de Pós-graduação em Ecologia e Conservação da Biodiversidade, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<mods:namePart id="99E4C6A144F5A909706C9F20532F9DE4">Vaz-De, Fernando Z.</mods:namePart>
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<mods:affiliation id="DB1E06148A6B84ED6BB28ACDC645744B">Instituto de Biociências, Departamento de Biologia e Zoologia, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<heading id="D0AA81EFDC5AFFB4A5EEA6D7FB4DFE59" box="[436,1152,388,415]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="93" reason="2">
<taxonomicName id="4C5D4D00DC5AFFB4A5EEA6D7FB4DFE59" ID-CoL="7VN7Q" authority="Gonzalez-Alvarado &amp; Vaz-de-Mello, 2021" authorityName="Gonzalez-Alvarado &amp; Vaz-de-Mello" authorityYear="2021" box="[436,1152,388,415]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="93" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B929EA91DC5AFFB4A5EEA6D7FD4EFE58" bold="true" box="[436,643,388,415]" italics="true" pageId="7" pageNumber="93">Deltochilum gilli</emphasis>
<bibRefCitation id="EFCC4B72DC5AFFB4A6D0A6D7FB4DFE59" author="Gonzalez-Alvarado A. &amp; Vaz-de-Mello F. Z." box="[650,1152,388,414]" pageId="7" pageNumber="93" pagination="0244657" refId="ref7969" refString="Gonzalez-Alvarado A. &amp; Vaz-de-Mello F. Z. 2021. Towards a comprehensive taxonomic revision of the Neotropical dung beetle subgenus Deltochilum (Deltohyboma) Lane, 1946 (Coleoptera: Scarabaeidae: Scarabaeinae): Division into species groups. PLOS ONE 16: e 0244657. https: // doi. org / 10.1371 / journal. pone. 0244657" type="journal article" year="2021">González-Alvarado &amp; Vaz-de-Mello, 2021</bibRefCitation>
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<subSubSection id="C3476508DC5AFFB4A5A0A6F4FBF4FE05" box="[506,1081,423,450]" pageId="7" pageNumber="93" type="description">
<paragraph id="8BE23683DC5AFFB4A5A0A6F4FBF4FE05" blockId="7.[436,1152,388,450]" box="[506,1081,423,450]" pageId="7" pageNumber="93">
<figureCitation id="13662A06DC5AFFB4A5A0A6F4FD97FE05" box="[506,602,423,450]" captionStart="Fig" captionStartId="3.[189,232,1892,1918]" captionTargetBox="[222,1362,786,1773]" captionTargetId="figure-203@3.[219,1368,777,1781]" captionTargetPageId="3" captionText="Fig. 1. Habitus of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589691" httpUri="https://zenodo.org/record/5589691/files/figure.png" pageId="7" pageNumber="93">Figs 1A</figureCitation>
,
<figureCitation id="13662A06DC5AFFB4A63CA6FBFD41FE05" box="[614,652,424,450]" captionStart="Fig" captionStartId="4.[189,232,1892,1918]" captionTargetBox="[237,1351,525,1826]" captionTargetId="figure-106@4.[237,1351,525,1826]" captionTargetPageId="4" captionText="Fig. 2. Head of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589695" httpUri="https://zenodo.org/record/5589695/files/figure.png" pageId="7" pageNumber="93">2A</figureCitation>
,
<figureCitation id="13662A06DC5AFFB4A6C3A6FBFD73FE05" box="[665,702,424,450]" captionStart="Fig" captionStartId="5.[189,232,1822,1848]" captionTargetBox="[217,1370,471,1774]" captionTargetId="figure-147@5.[217,1370,471,1774]" captionTargetPageId="5" captionText="Fig. 3. Pronotum of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744 E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Miscrosculpture of pronotal disc, Holotype of D. jocelynae sp. nov. G. Miscrosculpture pronotal disc, Holotype of D. nonstriatum sp. nov. White arrows show the shiny points." figureDoi="http://doi.org/10.5281/zenodo.5589697" httpUri="https://zenodo.org/record/5589697/files/figure.png" pageId="7" pageNumber="93">3A</figureCitation>
,
<figureCitation id="13662A06DC5AFFB4A691A6FBFD3CFE05" box="[715,753,424,450]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="7" pageNumber="93">4A</figureCitation>
,
<figureCitation id="13662A06DC5AFFB4A6A4A6F4FCE9FE05" box="[766,804,423,450]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="7" pageNumber="93">5A</figureCitation>
,
<figureCitation id="13662A06DC5AFFB4A76AA6F4FC9AFE06" box="[816,855,423,449]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[230,1363,815,1803]" captionTargetId="figure-152@8.[220,1368,814,1813]" captionTargetPageId="8" captionText="Fig. 6. Ventral view of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589707" httpUri="https://zenodo.org/record/5589707/files/figure.png" pageId="7" pageNumber="93">6A</figureCitation>
,
<figureCitation id="13662A06DC5AFFB4A739A6FBFC44FE05" box="[867,905,424,450]" captionStart="Fig" captionStartId="9.[189,232,1822,1848]" captionTargetBox="[206,1383,744,1716]" captionTargetId="figure-193@9.[203,1383,744,1716]" captionTargetPageId="9" captionText="Fig. 7.Aedeagus of members of thegilli species group.A–E. Lateral view.F–G. Dorsal view of paramera. Arrows show the apical-dorsal notches of the paramera. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov. CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745. F. Holotype of D. jocelynae sp. nov. G. Holotype of D. nonstriatum sp. nov., CMNEN WDS00041746." figureDoi="http://doi.org/10.5281/zenodo.5589711" httpUri="https://zenodo.org/record/5589711/files/figure.png" pageId="7" pageNumber="93">7A</figureCitation>
,
<figureCitation id="13662A06DC5AFFB4A7CFA6FBFC76FE05" box="[917,955,424,450]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[233,1362,650,1777]" captionTargetId="figure-181@10.[218,1370,644,1787]" captionTargetPageId="10" captionText="Fig. 8. Endophallus of members of thegilli species group.Arrows show the scales on the sub-medial area of the endophallus. A. Elongate scales. C. Regular scales. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589713" httpUri="https://zenodo.org/record/5589713/files/figure.png" pageId="7" pageNumber="93">8A</figureCitation>
,
<figureCitation id="13662A06DC5AFFB4A792A6FBFBDDFE05" box="[968,1040,424,450]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[268,1315,559,1863]" captionTargetId="figure-129@12.[268,1320,553,1873]" captionTargetPageId="12" captionText="Fig. 9. Habitus A, C and ventral view B, D of the only two females known of the gilli species group. A–B. Paratype of Deltochilum gilli González-Alvarado &amp; Vaz-de-Mello, 2021. C–D. Paratype D. jocelynae sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589715" httpUri="https://zenodo.org/record/5589715/files/figure.png" pageId="7" pageNumber="93">9A–B</figureCitation>
,
<figureCitation id="13662A06DC5AFFB4A046A6FBFBF4FE05" box="[1052,1081,424,450]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[336,1251,578,1870]" captionTargetId="figure-135@14.[336,1251,578,1870]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of thegilli species group. Blue circle =Deltochilum gilli González-Alvarado &amp; Vaz-de-Mello, 2021; yellow square = D. jocelynae sp. nov.; red star = D. nonstriatum sp. nov.; green triangle = D. quasistriatum sp. nov.; white eight-point star = D. tenuistriatum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="7" pageNumber="93">10</figureCitation>
</paragraph>
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<treatmentCitationGroup id="AB4D11ADDC5AFFB4A4E7A6BAFB83FDC4" box="[189,1102,489,516]" pageId="7" pageNumber="93">
<treatmentCitation id="0AFC1092DC5AFFB4A4E7A6BAFC7EFDC3" ID-DOI="http://doi.org/10.5281/zenodo.5590321" ID-Zenodo-Dep="5590321" author="Gonzalez-Alvarado A. &amp; Vaz-de-Mello F. Z." box="[189,947,489,516]" httpUri="http://treatment.plazi.org/id/03F48795DC4CFFA1A4E7A106FABAFA9A" page="50" pageId="7" pageNumber="93" year="2021">
<taxonomicName id="4C5D4D00DC5AFFB4A4E7A6BAFC7EFDC3" ID-CoL="7VN7Q" authority="Gonzalez-Alvarado &amp; Vaz-de-Mello, 2021: 50" authorityName="Gonzalez-Alvarado &amp; Vaz-de-Mello" authorityYear="2021" box="[189,947,489,516]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="93" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B929EA91DC5AFFB4A4E7A6BAFE4AFDC4" box="[189,391,489,515]" italics="true" pageId="7" pageNumber="93">Deltochilum gilli</emphasis>
<bibRefCitation id="EFCC4B72DC5AFFB4A5D4A6BAFC7EFDC3" author="Gonzalez-Alvarado A. &amp; Vaz-de-Mello F. Z." box="[398,947,489,516]" pageId="7" pageNumber="93" pagination="0244657" refId="ref7969" refString="Gonzalez-Alvarado A. &amp; Vaz-de-Mello F. Z. 2021. Towards a comprehensive taxonomic revision of the Neotropical dung beetle subgenus Deltochilum (Deltohyboma) Lane, 1946 (Coleoptera: Scarabaeidae: Scarabaeinae): Division into species groups. PLOS ONE 16: e 0244657. https: // doi. org / 10.1371 / journal. pone. 0244657" type="journal article" year="2021">González-Alvarado &amp; Vaz-de-Mello, 2021: 50</bibRefCitation>
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, figs 26c, 28.
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<subSubSection id="C3476508DC5AFFBBA4E7A579FD26FE81" lastPageId="8" lastPageNumber="94" pageId="7" pageNumber="93" type="diagnosis">
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<emphasis id="B929EA91DC5AFFB4A4E7A579FEFBFD83" bold="true" box="[189,310,554,580]" pageId="7" pageNumber="93">Diagnosis</emphasis>
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<paragraph id="8BE23683DC5AFFBBA4E7A50AFD26FE81" blockId="7.[189,1399,601,663]" lastBlockId="8.[189,1398,265,327]" lastPageId="8" lastPageNumber="94" pageId="7" pageNumber="93">
Elytra with apical tubercles on interstriae III–VII (
<figureCitation id="13662A06DC5AFFB4A77FA50AFC4AFDB3" box="[805,903,601,628]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[201,1386,717,1836]" captionTargetId="figure-170@7.[201,1386,717,1836]" captionTargetPageId="7" captionText="Fig. 5. Caudal view of mmbers of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="7" pageNumber="93">Fig. 5A</figureCitation>
), first stria almost twice as wide as than second stria, striae III–VII subequal in width and conspicuous (
<figureCitation id="13662A06DC5AFFB4A7E3A52FFBD0FD51" box="[953,1053,636,662]" captionStart="Fig" captionStartId="3.[189,232,1892,1918]" captionTargetBox="[222,1362,786,1773]" captionTargetId="figure-203@3.[219,1368,777,1781]" captionTargetPageId="3" captionText="Fig. 1. Habitus of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589691" httpUri="https://zenodo.org/record/5589691/files/figure.png" pageId="7" pageNumber="93">Figs 1A</figureCitation>
,
<figureCitation id="13662A06DC5AFFB4A076A52FFB9CFD51" box="[1068,1105,636,662]" captionStart="Fig" captionStartId="6.[189,232,1892,1918]" captionTargetBox="[219,1369,314,1798]" captionTargetId="figure-77@6.[219,1369,314,1798]" captionTargetPageId="6" captionText="Fig. 4. Elytra of members of the gilli species group. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype of D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589701" httpUri="https://zenodo.org/record/5589701/files/figure.png" pageId="7" pageNumber="93">4A</figureCitation>
) and sub-medial area of endophallus with elongate scales (
<figureCitation id="13662A06DC55FFBBA60FA65AFD79FEE4" box="[597,692,265,291]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[233,1362,650,1777]" captionTargetId="figure-181@10.[218,1370,644,1787]" captionTargetPageId="10" captionText="Fig. 8. Endophallus of members of thegilli species group.Arrows show the scales on the sub-medial area of the endophallus. A. Elongate scales. C. Regular scales. A. Holotype of Deltochilum gilli González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888. B. Holotype of D. jocelynae sp. nov., CNCI 379889. C. Holotype of D. nonstriatum sp. nov., CMNEN WSD00041746. D. Holotype of D. quasistriatum sp. nov., CMNEN WSD00041744. E. Holotype D. tenuistriatum sp. nov., CMNEN WSD00041745." figureDoi="http://doi.org/10.5281/zenodo.5589713" httpUri="https://zenodo.org/record/5589713/files/figure.png" pageId="8" pageNumber="94">Fig. 8A</figureCitation>
, arrow) distinguish this species (for a complete description see
<bibRefCitation id="EFCC4B72DC55FFBBA4B1A67FFD13FE81" author="Gonzalez-Alvarado A. &amp; Vaz-de-Mello F. Z." box="[235,734,300,326]" pageId="8" pageNumber="94" pagination="0244657" refId="ref7969" refString="Gonzalez-Alvarado A. &amp; Vaz-de-Mello F. Z. 2021. Towards a comprehensive taxonomic revision of the Neotropical dung beetle subgenus Deltochilum (Deltohyboma) Lane, 1946 (Coleoptera: Scarabaeidae: Scarabaeinae): Division into species groups. PLOS ONE 16: e 0244657. https: // doi. org / 10.1371 / journal. pone. 0244657" type="journal article" year="2021">González-Alvarado &amp; Vaz-de-Mello 2021</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF22660BDC5AFFB4A4E7A037FAB8F82E" ID-DOI="http://doi.org/10.5281/zenodo.5589705" ID-Zenodo-Dep="5589705" httpUri="https://zenodo.org/record/5589705/files/figure.png" pageId="7" pageNumber="93" startId="7.[189,232,1892,1918]" targetBox="[201,1386,717,1836]" targetPageId="7">
<paragraph id="8BE23683DC5AFFB4A4E7A037FAB8F82E" blockId="7.[189,1399,1892,2025]" pageId="7" pageNumber="93">
<emphasis id="B929EA91DC5AFFB4A4E7A037FEC1F8B8" bold="true" box="[189,268,1892,1919]" pageId="7" pageNumber="93">Fig. 5.</emphasis>
Caudal view of mmbers of the
<taxonomicName id="4C5D4D00DC5AFFB4A6DDA036FD7AF8B8" authorityName="Gonzalez-Alvarado &amp; Vaz-de-Mello" authorityYear="2021" box="[647,695,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="93" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B929EA91DC5AFFB4A6DDA036FD7AF8B8" box="[647,695,1893,1919]" italics="true" pageId="7" pageNumber="93">gilli</emphasis>
</taxonomicName>
species group.
<emphasis id="B929EA91DC5AFFB4A72AA037FC49F8B9" bold="true" box="[880,900,1892,1918]" pageId="7" pageNumber="93">A</emphasis>
. Holotype of
<taxonomicName id="4C5D4D00DC5AFFB4A071A036FB38F8B8" authorityName="Gonzalez-Alvarado &amp; Vaz-de-Mello" authorityYear="2021" box="[1067,1269,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="93" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B929EA91DC5AFFB4A071A036FB38F8B8" box="[1067,1269,1893,1919]" italics="true" pageId="7" pageNumber="93">Deltochilum gilli</emphasis>
</taxonomicName>
González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888.
<emphasis id="B929EA91DC5AFFB4A75FA0DBFCD5F865" bold="true" box="[773,792,1928,1954]" pageId="7" pageNumber="93">B</emphasis>
. Holotype of
<taxonomicName id="4C5D4D00DC5AFFB4A7E5A0DAFB9BF865" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[959,1110,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="93" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
<emphasis id="B929EA91DC5AFFB4A7E5A0DAFB9BF865" box="[959,1110,1928,1954]" italics="true" pageId="7" pageNumber="93">D. jocelynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC5AFFB4A007A0DBFB71F865" box="[1117,1212,1928,1954]" pageId="7" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
, CNCI 379889.
<emphasis id="B929EA91DC5AFFB4A4E4A0F8FF1FF802" bold="true" box="[190,210,1963,1989]" pageId="7" pageNumber="93">C</emphasis>
. Holotype of
<taxonomicName id="4C5D4D00DC5AFFB4A524A0FFFDF8F802" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[382,565,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="93" phylum="Arthropoda" rank="species" species="nonstriatum" status="sp. nov.">
<emphasis id="B929EA91DC5AFFB4A524A0FFFE5EF802" box="[382,403,1964,1989]" italics="true" pageId="7" pageNumber="93">D</emphasis>
.
<emphasis id="B929EA91DC5AFFB4A5FCA0FFFDF8F802" box="[422,565,1964,1989]" italics="true" pageId="7" pageNumber="93">nonstriatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC5AFFB4A61AA0FFFD6DF801" box="[576,672,1964,1990]" pageId="7" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
, CMNEN WSD00041746.
<emphasis id="B929EA91DC5AFFB4A7B1A0F8FBCDF802" bold="true" box="[1003,1024,1963,1989]" pageId="7" pageNumber="93">D</emphasis>
. Holotype of
<taxonomicName id="4C5D4D00DC5AFFB4A0F6A0FFFABBF802" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[1196,1398,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="93" phylum="Arthropoda" rank="species" species="quasistriatum" status="sp. nov.">
<emphasis id="B929EA91DC5AFFB4A0F6A0FFFB0CF802" box="[1196,1217,1964,1989]" italics="true" pageId="7" pageNumber="93">D</emphasis>
.
<emphasis id="B929EA91DC5AFFB4A089A0FFFABBF802" box="[1235,1398,1964,1989]" italics="true" pageId="7" pageNumber="93">quasistriatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC5AFFB4A4E7A09CFED7F82E" box="[189,282,1999,2025]" pageId="7" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
, CMNEN WSD00041744.
<emphasis id="B929EA91DC5AFFB4A606A09DFDBDF82F" bold="true" box="[604,624,1998,2024]" pageId="7" pageNumber="93">E</emphasis>
. Holotype of
<taxonomicName id="4C5D4D00DC5AFFB4A748A09CFC18F82F" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[786,981,1999,2025]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="93" phylum="Arthropoda" rank="species" species="tenuistriatum" status="sp. nov.">
<emphasis id="B929EA91DC5AFFB4A748A09CFCEAF82F" box="[786,807,1999,2024]" italics="true" pageId="7" pageNumber="93">D</emphasis>
.
<emphasis id="B929EA91DC5AFFB4A76DA09CFC18F82F" box="[823,981,1999,2024]" italics="true" pageId="7" pageNumber="93">tenuistriatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC5AFFB4A787A09CFBF7F82E" box="[989,1082,1999,2025]" pageId="7" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
, CMNEN WSD00041745.
</paragraph>
</caption>
<subSubSection id="C3476508DC55FFBBA4E7A63EFC65FD43" pageId="8" pageNumber="94" type="materials_examined">
<paragraph id="8BE23683DC55FFBBA4E7A63EFE67FE40" blockId="8.[189,426,365,391]" box="[189,426,365,391]" pageId="8" pageNumber="94">
<heading id="D0AA81EFDC55FFBBA4E7A63EFE67FE40" bold="true" box="[189,426,365,391]" fontSize="11" level="3" pageId="8" pageNumber="94" reason="3">
<emphasis id="B929EA91DC55FFBBA4E7A63EFE67FE40" bold="true" box="[189,426,365,391]" pageId="8" pageNumber="94">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BE23683DC55FFBBA4B0A6CFFDB7FDE7" blockId="8.[189,1399,412,544]" pageId="8" pageNumber="94">
<materialsCitation id="3B353CDEDC55FFBBA4B0A6CFFDB7FDE7" ID-GBIF-Occurrence="3391047308" collectedFrom="primary forest" collectingDate="2004-10-27" collectingDateMax="2004-11-01" collectingDateMin="2004-10-27" collectingMethod="pitfall trap" collectionCode="BDGC" collectorName="B. Hubley" country="Guyana" elevation="1234" latitude="5.1096663" location="District 8" longLatPrecision="1" longitude="-59.820915" municipality="Mount Wokomung" pageId="8" pageNumber="94" specimenCode="CNC 379888" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B929EA91DC55FFBBA4B0A6CFFE91FE71" bold="true" box="[234,348,412,438]" pageId="8" pageNumber="94">
<typeStatus id="54E68821DC55FFBBA4B0A6CFFE91FE71" box="[234,348,412,438]" pageId="8" pageNumber="94" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F34A7613DC55FFBBA4E7A6ECFEF4FE1E" box="[189,313,447,473]" name="Guyana" pageId="8" pageNumber="94">GUYANA</collectingCountry>
•
<specimenCount id="9D5BFD0ADC55FFBBA517A6ECFEBAFE1B" box="[333,375,447,476]" pageId="8" pageNumber="94" type="male">
1
<emphasis id="B929EA91DC55FFBBA53BA693FEBAFE1B" bold="true" box="[353,375,448,476]" pageId="8" pageNumber="94">♂</emphasis>
</specimenCount>
;
<location id="8E826058DC55FFBBA5DEA6ECFE3CFE1E" LSID="urn:lsid:plazi:treatment:03F48795DC5AFFBAA5EEA6D7FC4AFDEB:8E826058DC55FFBBA5DEA6ECFE3CFE1E" box="[388,497,447,473]" country="Guyana" latitude="5.1096663" longLatPrecision="1" longitude="-59.820915" municipality="Mount Wokomung" name="District 8" pageId="8" pageNumber="94">District 8</location>
,
<collectingMunicipality id="6B86ACF9DC55FFBBA5A6A693FD17FE1E" box="[508,730,447,474]" pageId="8" pageNumber="94">Mount Wokomung</collectingMunicipality>
;
<geoCoordinate id="EE695044DC55FFBBA6BEA6EDFCB1FE1D" box="[740,892,446,474]" degrees="5" direction="north" minutes="06" orientation="latitude" pageId="8" pageNumber="94" precision="1" seconds="34.8" value="5.1096663">5º06′34.8″ N</geoCoordinate>
,
<geoCoordinate id="EE695044DC55FFBBA7DFA6EDFBFCFE1D" box="[901,1073,446,474]" degrees="59" direction="west" minutes="49" orientation="longitude" pageId="8" pageNumber="94" precision="1" seconds="15.3" value="-59.820915">59º49′15.3″ W</geoCoordinate>
; alt.
<quantity id="4CA59B66DC55FFBBA03FA6ECFB73FE1D" box="[1125,1214,447,474]" metricMagnitude="3" metricUnit="m" metricValue="1.234" pageId="8" pageNumber="94" unit="m" value="1234.0">
<elevation id="0070D1B0DC55FFBBA03FA6ECFB73FE1D" box="[1125,1214,447,474]" metricMagnitude="3" metricUnit="m" metricValue="1.234" pageId="8" pageNumber="94" unit="m" value="1234.0">1234 m</elevation>
</quantity>
;
<date id="FFE31043DC55FFBBA092A6ECFF36FE3A" pageId="8" pageNumber="94" value="2004-10-27" valueMax="2004-11-01" valueMin="2004-10-27">
<collectingDate id="EFA7E9ABDC55FFBBA092A6ECFF36FE3A" pageId="8" pageNumber="94" value="2004-10-27" valueMax="2004-11-01" valueMin="2004-10-27">27 Oct.–1 Nov. 2004</collectingDate>
</date>
;
<collectorName id="26A85355DC55FFBBA55DA6B0FEB2FE3B" box="[263,383,482,509]" pageId="8" pageNumber="94">B. Hubley</collectorName>
leg.;
<collectingMethod id="521C4E94DC55FFBBA598A6B1FDF0FE3A" box="[450,573,482,509]" pageId="8" pageNumber="94">pitfall trap</collectingMethod>
(human dung),
<collectedFrom id="3841757EDC55FFBBA6A2A6B1FC69FE3B" box="[760,932,482,508]" pageId="8" pageNumber="94">primary forest</collectedFrom>
; ROM 2004516;
<collectionCode id="ED4CAE46DC55FFBBA035A6B0FB09FE3A" box="[1135,1220,483,509]" pageId="8" pageNumber="94">BDGC</collectionCode>
. [aedeagus and endophallus extracted];
<specimenCode id="DBFB9EF8DC55FFBBA58DA555FDBAFDE7" box="[471,631,518,544]" pageId="8" pageNumber="94">CNC 379888</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE23683DC55FFBBA4B0A514FC65FD43" blockId="8.[189,936,583,647]" pageId="8" pageNumber="94">
<materialsCitation id="3B353CDEDC55FFBBA4B0A514FC69FD43" ID-GBIF-Occurrence="3391047305" collectionCode="BDGC" country="Guyana" location="GUYANA" pageId="8" pageNumber="94" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B929EA91DC55FFBBA4B0A514FE91FDA6" bold="true" box="[234,348,583,609]" pageId="8" pageNumber="94">
<typeStatus id="54E68821DC55FFBBA4B0A514FE91FDA6" box="[234,348,583,609]" pageId="8" pageNumber="94" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="F34A7613DC55FFBBA4E7A539FEFAFD43" box="[189,311,618,644]" name="Guyana" pageId="8" pageNumber="94">GUYANA</collectingCountry>
•
<specimenCount id="9D5BFD0ADC55FFBBA50AA539FEB1FD40" box="[336,380,618,647]" pageId="8" pageNumber="94" type="female">
1
<emphasis id="B929EA91DC55FFBBA53CA538FEB1FD40" bold="true" box="[358,380,619,647]" pageId="8" pageNumber="94">♀</emphasis>
</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED4CAE46DC55FFBBA715A539FC69FD43" box="[847,932,618,644]" pageId="8" pageNumber="94">BDGC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF22660BDC55FFBBA4E7A037FAB8F82E" ID-DOI="http://doi.org/10.5281/zenodo.5589707" ID-Zenodo-Dep="5589707" httpUri="https://zenodo.org/record/5589707/files/figure.png" pageId="8" pageNumber="94" startId="8.[189,232,1892,1918]" targetBox="[230,1363,815,1803]" targetPageId="8">
<paragraph id="8BE23683DC55FFBBA4E7A037FAB8F82E" blockId="8.[189,1399,1892,2025]" pageId="8" pageNumber="94">
<emphasis id="B929EA91DC55FFBBA4E7A037FEC6F8B8" bold="true" box="[189,267,1892,1919]" pageId="8" pageNumber="94">Fig. 6.</emphasis>
Ventral view of members of the
<taxonomicName id="4C5D4D00DC55FFBBA6D7A036FD70F8B8" authorityName="Gonzalez-Alvarado &amp; Vaz-de-Mello" authorityYear="2021" box="[653,701,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="94" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B929EA91DC55FFBBA6D7A036FD70F8B8" box="[653,701,1893,1919]" italics="true" pageId="8" pageNumber="94">gilli</emphasis>
</taxonomicName>
species group.
<emphasis id="B929EA91DC55FFBBA72EA037FC44F8B9" bold="true" box="[884,905,1892,1918]" pageId="8" pageNumber="94">A</emphasis>
. Holotype of
<taxonomicName id="4C5D4D00DC55FFBBA076A036FB3BF8B8" authorityName="Gonzalez-Alvarado &amp; Vaz-de-Mello" authorityYear="2021" box="[1068,1270,1893,1919]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="94" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B929EA91DC55FFBBA076A036FB3BF8B8" box="[1068,1270,1893,1919]" italics="true" pageId="8" pageNumber="94">Deltochilum gilli</emphasis>
</taxonomicName>
González- Alvarado &amp; Vaz-de-Mello, 2021, CNCI 379888.
<emphasis id="B929EA91DC55FFBBA75FA0DBFCD5F865" bold="true" box="[773,792,1928,1954]" pageId="8" pageNumber="94">B</emphasis>
. Holotype of
<taxonomicName id="4C5D4D00DC55FFBBA7E5A0DAFB98F865" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[959,1109,1928,1954]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="94" phylum="Arthropoda" rank="species" species="jocelynae" status="sp. nov.">
<emphasis id="B929EA91DC55FFBBA7E5A0DAFB98F865" box="[959,1109,1928,1954]" italics="true" pageId="8" pageNumber="94">D. jocelynae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC55FFBBA007A0DBFB71F865" box="[1117,1212,1928,1954]" pageId="8" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
, CNCI 379889.
<emphasis id="B929EA91DC55FFBBA4E4A0F8FF1FF802" bold="true" box="[190,210,1963,1989]" pageId="8" pageNumber="94">C</emphasis>
. Holotype of
<taxonomicName id="4C5D4D00DC55FFBBA524A0FFFDF8F802" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[382,565,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="94" phylum="Arthropoda" rank="species" species="nonstriatum" status="sp. nov.">
<emphasis id="B929EA91DC55FFBBA524A0FFFE5EF802" box="[382,403,1964,1989]" italics="true" pageId="8" pageNumber="94">D</emphasis>
.
<emphasis id="B929EA91DC55FFBBA5FCA0FFFDF8F802" box="[422,565,1964,1989]" italics="true" pageId="8" pageNumber="94">nonstriatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC55FFBBA61AA0FFFD6DF801" box="[576,672,1964,1990]" pageId="8" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
, CMNEN WSD00041746.
<emphasis id="B929EA91DC55FFBBA7B1A0F8FBCDF802" bold="true" box="[1003,1024,1963,1989]" pageId="8" pageNumber="94">D</emphasis>
. Holotype of
<taxonomicName id="4C5D4D00DC55FFBBA0F6A0FFFABBF802" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[1196,1398,1964,1990]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="94" phylum="Arthropoda" rank="species" species="quasistriatum" status="sp. nov.">
<emphasis id="B929EA91DC55FFBBA0F6A0FFFB0CF802" box="[1196,1217,1964,1989]" italics="true" pageId="8" pageNumber="94">D</emphasis>
.
<emphasis id="B929EA91DC55FFBBA089A0FFFABBF802" box="[1235,1398,1964,1989]" italics="true" pageId="8" pageNumber="94">quasistriatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC55FFBBA4E7A09CFED7F82E" box="[189,282,1999,2025]" pageId="8" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
, CMNEN WSD00041744.
<emphasis id="B929EA91DC55FFBBA606A09DFDBDF82F" bold="true" box="[604,624,1998,2024]" pageId="8" pageNumber="94">E</emphasis>
. Holotype of
<taxonomicName id="4C5D4D00DC55FFBBA748A09CFC18F82F" authority="González-Alvarado &amp; Vaz-De, 2021" authorityName="González-Alvarado &amp; Vaz-De" authorityYear="2021" box="[786,981,1999,2025]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="94" phylum="Arthropoda" rank="species" species="tenuistriatum" status="sp. nov.">
<emphasis id="B929EA91DC55FFBBA748A09CFCEAF82F" box="[786,807,1999,2024]" italics="true" pageId="8" pageNumber="94">D</emphasis>
.
<emphasis id="B929EA91DC55FFBBA76DA09CFC18F82F" box="[823,981,1999,2024]" italics="true" pageId="8" pageNumber="94">tenuistriatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A57EADC55FFBBA787A09CFBF7F82E" box="[989,1082,1999,2025]" pageId="8" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
, CMNEN WSD00041745.
</paragraph>
</caption>
<subSubSection id="C3476508DC54FFBAA4E7A65AFD19FE7C" pageId="9" pageNumber="95" type="discussion">
<paragraph id="8BE23683DC54FFBAA4E7A65AFEE2FEE4" blockId="9.[189,303,265,291]" box="[189,303,265,291]" pageId="9" pageNumber="95">
<heading id="D0AA81EFDC54FFBAA4E7A65AFEE2FEE4" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="9" pageNumber="95" reason="3">
<emphasis id="B929EA91DC54FFBAA4E7A65AFEE2FEE4" bold="true" box="[189,303,265,291]" pageId="9" pageNumber="95">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BE23683DC54FFBAA4E7A664FD19FE7C" blockId="9.[189,1399,311,443]" pageId="9" pageNumber="95">
Within
<emphasis id="B929EA91DC54FFBAA542A66BFEE0FE96" box="[280,301,312,337]" italics="true" pageId="9" pageNumber="95">D</emphasis>
. (
<taxonomicName id="4C5D4D00DC54FFBAA510A664FE27FE96" authorityName="Lane" authorityYear="1946" box="[330,490,311,337]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="104" phylum="Arthropoda" rank="subGenus" subGenus="Deltohyboma">
<emphasis id="B929EA91DC54FFBAA510A664FE27FE96" box="[330,490,311,337]" italics="true" pageId="9" pageNumber="95">Deltohyboma</emphasis>
</taxonomicName>
) only few species which possess the anterior margin of clypeus expanded posteriorly into a triangular shape have more than four elytral apical tubercles. Frequently, in species bearing over four apical tubercles, the anterior margin of the clypeus is not expanded into a triangular shape.
<taxonomicName id="4C5D4D00DC54FFBAA554A6F2FEAFFE7C" authorityName="Gonzalez-Alvarado &amp; Vaz-de-Mello" authorityYear="2021" box="[270,354,417,443]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="95" phylum="Arthropoda" rank="species" species="gilli">
<emphasis id="B929EA91DC54FFBAA554A6F2FEEEFE7D" box="[270,291,417,442]" italics="true" pageId="9" pageNumber="95">D</emphasis>
.
<emphasis id="B929EA91DC54FFBAA568A6F2FEAFFE7C" box="[306,354,417,443]" italics="true" pageId="9" pageNumber="95">gilli</emphasis>
</taxonomicName>
is one of those few exceptions.
</paragraph>
</subSubSection>
<subSubSection id="C3476508DC54FFBAA4E7A6B1FC4AFDEB" pageId="9" pageNumber="95" type="distribution">
<paragraph id="8BE23683DC54FFBAA4E7A6B1FE79FE3B" blockId="9.[189,436,482,508]" box="[189,436,482,508]" pageId="9" pageNumber="95">
<heading id="D0AA81EFDC54FFBAA4E7A6B1FE79FE3B" bold="true" box="[189,436,482,508]" fontSize="11" level="3" pageId="9" pageNumber="95" reason="3">
<emphasis id="B929EA91DC54FFBAA4E7A6B1FE79FE3B" bold="true" box="[189,436,482,508]" pageId="9" pageNumber="95">Known distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BE23683DC54FFBAA4E7A542FC4AFDEB" blockId="9.[189,903,529,556]" box="[189,903,529,556]" pageId="9" pageNumber="95">
<collectingCountry id="F34A7613DC54FFBAA4E7A542FED1FDEC" box="[189,284,529,555]" name="Guyana" pageId="9" pageNumber="95">Guyana</collectingCountry>
. District 8, Mount Wokomung. (
<figureCitation id="13662A06DC54FFBAA6C2A542FD22FDEC" box="[664,751,529,555]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[336,1251,578,1870]" captionTargetId="figure-135@14.[336,1251,578,1870]" captionTargetPageId="14" captionText="Fig. 10. Distribution map of thegilli species group. Blue circle =Deltochilum gilli González-Alvarado &amp; Vaz-de-Mello, 2021; yellow square = D. jocelynae sp. nov.; red star = D. nonstriatum sp. nov.; green triangle = D. quasistriatum sp. nov.; white eight-point star = D. tenuistriatum sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5589717" httpUri="https://zenodo.org/record/5589717/files/figure.png" pageId="9" pageNumber="95">Fig. 10</figureCitation>
, blue circle).
</paragraph>
</subSubSection>
</treatment>
</document>
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