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<document id="5BFE059CACD3E786060EDFC3507ACA0F" ID-DOI="10.11646/zootaxa.3861.5.1" ID-GBIF-Dataset="0b368704-d9cc-4a82-8248-bc4c06dcf781" ID-ISSN="1175-5326" ID-Zenodo-Dep="252462" ID-ZooBank="E62DB6C3-0C5F-4898-99C4-1BEC70DD1734" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460912037909" checkinUser="plazi" docAuthor="Kaim, Andrzej, Jenkins, Robert G., Tanabe, Kazushige &amp; Kiel, Steffen" docDate="2014" docId="03F4372EFFFF6855FF340611FC18FEAC" docLanguage="en" docName="zt03861p440.pdf" docOrigin="Zootaxa 3861 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lyriochlamys complexicostata Gabb 1869" docType="treatment" docVersion="10" lastPageNumber="427" masterDocId="FFCD4F56FFE6684EFFA30151FFDCFFC8" masterDocTitle="Mollusks from late Mesozoic seep deposits, chiefly in California" masterLastPageNumber="440" masterPageNumber="401" pageNumber="426" updateTime="1698361398619" updateUser="plazi">
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<mods:title id="AE0D6AF5A0ED00A9568BF01DA0600F9A">Mollusks from late Mesozoic seep deposits, chiefly in California</mods:title>
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<mods:namePart id="F81ECBCC2712B4880FEBFF2BE3AFA8C4">Kaim, Andrzej</mods:namePart>
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<mods:namePart id="3D2ABEEF93BEB8AC918039ACE10EAB45">Jenkins, Robert G.</mods:namePart>
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<mods:namePart id="5B2C77972BEE220050BF1F3C30821840">Tanabe, Kazushige</mods:namePart>
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<mods:namePart id="E8DFCE48E4C441F3E966C7B463FE3467">Kiel, Steffen</mods:namePart>
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<mods:date id="BA1BDE2EE4A15330976A98DC91EE62C1">2014</mods:date>
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<treatment id="03F4372EFFFF6855FF340611FC18FEAC" ID-DOI="http://doi.org/10.5281/zenodo.6127285" ID-GBIF-Taxon="119588565" ID-Zenodo-Dep="6127285" LSID="urn:lsid:plazi:treatment:03F4372EFFFF6855FF340611FC18FEAC" httpUri="http://treatment.plazi.org/id/03F4372EFFFF6855FF340611FC18FEAC" lastPageId="27" lastPageNumber="427" pageId="25" pageNumber="426">
<subSubSection id="C347D5B3FFFF6857FF340611FF25F8B3" pageId="25" pageNumber="426" type="nomenclature">
<paragraph id="8BE28638FFFF6857FF340611FD70F892" blockId="25.[151,684,1856,1916]" box="[151,684,1856,1882]" pageId="25" pageNumber="426">
<heading id="D0AA3154FFFF6857FF340611FD70F892" bold="true" box="[151,684,1856,1882]" fontSize="11" level="1" pageId="25" pageNumber="426" reason="1">
<taxonomicName id="4C5DFDBBFFFF6857FF340611FD70F892" authority="Gabb, 1869" authorityName="Gabb" authorityYear="1869" box="[151,684,1856,1882]" class="Bivalvia" family="Pectinidae" genus="Lyriochlamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreoida" pageId="25" pageNumber="426" phylum="Mollusca" rank="species" species="complexicostata">
<emphasis id="B9295A2AFFFF6857FF340611FD70F892" bold="true" box="[151,684,1856,1882]" pageId="25" pageNumber="426">
<emphasis id="B9295A2AFFFF6857FF340611FDDEF892" bold="true" box="[151,514,1856,1882]" italics="true" pageId="25" pageNumber="426">Lyriochlamys complexicostata</emphasis>
(
<bibRefCitation id="EFCCFBC9FFFF6857FDB10611FD78F892" author="Gabb" box="[530,676,1856,1882]" pageId="25" pageNumber="426" refString="Gabb, W. M. (1869) Cretaceous and Tertiary fossils. California Geological Survey, Paleontology, 2, 1 - 299." type="journal article" year="1869">Gabb, 1869</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BE28638FFFF6857FF340632FF25F8B3" blockId="25.[151,684,1856,1916]" box="[151,249,1891,1916]" pageId="25" pageNumber="426">
(
<figureCitation id="13669ABDFFFF6857FF3C0632FF2DF8B3" box="[159,241,1891,1916]" captionStart="FIGURE 14" captionStartId="27.[151,250,1796,1818]" captionTargetBox="[151,1436,411,1774]" captionTargetId="figure@27.[151,1436,411,1775]" captionTargetPageId="27" captionText="FIGURE 14. Lyriochlamys complexicostata (Gabb, 1869) from Cretaceous seep carbonates in California, showing various stages of defoliation; all specimens from Hauterivian of Wilbur Springs except for that in fig B, which are from Rocky Creek (probably Valanginian). A. Small specimen with fine radial sculpture (USNM 597562). B. Large specimen in whom the main ribs appear sharp, note fragments of the same species on the far and lower left (GZG. INV. 85002). C. Specimen with almost no shell preserved (USNM 597563). D. Specimen with fine radial ribs and rough commarginal wrinkles on the ears (USNM 597564). E. specimen in which the main ribs appear blunt (USNM 605206). F. Specimen with blunt vertical ribs on the ear; note abundant fragments of the brachiopod Peregrinella whiteyi at the lower side (USNM 597565)." httpUri="https://zenodo.org/record/252476/files/figure.png" pageId="25" pageNumber="426">Fig. 14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C347D5B3FFFF6857FF3406FDFCB2F817" pageId="25" pageNumber="426" type="reference_group">
<paragraph id="8BE28638FFFF6857FF3406FDFD37F809" blockId="25.[151,878,1963,2015]" box="[151,747,1963,1985]" pageId="25" pageNumber="426">
<taxonomicName id="4C5DFDBBFFFF6857FF3406FDFDD9F809" authority="Gabb, 1869" authorityName="Gabb" authorityYear="1869" box="[151,517,1963,1985]" class="Bivalvia" family="Pectinidae" genus="Pecten" kingdom="Animalia" order="Pectinoida" pageId="25" pageNumber="426" phylum="Mollusca" rank="species" species="complexicostata">
<emphasis id="B9295A2AFFFF6857FF3406FDFE5BF809" box="[151,391,1963,1985]" italics="true" pageId="25" pageNumber="426">Pecten complexicostata</emphasis>
<bibRefCitation id="EFCCFBC9FFFF6857FE2E06FAFDD9F809" author="Gabb" box="[397,517,1963,1985]" pageId="25" pageNumber="426" refString="Gabb, W. M. (1869) Cretaceous and Tertiary fossils. California Geological Survey, Paleontology, 2, 1 - 299." type="journal article" year="1869">Gabb, 1869</bibRefCitation>
</taxonomicName>
, p. 199, pl. 33, fig. 97.
</paragraph>
<paragraph id="8BE28638FFFF6857FF34069BFCB2F817" blockId="25.[151,878,1963,2015]" box="[151,878,1993,2015]" pageId="25" pageNumber="426">
<taxonomicName id="4C5DFDBBFFFF6857FF34069BFDB9F817" authority="Gabb-Stanton 1895" authorityName="Gabb-Stanton" authorityYear="1895" box="[151,613,1993,2015]" class="Bivalvia" family="Pectinidae" genus="Pecten" kingdom="Animalia" order="Pectinoida" pageId="25" pageNumber="426" phylum="Mollusca" rank="species" species="complexicostata">
<emphasis id="B9295A2AFFFF6857FF34069BFE5BF817" box="[151,391,1993,2015]" italics="true" pageId="25" pageNumber="426">Pecten complexicostata</emphasis>
Gabb—Stanton 1895
</taxonomicName>
, p. 3738, pl. 2, figs. 79.
</paragraph>
</subSubSection>
<caption id="DF22D6B0FFFC6854FF340777FE89F9B1" httpUri="https://zenodo.org/record/252475/files/figure.png" pageId="26" pageNumber="427" targetBox="[340,1247,195,1552]" targetPageId="26">
<paragraph id="8BE28638FFFC6854FF340777FE89F9B1" blockId="26.[151,1435,1574,1657]" pageId="26" pageNumber="427">
<emphasis id="B9295A2AFFFC6854FF340777FEC3F9F4" bold="true" box="[151,287,1574,1596]" pageId="26" pageNumber="427">FIGURE 13.</emphasis>
<taxonomicName id="4C5DFDBBFFFC6854FE850777FD90F9F4" authority="Stanton, 1895" authorityName="Stanton" authorityYear="1895" box="[294,588,1574,1596]" class="Bivalvia" family="Nuculidae" genus="Nucula" kingdom="Animalia" order="Nuculida" pageId="26" pageNumber="427" phylum="Mollusca" rank="species" species="storrsi">
<emphasis id="B9295A2AFFFC6854FE850777FE64F9F4" box="[294,440,1574,1596]" italics="true" pageId="26" pageNumber="427">Nucula storrsi</emphasis>
Stanton, 1895
</taxonomicName>
from Tithonian seep carbonates of Paskenta.
<emphasis id="B9295A2AFFFC6854FBB80777FB8DF9F4" bold="true" box="[1051,1105,1574,1596]" pageId="26" pageNumber="427">A, B.</emphasis>
Specimen showing shell outline and smooth posterior part (
<emphasis id="B9295A2AFFFC6854FE080715FE61F992" bold="true" box="[427,445,1604,1626]" pageId="26" pageNumber="427">A</emphasis>
) and the taxodont hinge dentition (
<emphasis id="B9295A2AFFFC6854FC870715FCE9F992" bold="true" box="[804,821,1604,1626]" pageId="26" pageNumber="427">B</emphasis>
) (GZG.INV.85000).
<emphasis id="B9295A2AFFFC6854FBAA0715FBFEF992" bold="true" box="[1033,1058,1604,1626]" pageId="26" pageNumber="427">C.</emphasis>
Interior view on an articulated hinge (GZG.INV.85001).
</paragraph>
</caption>
<subSubSection id="C347D5B3FFFC6854FF3407F1FC8EF881" pageId="26" pageNumber="427" type="materials_examined">
<paragraph id="8BE28638FFFC6854FF3407F1FC8EF881" blockId="26.[151,1437,1696,2009]" pageId="26" pageNumber="427">
<emphasis id="B9295A2AFFFC6854FF3407F1FE14F971" bold="true" box="[151,456,1696,1721]" pageId="26" pageNumber="427">Material and occurrence.</emphasis>
Six specimens from Rocky Creek (probably Valanginian, Early Cretaceous), up to
<quantity id="4CA52BDDFFFC6854FA2307F1FF1FF915" metricMagnitude="-2" metricUnit="m" metricValue="6.8" pageId="26" pageNumber="427" unit="mm" value="68.0">68 mm</quantity>
in height. Stantons (1895) figured specimens (
<collectionCode id="ED4C1EFDFFFC6854FD420794FCE8F915" LSID="urn:lsid:biocol.org:col:34871" box="[737,820,1733,1757]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="427">USNM</collectionCode>
23028 and 23102) were not found in the
<collectionCode id="ED4C1EFDFFFC6854FAB00794FAB8F915" LSID="urn:lsid:biocol.org:col:34871" box="[1299,1380,1733,1757]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="26" pageNumber="427">USNM</collectionCode>
<typeStatus id="54E6389AFFFC6854FACF0797FA40F916" box="[1388,1436,1734,1758]" pageId="26" pageNumber="427">type</typeStatus>
collection during a visit in
<date id="FFE3A0F8FFFC6854FE7B07B9FD48F8C9" box="[472,660,1768,1793]" pageId="26" pageNumber="427" value="2012-11">November 2012</date>
, but many specimens from Hauterivian (Early Cretaceous) Wilbur Springs seep site (which is located near the Bear Creek seep site reported here; see description of locality #
<quantity id="4CA52BDDFFFC6854FA9E065CFABAF8EE" box="[1341,1382,1805,1830]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="26" pageNumber="427" unit="in" value="5.0">5 in</quantity>
<bibRefCitation id="EFCCFBC9FFFC6854FACF065CFEF1F881" pageId="26" pageNumber="427" refString="Kiel, S., Amano, K. &amp; Jenkins, R. G. (2008 a) Bivalves from Cretaceous cold-seep deposits on Hokkaido, Japan. Acta Palaeontologica Polonica, 53, 525 - 537. http: // dx. doi. org / 10.4202 / app. 2008.0310" type="journal article">
Kiel
<emphasis id="B9295A2AFFFC6854FF340663FF13F881" box="[151,207,1840,1865]" italics="true" pageId="26" pageNumber="427">et al.</emphasis>
(2008a)
</bibRefCitation>
for details) were seen in the non-type collection.
</paragraph>
</subSubSection>
<subSubSection id="C347D5B3FFFC6855FF640604FC18FEAC" lastPageId="27" lastPageNumber="428" pageId="26" pageNumber="427" type="discussion">
<paragraph id="8BE28638FFFC6855FF640604FC18FEAC" blockId="26.[151,1437,1696,2009]" lastBlockId="27.[151,1436,151,357]" lastPageId="27" lastPageNumber="428" pageId="26" pageNumber="427">
<emphasis id="B9295A2AFFFC6854FF640604FEE7F8A6" bold="true" box="[199,315,1877,1902]" pageId="26" pageNumber="427">Remarks.</emphasis>
Gabbs (1869) specimens are from Morgan Valley south of Clear Lake (
<bibRefCitation id="EFCCFBC9FFFC6854FB060604FA53F8A5" author="Stanton" box="[1189,1423,1877,1902]" pageId="26" pageNumber="427" refString="Stanton, T. W. (1895) Contributions to the Cretaceous paleontology of the Pacific coast: the fauna of the Knoxville beds. Bulletin of the United States Geological Survey, 133, 1 - 132." type="journal article" year="1895" yearSuffix="p">Stanton 1895, p. 37</bibRefCitation>
), which is a wide area that also encompasses the Rocky Creek site where the present material was collected. Stantons (1895) specimens are from Wilbur Springs, thus the species is widely distributed among Early Cretaceous seep deposits in California. Both at Rocky Creek and at Wilbur Springs the specimens occur in clusters in which several specimens were stacked upon each other in random orientation. At Wilbur Springs they were also closely associated with the brachiopod
<taxonomicName id="4C5DFDBBFFFD6855FDEF01EDFCE2FF1D" box="[588,830,188,213]" class="Rhynchonellata" family="Peregrinellidae" genus="Peregrinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="27" pageNumber="428" phylum="Brachiopoda" rank="species" species="whitneyi">
<emphasis id="B9295A2AFFFD6855FDEF01EDFCE2FF1D" box="[588,830,188,213]" italics="true" pageId="27" pageNumber="428">Peregrinella whitneyi</emphasis>
</taxonomicName>
. Other pectinids at Cretaceous seep deposits include a rare
<taxonomicName id="4C5DFDBBFFFD6855FF6A01B0FEA0FF30" box="[201,380,225,248]" class="Bivalvia" family="Propeamussiidae" genus="Propeamussium" kingdom="Animalia" order="Pectinoida" pageId="27" pageNumber="428" phylum="Mollusca" rank="genus">
<emphasis id="B9295A2AFFFD6855FF6A01B0FEA0FF30" box="[201,380,225,248]" italics="true" pageId="27" pageNumber="428">Propeamussium</emphasis>
</taxonomicName>
at the Early Cretaceous seeps deposits in the Kuhnpasset beds in
<collectingCountry id="F34AC6A8FFFD6855FBF7018EFB15FF30" box="[1108,1225,223,248]" name="Greenland" pageId="27" pageNumber="428">Greenland</collectingCountry>
(
<bibRefCitation id="EFCCFBC9FFFD6855FB74018EFA49FF30" author="Kelly" box="[1239,1429,223,248]" pageId="27" pageNumber="428" refString="Kelly, S. R. A., Blanc, E., Price, S. P. &amp; Whitham, A. G. (2000) Early Cretaceous giant bivalves from seep-related limestone mounds, Wollaston Forland, Northeast Greenland. Geological Society, London, Special Publications, 177, 227 - 246. http: // dx. doi. org / 10.1144 / gsl. sp. 2000.177.01.13" type="journal article" year="2000">
Kelly
<emphasis id="B9295A2AFFFD6855FABE01B0FA88FF30" box="[1309,1364,223,248]" italics="true" pageId="27" pageNumber="428">et al.</emphasis>
2000
</bibRefCitation>
) and a rare
<taxonomicName id="4C5DFDBBFFFD6855FEBB0054FEB8FED4" box="[280,356,261,284]" class="Bivalvia" family="Pectinidae" genus="Pecten" kingdom="Animalia" order="Pectinoida" pageId="27" pageNumber="428" phylum="Mollusca" rank="genus">
<emphasis id="B9295A2AFFFD6855FEBB0054FEB8FED4" box="[280,356,261,284]" italics="true" pageId="27" pageNumber="428">Pecten</emphasis>
</taxonomicName>
sp. at the potential seep site Koniakauer Schloß in
<collectingCountry id="F34AC6A8FFFD6855FC640055FBA6FED5" box="[967,1146,260,285]" name="Czech Republic" pageId="27" pageNumber="428">Czech Republic</collectingCountry>
(
<bibRefCitation id="EFCCFBC9FFFD6855FB290055FAC1FED5" author="Ascher" box="[1162,1309,260,285]" pageId="27" pageNumber="428" refString="Ascher, E. (1906) Die Gastropoden, Bivalven und Brachiopoden der Grodischter Schichten. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients, 19, 135 - 167." type="journal article" year="1906">Ascher 1906</bibRefCitation>
, cf.
<bibRefCitation id="EFCCFBC9FFFD6855FAED0055FE93FE88" author="Kiel" pageId="27" pageNumber="428" refString="Kiel, S. &amp; Peckmann, J. (2008) Paleoecology and evolutionary significance of an Early Cretaceous Peregrinella - dominated hydrocarbon-seep deposit on the Crimean Peninsula. Palaios, 23, 751 - 759." type="journal article" year="2008">Kiel &amp; Peckmann 2008</bibRefCitation>
,
<bibRefCitation id="EFCCFBC9FFFD6855FEF80076FDF6FE88" author="Kaim" box="[347,554,295,320]" pageId="27" pageNumber="428" refString="Kaim, A., Skupien, P. &amp; Jenkins, R. G. (2013 b) A new Lower Cretaceous hydrocarbon seep locality from the Czech Carpathians and its fauna. Palaeogeography, Palaeoclimatology, Palaeoecology, 390, 42 - 51. http: // dx. doi. org / 10.1016 / j. palaeo. 2013.03.010" type="journal article" year="2013" yearSuffix="b">
Kaim
<emphasis id="B9295A2AFFFD6855FE020078FE07FE88" box="[417,475,295,320]" italics="true" pageId="27" pageNumber="428">et al.</emphasis>
2013b
</bibRefCitation>
) which appears to have finer, more numerous, and more closely spaces radial ribs than
<taxonomicName id="4C5DFDBBFFFD6855FF5C001DFD8FFEAD" box="[255,595,332,357]" class="Bivalvia" family="Pectinidae" genus="Lyriochlamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreoida" pageId="27" pageNumber="428" phylum="Mollusca" rank="species" species="complexicostata">
<emphasis id="B9295A2AFFFD6855FF5C001DFD8FFEAD" box="[255,595,332,357]" italics="true" pageId="27" pageNumber="428">Lyriochlamys complexicostata</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFCCFBC9FFFD6855FD2C001DFCFCFEAD" author="Ascher" box="[655,800,332,357]" pageId="27" pageNumber="428" refString="Ascher, E. (1906) Die Gastropoden, Bivalven und Brachiopoden der Grodischter Schichten. Beitrage zur Palaontologie und Geologie Osterreich-Ungarns und des Orients, 19, 135 - 167." type="journal article" year="1906">Ascher 1906</bibRefCitation>
, pl. 14, fig. 1).
</paragraph>
</subSubSection>
</treatment>
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