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<document id="F3E8ACE00F33B1F5475B743680FBA8B3" ID-DOI="10.11646/zootaxa.3367.1.14" ID-ISSN="1175-5326" ID-Zenodo-Dep="5252000" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629787795747" checkinUser="felipe" docAuthor="Komai, Tomoyuki &amp; Osawa, Masayuki" docDate="2012" docId="F95087D8FFE0FFA6269BFCA0BD673E1C" docLanguage="en" docName="zt03367p154.pdf" docOrigin="Zootaxa 3367 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3367.1.14" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Kumepagurus cavernicolus Komai &amp; Osawa 2012, n. sp." docType="treatment" docVersion="3" lastPageNumber="153" masterDocId="0569FFA0FFE1FFAE260CFFBFBF6B3A0C" masterDocTitle="A new genus and new species of pagurid hermit crab (Crustacea: Decapoda: Anomura: Paguroidea) from a submarine cave in Kume Island, Ryukyu Islands *" masterLastPageNumber="154" masterPageNumber="145" pageNumber="146" updateTime="1699111969742" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="A0BF69C68A4F46952A1E3D0C38662982" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6E5AAF6F08F9DF1D09D275DECECBE2B7">
<mods:title id="B72CD579286B00924D27977486F7BE8F">A new genus and new species of pagurid hermit crab (Crustacea: Decapoda: Anomura: Paguroidea) from a submarine cave in Kume Island, Ryukyu Islands *</mods:title>
</mods:titleInfo>
<mods:name id="D43E6C95FC70D45F6AFC48F08E3EB269" type="personal">
<mods:role id="6D3E43327A46FE7B20FF5A1F706C74A2">
<mods:roleTerm id="E146552C00FBF13B69560C58875683E3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FDDA6E4358DB4164F386EBE07F38598E">Komai, Tomoyuki</mods:namePart>
<mods:affiliation id="A0ECBD4E5CC87D4B800015ED366E6535">Natural History Museum and Institute, Chiba, 955 - 2 Aoba-cho, Chuo-ku, Chiba, 260 - 8682 Japan. E-mail: komai @ chiba-muse. or. jp Research Center for Coastal Lagoon Environments, Shimane University, 1060 Nishikawatsu-cho, Matsue, Shimane, 690 - 8504, Japan. E-mail: osawam @ soc. shimane-u. ac. jp</mods:affiliation>
</mods:name>
<mods:name id="F1FA94834196D22040F369ABD4307B66" type="personal">
<mods:role id="957C194418CB5AA04E61CCE0EF54B618">
<mods:roleTerm id="9D3BB437BD554AE14E8E5F1803516193">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FF5AA8ADD2134CE3AF3E30803EF774A9">Osawa, Masayuki</mods:namePart>
</mods:name>
<mods:typeOfResource id="0D13589FDB4C4A1D2F66D3BCED993F77">text</mods:typeOfResource>
<mods:relatedItem id="B97860B5C56F8E3C0442EC7558A9702E" type="host">
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<mods:title id="5FFF267E526AAAFCBB031B4D9BFEEAD5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B802974B6E43856525ACAF70A740D5F5">
<mods:date id="B62CAAA8729ADD105073187B8031737C">2012</mods:date>
<mods:detail id="C2469AF30CA49370FFA182777D6231DC" type="pubDate">
<mods:number id="806676F8E241602E57C302AE7225007E">2012-07-04</mods:number>
</mods:detail>
<mods:detail id="00E005B170812297F4F0A99C0B6F53E8" type="volume">
<mods:number id="4783FF30F74777CB49C7CF9A71FD9482">3367</mods:number>
</mods:detail>
<mods:detail id="93B7866F1F99DC537E3D422FDEEBD58B" type="issue">
<mods:number id="BA40C5FD63811FDE21BC12157D3EF64E">1</mods:number>
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<mods:url id="5EC3CB4C6DD0EA7A9CCB5007B7D04B7F">https://biotaxa.org/Zootaxa/article/view/zootaxa.3367.1.14</mods:url>
</mods:location>
<mods:classification id="24FF6C7E8A1B925CCBDA3478CD12DCC9">journal article</mods:classification>
<mods:identifier id="73C91F0E6DE79E176781DA6B8F9CF400" type="DOI">10.11646/zootaxa.3367.1.14</mods:identifier>
<mods:identifier id="62EF7C2382C9F5DD4253417940E7CABA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B0C3525604E4F2A6881EFC2EAFC90CBF" type="Zenodo-Dep">5252000</mods:identifier>
</mods:mods>
<treatment id="F95087D8FFE0FFA6269BFCA0BD673E1C" LSID="urn:lsid:plazi:treatment:F95087D8FFE0FFA6269BFCA0BD673E1C" httpUri="http://treatment.plazi.org/id/F95087D8FFE0FFA6269BFCA0BD673E1C" lastPageId="8" lastPageNumber="153" pageId="1" pageNumber="146">
<subSubSection id="39E36545FFE0FFAF269BFCA0BD8C3956" pageId="1" pageNumber="146" type="nomenclature">
<paragraph id="714636CEFFE0FFAF269BFCA0BD473935" blockId="1.[151,743,799,894]" box="[151,556,799,825]" pageId="1" pageNumber="146">
<heading id="2A0E81A2FFE0FFAF269BFCA0BD473935" bold="true" box="[151,556,799,825]" fontSize="11" level="1" pageId="1" pageNumber="146" reason="1">
<emphasis id="438DEADCFFE0FFAF269BFCA0BD473935" bold="true" box="[151,556,799,825]" pageId="1" pageNumber="146">
<taxonomicName id="B6F94D4DFFE0FFAF269BFCA0BE893935" ID-CoL="95b77d9b-c8a3-42f6-93b5-6d6f1248c83e" authority="Komai &amp; Osawa, 2012" authorityName="Komai &amp; Osawa" authorityYear="2012" box="[151,482,799,825]" class="Malacostraca" family="Paguridae" genus="Kumepagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="146" phylum="Arthropoda" rank="species" species="cavernicolus" status="sp. nov.">
<emphasis id="438DEADCFFE0FFAF269BFCA0BE893935" bold="true" box="[151,482,799,825]" italics="true" pageId="1" pageNumber="146">Kumepagurus cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58BE57A7FFE0FFAF27E5FCA0BD473935" box="[489,556,799,825]" pageId="1" pageNumber="146" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="714636CEFFE0FFAF269BFCFCBD8C3956" blockId="1.[151,743,799,894]" box="[151,743,834,858]" pageId="1" pageNumber="146">[new Japanese name: Kumejima-doukutsu-yadokari]</paragraph>
</subSubSection>
<subSubSection id="39E36545FFE0FFAF269BFCD9BE793972" box="[151,274,870,894]" pageId="1" pageNumber="146" type="description">
<paragraph id="714636CEFFE0FFAF269BFCD9BE793972" blockId="1.[151,743,799,894]" box="[151,274,870,894]" pageId="1" pageNumber="146">
(
<figureCitation id="E9C22A4BFFE0FFAF2693FCD9BE623972" box="[159,265,870,894]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,250,1838,1861]" captionStartId-1="3.[151,250,1464,1487]" captionStartId-2="4.[151,250,1944,1967]" captionStartId-3="5.[151,250,1882,1905]" captionStartId-4="6.[151,250,1940,1963]" captionStartId-5="7.[151,250,1929,1952]" captionTargetBox-0="[251,1348,185,1803]" captionTargetBox-1="[171,1406,552,1420]" captionTargetBox-2="[221,1358,216,1899]" captionTargetBox-3="[179,1406,783,1828]" captionTargetBox-4="[184,1404,760,1894]" captionTargetBox-5="[348,1239,1237,1908]" captionTargetId-0="figure-131@2.[228,1370,179,1835]" captionTargetId-1="figure-395@3.[151,1436,525,1451]" captionTargetId-2="figure-72@4.[200,1386,193,1931]" captionTargetId-3="figure-321@5.[151,1436,769,1861]" captionTargetId-4="figure-325@6.[151,1436,739,1919]" captionTargetId-5="figure-460@7.[348,1239,1237,1908]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." captionText-1="FIGURE 2. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Left mouthparts. A, mandible, outer view; B, maxillule, outer view (proximal endite broken off); inset, endopod, lateral view; C, maxilla, outer view; D, endopod of maxilla, inner view; E, first maxilliped, outer view (proximal endite broken); F, second maxilliped, outer view. Scale = 0.5 mm." captionText-2="FIGURE 3. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Right cheliped. A, chela, dorsal view; B, same, mesial view; C, same, lateral (perpendicular) view; D, same, ventral view; E, carpus, dorsal view; F, carpus and merus, lateral view; G, same, mesial view. Scale = 1 mm." captionText-3="FIGURE 4. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Left cheliped. A, chela and carpus, dorsal view; B, entire left cheliped, mesial view; C, same,, lateral view; D, tips of fingers, dorsal view. Scales: A–C = 1 mm; D = 0.5 mm." captionText-4="FIGURE 5. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Ambulatory legs. A, right second pereopod, lateral view; B, same, dactylus, mesial view; C, left third pereopod, lateral view; D, same, dactylus, mesial view. Scales = 1mm." captionText-5="FIGURE 6. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Entire animal in dorsal view, showing colouration in life. Photo: Tin-Yam Chan." figureDoi-0="http://doi.org/10.5281/zenodo.5252004" figureDoi-1="http://doi.org/10.5281/zenodo.5252008" figureDoi-2="http://doi.org/10.5281/zenodo.5252012" figureDoi-3="http://doi.org/10.5281/zenodo.5252016" figureDoi-4="http://doi.org/10.5281/zenodo.5252020" figureDoi-5="http://doi.org/10.5281/zenodo.5252022" httpUri-0="https://zenodo.org/record/5252004/files/figure.png" httpUri-1="https://zenodo.org/record/5252008/files/figure.png" httpUri-2="https://zenodo.org/record/5252012/files/figure.png" httpUri-3="https://zenodo.org/record/5252016/files/figure.png" httpUri-4="https://zenodo.org/record/5252020/files/figure.png" httpUri-5="https://zenodo.org/record/5252022/files/figure.png" pageId="1" pageNumber="146">Figs. 1–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="39E36545FFE0FFAF269BFC11BFBD3E02" pageId="1" pageNumber="146" type="materials_examined">
<paragraph id="714636CEFFE0FFAF269BFC11BFBD3E02" blockId="1.[151,1436,942,1938]" pageId="1" pageNumber="146">
<emphasis id="438DEADCFFE0FFAF269BFC11BEE939CA" box="[151,386,942,967]" inLineHeading="true" pageId="1" pageNumber="146" reason="1">
<emphasis id="438DEADCFFE0FFAF269BFC11BE1039CB" bold="true" box="[151,379,942,967]" pageId="1" pageNumber="146">Material examined</emphasis>
.
</emphasis>
<materialsCitation id="C1913C93FFE0FFAF2786FC11BFBD3E02" collectingDate="2009-11-15" collectorName="Y. Fujita &amp; S. Ogawa" elevation="50" latitude="26.332733" location="Kume Island" longLatPrecision="1" longitude="126.731766" pageId="1" pageNumber="146" specimenCount="1" typeStatus="holotype">
<typeStatus id="AE42886CFFE0FFAF2786FC11BE9C39CA" box="[394,503,942,966]" pageId="1" pageNumber="146" type="holotype">Holotype</typeStatus>
: male (sl
<quantity id="B6019B2BFFE0FFAF246BFC11BDA939CA" box="[615,706,942,966]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="1" pageNumber="146" unit="mm" value="2.4">2.4 mm</quantity>
),
<quantity id="B6019B2BFFE0FFAF24DBFC11BC7939CA" box="[727,786,942,966]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="146" unit="m" value="50.0">
<elevation id="FAD4D1FDFFE0FFAF24DBFC11BC7939CA" box="[727,786,942,966]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="146" unit="m" value="50.0">50 m</elevation>
</quantity>
from entrance inside a submarine limestone cave,
<location id="74266015FFE0FFAF235BFC10BFB539E6" LSID="urn:lsid:plazi:treatment:F95087D8FFE0FFA6269BFCA0BD673E1C:74266015FFE0FFAF235BFC10BFB539E6" latitude="26.332733" longLatPrecision="1" longitude="126.731766" name="Kume Island" pageId="1" pageNumber="146">Kume Island</location>
,
<geoCoordinate id="14CD5009FFE0FFAF26E7FC6DBE1739E6" box="[235,380,978,1002]" degrees="26" direction="north" minutes="19.964" orientation="latitude" pageId="1" pageNumber="146" precision="1" value="26.332733">26°19.964ʹN</geoCoordinate>
,
<geoCoordinate id="14CD5009FFE0FFAF2785FC6DBD4E39E6" box="[393,549,978,1002]" degrees="126" direction="east" minutes="43.906" orientation="longitude" pageId="1" pageNumber="146" precision="1" value="126.731766">126°43.906ʹE</geoCoordinate>
,
<quantity id="B6019B2BFFE0FFAF243EFC6DBD0439E6" box="[562,623,978,1002]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="146" unit="m" value="38.0">
<elevation id="FAD4D1FDFFE0FFAF243EFC6DBD0439E6" box="[562,623,978,1002]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="146" unit="m" value="38.0">38 m</elevation>
</quantity>
,
<date id="0547100EFFE0FFAF2470FC6DBC3139E6" box="[636,858,978,1002]" pageId="1" pageNumber="146" value="2009-11-15">
<collectingDate id="1503E9E6FFE0FFAF2470FC6DBC3139E6" box="[636,858,978,1002]" pageId="1" pageNumber="146" value="2009-11-15">15 November 2009</collectingDate>
</date>
,
<collectingMethod id="A8B84ED9FFE0FFAF256BFC6DBCA339E6" box="[871,968,978,1002]" pageId="1" pageNumber="146">bait trap</collectingMethod>
, coll.
<collectorName id="DC0C5318FFE0FFAF2203FC6CBB1A39E6" box="[1039,1137,978,1002]" pageId="1" pageNumber="146">Y. Fujita</collectorName>
&amp;
<collectorName id="DC0C5318FFE0FFAF2294FC6DBA6239E6" box="[1176,1289,978,1002]" pageId="1" pageNumber="146">S. Ogawa</collectorName>
, RUMF-ZC- 1518.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="39E36545FFE0FFA626CBFBA5BCBE3BF8" lastPageId="8" lastPageNumber="153" pageId="1" pageNumber="146" type="description">
<paragraph id="714636CEFFE0FFAF26CBFBA5BD3B3F06" blockId="1.[151,1436,942,1938]" pageId="1" pageNumber="146">
<emphasis id="438DEADCFFE0FFAF26CBFBA5BE333E3E" box="[199,344,1050,1075]" inLineHeading="true" pageId="1" pageNumber="146" reason="1">
<emphasis id="438DEADCFFE0FFAF26CBFBA5BE3A3E3F" bold="true" box="[199,337,1050,1075]" pageId="1" pageNumber="146">Description</emphasis>
.
</emphasis>
Shield (
<figureCitation id="E9C22A4BFFE0FFAF27B6FBA5BD5D3E3E" box="[442,566,1050,1074]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="1" pageNumber="146">Fig. 1A, B</figureCitation>
) as long as broad; anterior margins between rostrum and lateral projections concave; anterolateral margins slightly terraced, posterior margin roundly truncate; dorsal surface with few tufts of short setae laterally. Rostrum roundly triangular, terminating in tiny sharp spine. Lateral projections obtusely triangular, not reaching level of rostral tip; each with small submarginal spine. Posterior carapace (
<figureCitation id="E9C22A4BFFE0FFAF2334FB39BAF83E92" box="[1336,1427,1158,1182]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="1" pageNumber="146">Fig. 1B</figureCitation>
) membranous; with pair of tufts of short setae at level of midlength; posteromedian plate moderately wide, with slightly convex lateral margins; sulci cardiobranchiales parallel, extending to midlength between cervical groove and posteromedian margin of carapace.
</paragraph>
<paragraph id="714636CEFFE0FFAF26CBFAA9BBD63F96" blockId="1.[151,1436,942,1938]" pageId="1" pageNumber="146">
Ocular peduncles (
<figureCitation id="E9C22A4BFFE0FFAF2795FAA9BE9B3F22" box="[409,496,1302,1326]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="1" pageNumber="146">Fig. 1A</figureCitation>
) about 0.6 times as long as shield, moderately stout, not tapering distally or inflated basally, each with prominent tuft of stiff setae at base of cornea and few short setae dorsomesially; corneas not dilated, corneal width about 0.3 times peduncular length. Ocular acicles narrowly subtriangular, directed slightly mesially, terminating in subacute marginal spine. Interocular lobe small, visible in dorsal view.
</paragraph>
<paragraph id="714636CEFFE0FFAF26CBFA19BD043C42" blockId="1.[151,1436,942,1938]" pageId="1" pageNumber="146">
Antennular peduncles (
<figureCitation id="E9C22A4BFFE0FFAF27DEFA19BD383FB2" box="[466,595,1446,1470]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="1" pageNumber="146">Fig. 1A, C</figureCitation>
), when fully extended, overreaching ocular peduncles by full lengths of ultimate segments. Basal segment with small spine on lateral margin located slightly distal to midlength; distolateral angle produced in small lobe. Ultimate segment elongate, 0.9 times as long as shield, slightly broadened distally in lateral view, with few short setae dorsally. Dorsal flagellum with distal portion much longer than proximal, aesthetasc-bearing portion.
</paragraph>
<paragraph id="714636CEFFE0FFAF26CBF9E5BD473D9E" blockId="1.[151,1436,942,1938]" pageId="1" pageNumber="146">
Antennal peduncles (
<figureCitation id="E9C22A4BFFE0FFAF27B1F9E5BD733C7E" box="[445,536,1626,1650]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="1" pageNumber="146">Fig. 1A</figureCitation>
) relatively long, overreaching distal corneal margins by full lengths of fifth segments. First segment unarmed on lateral face, ventromesial distal margin produced, with bifid spinule just lateral to antennal gland opening. Second segment with dorsolateral distal angle produced, reaching midlength of third segment, terminating in bifid spine; dorsomesial distal angle with small spine. Third segment with 1 small spine at ventrodistal margin. Fourth and fifth segments with few short setae mesially, fourth segment slightly shorter than fifth segment. Antennal acicle slender, elongate, overreaching distal corneal margin by about half own length, not reaching distal end of fifth peduncular segment, terminating in small spine, bearing prominent tuft of long setae distally and row of sparse short setae dorsomesially. Antennal flagellum (
<figureCitation id="E9C22A4BFFE0FFAF2233F8E9BBFE3D62" box="[1087,1173,1878,1902]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="1" pageNumber="146">Fig. 1D</figureCitation>
) very long, each article with few short to long setae distally.
</paragraph>
<caption id="25866646FFE3FFAC269BF891BE5E3DE1" ID-DOI="http://doi.org/10.5281/zenodo.5252004" ID-Zenodo-Dep="5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="2" pageNumber="147" startId="2.[151,250,1838,1861]" targetBox="[251,1348,185,1803]" targetPageId="2">
<paragraph id="714636CEFFE3FFAC269BF891BE5E3DE1" blockId="2.[151,1436,1838,2029]" pageId="2" pageNumber="147">
<emphasis id="438DEADCFFE3FFAC269BF891BE7F3D49" bold="true" box="[151,276,1838,1861]" pageId="2" pageNumber="147">FIGURE 1.</emphasis>
<taxonomicName id="B6F94D4DFFE3FFAC2717F88FBD583D49" authority="Komai &amp; Osawa, 2012" authorityName="Komai &amp; Osawa" authorityYear="2012" box="[283,563,1839,1861]" class="Malacostraca" family="Paguridae" genus="Kumepagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="147" phylum="Arthropoda" rank="species" species="cavernicolus" status="n. gen., n. sp.">
<emphasis id="438DEADCFFE3FFAC2717F88FBD583D49" box="[283,563,1839,1861]" italics="true" pageId="2" pageNumber="147">Kumepagurus cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58BE57A7FFE3FFAC2436F890BDA13D48" box="[570,714,1839,1860]" pageId="2" pageNumber="147" rank="species">
<emphasis id="438DEADCFFE3FFAC2436F890BDE83D48" bold="true" box="[570,643,1839,1860]" pageId="2" pageNumber="147">n. gen.</emphasis>
,
<emphasis id="438DEADCFFE3FFAC249CF890BDA13D48" bold="true" box="[656,714,1839,1860]" pageId="2" pageNumber="147">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm.
</paragraph>
</caption>
<paragraph id="714636CEFFE2FFAD26CBFF28BE3C3BF8" blockId="3.[151,1436,151,500]" pageId="3" pageNumber="148">
Mandible (
<figureCitation id="E9C22A4BFFE2FFAD274EFF28BEF03AA3" box="[322,411,151,175]" captionStart="FIGURE 2" captionStartId="3.[151,250,1464,1487]" captionTargetBox="[171,1406,552,1420]" captionTargetId="figure-395@3.[151,1436,525,1451]" captionTargetPageId="3" captionText="FIGURE 2. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Left mouthparts. A, mandible, outer view; B, maxillule, outer view (proximal endite broken off); inset, endopod, lateral view; C, maxilla, outer view; D, endopod of maxilla, inner view; E, first maxilliped, outer view (proximal endite broken); F, second maxilliped, outer view. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252008" httpUri="https://zenodo.org/record/5252008/files/figure.png" pageId="3" pageNumber="148">Fig. 2A</figureCitation>
) without distinctive features. Maxillule (
<figureCitation id="E9C22A4BFFE2FFAD2565FF28BCAB3AA3" box="[873,960,151,175]" captionStart="FIGURE 2" captionStartId="3.[151,250,1464,1487]" captionTargetBox="[171,1406,552,1420]" captionTargetId="figure-395@3.[151,1436,525,1451]" captionTargetPageId="3" captionText="FIGURE 2. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Left mouthparts. A, mandible, outer view; B, maxillule, outer view (proximal endite broken off); inset, endopod, lateral view; C, maxilla, outer view; D, endopod of maxilla, inner view; E, first maxilliped, outer view (proximal endite broken); F, second maxilliped, outer view. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252008" httpUri="https://zenodo.org/record/5252008/files/figure.png" pageId="3" pageNumber="148">Fig. 2B</figureCitation>
) with endopod slender, weakly developed inner lobe with 1 apical spiniform seta, outer lobe elongate, with 1 terminal and 1 basal seta. Maxilla (
<figureCitation id="E9C22A4BFFE2FFAD230AFF03BA303AD8" box="[1286,1371,188,212]" captionStart="FIGURE 2" captionStartId="3.[151,250,1464,1487]" captionTargetBox="[171,1406,552,1420]" captionTargetId="figure-395@3.[151,1436,525,1451]" captionTargetPageId="3" captionText="FIGURE 2. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Left mouthparts. A, mandible, outer view; B, maxillule, outer view (proximal endite broken off); inset, endopod, lateral view; C, maxilla, outer view; D, endopod of maxilla, inner view; E, first maxilliped, outer view (proximal endite broken); F, second maxilliped, outer view. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252008" httpUri="https://zenodo.org/record/5252008/files/figure.png" pageId="3" pageNumber="148">Fig. 2C</figureCitation>
) with relatively short scaphognathite; endopod slender, gradually tapering distally (
<figureCitation id="E9C22A4BFFE2FFAD220AFF60BB083AFB" box="[1030,1123,223,247]" captionStart="FIGURE 2" captionStartId="3.[151,250,1464,1487]" captionTargetBox="[171,1406,552,1420]" captionTargetId="figure-395@3.[151,1436,525,1451]" captionTargetPageId="3" captionText="FIGURE 2. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Left mouthparts. A, mandible, outer view; B, maxillule, outer view (proximal endite broken off); inset, endopod, lateral view; C, maxilla, outer view; D, endopod of maxilla, inner view; E, first maxilliped, outer view (proximal endite broken); F, second maxilliped, outer view. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252008" httpUri="https://zenodo.org/record/5252008/files/figure.png" pageId="3" pageNumber="148">Fig. 2D</figureCitation>
). First maxilliped (
<figureCitation id="E9C22A4BFFE2FFAD2331FF60BAF83AFB" box="[1341,1427,223,247]" captionStart="FIGURE 2" captionStartId="3.[151,250,1464,1487]" captionTargetBox="[171,1406,552,1420]" captionTargetId="figure-395@3.[151,1436,525,1451]" captionTargetPageId="3" captionText="FIGURE 2. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Left mouthparts. A, mandible, outer view; B, maxillule, outer view (proximal endite broken off); inset, endopod, lateral view; C, maxilla, outer view; D, endopod of maxilla, inner view; E, first maxilliped, outer view (proximal endite broken); F, second maxilliped, outer view. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252008" httpUri="https://zenodo.org/record/5252008/files/figure.png" pageId="3" pageNumber="148">Fig. 2E</figureCitation>
) with distal endite subovate; endopod reaching beyond distal margin of distal endite; exopod consisting of 2 segments and flagellum, distal segment strongly flexed laterally, proximal segment with slightly expanded lateral margin, flagellum shorter than distal segment. Second maxilliped (
<figureCitation id="E9C22A4BFFE2FFAD258BFEF3BCB03B68" box="[903,987,332,356]" captionStart="FIGURE 2" captionStartId="3.[151,250,1464,1487]" captionTargetBox="[171,1406,552,1420]" captionTargetId="figure-395@3.[151,1436,525,1451]" captionTargetPageId="3" captionText="FIGURE 2. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Left mouthparts. A, mandible, outer view; B, maxillule, outer view (proximal endite broken off); inset, endopod, lateral view; C, maxilla, outer view; D, endopod of maxilla, inner view; E, first maxilliped, outer view (proximal endite broken); F, second maxilliped, outer view. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252008" httpUri="https://zenodo.org/record/5252008/files/figure.png" pageId="3" pageNumber="148">Fig. 2F</figureCitation>
) relatively large, with elongate exopod; flagellum well developed, flexed at proximal 0.3. Third maxilliped (
<figureCitation id="E9C22A4BFFE2FFAD25BEFED0BB623B8B" box="[946,1033,367,391]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="3" pageNumber="148">Fig. 1E</figureCitation>
) with endopod relatively long and slender; carpus and merus unarmed; ischium and basis incompletely fused; crista dentata on ischium consisting of row of about 10 rather widely separated, acute teeth, accessory tooth absent; basis with 1 tiny denticle on mesial margin (
<figureCitation id="E9C22A4BFFE2FFAD26F9FE63BE223BF8" box="[245,329,476,500]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="3" pageNumber="148">Fig. 1F</figureCitation>
).
</paragraph>
<caption id="25866646FFE2FFAD269BFA07BE123C26" ID-DOI="http://doi.org/10.5281/zenodo.5252008" ID-Zenodo-Dep="5252008" httpUri="https://zenodo.org/record/5252008/files/figure.png" pageId="3" pageNumber="148" startId="3.[151,250,1464,1487]" targetBox="[171,1406,552,1420]" targetPageId="3">
<paragraph id="714636CEFFE2FFAD269BFA07BE123C26" blockId="3.[151,1436,1464,1578]" pageId="3" pageNumber="148">
<emphasis id="438DEADCFFE2FFAD269BFA07BE7E3FC3" bold="true" box="[151,277,1464,1487]" pageId="3" pageNumber="148">FIGURE 2.</emphasis>
<taxonomicName id="B6F94D4DFFE2FFAD2713FA06BD5C3FC2" authority="Komai &amp; Osawa, 2012" authorityName="Komai &amp; Osawa" authorityYear="2012" box="[287,567,1464,1486]" class="Malacostraca" family="Paguridae" genus="Kumepagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="cavernicolus" status="n. gen., n. sp.">
<emphasis id="438DEADCFFE2FFAD2713FA06BD5C3FC2" box="[287,567,1464,1486]" italics="true" pageId="3" pageNumber="148">Kumepagurus cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58BE57A7FFE2FFAD244DFA06BDBD3FC2" box="[577,726,1465,1486]" pageId="3" pageNumber="148" rank="species">
<emphasis id="438DEADCFFE2FFAD244DFA06BDEF3FC2" bold="true" box="[577,644,1465,1486]" pageId="3" pageNumber="148">n. gen</emphasis>
.,
<emphasis id="438DEADCFFE2FFAD2496FA06BDBD3FC2" bold="true" box="[666,726,1465,1486]" pageId="3" pageNumber="148">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (sl 2.4 mm), RUMF-ZC-1518. Left mouthparts. A, mandible, outer view; B, maxillule, outer view (proximal endite broken off); inset, endopod, lateral view; C, maxilla, outer view; D, endopod of maxilla, inner view; E, first maxilliped, outer view (proximal endite broken); F, second maxilliped, outer view. Scale = 0.5 mm.
</paragraph>
</caption>
<paragraph id="714636CEFFE2FFAB26CBF9E8BE8438C0" blockId="3.[151,1436,1623,2007]" lastBlockId="5.[151,1436,151,716]" lastPageId="5" lastPageNumber="150" pageId="3" pageNumber="148">
Right cheliped (
<figureCitation id="E9C22A4BFFE2FFAD278FF9E8BD693C63" box="[387,514,1623,1647]" captionStart="FIGURE 3" captionStartId="4.[151,250,1944,1967]" captionTargetBox="[221,1358,216,1899]" captionTargetId="figure-72@4.[200,1386,193,1931]" captionTargetPageId="4" captionText="FIGURE 3. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Right cheliped. A, chela, dorsal view; B, same, mesial view; C, same, lateral (perpendicular) view; D, same, ventral view; E, carpus, dorsal view; F, carpus and merus, lateral view; G, same, mesial view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5252012" httpUri="https://zenodo.org/record/5252012/files/figure.png" pageId="3" pageNumber="148">Fig. 3A–G</figureCitation>
) massive. Chela operculate, subrectangular, about 1.2 times longer than wide; propodal-carpal articulation rotated clockwise about 20° from perpendicular. Dactylus subequal in length to palm, articulating obliquely; dorsal surface with scattered blunt granules, dorsomesial margin produced, forming sharp crest with granulated margin; ventral face convex along midline, shallowly concave proximolaterally, with scattered small, blister-like tubercles; cutting edge with 3 bluntly triangular calcareous teeth, terminating in tiny, sharp corneous claw. Palm longer than carpus, widened distally, about 1.2 times wider than long; dorsal surface slightly convex in general, with scattered, occasionally corneous-tipped, granules; dorsolateral margin delimited by row of minute granules not extending onto fixed finger, distal end of dorsolateral margin forming low, but rather prominent short ridge; dorsomesial margin delimited proximally by distinct crest with slightly upturned edge, distal part of dorsomesial portion rounded; lateral, mesial, ventral surfaces (including fixed finger) densely covered with low, rounded granules (granules largest and rather blister-like on ventral surface of fixed finger and distal part of palm), lateral surface with scattered short to long setae dorsally, ventral surface having blunt transverse ridge extending from base of dactylus to ventrolateral distal margin of fixed finger, thus ventral face of fixed finger shallowly concave. Fixed finger with markedly convex lateral face; cutting edge with 3 triangular calcareous teeth (middle tooth strongest), terminating in minute corneous claw; scattered short to moderately long setae present on dorsal surface and dorsal part of mesial face. Carpus subequal in length to merus, widened distally; dorsal surface elevated in midline, with row of small spines, sloping lateral and mesial sides with scattered granules, latter with long stiff setae adjacent to mesial margin; lateral margin faintly delimited, with 1 small distal spine, mesial margin sharply delimited by spinulose carina (spinules strongest distally) bearing long stiff setae; ventrolateral surface with scattered granules; ventromesial face nearly smooth but with scattered stiff setae; ventral surface strongly convex, with scattered granules and stiff setae, proximal part only weakly calcified. Merus with few low, transverse ridges in dorsodistal part, distalmost ridge longest, with row of short stiff setae; lateral face covered with minute granules and with iridescent sheen; ventrolateral margin elevated, strongly convex in lateral view, with row of spinules in distal half; mesial face nearly flat, smooth; ventromesial margin crested, strongly convex in lateral view, with row of spinules and stiff setae; only weakly calcified distal part of ventral surface forming deep concavity to accommodate flexed carpus, proximal part well calcified and almost flat. Ischium unarmed, with iridescent sheen on lateral face. Coxa unarmed.
</paragraph>
<caption id="25866646FFE5FFAA269BF827BC263DFF" ID-DOI="http://doi.org/10.5281/zenodo.5252012" ID-Zenodo-Dep="5252012" httpUri="https://zenodo.org/record/5252012/files/figure.png" pageId="4" pageNumber="149" startId="4.[151,250,1944,1967]" targetBox="[221,1358,216,1899]" targetPageId="4">
<paragraph id="714636CEFFE5FFAA269BF827BC263DFF" blockId="4.[151,1436,1944,2035]" pageId="4" pageNumber="149">
<emphasis id="438DEADCFFE5FFAA269BF827BE793DA3" bold="true" box="[151,274,1944,1967]" pageId="4" pageNumber="149">FIGURE 3.</emphasis>
<taxonomicName id="B6F94D4DFFE5FFAA2714F825BD463DA3" authority="Komai &amp; Osawa, 2012" authorityName="Komai &amp; Osawa" authorityYear="2012" box="[280,557,1945,1967]" class="Malacostraca" family="Paguridae" genus="Kumepagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="149" phylum="Arthropoda" rank="species" species="cavernicolus" status="n. gen., n. sp.">
<emphasis id="438DEADCFFE5FFAA2714F825BD463DA3" box="[280,557,1945,1967]" italics="true" pageId="4" pageNumber="149">Kumepagurus cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58BE57A7FFE5FFAA2438F826BDAB3DA2" box="[564,704,1945,1966]" pageId="4" pageNumber="149" rank="species">
<emphasis id="438DEADCFFE5FFAA2438F826BD103DA2" bold="true" box="[564,635,1945,1966]" pageId="4" pageNumber="149">n. gen.</emphasis>
,
<emphasis id="438DEADCFFE5FFAA248BF826BDAB3DA2" bold="true" box="[647,704,1945,1966]" pageId="4" pageNumber="149">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (sl 2.4 mm), RUMF-ZC-1518. Right cheliped. A, chela, dorsal view; B, same, mesial view; C, same, lateral (perpendicular) view; D, same, ventral view; E, carpus, dorsal view; F, carpus and merus, lateral view; G, same, mesial view. Scale = 1 mm.
</paragraph>
</caption>
<caption id="25866646FFE4FFAB269BF8E5BE1E3DA2" ID-DOI="http://doi.org/10.5281/zenodo.5252016" ID-Zenodo-Dep="5252016" httpUri="https://zenodo.org/record/5252016/files/figure.png" pageId="5" pageNumber="150" startId="5.[151,250,1882,1905]" targetBox="[179,1406,783,1828]" targetPageId="5">
<paragraph id="714636CEFFE4FFAB269BF8E5BE1E3DA2" blockId="5.[151,1436,1882,1966]" pageId="5" pageNumber="150">
<emphasis id="438DEADCFFE4FFAB269BF8E5BE783D7D" bold="true" box="[151,275,1882,1905]" pageId="5" pageNumber="150">FIGURE 4.</emphasis>
<taxonomicName id="B6F94D4DFFE4FFAB2717F8E3BD5A3D7D" authority="Komai &amp; Osawa, 2012" authorityName="Komai &amp; Osawa" authorityYear="2012" box="[283,561,1883,1905]" class="Malacostraca" family="Paguridae" genus="Kumepagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="150" phylum="Arthropoda" rank="species" species="cavernicolus" status="n. gen., n. sp.">
<emphasis id="438DEADCFFE4FFAB2717F8E3BD5A3D7D" box="[283,561,1883,1905]" italics="true" pageId="5" pageNumber="150">Kumepagurus cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58BE57A7FFE4FFAB2434F8E4BDA33D7C" box="[568,712,1883,1905]" pageId="5" pageNumber="150" rank="species">
<emphasis id="438DEADCFFE4FFAB2434F8E4BDEA3D7C" bold="true" box="[568,641,1883,1904]" pageId="5" pageNumber="150">n. gen.</emphasis>
,
<emphasis id="438DEADCFFE4FFAB2482F8E4BDA33D7C" bold="true" box="[654,712,1883,1904]" pageId="5" pageNumber="150">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (sl 2.4 mm), RUMF-ZC-1518. Left cheliped. A, chela and carpus, dorsal view; B, entire left cheliped, mesial view; C, same,, lateral view; D, tips of fingers, dorsal view. Scales: A–C = 1 mm; D = 0.5 mm.
</paragraph>
</caption>
<paragraph id="714636CEFFE7FFA826CBFF28BEE238C0" blockId="6.[151,1437,151,716]" pageId="6" pageNumber="151">
Left cheliped (
<figureCitation id="E9C22A4BFFE7FFA82778FF28BE9A3AA3" box="[372,497,151,175]" captionStart="FIGURE 4" captionStartId="5.[151,250,1882,1905]" captionTargetBox="[179,1406,783,1828]" captionTargetId="figure-321@5.[151,1436,769,1861]" captionTargetPageId="5" captionText="FIGURE 4. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Left cheliped. A, chela and carpus, dorsal view; B, entire left cheliped, mesial view; C, same,, lateral view; D, tips of fingers, dorsal view. Scales: A–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252016" httpUri="https://zenodo.org/record/5252016/files/figure.png" pageId="6" pageNumber="151">Fig. 4A–C</figureCitation>
) moderately slender, reaching nearly to distal end of right cheliped, with slight degree of counterclockwise rotation from perpendicular. Chela about 3.1 times longer than wide and 1.4 times longer than carpus; dactylus and fixed finger only slightly curved ventrally, without hiatus. Dactylus about 1.2 times longer than palm, unarmed but with sparse tufts of short to moderately long setae, terminating in small corneous claw (
<figureCitation id="E9C22A4BFFE7FFA82740FE98BEC33B33" box="[332,424,295,319]" captionStart="FIGURE 4" captionStartId="5.[151,250,1882,1905]" captionTargetBox="[179,1406,783,1828]" captionTargetId="figure-321@5.[151,1436,769,1861]" captionTargetPageId="5" captionText="FIGURE 4. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Left cheliped. A, chela and carpus, dorsal view; B, entire left cheliped, mesial view; C, same,, lateral view; D, tips of fingers, dorsal view. Scales: A–C = 1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252016" httpUri="https://zenodo.org/record/5252016/files/figure.png" pageId="6" pageNumber="151">Fig. 4D</figureCitation>
); dorsomesial margin not delimited; cutting edge with row of closely set corneous teeth over entire length. Palm 0.6 length of carpus; dorsal surface slightly convex, with scattered granules and sparse setae and few low protuberances mesially, without delineation of dorsolateral and dorsomesial margins; mesial and ventral faces generally smooth, with scattered setae. Fixed finger bearing 2 acute corneous spines at tip (
<figureCitation id="E9C22A4BFFE7FFA8233FFE2BBAE43BA0" box="[1331,1423,404,428]" captionStart="FIGURE 2" captionStartId="3.[151,250,1464,1487]" captionTargetBox="[171,1406,552,1420]" captionTargetId="figure-395@3.[151,1436,525,1451]" captionTargetPageId="3" captionText="FIGURE 2. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Left mouthparts. A, mandible, outer view; B, maxillule, outer view (proximal endite broken off); inset, endopod, lateral view; C, maxilla, outer view; D, endopod of maxilla, inner view; E, first maxilliped, outer view (proximal endite broken); F, second maxilliped, outer view. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252008" httpUri="https://zenodo.org/record/5252008/files/figure.png" pageId="6" pageNumber="151">Fig. 2D</figureCitation>
); cutting edge with few tiny acute calcareous teeth and row of sparse minute, corneous teeth. Carpus widened distally, length about 3.0 of distal width; dorsal surface sloping to lateral face, having tufts of short to long stiff setae and row of tiny spines mesially; lateral face with scattered granules, distal margin with small spine; mesial face with long stiff setae dorsally; ventral surface slightly convex, with sparse setae. Merus with sparse row of short setae on dorsal surface, dorsodistal margin unarmed; lateral face with scattered granules, ventrolateral margin with row of small spines in distal half; mesial face smooth, with few short setae, ventromesial margin with 1 distal spine, otherwise unarmed; ventral surface with sparse tufts of long setae. Ischium unarmed, with scattered short setae. Coxa unarmed.
</paragraph>
<caption id="25866646FFE7FFA8269BF82BBED93DE3" ID-DOI="http://doi.org/10.5281/zenodo.5252020" ID-Zenodo-Dep="5252020" httpUri="https://zenodo.org/record/5252020/files/figure.png" pageId="6" pageNumber="151" startId="6.[151,250,1940,1963]" targetBox="[184,1404,760,1894]" targetPageId="6">
<paragraph id="714636CEFFE7FFA8269BF82BBED93DE3" blockId="6.[151,1436,1940,2031]" pageId="6" pageNumber="151">
<emphasis id="438DEADCFFE7FFA8269BF82BBE7E3DA6" bold="true" box="[151,277,1940,1963]" pageId="6" pageNumber="151">FIGURE 5.</emphasis>
<taxonomicName id="B6F94D4DFFE7FFA82712F82ABD5D3DA7" authority="Komai &amp; Osawa, 2012" authorityName="Komai &amp; Osawa" authorityYear="2012" box="[286,566,1941,1963]" class="Malacostraca" family="Paguridae" genus="Kumepagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="151" phylum="Arthropoda" rank="species" species="cavernicolus" status="n. gen., n. sp.">
<emphasis id="438DEADCFFE7FFA82712F82ABD5D3DA7" box="[286,566,1941,1963]" italics="true" pageId="6" pageNumber="151">Kumepagurus cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58BE57A7FFE7FFA82433F82ABDB83DA6" box="[575,723,1941,1962]" pageId="6" pageNumber="151" rank="species">
<emphasis id="438DEADCFFE7FFA82433F82ABDE23DA6" bold="true" box="[575,649,1941,1962]" pageId="6" pageNumber="151">n. gen.</emphasis>
,
<emphasis id="438DEADCFFE7FFA8249BF82ABDB83DA6" bold="true" box="[663,723,1941,1962]" pageId="6" pageNumber="151">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (sl 2.4 mm), RUMF-ZC-1518. Ambulatory legs. A, right second pereopod, lateral view; B, same, dactylus, mesial view; C, left third pereopod, lateral view; D, same, dactylus, mesial view. Scales = 1mm.
</paragraph>
</caption>
<paragraph id="714636CEFFE6FFA926CBFF28BDA93830" blockId="7.[151,1436,151,1184]" pageId="7" pageNumber="152">
Ambulatory legs (
<figureCitation id="E9C22A4BFFE6FFA92798FF28BD643AA3" box="[404,527,151,175]" captionStart="FIGURE 5" captionStartId="6.[151,250,1940,1963]" captionTargetBox="[184,1404,760,1894]" captionTargetId="figure-325@6.[151,1436,739,1919]" captionTargetPageId="6" captionText="FIGURE 5. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Ambulatory legs. A, right second pereopod, lateral view; B, same, dactylus, mesial view; C, left third pereopod, lateral view; D, same, dactylus, mesial view. Scales = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5252020" httpUri="https://zenodo.org/record/5252020/files/figure.png" pageId="7" pageNumber="152">Fig. 5A, C</figureCitation>
) relatively slender, slightly overreaching tip of right cheliped. Dactyli about 1.3 (second) or 1.7 (third) length of propodi, about 11.0 times longer than wide, slightly curved ventrally in lateral view, nearly straight in dorsal view, terminating in long, slender corneous claws; dorsal margins each with row of moderately long setae; lateral faces each with very shallow longitudinal sulcus; mesial faces each with very shallow longitudinal sulcus and row of 8 (second) or 9–11 (third) corneous spinules adjacent to dorsal margin; ventral margins each with 10–12 slender corneous spines increasing in size distally (
<figureCitation id="E9C22A4BFFE6FFA92259FEF3BBB83B68" box="[1109,1235,332,356]" captionStart="FIGURE 5" captionStartId="6.[151,250,1940,1963]" captionTargetBox="[184,1404,760,1894]" captionTargetId="figure-325@6.[151,1436,739,1919]" captionTargetPageId="6" captionText="FIGURE 5. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Ambulatory legs. A, right second pereopod, lateral view; B, same, dactylus, mesial view; C, left third pereopod, lateral view; D, same, dactylus, mesial view. Scales = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5252020" httpUri="https://zenodo.org/record/5252020/files/figure.png" pageId="7" pageNumber="152">Fig. 5B, D</figureCitation>
). Propodi slightly curved ventrally, those of second longer than those of third; dorsal surfaces each with sparse short and long setae; lateral and mesial faces smooth; ventral surfaces with sparse short setae, ventrodistal margins each armed with 1 or 2 slender corneous spines. Carpi with or without spinule at dorsodistal angle and with few setae on dorsal surface; lateral and mesial faces smooth. Meri unarmed, with sparse setae on dorsal and ventral surfaces; ventrolateral margins shallowly concave in distal 0.2. Ischia unarmed. Distinct concavity on ventral margin around articulation between merus and ischium in second pereopods.
</paragraph>
<paragraph id="714636CEFFE6FFA926CBFDF8BA5138A4" blockId="7.[151,1436,151,1184]" pageId="7" pageNumber="152">
Fourth pereopods (
<figureCitation id="E9C22A4BFFE6FFA92790FDF8BE983853" box="[412,499,583,607]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="7" pageNumber="152">Fig. 1G</figureCitation>
) simple. Dactyli each with row of corneous teeth on ventral margin; dorsal margin with prominent tuft of stiff setae. Propodi each with tuft of setae distally on dorsal margin; rasp consisting of single row of corneous scales increasing in size distally; carpus with tufts of setae in distal half of dorsal margin.
</paragraph>
<paragraph id="714636CEFFE6FFA9269BFD0BBEB138C0" blockId="7.[151,1436,151,1184]" box="[151,474,692,716]" pageId="7" pageNumber="152">Fifth pereopods semichelate.</paragraph>
<paragraph id="714636CEFFE6FFA926CBFD67BEDC3918" blockId="7.[151,1436,151,1184]" pageId="7" pageNumber="152">
Anterior lobe of sixth thoracic sternite (
<figureCitation id="E9C22A4BFFE6FFA92487FD67BD8938FC" box="[651,738,728,752]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="7" pageNumber="152">Fig. 1H</figureCitation>
) subsemicircular, with row of setae on anterior margin, setae longest in medial portion.
</paragraph>
<paragraph id="714636CEFFE6FFA926CBFC9FBEB939E7" blockId="7.[151,1436,151,1184]" pageId="7" pageNumber="152">
Right sexual tube (
<figureCitation id="E9C22A4BFFE6FFA92790FC9FBE823934" box="[412,489,800,824]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="7" pageNumber="152">Fig. 1I</figureCitation>
) medium in length, coiled, oriented laterally, tapering distally; tip roundly truncate; basal portion with covering of numerous short setae. Left sexual tube (
<figureCitation id="E9C22A4BFFE6FFA925D0FCFBBB473950" box="[988,1068,836,860]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="7" pageNumber="152">Fig. 1I</figureCitation>
) short, but distinctly extending beyond posterior margin of coxa, somewhat flattened, directed posterolaterally, tapering distally to acute tip; numerous short setae present on dorsal side. Coxae of fifth pereopods (
<figureCitation id="E9C22A4BFFE6FFA925A3FC33BC9139A8" box="[943,1018,908,932]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="7" pageNumber="152">Fig. 1I</figureCitation>
) unequal (left larger than right), each with transverse row of setae on anterior margin. Eighth thoracic sternite (
<figureCitation id="E9C22A4BFFE6FFA925D3FC0FBB4739C4" box="[991,1068,944,968]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="7" pageNumber="152">Fig. 1I</figureCitation>
) with 2 compressed lobes, each lobe bearing marginal setae.
</paragraph>
<paragraph id="714636CEFFE6FFA926CBFC47BD243E38" blockId="7.[151,1436,151,1184]" pageId="7" pageNumber="152">
Pleon dextrally twisted (
<figureCitation id="E9C22A4BFFE6FFA927E6FC47BD5E3E1C" box="[490,565,1016,1040]" captionStart="FIGURE 6" captionStartId="7.[151,250,1929,1952]" captionTargetBox="[348,1239,1237,1908]" captionTargetId="figure-460@7.[348,1239,1237,1908]" captionTargetPageId="7" captionText="FIGURE 6. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Entire animal in dorsal view, showing colouration in life. Photo: Tin-Yam Chan." figureDoi="http://doi.org/10.5281/zenodo.5252022" httpUri="https://zenodo.org/record/5252022/files/figure.png" pageId="7" pageNumber="152">Fig. 6</figureCitation>
), with relatively long, unequally biramous third to fifth pleopods; fourth pleopod longest, with elongate exopod.
</paragraph>
<paragraph id="714636CEFFE6FFA926CBFBFFBEF63E93" blockId="7.[151,1436,151,1184]" pageId="7" pageNumber="152">
Telson (
<figureCitation id="E9C22A4BFFE6FFA9272EFBFFBEFE3E54" box="[290,405,1088,1112]" captionStart="FIGURE 1" captionStartId="2.[151,250,1838,1861]" captionTargetBox="[251,1348,185,1803]" captionTargetId="figure-131@2.[228,1370,179,1835]" captionTargetPageId="2" captionText="FIGURE 1. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. A, shield and cephalic appendages, dorsal view; B, carapace, dorsal view; C, basal segment of left antennular peduncle, ventral view; D, proximal part of left antennal flagellum, dorsal view; E, left third maxilliped, lateral view; F, same, basis and ischium, ventral view; G, left fourth pereopod, lateral view; H, sixth thoracic sternite, ventral view; I, eighth thoracic sternite and coxae of fifth pereopods with sexual tubes, ventral view; J, telson, dorsal view; K, posterior lobe of telson, posterior view. Scales: A, B, D = 1 mm; C, E–K = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5252004" httpUri="https://zenodo.org/record/5252004/files/figure.png" pageId="7" pageNumber="152">Fig. 1J, K</figureCitation>
) with slightly asymmetrical posterior lobes separated by very small median cleft; terminal margins strongly oblique, microscopically denticulate; transverse indentations weakly delimited, each with prominent tuft of setae.
</paragraph>
<caption id="25866646FFE6FFA9269BF836BDBD3DB2" ID-DOI="http://doi.org/10.5281/zenodo.5252022" ID-Zenodo-Dep="5252022" httpUri="https://zenodo.org/record/5252022/files/figure.png" pageId="7" pageNumber="152" startId="7.[151,250,1929,1952]" targetBox="[348,1239,1237,1908]" targetPageId="7">
<paragraph id="714636CEFFE6FFA9269BF836BDBD3DB2" blockId="7.[151,1436,1929,1982]" pageId="7" pageNumber="152">
<emphasis id="438DEADCFFE6FFA9269BF836BE783DAC" bold="true" box="[151,275,1929,1952]" pageId="7" pageNumber="152">FIGURE 6.</emphasis>
<taxonomicName id="B6F94D4DFFE6FFA92717F834BD5A3DAC" authority="Komai &amp; Osawa, 2012" authorityName="Komai &amp; Osawa" authorityYear="2012" box="[283,561,1930,1952]" class="Malacostraca" family="Paguridae" genus="Kumepagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="152" phylum="Arthropoda" rank="species" species="cavernicolus" status="n. gen., n. sp.">
<emphasis id="438DEADCFFE6FFA92717F834BD5A3DAC" box="[283,561,1930,1952]" italics="true" pageId="7" pageNumber="152">Kumepagurus cavernicolus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="58BE57A7FFE6FFA92434F835BDA33D93" box="[568,712,1930,1951]" pageId="7" pageNumber="152" rank="species">
<emphasis id="438DEADCFFE6FFA92434F835BDEB3D93" bold="true" box="[568,640,1930,1951]" pageId="7" pageNumber="152">n. gen.</emphasis>
,
<emphasis id="438DEADCFFE6FFA92482F835BDA33D93" bold="true" box="[654,712,1930,1951]" pageId="7" pageNumber="152">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype, male (sl 2.4 mm), RUMF-ZC-1518. Entire animal in dorsal view, showing colouration in life. Photo: Tin-Yam Chan.
</paragraph>
</caption>
<paragraph id="714636CEFFE9FFA626CBFF28BCBE3BF8" blockId="8.[151,1437,151,1040]" pageId="8" pageNumber="153">
<emphasis id="438DEADCFFE9FFA626CBFF28BD6C3AA3" box="[199,519,151,176]" inLineHeading="true" pageId="8" pageNumber="153" reason="1">
<emphasis id="438DEADCFFE9FFA626CBFF28BEC83ABC" bold="true" box="[199,419,151,176]" pageId="8" pageNumber="153">Colouration in life</emphasis>
(
<figureCitation id="E9C22A4BFFE9FFA627B8FF28BE923AA3" box="[436,505,151,175]" captionStart="FIGURE 6" captionStartId="7.[151,250,1929,1952]" captionTargetBox="[348,1239,1237,1908]" captionTargetId="figure-460@7.[348,1239,1237,1908]" captionTargetPageId="7" captionText="FIGURE 6. Kumepagurus cavernicolus n. gen., n. sp., holotype, male (sl 2.4 mm), RUMF-ZC-1518. Entire animal in dorsal view, showing colouration in life. Photo: Tin-Yam Chan." figureDoi="http://doi.org/10.5281/zenodo.5252022" httpUri="https://zenodo.org/record/5252022/files/figure.png" pageId="8" pageNumber="153">Fig. 6</figureCitation>
).
</emphasis>
Shield generally white, reddish gut inside visible; posterior carapace translucent. Ocular peduncle generally white, with reddish-brown longitudinal stripe mesially and laterally on dorsal surface; cornea gray. Antennular peduncle translucent, with tinge of reddish-brown. Antennal peduncle with white first to third segments and reddish-brown fourth and fifth segments; antennal acicle white. Right chela generally pale brown, darker mesially and laterally, mesial crest white; carpus light brown, with white patches laterally and mesially on dorsal surface; merus generally light brown, lateral face with large white patch. Left chela generally light brown, whitish on lateral side of dorsal surface of palm. Dactyli of ambulatory legs white in distal 0.6–0.7, red in proximal 0.3–0.4; propodi and carpi white (on the photo, the area appears blue, but this is due to the lighting), lateral faces with red longitudinal stripes dorsally and ventrally; meri with distal and middle red bands and small proximal red spot, remainder white; ischia generally white. Pleon reddish.
</paragraph>
</subSubSection>
<subSubSection id="39E36545FFE9FFA626CBFE40BA51381B" box="[199,1338,511,536]" pageId="8" pageNumber="153" type="distribution">
<paragraph id="714636CEFFE9FFA626CBFE40BA51381B" blockId="8.[151,1437,151,1040]" box="[199,1338,511,536]" pageId="8" pageNumber="153">
<emphasis id="438DEADCFFE9FFA626CBFE40BE0B381B" box="[199,352,511,536]" inLineHeading="true" pageId="8" pageNumber="153" reason="1">
<emphasis id="438DEADCFFE9FFA626CBFE40BE323814" bold="true" box="[199,345,511,536]" pageId="8" pageNumber="153">Distribution</emphasis>
.
</emphasis>
Known only from the
<typeStatus id="AE42886CFFE9FFA6246CFDBFBDFB3814" box="[608,656,512,536]" pageId="8" pageNumber="153">type</typeStatus>
locality, submarine cave in Kume Island, at a depth of
<specimenCount id="67FFFD47FFE9FFA622F6FE40BA51381B" box="[1274,1338,511,535]" pageId="8" pageNumber="153" type="generic">
<quantity id="B6019B2BFFE9FFA622F6FE40BA5D381B" box="[1274,1334,511,535]" metricMagnitude="1" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="153" unit="m" value="38.0">38 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="39E36545FFE9FFA626CBFD9BBD9B39C4" pageId="8" pageNumber="153" type="discussion">
<paragraph id="714636CEFFE9FFA626CBFD9BBD9B39C4" blockId="8.[151,1437,151,1040]" pageId="8" pageNumber="153">
<emphasis id="438DEADCFFE9FFA626CBFD9BBE503837" box="[199,315,548,572]" inLineHeading="true" pageId="8" pageNumber="153" reason="1">
<emphasis id="438DEADCFFE9FFA626CBFD9BBE5F3830" bold="true" box="[199,308,548,572]" pageId="8" pageNumber="153">Remarks</emphasis>
.
</emphasis>
To date, the following 11 hermit crab species have been recorded from marine caves in the Ryukyu Islands:
<taxonomicName id="B6F94D4DFFE9FFA626FAFDF7BD333853" authority="Morgan, 1992" authorityName="Morgan" authorityYear="1992" box="[246,600,583,608]" class="Malacostraca" family="Diogenidae" genus="Paguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="jalur">
<emphasis id="438DEADCFFE9FFA626FAFDF7BEC6386C" box="[246,429,584,608]" italics="true" pageId="8" pageNumber="153">Paguristes jalur</emphasis>
<bibRefCitation id="15684B3FFFE9FFA627BAFDF7BD333853" author="Morgan, G. J." box="[438,600,583,608]" pageId="8" pageNumber="153" pagination="163 - 174" refId="ref4502" refString="Morgan, G. J. (1992) The hermit crabs (Crustacea: Decapoda: Coenobitidae, Diogenidae, Paguridae) of Christmas and Cocos (Keeling) Islands, Indian Ocean, with description of a new species of Paguristes. Raffles Bulletin of Zoology, 40, 163 - 174." type="journal article" year="1992">Morgan, 1992</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="B6F94D4DFFE9FFA6246AFDF7BB283853" authority="McLaughlin, 2002" authorityName="McLaughlin" authorityYear="2002" box="[614,1091,583,608]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="foresti">
<emphasis id="438DEADCFFE9FFA6246AFDF7BC03386C" box="[614,872,584,608]" italics="true" pageId="8" pageNumber="153">Catapaguroides foresti</emphasis>
<bibRefCitation id="15684B3FFFE9FFA6257DFDF8BB283853" author="McLaughlin, P. A." box="[881,1091,583,607]" pageId="8" pageNumber="153" pagination="495 - 504" refId="ref4308" refString="McLaughlin, P. A. (2002) A new species of the genus Catapaguroides (Decapoda, Anomura, Paguroidea, Paguridae) from Guam, Micronesia. Crustaceana, 75, 495 - 504." type="journal article" year="2002">McLaughlin, 2002</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="B6F94D4DFFE9FFA62261FDF7BFBF3888" authority="Osawa &amp; Takeda, 2004" authorityName="Osawa &amp; Takeda" authorityYear="2004" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="kasei">
<emphasis id="438DEADCFFE9FFA62261FDF7BBA3386C" box="[1133,1224,584,608]" italics="true" pageId="8" pageNumber="153">C. kasei</emphasis>
<bibRefCitation id="15684B3FFFE9FFA622DDFDF8BFBF3888" author="Osawa, M. &amp; Takeda, M." pageId="8" pageNumber="153" pagination="1097 - 1132" refId="ref4659" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38, 1097 - 1132." type="journal article" year="2004">Osawa &amp; Takeda, 2004</bibRefCitation>
</taxonomicName>
);
<taxonomicName id="B6F94D4DFFE9FFA626E1FDD3BDC53888" authority="Osawa &amp; Takeda, 2004" authorityName="Osawa &amp; Takeda" authorityYear="2004" box="[237,686,620,644]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="iejimensis">
<emphasis id="438DEADCFFE9FFA626E1FDD3BEEF3888" box="[237,388,620,644]" italics="true" pageId="8" pageNumber="153">C. iejimensis</emphasis>
<bibRefCitation id="15684B3FFFE9FFA6279EFDD3BDC53888" author="Osawa, M. &amp; Takeda, M." box="[402,686,620,644]" pageId="8" pageNumber="153" pagination="1097 - 1132" refId="ref4659" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38, 1097 - 1132." type="journal article" year="2004">Osawa &amp; Takeda, 2004</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="B6F94D4DFFE9FFA624CCFDD3BB923888" authority="Komai, Yamada &amp; Shirakawa, 2010" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[704,1273,620,644]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="438DEADCFFE9FFA624CCFDD3BC523888" box="[704,825,620,644]" italics="true" pageId="8" pageNumber="153">C. longior</emphasis>
<bibRefCitation id="15684B3FFFE9FFA6254BFDD3BB923888" author="Komai, T. &amp; Yamada, Y. &amp; Shirakawa, N." box="[839,1273,620,644]" pageId="8" pageNumber="153" pagination="32 - 42" refId="ref4246" refString="Komai, T., Yamada, Y. &amp; Shirakawa, N. (2010) A new hermit crab species of the genus Catapaguroides A. Milne-Edwards &amp; Bouvier (Decapoda: Anomura: Paguridae) from the Ryukyu Islands, Japan, and additional record of C. foresti McLaughlin. Zootaxa, 2690, 32 - 42." type="journal article" year="2010">Komai, Yamada &amp; Shirakawa, 2010</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="B6F94D4DFFE9FFA62306FDD3BE8338A4" authority="(Asakura, 1999)" baseAuthorityName="Asakura" baseAuthorityYear="1999" class="Malacostraca" family="Paguridae" genus="Catapagurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="tuberculosus">
<emphasis id="438DEADCFFE9FFA62306FDD3BE4E38A4" italics="true" pageId="8" pageNumber="153">Catapagurus tuberculosus</emphasis>
(
<bibRefCitation id="15684B3FFFE9FFA62739FD2FBE8B38A4" author="Asakura, A." box="[309,480,656,680]" pageId="8" pageNumber="153" pagination="381 - 395" refId="ref4113" refString="Asakura, A. (1999) Icelopagurus tuberculosus, a distinct new hermit crab species (Crustacea: Decapoda: Paguridae) form Japan. Proceedings of the Biological Society of Washington, 112, 381 - 395." type="journal article" year="1999">Asakura, 1999</bibRefCitation>
)
</taxonomicName>
;
<taxonomicName id="B6F94D4DFFE9FFA627F9FD2FBB5F38A4" authority="Osawa, Fujita &amp; Okuno, 2006" authorityName="Osawa, Fujita &amp; Okuno" authorityYear="2006" box="[501,1076,656,680]" class="Malacostraca" family="Paguridae" genus="Pagurixus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="438DEADCFFE9FFA627F9FD2FBDBB38A4" box="[501,720,656,680]" italics="true" pageId="8" pageNumber="153">Pagurixus longipes</emphasis>
<bibRefCitation id="15684B3FFFE9FFA624D4FD2FBB5F38A4" author="Osawa, M. &amp; Fujita, Y. &amp; Okuno, J." box="[728,1076,656,680]" pageId="8" pageNumber="153" pagination="27 - 45" refId="ref4557" refString="Osawa, M., Fujita, Y. &amp; Okuno, J. (2006) Two new species of Pagurixus (Crustacea: Decapoda: Anomura: Paguridae) from submarine caves of the Ryukyu Islands, southwestern Japan. Zootaxa, 1148, 27 - 45." type="journal article" year="2006">Osawa, Fujita &amp; Okuno, 2006</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="B6F94D4DFFE9FFA6224CFD2EBFB938C0" authority="Komai &amp; Asakura, 1995" authorityName="Komai &amp; Asakura" authorityYear="1995" class="Malacostraca" family="Paguridae" genus="Pagurixus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="nomurai">
<emphasis id="438DEADCFFE9FFA6224CFD2EBBD738AB" box="[1088,1212,656,680]" italics="true" pageId="8" pageNumber="153">P. nomurai</emphasis>
<bibRefCitation id="15684B3FFFE9FFA622C8FD2FBFB938C0" author="Komai, T. &amp; Asakura, A." pageId="8" pageNumber="153" pagination="341 - 354" refId="ref4187" refString="Komai, T. &amp; Asakura, A. (1995) Pagurixus nomurai, new species, and additional record of Pagurixus maorus (Nobili, 1906), hermit crabs from Kume-jima Island, the Ryukyus, Japan (Decapoda: Anomura: Paguridae). Journal of Crustacean Biology, 15, 341 - 354." type="journal article" year="1995">Komai &amp; Asakura, 1995</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="B6F94D4DFFE9FFA626EEFD0ABDA038C0" authority="Osawa, Fujita &amp; Okuno, 2006" authorityName="Osawa, Fujita &amp; Okuno" authorityYear="2006" box="[226,715,692,716]" class="Malacostraca" family="Paguridae" genus="Pagurixus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="pulcher">
<emphasis id="438DEADCFFE9FFA626EEFD0ABE3338C0" box="[226,344,692,716]" italics="true" pageId="8" pageNumber="153">P. pulcher</emphasis>
<bibRefCitation id="15684B3FFFE9FFA62768FD0BBDA038C0" author="Osawa, M. &amp; Fujita, Y. &amp; Okuno, J." box="[356,715,692,716]" pageId="8" pageNumber="153" pagination="27 - 45" refId="ref4557" refString="Osawa, M., Fujita, Y. &amp; Okuno, J. (2006) Two new species of Pagurixus (Crustacea: Decapoda: Anomura: Paguridae) from submarine caves of the Ryukyu Islands, southwestern Japan. Zootaxa, 1148, 27 - 45." type="journal article" year="2006">Osawa, Fujita &amp; Okuno, 2006</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="B6F94D4DFFE9FFA624D0FD0BBA6838C0" authority="Osawa &amp; Okuno, 2007" authorityName="Osawa &amp; Okuno" authorityYear="2007" box="[732,1283,692,716]" class="Malacostraca" family="Paguridae" genus="Pylopaguropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="bellula">
<emphasis id="438DEADCFFE9FFA624D0FD0BBC8F38C0" box="[732,996,692,716]" italics="true" pageId="8" pageNumber="153">Pylopaguropsis bellula</emphasis>
<bibRefCitation id="15684B3FFFE9FFA625FCFD0BBA6838C0" author="Osawa, M. &amp; Okuno, J." box="[1008,1283,692,716]" pageId="8" pageNumber="153" pagination="29 - 46" refId="ref4607" refString="Osawa, M. &amp; Okuno, J. (2007) New species of the genus Pylopaguropsis (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan, with notes on two poorly known pagurids. Species Diversity, 12, 29 - 46." type="journal article" year="2007">Osawa &amp; Okuno, 2007</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="B6F94D4DFFE9FFA62318FD0ABEA638FC" authority="McLaughlin &amp; Haig, 1989" authorityName="McLaughlin &amp; Haig" authorityYear="1989" class="Malacostraca" family="Paguridae" genus="Pylopaguropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="fimbriata">
<emphasis id="438DEADCFFE9FFA62318FD0ABAF738C0" box="[1300,1436,692,716]" italics="true" pageId="8" pageNumber="153">P. fimbriata</emphasis>
<bibRefCitation id="15684B3FFFE9FFA6269BFD67BEA638FC" author="McLaughlin, P. A. &amp; Haig, J." box="[151,461,728,752]" pageId="8" pageNumber="153" pagination="123 - 171" refId="ref4391" refString="McLaughlin, P. A. &amp; Haig, J. (1989) On the status of Pylopaguropsis zebra (Henderson), P. magnimanus (Henderson), and Galapagurus teevanus Boone, with descriptions of seven new species of Pylopaguropsis (Crustacea: Anomura: Paguridae). Micronesica, 22, 123 - 171." type="journal article" year="1989">McLaughlin &amp; Haig, 1989</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="B6F94D4DFFE9FFA627D7FD66BCFE38FC" authority="McLaughlin &amp; Haig, 1989" authorityName="McLaughlin &amp; Haig" authorityYear="1989" box="[475,917,728,752]" class="Malacostraca" family="Paguridae" genus="Pylopaguropsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="438DEADCFFE9FFA627D7FD66BD3238E3" box="[475,601,728,752]" italics="true" pageId="8" pageNumber="153">P. speciosa</emphasis>
<bibRefCitation id="15684B3FFFE9FFA6246FFD67BCFE38FC" author="McLaughlin, P. A. &amp; Haig, J." box="[611,917,728,752]" pageId="8" pageNumber="153" pagination="123 - 171" refId="ref4391" refString="McLaughlin, P. A. &amp; Haig, J. (1989) On the status of Pylopaguropsis zebra (Henderson), P. magnimanus (Henderson), and Galapagurus teevanus Boone, with descriptions of seven new species of Pylopaguropsis (Crustacea: Anomura: Paguridae). Micronesica, 22, 123 - 171." type="journal article" year="1989">McLaughlin &amp; Haig, 1989</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="15684B3FFFE9FFA625ABFD67BBD838FC" author="Osawa, M. &amp; Takeda, M." box="[935,1203,728,752]" pageId="8" pageNumber="153" pagination="1097 - 1132" refId="ref4659" refString="Osawa, M. &amp; Takeda, M. (2004) Hermit crabs (Crustacea: Decapoda: Anomura: Paguroidea) from submarine caves in the Ryukyu Islands, south-western Japan. Journal of Natural History, 38, 1097 - 1132." type="journal article" year="2004">Osawa &amp; Takeda 2004</bibRefCitation>
;
<bibRefCitation id="15684B3FFFE9FFA622CEFD67BAFC38FC" author="Osawa, M. &amp; Fujita, Y. &amp; Okuno, J." box="[1218,1431,728,752]" pageId="8" pageNumber="153" pagination="27 - 45" refId="ref4557" refString="Osawa, M., Fujita, Y. &amp; Okuno, J. (2006) Two new species of Pagurixus (Crustacea: Decapoda: Anomura: Paguridae) from submarine caves of the Ryukyu Islands, southwestern Japan. Zootaxa, 1148, 27 - 45." type="journal article" year="2006">
Osawa
<emphasis id="438DEADCFFE9FFA62314FD66BA2038FC" box="[1304,1355,728,752]" italics="true" pageId="8" pageNumber="153">et al</emphasis>
. 2006
</bibRefCitation>
;
<bibRefCitation id="15684B3FFFE9FFA6269BFD43BEF23918" author="Osawa, M. &amp; Okuno, J." box="[151,409,764,788]" pageId="8" pageNumber="153" pagination="29 - 46" refId="ref4607" refString="Osawa, M. &amp; Okuno, J. (2007) New species of the genus Pylopaguropsis (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan, with notes on two poorly known pagurids. Species Diversity, 12, 29 - 46." type="journal article" year="2007">Osawa &amp; Okuno 2007</bibRefCitation>
;
<bibRefCitation id="15684B3FFFE9FFA627A9FD43BD133918" author="Komai, T. &amp; Yamada, Y. &amp; Shirakawa, N." box="[421,632,764,788]" pageId="8" pageNumber="153" pagination="32 - 42" refId="ref4246" refString="Komai, T., Yamada, Y. &amp; Shirakawa, N. (2010) A new hermit crab species of the genus Catapaguroides A. Milne-Edwards &amp; Bouvier (Decapoda: Anomura: Paguridae) from the Ryukyu Islands, Japan, and additional record of C. foresti McLaughlin. Zootaxa, 2690, 32 - 42." type="journal article" year="2010">
Komai
<emphasis id="438DEADCFFE9FFA627F6FD42BD473918" box="[506,556,764,788]" italics="true" pageId="8" pageNumber="153">et al</emphasis>
. 2010
</bibRefCitation>
). However, all but
<taxonomicName id="B6F94D4DFFE9FFA6255CFD43BB353918" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[848,1118,764,788]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="438DEADCFFE9FFA6255CFD43BB353918" box="[848,1118,764,788]" italics="true" pageId="8" pageNumber="153">Catapaguroides longior</emphasis>
</taxonomicName>
occur in cryptic but normal non-cave habitats (
<bibRefCitation id="15684B3FFFE9FFA62767FC9FBDFE3934" author="McLaughlin, P. A. &amp; Haig, J." box="[363,661,800,824]" pageId="8" pageNumber="153" pagination="123 - 171" refId="ref4391" refString="McLaughlin, P. A. &amp; Haig, J. (1989) On the status of Pylopaguropsis zebra (Henderson), P. magnimanus (Henderson), and Galapagurus teevanus Boone, with descriptions of seven new species of Pylopaguropsis (Crustacea: Anomura: Paguridae). Micronesica, 22, 123 - 171." type="journal article" year="1989">McLaughlin &amp; Haig 1989</bibRefCitation>
;
<bibRefCitation id="15684B3FFFE9FFA624ADFC9FBC573934" author="Morgan, G. J." box="[673,828,800,824]" pageId="8" pageNumber="153" pagination="163 - 174" refId="ref4502" refString="Morgan, G. J. (1992) The hermit crabs (Crustacea: Decapoda: Coenobitidae, Diogenidae, Paguridae) of Christmas and Cocos (Keeling) Islands, Indian Ocean, with description of a new species of Paguristes. Raffles Bulletin of Zoology, 40, 163 - 174." type="journal article" year="1992">Morgan 1992</bibRefCitation>
;
<bibRefCitation id="15684B3FFFE9FFA62544FC9FBB323934" author="Komai, T. &amp; Asakura, A." box="[840,1113,800,824]" pageId="8" pageNumber="153" pagination="341 - 354" refId="ref4187" refString="Komai, T. &amp; Asakura, A. (1995) Pagurixus nomurai, new species, and additional record of Pagurixus maorus (Nobili, 1906), hermit crabs from Kume-jima Island, the Ryukyus, Japan (Decapoda: Anomura: Paguridae). Journal of Crustacean Biology, 15, 341 - 354." type="journal article" year="1995">Komai &amp; Asakura 1995</bibRefCitation>
;
<bibRefCitation id="15684B3FFFE9FFA6226AFC9FBA6D3934" author="Asakura, A." box="[1126,1286,800,824]" pageId="8" pageNumber="153" pagination="381 - 395" refId="ref4113" refString="Asakura, A. (1999) Icelopagurus tuberculosus, a distinct new hermit crab species (Crustacea: Decapoda: Paguridae) form Japan. Proceedings of the Biological Society of Washington, 112, 381 - 395." type="journal article" year="1999">Asakura 1999</bibRefCitation>
;
<bibRefCitation id="15684B3FFFE9FFA6231EFC9FBFB93950" author="McLaughlin, P. A." pageId="8" pageNumber="153" pagination="495 - 504" refId="ref4308" refString="McLaughlin, P. A. (2002) A new species of the genus Catapaguroides (Decapoda, Anomura, Paguroidea, Paguridae) from Guam, Micronesia. Crustaceana, 75, 495 - 504." type="journal article" year="2002">McLaughlin 2002</bibRefCitation>
;
<bibRefCitation id="15684B3FFFE9FFA626D2FCFBBEB53950" author="Osawa, M. &amp; Okuno, J." box="[222,478,836,860]" pageId="8" pageNumber="153" pagination="29 - 46" refId="ref4607" refString="Osawa, M. &amp; Okuno, J. (2007) New species of the genus Pylopaguropsis (Crustacea: Decapoda: Anomura: Paguridae) from the Ryukyu Islands, southwestern Japan, with notes on two poorly known pagurids. Species Diversity, 12, 29 - 46." type="journal article" year="2007">Osawa &amp; Okuno 2007</bibRefCitation>
; unpublished data), and therefore it is considered that the occurrence of these species in cave environments is rather opportunistic.
<taxonomicName id="B6F94D4DFFE9FFA6249DFCD7BCF5398C" authorityName="Komai, Yamada &amp; Shirakawa" authorityYear="2010" box="[657,926,872,896]" class="Malacostraca" family="Paguridae" genus="Catapaguroides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="153" phylum="Arthropoda" rank="species" species="longior">
<emphasis id="438DEADCFFE9FFA6249DFCD7BCF5398C" box="[657,926,872,896]" italics="true" pageId="8" pageNumber="153">Catapaguroides longior</emphasis>
</taxonomicName>
and the present new species have been found only in deep chambers of submarine caves, despite extensive collecting efforts in recent years. It is possible that these two species are restricted to cave environments.
</paragraph>
</subSubSection>
<subSubSection id="39E36545FFE9FFA626CBFC6BBD673E1C" pageId="8" pageNumber="153" type="etymology">
<paragraph id="714636CEFFE9FFA626CBFC6BBD673E1C" blockId="8.[151,1437,151,1040]" pageId="8" pageNumber="153">
<emphasis id="438DEADCFFE9FFA626CBFC6BBE2639E7" box="[199,333,980,1004]" inLineHeading="true" pageId="8" pageNumber="153" reason="1">
<emphasis id="438DEADCFFE9FFA626CBFC6BBE2D39E0" bold="true" box="[199,326,980,1004]" pageId="8" pageNumber="153">Etymology</emphasis>
.
</emphasis>
The species name is a combination of the Latin,
<emphasis id="438DEADCFFE9FFA6257FFC6ABCA639E0" box="[883,973,981,1004]" italics="true" pageId="8" pageNumber="153">caverna</emphasis>
(= cave) and
<emphasis id="438DEADCFFE9FFA6226BFC6BBBFC39E0" box="[1127,1175,980,1004]" italics="true" pageId="8" pageNumber="153">cola</emphasis>
(= living), in reference to the habitat of this new species.
</paragraph>
</subSubSection>
</treatment>
</document>
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