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<mods:title id="A4585D93997ACAF22700F45F5E1E0E14">Revision of the new world species of Hylurgops LeConte, 1876 with the description of a new genus in the Hylastini (Coleoptera: Scolytinae) and comments on some Palearctic species</mods:title>
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<mods:namePart id="ECA445A8647F86D415B280F8CC9CBFC6">Mercado-Vélez, Javier E.</mods:namePart>
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<treatment id="03EADA36FFFB332603E5FCA85C97F84C" ID-DOI="http://doi.org/10.5281/zenodo.5691413" ID-GBIF-Taxon="119421844" ID-Zenodo-Dep="5691413" LSID="urn:lsid:plazi:treatment:03EADA36FFFB332603E5FCA85C97F84C" httpUri="http://treatment.plazi.org/id/03EADA36FFFB332603E5FCA85C97F84C" lastPageId="27" lastPageNumber="328" pageId="25" pageNumber="326">
<subSubSection id="C35938ABFFFB332403E5FCA85EA1FC64" pageId="25" pageNumber="326" type="nomenclature">
<paragraph id="8BFC6B20FFFB332403E5FCA85D3EFC45" blockId="25.[151,639,871,931]" box="[151,639,871,898]" pageId="25" pageNumber="326">
<heading id="D0B4DC4CFFFB332403E5FCA85D3EFC45" bold="true" box="[151,639,871,898]" fontSize="11" level="1" pageId="25" pageNumber="326" reason="1">
<taxonomicName id="4C4310A3FFFB332403E5FCA85D3EFC45" ID-CoL="3NGZG" authority="Blandford, 1897" authorityName="Blandford" authorityYear="1897" box="[151,639,871,898]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="incomptus">
<emphasis id="B937B732FFFB332403E5FCA85D3EFC45" bold="true" box="[151,639,871,898]" pageId="25" pageNumber="326">
<emphasis id="B937B732FFFB332403E5FCA85EDBFC46" bold="true" box="[151,410,871,897]" italics="true" pageId="25" pageNumber="326">Hylurgops incomptus</emphasis>
(
<bibRefCitation id="EFD216D1FFFB332402DBFCA75D36FC45" author="Blandford" box="[425,631,872,898]" pageId="25" pageNumber="326" refString="Blandford, W. F. H. (1897) Scolytidae. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali-Americana. Insecta, Coleoptera, 4 (6), pp. 145 - 184." type="journal article" year="1897">Blandford, 1897</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFC6B20FFFB332403E5FC455EA1FC64" blockId="25.[151,639,871,931]" box="[151,480,906,931]" pageId="25" pageNumber="326">
(
<figureCitation id="137877A5FFFB332403ECFC455E4BFC65" box="[158,266,906,931]" captionStart="FIGURE 7" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[199,1387,194,1893]" captionTargetId="figure@9.[190,1396,193,1897]" captionTargetPageId="9" captionText="FIGURE 7. Aedeagus of the New World Hylurgops and Pachysquamus: a) Pachysquamus subcostulatus, b) Hylurgops planirostris, c) H. rugipennis, d) H. pinifex, e) H. incomptus, f) H. longipennis, g) H. knausi, h) H. reticulatus, and i) H. porosus." httpUri="https://zenodo.org/record/252621/files/figure.png" pageId="25" pageNumber="326">Figures 7</figureCitation>
e, 15e, 17e, 20, 21)
</paragraph>
</subSubSection>
<subSubSection id="C35938ABFFFB332403E5FC1D5EFBFB45" pageId="25" pageNumber="326" type="reference_group">
<paragraph id="8BFC6B20FFFB332403E5FC1D5D33FBC0" blockId="25.[151,1435,978,1154]" pageId="25" pageNumber="326">
<treatmentCitationGroup id="AB534C0EFFFB332403E5FC1D5D33FBC0" pageId="25" pageNumber="326">
<taxonomicName id="4C4310A3FFFB332403E5FC1D5D0BFC2F" ID-CoL="6MMVV" authority="Blandford, 1897:145" authorityName="Blandford" authorityPageNumber="145" authorityYear="1897" box="[151,586,978,1000]" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="incomptus">
<emphasis id="B937B732FFFB332403E5FC1D5E26FC2F" bold="true" box="[151,359,978,1000]" italics="true" pageId="25" pageNumber="326">Hylastes incomptus</emphasis>
<treatmentCitation id="0AE24D31FFFB33240200FC1D5D0BFC2F" author="Blandford" box="[370,586,978,1000]" page="145" pageId="25" pageNumber="326" year="1897">
<bibRefCitation id="EFD216D1FFFB33240200FC1D5D61FC2F" author="Blandford" box="[370,544,978,1000]" pageId="25" pageNumber="326" refString="Blandford, W. F. H. (1897) Scolytidae. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali-Americana. Insecta, Coleoptera, 4 (6), pp. 145 - 184." type="journal article" year="1897">Blandford, 1897</bibRefCitation>
:145
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</taxonomicName>
(
<typeStatus id="54F8D582FFFB3324012FFC1D5DFEFC2F" box="[605,703,978,1000]" pageId="25" pageNumber="326" type="syntype">Syntypes</typeStatus>
: Andres Chalchicomula, now Ciudad Serdan in Puebla,
<collectingCountry id="F3542BB0FFFB3324066DFC1D5A2DFC2F" box="[1311,1388,978,1000]" name="Mexico" pageId="25" pageNumber="326">Mexico</collectingCountry>
(1), Salazar (1), Chilpancingo in Guerrero (1))
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC6B20FFFB332403E5FBDF5D38FBE1" blockId="25.[151,1435,978,1154]" box="[151,633,1039,1062]" pageId="25" pageNumber="326">
<treatmentCitationGroup id="AB534C0EFFFB332403E5FBDF5D38FBE1" box="[151,633,1039,1062]" pageId="25" pageNumber="326">
<taxonomicName id="4C4310A3FFFB332403E5FBDF5D38FBE1" ID-CoL="6MMVV" authority="Hagedorn, 1910:45" authorityName="Hagedorn" authorityPageNumber="45" authorityYear="1910" box="[151,633,1039,1062]" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="incomptus" subGenus="Hylurgops">
<emphasis id="B937B732FFFB332403E5FBDF5EE9FBE2" bold="true" box="[151,424,1039,1062]" pageId="25" pageNumber="326">
<emphasis id="B937B732FFFB332403E5FBDF5FF0FBE2" bold="true" box="[151,177,1040,1061]" italics="true" pageId="25" pageNumber="326">H.</emphasis>
(
<emphasis id="B937B732FFFB332403CDFBC05E6CFBE2" bold="true" box="[191,301,1039,1061]" italics="true" pageId="25" pageNumber="326">Hylurgops</emphasis>
)
<emphasis id="B937B732FFFB3324024EFBC05EE9FBE2" bold="true" box="[316,424,1039,1061]" italics="true" pageId="25" pageNumber="326">incomptus</emphasis>
</emphasis>
,
<treatmentCitation id="0AE24D31FFFB332402C7FBDF5D38FBE1" author="Hagedorn" box="[437,633,1040,1062]" page="45" pageId="25" pageNumber="326" year="1910">
<bibRefCitation id="EFD216D1FFFB332402C7FBDF5D1BFBE1" author="Hagedorn" box="[437,602,1040,1062]" pageId="25" pageNumber="326" refString="Hagedorn, M. (1910) Coleoptera Fam. Ipidae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 111. Brussels, pp. 1 - 178 + XIV plts." type="book chapter" year="1910">Hagedorn, 1910</bibRefCitation>
:45
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</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC6B20FFFB332403E5FBE15D72FB83" blockId="25.[151,1435,978,1154]" box="[151,563,1070,1092]" pageId="25" pageNumber="326">
<treatmentCitationGroup id="AB534C0EFFFB332403E5FBE15D72FB83" box="[151,563,1070,1092]" pageId="25" pageNumber="326">
<taxonomicName id="4C4310A3FFFB332403E5FBE15D72FB83" ID-CoL="3NGZG" authority="Kleine, 1912:166" authorityName="Kleine" authorityPageNumber="166" authorityYear="1912" box="[151,563,1070,1092]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="incomptus">
<emphasis id="B937B732FFFB332403E5FBE15E36FB83" bold="true" box="[151,375,1070,1092]" italics="true" pageId="25" pageNumber="326">Hylurgops incomptus</emphasis>
,
<treatmentCitation id="0AE24D31FFFB332402F6FBE15D72FB83" author="Kleine" box="[388,563,1070,1092]" page="166" pageId="25" pageNumber="326" year="1912">
<bibRefCitation id="EFD216D1FFFB332402F6FBE15D48FB83" author="Kleine" box="[388,521,1070,1092]" pageId="25" pageNumber="326" refString="Kleine, R. (1912) Die geographische verbreitung der Ipiden-Genera orbis terrarum. (Col.). Berliner Entomololgische Zeitschrift, 57, 155 - 192." type="journal article" year="1912">Kleine, 1912</bibRefCitation>
:166
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC6B20FFFB332403E5FB815C80FBA4" blockId="25.[151,1435,978,1154]" box="[151,961,1101,1123]" pageId="25" pageNumber="326">
<treatmentCitationGroup id="AB534C0EFFFB332403E5FB815C80FBA4" box="[151,961,1101,1123]" pageId="25" pageNumber="326">
<taxonomicName id="4C4310A3FFFB332403E5FB815EA2FBA4" ID-CoL="3NGZD" authority="Swaine, 1917:17" authorityName="Swaine" authorityPageNumber="17" authorityYear="1917" box="[151,483,1101,1123]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="grandicollis">
<emphasis id="B937B732FFFB332403E5FB815E72FBA4" bold="true" box="[151,307,1101,1123]" italics="true" pageId="25" pageNumber="326">H. grandicollis</emphasis>
<treatmentCitation id="0AE24D31FFFB3324024BFB825EA2FBA4" author="Swaine" box="[313,483,1101,1123]" page="17" pageId="25" pageNumber="326" year="1917">
<bibRefCitation id="EFD216D1FFFB3324024BFB825E86FBA4" author="Swaine" box="[313,455,1101,1123]" pageId="25" pageNumber="326" refString="Swaine, J. M. (1917) Canadian bark beetles, Part 1. Description of new species. Canada Department of Agriculture, Entomological Branch, Bulletin, 14 (1), 1 - 32." type="journal article" year="1917">Swaine, 1917</bibRefCitation>
:17
</treatmentCitation>
</taxonomicName>
(Cloudcroft and Santa Fe, New
<collectingCountry id="F3542BB0FFFB33240059FB825C3AFBA4" box="[811,891,1101,1123]" name="Mexico" pageId="25" pageNumber="326">Mexico</collectingCountry>
,
<collectingCountry id="F3542BB0FFFB332400F7FB825CF8FBA4" box="[901,953,1101,1123]" name="United States of America" pageId="25" pageNumber="326">USA</collectingCountry>
)
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC6B20FFFB332403E5FBA35EFBFB45" blockId="25.[151,1435,978,1154]" box="[151,442,1132,1154]" pageId="25" pageNumber="326">
Synonymy:
<bibRefCitation id="EFD216D1FFFB33240260FBA35ECEFB45" author="Wood" box="[274,399,1132,1154]" pageId="25" pageNumber="326" refString="Wood, S. L. (1957) Distributional Notes on and Synonymies of Some North American Scolytidae (Coleoptera). The Canadian Entomologist, 89 (9), 396 - 403. http: // dx. doi. org / 10.4039 / ent 89396 - 9" type="journal article" year="1957">Wood, 1957</bibRefCitation>
:397
</paragraph>
</subSubSection>
<subSubSection id="C35938ABFFFB332403E5FB615D28FAF4" pageId="25" pageNumber="326" type="diagnosis">
<paragraph id="8BFC6B20FFFB332403E5FB615D28FAF4" blockId="25.[151,1437,1198,2021]" pageId="25" pageNumber="326">
<emphasis id="B937B732FFFB332403E5FB615E50FB00" bold="true" box="[151,273,1198,1223]" pageId="25" pageNumber="326">Diagnosis.</emphasis>
<taxonomicName id="4C4310A3FFFB3324026AFB615D49FB01" box="[280,520,1198,1223]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="incomptus">
<emphasis id="B937B732FFFB3324026AFB615D49FB01" box="[280,520,1198,1223]" italics="true" pageId="25" pageNumber="326">Hylurgops incomptus</emphasis>
</taxonomicName>
is distinguished from all other
<taxonomicName id="4C4310A3FFFB33240015FB615C9DFB00" box="[871,988,1198,1223]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus">
<emphasis id="B937B732FFFB33240015FB615C9DFB00" box="[871,988,1198,1223]" italics="true" pageId="25" pageNumber="326">Hylurgops</emphasis>
</taxonomicName>
by the nearly complete acute margin of the sides of its pronotum (
<figureCitation id="137877A5FFFB332402CBFB1D5D4DFB2C" box="[441,524,1234,1259]" captionStart="FIGURE 20" captionStartId="26.[151,250,837,859]" captionTargetBox="[165,1428,375,815]" captionTargetId="figure@26.[156,1430,373,815]" captionTargetPageId="26" captionText="FIGURE 20. Hylurgops incomptus pronotum: a) prothoracic lateral margin, b) pronotum broadening from base (typical form), c) pronotum narrowing from base (Arizona and New Mexico variant)." httpUri="https://zenodo.org/record/252634/files/figure.png" pageId="25" pageNumber="326">Fig. 20</figureCitation>
a). Is distinguished from the sympatric
<taxonomicName id="4C4310A3FFFB332400CCFB1B5B22FB2C" box="[958,1123,1234,1259]" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis id="B937B732FFFB332400CCFB1B5B22FB2C" box="[958,1123,1234,1259]" italics="true" pageId="25" pageNumber="326">H. longipennis</emphasis>
</taxonomicName>
by its larger size, by having a pronotal base almost as broad as the elytral base, by the smaller and more abundant pronotal punctures, and by the long and yellowish hair-like vestiture.
</paragraph>
</subSubSection>
<subSubSection id="C35938ABFFFB332603B5FAF15D37FF3C" lastPageId="27" lastPageNumber="328" pageId="25" pageNumber="326" type="description">
<paragraph id="8BFC6B20FFFB332703B5FAF15C95FE87" blockId="25.[151,1437,1198,2021]" lastBlockId="26.[151,1436,151,320]" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326">
<emphasis id="B937B732FFFB332403B5FAF15ECEFA90" bold="true" box="[199,399,1342,1367]" pageId="25" pageNumber="326">Description. Size</emphasis>
. Length 4.35.5 (avg. 4.9 ± 0.4) mm long, 2.7× longer than wide.
<emphasis id="B937B732FFFB33240700FAF15BF4FA90" bold="true" box="[1138,1205,1342,1367]" pageId="25" pageNumber="326">Color</emphasis>
. Mature adult black.
<emphasis id="B937B732FFFB332403E5FAAD5F9DFABC" bold="true" box="[151,220,1378,1403]" pageId="25" pageNumber="326">Frons</emphasis>
. Transverse impression moderately strong; lower carina distinct, elevated from epistomal margin to convexity below frontal impression, where it bifurcates until reaching it; vestiture hair-like, longer below middle impression, 47× the length of a punctures width.
<emphasis id="B937B732FFFB332401AAFA655C10FA04" bold="true" box="[728,849,1450,1475]" pageId="25" pageNumber="326">Pronotum</emphasis>
. Slightly longer than wide 1.01.1 (1.0 ± 0), widest at base smoothly tapering to basal ⅔ (
<figureCitation id="137877A5FFFB3324012AFA015DEEFA20" box="[600,687,1486,1511]" captionStart="FIGURE 20" captionStartId="26.[151,250,837,859]" captionTargetBox="[165,1428,375,815]" captionTargetId="figure@26.[156,1430,373,815]" captionTargetPageId="26" captionText="FIGURE 20. Hylurgops incomptus pronotum: a) prothoracic lateral margin, b) pronotum broadening from base (typical form), c) pronotum narrowing from base (Arizona and New Mexico variant)." httpUri="https://zenodo.org/record/252634/files/figure.png" pageId="25" pageNumber="326">Fig. 20</figureCitation>
b), anterior ⅓ strongly, roundly narrowing anteriorly; basal ¾ of lateral margin sharply elevated, anterior ¼ less sharply elevated, broadly rounded; middle line indistinct, not raised, indicated by lack of punctures from base to anterior edge, interpuncture surfaces smooth; discal punctures small, deeply impressed, abundant, of two different sizes, large 22.5× size of small; vestiture distinct, length of erect discal setae 36× the width of a large discal puncture, 59× around margins.
<emphasis id="B937B732FFFB3324075CF9915B3BF9B0" bold="true" box="[1070,1146,1630,1655]" pageId="25" pageNumber="326">Elytra</emphasis>
. Anterior margin almost straight, with scattered medium-sized crenulations; strial punctures, small, round; interstriae 23× wider than striae, surface usually glossy but reticulate in some specimens; interstriae with two rows of punctate granules, each with a long, semi erect, hair-like setae from elytral disc to declivity, flanking an erect, thicker and longer (46× length discal puncture diameter) hair-like setae arising from uniseriate granules. Declivital vestiture more abundant than on disc, scale-like setae usually absent, in their place hair-like setae present.
<emphasis id="B937B732FFFB332407F1F8DA5BAAF8E9" bold="true" box="[1155,1259,1813,1838]" pageId="25" pageNumber="326">Declivity</emphasis>
. 2nd interstriae impressed, 3rd not distinctly wider than 2nd and not widened apically, all with pointed granules medially, height up to ½ width of declivital puncture, separated by 1.5× size of discal puncture; surface with four to five rows of minute oval punctures with acuminate serrations on their blunt apices; vestiture consisting of uniseriate row of long, erect, hair-like setae intercalated in two lateral rows of slightly shorter hair-like setae.
<emphasis id="B937B732FFFB332407A2F8675AD5F806" bold="true" box="[1232,1428,1960,1985]" pageId="25" pageNumber="326">Ventral sclerites</emphasis>
. Finely reticulate.
<emphasis id="B937B732FFFB33240212F8035ED9F822" bold="true" box="[352,408,1996,2021]" pageId="25" pageNumber="326">Legs</emphasis>
. Protibiae with 12 large socketed teeth before the angle, meso- and metatibiae with two large socketed teeth each, before the angle; third tarsal segments slightly broader than others.
<emphasis id="B937B732FFF833270656FF585AD5FF77" bold="true" box="[1316,1428,151,176]" pageId="26" pageNumber="327">Variation</emphasis>
. Specimens from southern Arizona and New
<collectingCountry id="F3542BB0FFF8332701D9FF735C45FF12" box="[683,772,188,213]" name="Mexico" pageId="26" pageNumber="327">Mexico</collectingCountry>
. Pronotum smoothly broadening on basal ⅔ (
<figureCitation id="137877A5FFF83327065BFF735AC3FF13" box="[1321,1410,188,213]" captionStart="FIGURE 20" captionStartId="26.[151,250,837,859]" captionTargetBox="[165,1428,375,815]" captionTargetId="figure@26.[156,1430,373,815]" captionTargetPageId="26" captionText="FIGURE 20. Hylurgops incomptus pronotum: a) prothoracic lateral margin, b) pronotum broadening from base (typical form), c) pronotum narrowing from base (Arizona and New Mexico variant)." httpUri="https://zenodo.org/record/252634/files/figure.png" pageId="26" pageNumber="327">Fig. 20</figureCitation>
c), interpuncture surfaces reticulate. Elytral disc always reticulate, the long hair-like setae bearing granulate lateral rows on interstriae and declivity absent, bearing scale-like setae on declivity. Specimens from Durango,
<collectingCountry id="F3542BB0FFF83327066CFECB5A32FEDA" box="[1310,1395,260,285]" name="Mexico" pageId="26" pageNumber="327">Mexico</collectingCountry>
are intermediate (see discussion).
<emphasis id="B937B732FFF833270298FEE85D1CFE87" bold="true" box="[490,605,295,320]" pageId="26" pageNumber="327">Aedeagus</emphasis>
. Lacking a ventral lobe (
<figureCitation id="137877A5FFF833270007FEE85CF8FE87" box="[885,953,295,320]" captionStart="FIGURE 7" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[199,1387,194,1893]" captionTargetId="figure@9.[190,1396,193,1897]" captionTargetPageId="9" captionText="FIGURE 7. Aedeagus of the New World Hylurgops and Pachysquamus: a) Pachysquamus subcostulatus, b) Hylurgops planirostris, c) H. rugipennis, d) H. pinifex, e) H. incomptus, f) H. longipennis, g) H. knausi, h) H. reticulatus, and i) H. porosus." httpUri="https://zenodo.org/record/252621/files/figure.png" pageId="26" pageNumber="327">Fig. 7</figureCitation>
e).
</paragraph>
<caption id="DF3C3BA8FFF8332703E5FC8A5C1CFCBE" httpUri="https://zenodo.org/record/252634/files/figure.png" pageId="26" pageNumber="327" targetBox="[165,1428,375,815]" targetPageId="26">
<paragraph id="8BFC6B20FFF8332703E5FC8A5C1CFCBE" blockId="26.[151,1436,836,889]" pageId="26" pageNumber="327">
<emphasis id="B937B732FFF8332703E5FC8A5E5EFC9D" bold="true" box="[151,287,837,859]" pageId="26" pageNumber="327">FIGURE 20.</emphasis>
<taxonomicName id="4C4310A3FFF833270257FC8B5EBFFC9C" box="[293,510,836,859]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="incomptus">
<emphasis id="B937B732FFF833270257FC8B5EBFFC9C" box="[293,510,836,859]" italics="true" pageId="26" pageNumber="327">Hylurgops incomptus</emphasis>
</taxonomicName>
pronotum: a) prothoracic lateral margin, b) pronotum broadening from base (typical form), c) pronotum narrowing from base (Arizona and New Mexico variant).
</paragraph>
</caption>
<caption id="DF3C3BA8FFF8332703E5F84A5EAFF81F" httpUri="https://zenodo.org/record/252635/files/figure.png" pageId="26" pageNumber="327" targetBox="[152,1436,940,1902]" targetPageId="26">
<paragraph id="8BFC6B20FFF8332703E5F84A5EAFF81F" blockId="26.[151,1436,1924,2008]" pageId="26" pageNumber="327">
<emphasis id="B937B732FFF8332703E5F84A5E5EF85D" bold="true" box="[151,287,1925,1947]" pageId="26" pageNumber="327">FIGURE 21.</emphasis>
<taxonomicName id="4C4310A3FFF833270254F84B5EBFF85C" box="[294,510,1924,1947]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="incomptus">
<emphasis id="B937B732FFF833270254F84B5EBFF85C" box="[294,510,1924,1947]" italics="true" pageId="26" pageNumber="327">Hylurgops incomptus</emphasis>
</taxonomicName>
distribution (circles) with main hosts:
<taxonomicName id="4C4310A3FFF8332700F5F8495B67F85C" box="[903,1062,1926,1947]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="26" pageNumber="327" phylum="Tracheophyta" rank="species" species="arizonica">
<emphasis id="B937B732FFF8332700F5F8495B67F85C" box="[903,1062,1926,1947]" italics="true" pageId="26" pageNumber="327">Pinus arizonica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4310A3FFF833270741F8495B95F85D" box="[1075,1236,1924,1947]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="26" pageNumber="327" phylum="Tracheophyta" rank="species" species="duranguensis">
<emphasis id="B937B732FFF833270741F8495B95F85D" box="[1075,1236,1924,1947]" italics="true" pageId="26" pageNumber="327">P. duranguensis</emphasis>
</taxonomicName>
,
<emphasis id="B937B732FFF833270793F84B5ADDF85D" box="[1249,1436,1924,1947]" italics="true" pageId="26" pageNumber="327">
and
<taxonomicName id="4C4310A3FFF83327067CF8495ADDF85D" box="[1294,1436,1924,1947]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="26" pageNumber="327" phylum="Tracheophyta" rank="species" species="ponderosae">P. ponderosae</taxonomicName>
</emphasis>
(central, AZ and NM).
<taxonomicName id="4C4310A3FFF8332702E4F86B5D71F87E" box="[406,560,1955,1977]" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="longipennis">
<emphasis id="B937B732FFF8332702E4F86B5D71F87E" box="[406,560,1955,1977]" italics="true" pageId="26" pageNumber="327">H. longipennis</emphasis>
</taxonomicName>
distribution (triangles) with main hosts:
<taxonomicName id="4C4310A3FFF83327009EF86B5B2DF87E" box="[1004,1132,1955,1977]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="26" pageNumber="327" phylum="Tracheophyta" rank="species" species="hartwegii">
<emphasis id="B937B732FFF83327009EF86B5B2DF87E" box="[1004,1132,1955,1977]" italics="true" pageId="26" pageNumber="327">P. hartwegii</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4310A3FFF8332707D5F86B5A47F87E" box="[1191,1286,1955,1977]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="26" pageNumber="327" phylum="Tracheophyta" rank="species" species="patula">
<emphasis id="B937B732FFF8332707D5F86B5A47F87E" box="[1191,1286,1955,1977]" italics="true" pageId="26" pageNumber="327">P. patula</emphasis>
</taxonomicName>
. Insert shows sympatric zone near Mexico City.
</paragraph>
</caption>
<paragraph id="8BFC6B20FFF9332603B5FF585D37FF3C" blockId="27.[151,1437,151,1931]" pageId="27" pageNumber="328">
<emphasis id="B937B732FFF9332603B5FF585E60FF77" bold="true" box="[199,289,151,176]" pageId="27" pageNumber="328">Gallery</emphasis>
: The maternal gallery is constructed near or below the root collar, upward from the entrance hole, longitudinal and slightly sinuate from
<quantity id="4CBBC6C5FFF933260131FF725D88FF12" box="[579,713,189,214]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="27" pageNumber="328" unit="mm" value="90.0" valueMax="120.0" valueMin="60.0">60120 mm</quantity>
long (
<bibRefCitation id="EFD216D1FFF933260062FF725CB9FF11" author="Schwerdtfeger" box="[784,1016,189,214]" pageId="27" pageNumber="328" refString="Schwerdtfeger, F. (1957) Scolytidae (Col.) an Pinus - Arten in Mittelamerika III. Die Gattungen Hylastes Erichson und Hylurgops Leconte: (6. Beitrag zur Forstentomologie Mittelamerikas). Deutsche Gesellschaft fur Angewandte Entomologie, 41 (2 - 3), 363 - 367. http: // dx. doi. org / 10.1111 / j. 1439 - 0418.1957. tb 01300. x" type="journal article" year="1957">Schwerdtfeger 1957</bibRefCitation>
). The larval tunnels extend from both sides of the maternal gallery (
<bibRefCitation id="EFD216D1FFF933260290FF2D5D28FF3C" author="Wood" box="[482,617,226,251]" pageId="27" pageNumber="328" refString="Wood, S. L. (1982) The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs, 6, 1 - 1359." type="journal article" year="1982">Wood 1982</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35938ABFFF9332603B5FEC85CD7FD18" pageId="27" pageNumber="328" type="materials_examined">
<paragraph id="8BFC6B20FFF9332603B5FEC85FB5FDB7" blockId="27.[151,1437,151,1931]" pageId="27" pageNumber="328">
<emphasis id="B937B732FFF9332603B5FEC85EEEFEE7" bold="true" box="[199,431,263,288]" pageId="27" pageNumber="328">Material examined</emphasis>
.
<specimenCount id="9D45A0A9FFF9332602B0FEC75D30FEE7" box="[450,625,263,288]" pageId="27" pageNumber="328" type="generic">108 specimens</specimenCount>
.
<collectingCountry id="F3542BB0FFF9332601F3FEC75DB2FEE7" box="[641,755,264,288]" name="Mexico" pageId="27" pageNumber="328">MEXICO</collectingCountry>
.
<emphasis id="B937B732FFF933260071FEC85C24FEE7" bold="true" box="[771,869,263,288]" pageId="27" pageNumber="328">Chiapas</emphasis>
:
<quantity id="4CBBC6C5FFF93326000BFEC85CF5FEE7" box="[889,948,263,288]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="27" pageNumber="328" unit="mi" value="5.0">5 mi</quantity>
. E
<collectingCountry id="F3542BB0FFF933260090FEC75B3EFEE7" box="[994,1151,263,288]" name="Saint Kitts and Nevis" pageId="27" pageNumber="328">San Cristobal</collectingCountry>
(
<collectionCode id="ED52F3E5FFF9332607E1FEC75B9AFEE7" box="[1171,1243,264,288]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
),
<quantity id="4CBBC6C5FFF933260786FEC85A71FEE7" box="[1268,1328,263,288]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="27" pageNumber="328" unit="mi" value="6.0">6 mi</quantity>
. NE
<collectingCountry id="F3542BB0FFF933260600FEC75FBFFE81" name="Saint Kitts and Nevis" pageId="27" pageNumber="328">San Cristobal</collectingCountry>
(
<collectionCode id="ED52F3E5FFF93326027EFEE15E15FE81" box="[268,340,302,326]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
),
<quantity id="4CBBC6C5FFF933260215FEE15EDCFE81" box="[359,413,301,326]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="27" pageNumber="328" unit="mi" value="7.0">7 mi</quantity>
. E
<collectingCountry id="F3542BB0FFF9332602CDFEE15D17FE81" box="[447,598,301,326]" name="Saint Kitts and Nevis" pageId="27" pageNumber="328">San Cristobal</collectingCountry>
(
<collectionCode id="ED52F3E5FFF933260116FEE15DEDFE81" box="[612,684,302,326]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
),
<quantity id="4CBBC6C5FFF9332601B2FEE15DB4FE81" box="[704,757,301,326]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="27" pageNumber="328" unit="mi" value="8.0">8 mi</quantity>
. E
<collectingCountry id="F3542BB0FFF933260065FEE15CEFFE81" box="[791,942,301,326]" name="Saint Kitts and Nevis" pageId="27" pageNumber="328">San Cristobal</collectingCountry>
(
<collectionCode id="ED52F3E5FFF9332600CFFEE15B44FE81" box="[957,1029,302,326]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
),
<quantity id="4CBBC6C5FFF93326076AFEE15B0FFE81" box="[1048,1102,301,326]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="27" pageNumber="328" unit="mi" value="8.0">8 mi</quantity>
. SE
<collectingCountry id="F3542BB0FFF93326070DFEE15A56FE81" box="[1151,1303,301,326]" name="Saint Kitts and Nevis" pageId="27" pageNumber="328">San Cristobal</collectingCountry>
(
<collectionCode id="ED52F3E5FFF933260657FEE15A2CFE81" box="[1317,1389,302,326]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
),
<quantity id="4CBBC6C5FFF9332606F2FEE25FF9FEAC" metricMagnitude="4" metricUnit="m" metricValue="4.02336" pageId="27" pageNumber="328" unit="mi" value="25.0">25 mi</quantity>
. SE Teopisca (
<collectionCode id="ED52F3E5FFF933260216FE9C5EEDFEAC" box="[356,428,339,363]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
),
<collectingCountry id="F3542BB0FFF9332602B1FE9C5D1FFEAC" box="[451,606,338,363]" name="Saint Kitts and Nevis" pageId="27" pageNumber="328">San Cristobal</collectingCountry>
(
<collectionCode id="ED52F3E5FFF93326011DFE9C5DF9FEAC" box="[623,696,339,363]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
).
<emphasis id="B937B732FFF9332601BFFE9D5C12FEAC" bold="true" box="[717,851,338,363]" pageId="27" pageNumber="328">Chihuahua</emphasis>
: Mesa del Huracan (
<collectionCode id="ED52F3E5FFF93326073AFE9C5BD0FEAC" box="[1096,1169,339,363]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
).
<emphasis id="B937B732FFF9332607D4FE9D5A51FEAC" bold="true" box="[1190,1296,338,363]" pageId="27" pageNumber="328">Durango</emphasis>
:
<quantity id="4CBBC6C5FFF933260650FE9C5A1AFEAC" box="[1314,1371,338,363]" metricMagnitude="4" metricUnit="m" metricValue="1.4484096" pageId="27" pageNumber="328" unit="mi" value="9.0">9 mi</quantity>
. E El Palmito (
<collectionCode id="ED52F3E5FFF933260273FEB75E08FE57" box="[257,329,376,400]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
),
<quantity id="4CBBC6C5FFF933260213FEB75EE8FE57" box="[353,425,375,400]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="27" pageNumber="328" unit="mi" value="10.0">10 mi</quantity>
. SW El Salto (
<collectionCode id="ED52F3E5FFF93326012FFEB75DE4FE57" box="[605,677,376,400]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
),
<quantity id="4CBBC6C5FFF9332601CFFEB75C44FE57" box="[701,773,375,400]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="27" pageNumber="328" unit="mi" value="10.0">10 mi</quantity>
. SW El Salto (
<collectionCode id="ED52F3E5FFF9332600C8FEB75B42FE57" box="[954,1027,376,400]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
).
<emphasis id="B937B732FFF93326076BFEB85BE0FE57" bold="true" box="[1049,1185,375,400]" pageId="27" pageNumber="328">
<collectingCountry id="F3542BB0FFF93326076BFEB85B31FE57" box="[1049,1136,375,400]" name="Mexico" pageId="27" pageNumber="328">Mexico</collectingCountry>
DF
</emphasis>
:
<quantity id="4CBBC6C5FFF9332607C6FEB85BBDFE57" box="[1204,1276,375,400]" metricMagnitude="4" metricUnit="m" metricValue="2.414016" pageId="27" pageNumber="328" unit="mi" value="15.0">15 mi</quantity>
. S El Guarda (
<collectionCode id="ED52F3E5FFF9332603EDFE515FA9FE71" box="[159,232,414,438]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
).
<emphasis id="B937B732FFF93326038FFE525ED8FE71" bold="true" box="[253,409,413,438]" pageId="27" pageNumber="328">
<collectingCountry id="F3542BB0FFF93326038FFE525E15FE71" box="[253,340,413,438]" name="Mexico" pageId="27" pageNumber="328">Mexico</collectingCountry>
State
</emphasis>
:
<quantity id="4CBBC6C5FFF9332602DEFE515EB2FE71" box="[428,499,413,438]" metricMagnitude="4" metricUnit="m" metricValue="6.437376" pageId="27" pageNumber="328" unit="mi" value="40.0">40 mi</quantity>
. W (
<collectionCode id="ED52F3E5FFF93326015FFE515D3FFE71" box="[557,638,414,438]" pageId="27" pageNumber="328">DEBC</collectionCode>
); Vivero San Cayetano (
<collectionCode id="ED52F3E5FFF9332600E8FE515CAAFE71" box="[922,1003,414,438]" pageId="27" pageNumber="328">DEBC</collectionCode>
).
<emphasis id="B937B732FFF933260773FE525BD5FE71" bold="true" box="[1025,1172,413,438]" pageId="27" pageNumber="328">Nuevo Leon</emphasis>
: Cerro Potosi (
<collectionCode id="ED52F3E5FFF933260635FE515AD1FE71" box="[1351,1424,414,438]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
).
<emphasis id="B937B732FFF9332603E5FE0D5FB0FE1C" bold="true" box="[151,241,450,475]" pageId="27" pageNumber="328">Oaxaca</emphasis>
:
<quantity id="4CBBC6C5FFF933260276FE0C5E25FE1C" box="[260,356,450,475]" metricMagnitude="4" metricUnit="m" metricValue="1.7899999999999998" pageId="27" pageNumber="328" unit="km" value="17.9">17.9 km</quantity>
E Teotitlan (
<collectionCode id="ED52F3E5FFF933260170FE0C5D13FE1C" box="[514,594,451,475]" pageId="27" pageNumber="328">DEBC</collectionCode>
), Hwy. 175,
<quantity id="4CBBC6C5FFF93326019EFE0D5C7AFE1C" box="[748,827,450,475]" metricMagnitude="3" metricUnit="m" metricValue="5.632704" pageId="27" pageNumber="328" unit="mi" value="3.5">3.5 mi</quantity>
. S Suchixtepec (
<collectionCode id="ED52F3E5FFF933260773FE0C5B0BFE1C" box="[1025,1098,451,475]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
).
<emphasis id="B937B732FFF933260712FE0D5BF3FE1C" bold="true" box="[1120,1202,450,475]" pageId="27" pageNumber="328">Puebla</emphasis>
:
<quantity id="4CBBC6C5FFF9332607B4FE0D5A41FE1C" box="[1222,1280,450,475]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="27" pageNumber="328" unit="mi" value="6.0">6 mi</quantity>
. W Teziutlan (
<collectionCode id="ED52F3E5FFF9332603EDFE275FA9FDC7" box="[159,232,488,512]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
).
<emphasis id="B937B732FFF933260389FE285E72FDC7" bold="true" box="[251,307,487,512]" pageId="27" pageNumber="328">Vera</emphasis>
c
<emphasis id="B937B732FFF93326024DFE285E26FDC7" bold="true" box="[319,359,487,512]" pageId="27" pageNumber="328">ruz</emphasis>
:
<quantity id="4CBBC6C5FFF933260205FE285E83FDC7" box="[375,450,487,512]" metricMagnitude="3" metricUnit="m" metricValue="5.7936384" pageId="27" pageNumber="328" unit="mi" value="3.6">3.6 mi</quantity>
. W Las Vigas (
<collectionCode id="ED52F3E5FFF93326011FFE275DF7FDC7" box="[621,694,488,512]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
).
<collectingCountry id="F3542BB0FFF9332601BBFE275C45FDC7" box="[713,772,488,512]" name="United States of America" pageId="27" pageNumber="328">USA</collectingCountry>
.
<emphasis id="B937B732FFF93326007DFE285C2EFDC7" bold="true" box="[783,879,487,512]" pageId="27" pageNumber="328">Arizona</emphasis>
: Cochise Co.: Rustler Pk., Chiricagua Mountains (
<collectionCode id="ED52F3E5FFF9332603EDFDC15FA9FDE1" box="[159,232,526,550]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
); Pima Co.: Sta.
<taxonomicName id="4C4310A3FFF9332602D1FDC25D3EFDE1" authority="Mountains" authorityName="Mountains" box="[419,639,525,550]" class="Insecta" family="Staphylinidae" genus="Catalina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="genus">Catalina Mountains</taxonomicName>
(
<collectionCode id="ED52F3E5FFF9332601FCFDC15D96FDE1" box="[654,727,526,550]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
); Sedona Co.: Oak Creek Canyon (
<collectionCode id="ED52F3E5FFF93326072DFDC15BE9FDE1" box="[1119,1192,526,550]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
).
<emphasis id="B937B732FFF9332607C9FDC25A0CFDE1" bold="true" box="[1211,1357,525,550]" pageId="27" pageNumber="328">
New
<collectingCountry id="F3542BB0FFF933260784FDC25A0CFDE1" box="[1270,1357,525,550]" name="Mexico" pageId="27" pageNumber="328">Mexico</collectingCountry>
</emphasis>
: Otero Co.: Cloudcroft (
<collectionCode id="ED52F3E5FFF933260213FDFC5EF5FD8C" box="[353,436,563,587]" pageId="27" pageNumber="328">RMRS</collectionCode>
), Cloudcroft
<typeStatus id="54F8D582FFF933260121FDFC5DF4FD8C" box="[595,693,563,587]" pageId="27" pageNumber="328" type="paratype">Paratype</typeStatus>
9241 (
<collectionCode id="ED52F3E5FFF93326007FFDFC5C17FD8C" box="[781,854,563,587]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
); Santa Fe Co.: Santa Fe Canyon
<typeStatus id="54F8D582FFF933260787FDFC5A16FD8C" box="[1269,1367,563,587]" pageId="27" pageNumber="328" type="paratype">Paratype</typeStatus>
9241 (
<collectionCode id="ED52F3E5FFF9332603EDFD975FA9FDB7" box="[159,232,600,624]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="328">CNCI</collectionCode>
).
</paragraph>
<paragraph id="8BFC6B20FFF9332603B5FDB25CD7FD18" blockId="27.[151,1437,151,1931]" pageId="27" pageNumber="328">
<emphasis id="B937B732FFF9332603B5FDB25E4BFD51" bold="true" box="[199,266,637,662]" pageId="27" pageNumber="328">Hosts</emphasis>
:
<taxonomicName id="4C4310A3FFF93326026DFDB15E96FD52" box="[287,471,638,661]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="27" pageNumber="328" phylum="Tracheophyta" rank="species" species="arizonica">
<emphasis id="B937B732FFF93326026DFDB15E96FD52" box="[287,471,638,661]" italics="true" pageId="27" pageNumber="328">Pinus arizonica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4310A3FFF93326029EFDB15DE4FD51" box="[492,677,637,662]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="27" pageNumber="328" phylum="Tracheophyta" rank="species" species="duranguensis">
<emphasis id="B937B732FFF93326029EFDB15DE4FD51" box="[492,677,637,662]" italics="true" pageId="27" pageNumber="328">P. duranguensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4310A3FFF9332601CBFDB15C26FD51" box="[697,871,637,662]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="27" pageNumber="328" phylum="Tracheophyta" rank="species" species="engelmannii">
<emphasis id="B937B732FFF9332601CBFDB15C26FD51" box="[697,871,637,662]" italics="true" pageId="27" pageNumber="328">P. engelmannii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4310A3FFF933260009FDB15B49FD51" box="[891,1032,637,662]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="27" pageNumber="328" phylum="Tracheophyta" rank="species" species="hartwegii">
<emphasis id="B937B732FFF933260009FDB15B49FD51" box="[891,1032,637,662]" italics="true" pageId="27" pageNumber="328">P. hartwegii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4310A3FFF93326076FFDB15BF1FD51" box="[1053,1200,637,662]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="27" pageNumber="328" phylum="Tracheophyta" rank="species" species="leiophylla">
<emphasis id="B937B732FFF93326076FFDB15BF1FD51" box="[1053,1200,637,662]" italics="true" pageId="27" pageNumber="328">P. leiophylla</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4310A3FFF9332607B6FDB15A32FD52" box="[1220,1395,638,661]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="27" pageNumber="328" phylum="Tracheophyta" rank="species" species="montezumae">
<emphasis id="B937B732FFF9332607B6FDB15A32FD52" box="[1220,1395,638,661]" italics="true" pageId="27" pageNumber="328">P. montezumae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4310A3FFF9332606F5FDB15E60FD7C" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="27" pageNumber="328" phylum="Tracheophyta" rank="species" species="tecunumanii">
<emphasis id="B937B732FFF9332606F5FDB15E60FD7C" italics="true" pageId="27" pageNumber="328">P. tecunumanii</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4310A3FFF933260244FD6B5EDEFD7C" box="[310,415,674,699]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="27" pageNumber="328" phylum="Tracheophyta" rank="species" species="patula">
<emphasis id="B937B732FFF933260244FD6B5EDEFD7C" box="[310,415,674,699]" italics="true" pageId="27" pageNumber="328">P. patula</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4310A3FFF9332602C6FD6B5D18FD7C" box="[436,601,674,699]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="27" pageNumber="328" phylum="Tracheophyta" rank="species" species="ponderosae">
<emphasis id="B937B732FFF9332602C6FD6B5D18FD7C" box="[436,601,674,699]" italics="true" pageId="27" pageNumber="328">P. ponderosae</emphasis>
</taxonomicName>
(included without revised information due to beetle records north of Mogollon Rim (the northern limit of
<taxonomicName id="4C4310A3FFF933260144FD065DFDFD27" box="[566,700,713,736]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="27" pageNumber="328" phylum="Tracheophyta" rank="species" species="arizonica">
<emphasis id="B937B732FFF933260144FD065DFDFD27" box="[566,700,713,736]" italics="true" pageId="27" pageNumber="328">P. arizonica</emphasis>
</taxonomicName>
),
<taxonomicName id="4C4310A3FFF9332601A1FD065CCEFD27" box="[723,911,711,736]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="27" pageNumber="328" phylum="Tracheophyta" rank="species" species="pseudostrobus">
<emphasis id="B937B732FFF9332601A1FD065CCEFD27" box="[723,911,711,736]" italics="true" pageId="27" pageNumber="328">P. pseudostrobus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35938ABFFF9332603B5FD225DE0FCEC" pageId="27" pageNumber="328" type="distribution">
<paragraph id="8BFC6B20FFF9332603B5FD225DE0FCEC" blockId="27.[151,1437,151,1931]" pageId="27" pageNumber="328">
<emphasis id="B937B732FFF9332603B5FD225E18FCC1" bold="true" box="[199,345,749,774]" pageId="27" pageNumber="328">Distribution</emphasis>
(
<figureCitation id="137877A5FFF93326021BFD225EFCFCC1" box="[361,445,749,774]" captionStart="FIGURE 21" captionStartId="26.[151,250,1925,1947]" captionTargetBox="[152,1436,940,1902]" captionTargetId="figure@26.[151,1436,939,1903]" captionTargetPageId="26" captionText="FIGURE 21. Hylurgops incomptus distribution (circles) with main hosts: Pinus arizonica, P. duranguensis, and P. ponderosae (central, AZ and NM). H. longipennis distribution (triangles) with main hosts: P. hartwegii and P. patula. Insert shows sympatric zone near Mexico City." httpUri="https://zenodo.org/record/252635/files/figure.png" pageId="27" pageNumber="328">Fig. 21</figureCitation>
). CENTRAL
<collectingCountry id="F3542BB0FFF933260129FD215DA0FCC1" box="[603,737,750,774]" name="United States of America" pageId="27" pageNumber="328">AMERICA</collectingCountry>
:
<collectingCountry id="F3542BB0FFF93326019EFD225C1DFCC1" box="[748,860,749,774]" name="Honduras" pageId="27" pageNumber="328">Honduras</collectingCountry>
,
<collectingCountry id="F3542BB0FFF933260015FD225CA1FCC1" box="[871,992,749,774]" name="Guatemala" pageId="27" pageNumber="328">Guatemala</collectingCountry>
and
<collectingCountry id="F3542BB0FFF93326076BFD225BE1FCC1" box="[1049,1184,749,774]" name="El Salvador" pageId="27" pageNumber="328">El Salvador</collectingCountry>
; NORTH
<collectingCountry id="F3542BB0FFF933260661FD215AD8FCC1" box="[1299,1433,750,774]" name="United States of America" pageId="27" pageNumber="328">AMERICA</collectingCountry>
: Arizona and New
<collectingCountry id="F3542BB0FFF933260211FCDD5EF9FCEC" box="[355,440,786,811]" name="Mexico" pageId="27" pageNumber="328">Mexico</collectingCountry>
to southern
<collectingCountry id="F3542BB0FFF933260136FCDD5DDCFCEC" box="[580,669,786,811]" name="Mexico" pageId="27" pageNumber="328">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35938ABFFF9332603B5FCF85C97F84C" pageId="27" pageNumber="328" type="discussion">
<paragraph id="8BFC6B20FFF9332603B5FCF85ED5FC78" blockId="27.[151,1437,151,1931]" pageId="27" pageNumber="328">
<emphasis id="B937B732FFF9332603B5FCF85E0DFC97" bold="true" box="[199,332,823,848]" pageId="27" pageNumber="328">Discussion.</emphasis>
<bibRefCitation id="EFD216D1FFF933260226FCF85D59FC97" author="Blandford" box="[340,536,823,848]" pageId="27" pageNumber="328" refString="Blandford, W. F. H. (1897) Scolytidae. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali-Americana. Insecta, Coleoptera, 4 (6), pp. 145 - 184." type="journal article" year="1897">Blandford (1897)</bibRefCitation>
described this species as
<taxonomicName id="4C4310A3FFF93326004EFCF85B56FC97" box="[828,1047,823,848]" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="incomptus">
<emphasis id="B937B732FFF93326004EFCF85B56FC97" box="[828,1047,823,848]" italics="true" pageId="27" pageNumber="328">Hylastes incomptus</emphasis>
</taxonomicName>
under Erichsons second division, and
<bibRefCitation id="EFD216D1FFF9332603B4FC925EC6FCB1" author="Hagedorn" box="[198,391,861,886]" pageId="27" pageNumber="328" refString="Hagedorn, M. (1910) Coleoptera Fam. Ipidae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 111. Brussels, pp. 1 - 178 + XIV plts." type="book chapter" year="1910">Hagedorn (1910)</bibRefCitation>
included
<taxonomicName id="4C4310A3FFF933260284FC915DCAFCB2" box="[502,651,862,885]" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="incomptus">
<emphasis id="B937B732FFF933260284FC915DCAFCB2" box="[502,651,862,885]" italics="true" pageId="27" pageNumber="328">H. incomptus</emphasis>
</taxonomicName>
under the subgenus
<taxonomicName id="4C4310A3FFF933260001FC925CA9FCB1" box="[883,1000,861,886]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B937B732FFF933260001FC925CA9FCB1" box="[883,1000,861,886]" italics="true" pageId="27" pageNumber="328">Hylurgops</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFD216D1FFF933260084FC925BD2FCB1" author="Kleine" box="[1014,1171,861,886]" pageId="27" pageNumber="328" refString="Kleine, R. (1912) Die geographische verbreitung der Ipiden-Genera orbis terrarum. (Col.). Berliner Entomololgische Zeitschrift, 57, 155 - 192." type="journal article" year="1912">Kleine (1912)</bibRefCitation>
considered Hagedorns species concept of
<taxonomicName id="4C4310A3FFF93326021DFC4D5E8EFC5C" box="[367,463,898,923]" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B937B732FFF93326021DFC4D5E8EFC5C" box="[367,463,898,923]" italics="true" pageId="27" pageNumber="328">Hylastes</emphasis>
</taxonomicName>
to be too broad and did not follow his subgeneric concept, placing
<taxonomicName id="4C4310A3FFF9332607A3FC4B5A26FC5C" box="[1233,1383,900,923]" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="incomptus">
<emphasis id="B937B732FFF9332607A3FC4B5A26FC5C" box="[1233,1383,900,923]" italics="true" pageId="27" pageNumber="328">H. incomptus</emphasis>
</taxonomicName>
as a member of
<taxonomicName id="4C4310A3FFF93326026AFC685ECCFC07" box="[280,397,935,960]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="genus">
<emphasis id="B937B732FFF93326026AFC685ECCFC07" box="[280,397,935,960]" italics="true" pageId="27" pageNumber="328">Hylurgops</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFC6B20FFF9332603B5FC025C40FB2C" blockId="27.[151,1437,151,1931]" pageId="27" pageNumber="328">
When
<bibRefCitation id="EFD216D1FFF933260266FC025EFBFC21" author="Swaine" box="[276,442,973,998]" pageId="27" pageNumber="328" refString="Swaine, J. M. (1917) Canadian bark beetles, Part 1. Description of new species. Canada Department of Agriculture, Entomological Branch, Bulletin, 14 (1), 1 - 32." type="journal article" year="1917">Swaine (1917)</bibRefCitation>
described
<taxonomicName id="4C4310A3FFF933260148FC015DA7FC21" box="[570,742,973,998]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="grandicollis">
<emphasis id="B937B732FFF933260148FC015DA7FC21" box="[570,742,973,998]" italics="true" pageId="27" pageNumber="328">H. grandicollis</emphasis>
</taxonomicName>
from three specimens collected in New
<collectingCountry id="F3542BB0FFF9332607CDFC025A56FC21" box="[1215,1303,973,998]" name="Mexico" pageId="27" pageNumber="328">Mexico</collectingCountry>
, he did not make any reference to Blandfords (1897) species.
<bibRefCitation id="EFD216D1FFF93326019AFC3D5CCEFBCC" author="Swaine" box="[744,911,1010,1035]" pageId="27" pageNumber="328" refString="Swaine, J. M. (1917) Canadian bark beetles, Part 1. Description of new species. Canada Department of Agriculture, Entomological Branch, Bulletin, 14 (1), 1 - 32." type="journal article" year="1917">Swaine (1917)</bibRefCitation>
mentioned having examined specimens from California but gave no specific locality. No specimens from California were examined during this study, and
<bibRefCitation id="EFD216D1FFF933260626FBD85E0BFB91" author="Bright" pageId="27" pageNumber="328" refString="Bright, D. E. &amp; Stark, R. W. (1973) The Bark and Ambrosia Beetles of California: Coleoptera: Scolytidae and Platypodidae. University of California Press, Berkeley, California, 169 pp." type="book" year="1973">Bright &amp; Stark (1973)</bibRefCitation>
did not include
<taxonomicName id="4C4310A3FFF933260166FBF15D82FB91" box="[532,707,1085,1110]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="grandicollis">
<emphasis id="B937B732FFF933260166FBF15D82FB91" box="[532,707,1085,1110]" italics="true" pageId="27" pageNumber="328">H. grandicollis</emphasis>
</taxonomicName>
from that state. In 1955, Schedl reported
<taxonomicName id="4C4310A3FFF9332607CDFBF15A18FB92" box="[1215,1369,1086,1109]" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="incomptus">
<emphasis id="B937B732FFF9332607CDFBF15A18FB92" box="[1215,1369,1086,1109]" italics="true" pageId="27" pageNumber="328">H. incomptus</emphasis>
</taxonomicName>
from
<collectingCountry id="F3542BB0FFF9332603E5FBAD5E52FBBC" box="[151,275,1122,1147]" name="Guatemala" pageId="27" pageNumber="328">Guatemala</collectingCountry>
, its southernmost distribution.
<bibRefCitation id="EFD216D1FFF933260105FBAD5C4FFBBC" author="Wood" box="[631,782,1122,1147]" pageId="27" pageNumber="328" refString="Wood, S. L. (1957) Distributional Notes on and Synonymies of Some North American Scolytidae (Coleoptera). The Canadian Entomologist, 89 (9), 396 - 403. http: // dx. doi. org / 10.4039 / ent 89396 - 9" type="journal article" year="1957">Wood (1957)</bibRefCitation>
synonymized
<taxonomicName id="4C4310A3FFF9332600C4FBAB5BFFFBBC" authority="Swaine" authorityName="Swaine" box="[950,1214,1122,1147]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="grandicollis">
<emphasis id="B937B732FFF9332600C4FBAB5B23FBBC" box="[950,1122,1122,1147]" italics="true" pageId="27" pageNumber="328">H. grandicollis</emphasis>
Swaine
</taxonomicName>
with
<taxonomicName id="4C4310A3FFF933260676FBAB5E48FB67" authority="Blandford" authorityName="Blandford" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="incomptus">
<emphasis id="B937B732FFF933260676FBAB5ADDFBBC" box="[1284,1436,1124,1147]" italics="true" pageId="27" pageNumber="328">H. incomptus</emphasis>
Blandford
</taxonomicName>
after comparing Swaines
<taxonomicName id="4C4310A3FFF933260144FB485DFCFB67" box="[566,701,1159,1184]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="grandicollis">
<emphasis id="B937B732FFF933260144FB485DFCFB67" box="[566,701,1159,1184]" italics="true" pageId="27" pageNumber="328">grandicollis</emphasis>
</taxonomicName>
<typeStatus id="54F8D582FFF9332601B6FB475DB5FB67" box="[708,756,1160,1184]" pageId="27" pageNumber="328">type</typeStatus>
with several of his and Eggers specimens from unspecified locations.
<bibRefCitation id="EFD216D1FFF933260278FB625EDFFB01" author="Wood" box="[266,414,1197,1222]" pageId="27" pageNumber="328" refString="Wood, S. L. (1982) The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs, 6, 1 - 1359." type="journal article" year="1982">Wood (1982)</bibRefCitation>
based his description on a homotype from Cerro Calel,
<collectingCountry id="F3542BB0FFF933260761FB625BCDFB01" box="[1043,1164,1197,1222]" name="Guatemala" pageId="27" pageNumber="328">Guatemala</collectingCountry>
and examined the
<typeStatus id="54F8D582FFF933260613FB615ADDFB01" box="[1377,1436,1198,1222]" pageId="27" pageNumber="328">types</typeStatus>
of
<taxonomicName id="4C4310A3FFF9332603C7FB1B5E0BFB2C" box="[181,330,1236,1259]" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="incomptus">
<emphasis id="B937B732FFF9332603C7FB1B5E0BFB2C" box="[181,330,1236,1259]" italics="true" pageId="27" pageNumber="328">H. incomptus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4310A3FFF9332602F3FB1B5D6BFB2C" box="[385,554,1234,1259]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="grandicollis">
<emphasis id="B937B732FFF9332602F3FB1B5D6BFB2C" box="[385,554,1234,1259]" italics="true" pageId="27" pageNumber="328">H. grandicollis</emphasis>
</taxonomicName>
among 151 others.
</paragraph>
<paragraph id="8BFC6B20FFF9332603B5FB385B7BF9D2" blockId="27.[151,1437,151,1931]" pageId="27" pageNumber="328">
Specimens from Arizona and New
<collectingCountry id="F3542BB0FFF933260127FB385DECFAD7" box="[597,685,1271,1296]" name="Mexico" pageId="27" pageNumber="328">Mexico</collectingCountry>
, including the Swaine
<taxonomicName id="4C4310A3FFF9332600C2FB365B1BFAD7" box="[944,1114,1271,1296]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="grandicollis">
<emphasis id="B937B732FFF9332600C2FB365B1BFAD7" box="[944,1114,1271,1296]" italics="true" pageId="27" pageNumber="328">H. grandicollis</emphasis>
</taxonomicName>
<typeStatus id="54F8D582FFF933260710FB375BE1FAD7" box="[1122,1184,1272,1296]" pageId="27" pageNumber="328">types</typeStatus>
, have a pronotum that gradually broadens from the base to the middle or to slightly anterior of the middle (
<figureCitation id="137877A5FFF933260710FAD25BF6FAF1" box="[1122,1207,1309,1334]" captionStart="FIGURE 20" captionStartId="26.[151,250,837,859]" captionTargetBox="[165,1428,375,815]" captionTargetId="figure@26.[156,1430,373,815]" captionTargetPageId="26" captionText="FIGURE 20. Hylurgops incomptus pronotum: a) prothoracic lateral margin, b) pronotum broadening from base (typical form), c) pronotum narrowing from base (Arizona and New Mexico variant)." httpUri="https://zenodo.org/record/252634/files/figure.png" pageId="27" pageNumber="328">Fig. 20</figureCitation>
b). These specimens also have a reticulate surface of the elytra. On the declivity, the hair-like setae are shorter, reddish-yellow, and less abundant due to the presence of a single row of medially arranged granules on the interstriae and the scale-like setae are more abundant. The pronotum of Mexican specimens from Chiapas gradually narrow from the base toward the apex (
<figureCitation id="137877A5FFF933260217FA7D5EFCFA0C" box="[357,445,1458,1483]" captionStart="FIGURE 20" captionStartId="26.[151,250,837,859]" captionTargetBox="[165,1428,375,815]" captionTargetId="figure@26.[156,1430,373,815]" captionTargetPageId="26" captionText="FIGURE 20. Hylurgops incomptus pronotum: a) prothoracic lateral margin, b) pronotum broadening from base (typical form), c) pronotum narrowing from base (Arizona and New Mexico variant)." httpUri="https://zenodo.org/record/252634/files/figure.png" pageId="27" pageNumber="328">Fig. 20</figureCitation>
c). The elytral surface varies from reticulate to smooth and glossy, and the declivital vestiture is light-yellowish with abundant, medium length, hair-like setae due to the presence of a pair of lateral granules in addition to the central interstrial row and less abundant scale-like setae.
</paragraph>
<paragraph id="8BFC6B20FFF9332603B5F9ED5C97F84C" blockId="27.[151,1437,151,1931]" pageId="27" pageNumber="328">
The occurrence of the species between the populations of southern Arizona and New
<collectingCountry id="F3542BB0FFF9332607D0F9ED5BB6F9FC" box="[1186,1271,1570,1595]" name="Mexico" pageId="27" pageNumber="328">Mexico</collectingCountry>
is fragmented possibly due to the host patchiness in the few mountains across the Sonoran and Chihuahuan deserts. The northernmost Mexican record reported by
<bibRefCitation id="EFD216D1FFF93326010CF9A25C55F941" author="Wood" box="[638,788,1645,1670]" pageId="27" pageNumber="328" refString="Wood, S. L. (1982) The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs, 6, 1 - 1359." type="journal article" year="1982">Wood (1982)</bibRefCitation>
is from Mesa del Huracan in Chihuahua, about
<quantity id="4CBBC6C5FFF933260637F9A15ADDF941" box="[1349,1436,1645,1670]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="27" pageNumber="328" unit="km" value="300.0">300 km</quantity>
southeast of the southernmost record in Arizona in the isolated Huachuca Mountains. Specimens from these two disjunct regions are indistinguishable. Specimens from the Sierra Madre del Sur in the Mexican state of Oaxaca also closely resemble the northern specimens, except that some are smaller. The greatest differences are found between specimens occurring north of Durango,
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and specimens from Chiapas in southern
<collectingCountry id="F3542BB0FFF93326064DF8CD5AD9F8DC" box="[1343,1432,1794,1819]" name="Mexico" pageId="27" pageNumber="328">Mexico</collectingCountry>
. However, individuals from extremes of the range share some characters, suggesting that the species probably exhibit clinal variation. Collecting other material for additional morphometric and molecular analysis would help determine if specimens from these populations represent different species.
</paragraph>
</subSubSection>
</treatment>
</document>