plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/EA/61/03EA61740E26CB15494BFAB9FCD6FD13.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

876 lines
117 KiB
XML
Raw Blame History

This file contains invisible Unicode characters

This file contains invisible Unicode characters that are indistinguishable to humans but may be processed differently by a computer. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="0EC696C8E604BC3581A8125FC9037465" ID-CLB-Dataset="5509" ID-DOI="10.11646/zootaxa.187.1.1" ID-GBIF-Dataset="71409b97-0770-4e5a-bb9c-4d44f65d7e06" ID-ISSN="1175­5334" ID-Zenodo-Dep="5014522" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624383239017" checkinUser="felipe" docAuthor="Acosta, Luis E." docDate="2003" docId="03EA61740E26CB15494BFAB9FCD6FD13" docLanguage="en" docName="zt00187.pdf.imf" docOrigin="Zootaxa 187 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.187.1.1" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Urophonius somuncura Acosta 2003, n. sp." docType="treatment" docUuid="5D2B4200-8470-41DC-BAAE-D754A7AD582A" docUuidSource="ZooBank" docVersion="8" lastPageNumber="10" masterDocId="FFD3190C0E27CB1F4843FFCFFFA2FFFD" masterDocTitle="Description of a new Patagonian species of Urophonius Pocock (Scorpiones, Bothriuridae), from Meseta de Somuncurá, Argentina" masterLastPageNumber="12" masterPageNumber="1" pageNumber="2" updateTime="1699020109422" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="A0E6EC92149AFD7D57EF705F4AD03F74" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AE0677B53284247098D6AA6540FD92AB">
<mods:title id="0779571650046A10415639F865EBF162">Description of a new Patagonian species of Urophonius Pocock (Scorpiones, Bothriuridae), from Meseta de Somuncurá, Argentina</mods:title>
</mods:titleInfo>
<mods:name id="A703B3BC5DC3ACDFFABC7B085CDFE9E4" type="personal">
<mods:role id="A9701900408FD00E680B1120E91FB517">
<mods:roleTerm id="999F5B9A6F0AFD74CDDA6A25A9BB7CD6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A2F6BF7CD6090F260E2FCCC5CA72C8F8">Acosta, Luis E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="912F4AC9C1CE6DCDF4F742F6EF75DF16">text</mods:typeOfResource>
<mods:relatedItem id="7ACFFA68737080F2FD019B2CDEF975B0" type="host">
<mods:titleInfo id="92D6DAF1DB0A66E908B0AD1F10DDF733">
<mods:title id="E56BDEFCA8930A2904C6094E59D4B5FA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B290AA0C866BD2150EF73C995CD6DE62">
<mods:date id="B12C6DA0DA081D9A8528DB974AC7A870">2003</mods:date>
<mods:detail id="C3F06427D8506E2A558B9B8ABD9946B4" type="pubDate">
<mods:number id="90E26BDB147BA9E502FFD90F295C5461">2003-04-24</mods:number>
</mods:detail>
<mods:detail id="FB9AC1C6F1A22409145D0EF872FE10C7" type="volume">
<mods:number id="B978E36E88472AF6109BE9DEB9A48AA6">187</mods:number>
</mods:detail>
<mods:detail id="16642BF1A279B358BCF2D76644EBFA03" type="issue">
<mods:number id="87D5D78E6A6B584C0D752A3ED782EB7F">1</mods:number>
</mods:detail>
<mods:extent id="DB38B6DF489A508C1BB22E525AC98C54" unit="page">
<mods:start id="A3F5014637D528FE825FD2C7F5925A72">1</mods:start>
<mods:end id="0CA3644C86CB0D2DB02CE19C2512B86D">12</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="5631A3FA97F526861F2317295CA2692E">
<mods:url id="1F15DE6AB1F22365E152CDC505F2C392">https://biotaxa.org/Zootaxa/article/view/zootaxa.187.1.1</mods:url>
</mods:location>
<mods:classification id="03BFCB0F5BB3474C9E925C4378B88275">journal article</mods:classification>
<mods:identifier id="48E21C7F11137DCEF2A5B823A60271D8" type="CLB-Dataset">5509</mods:identifier>
<mods:identifier id="076381B855AECA001610669FC0BADE05" type="DOI">10.11646/zootaxa.187.1.1</mods:identifier>
<mods:identifier id="C0E7A2234D19148BDF5849EA220B95FC" type="GBIF-Dataset">71409b97-0770-4e5a-bb9c-4d44f65d7e06</mods:identifier>
<mods:identifier id="5B57D5E408D2CAA9758F6254059B8688" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="40EA4A20AD7BF89D80612D217EE76FB1" type="Zenodo-Dep">5014522</mods:identifier>
<mods:identifier id="C964B7556A478A833CFAE54B3624FA88" type="ZooBank">1ABEFCE0-C401-47AD-AAB8-81C5A327F41B</mods:identifier>
</mods:mods>
<treatment id="03EA61740E26CB15494BFAB9FCD6FD13" ID-DOI="http://doi.org/10.5281/zenodo.5014553" ID-GBIF-Taxon="183501877" ID-Zenodo-Dep="5014553" LSID="urn:lsid:zoobank.org:act:5D2B4200-8470-41DC-BAAE-D754A7AD582A" httpUri="http://treatment.plazi.org/id/03EA61740E26CB15494BFAB9FCD6FD13" lastPageId="10" lastPageNumber="10" pageId="1" pageNumber="2">
<subSubSection id="C35983E90E26CB1E494BFAB9FDCDFA6C" box="[264,623,1398,1425]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8BFCD0620E26CB1E494BFAB9FDCDFA6C" blockId="1.[264,623,1398,1465]" box="[264,623,1398,1425]" pageId="1" pageNumber="2">
<heading id="D0B4670E0E26CB1E494BFAB9FDCDFA6C" box="[264,623,1398,1425]" level="2" pageId="1" pageNumber="2">
<taxonomicName id="4C43ABE10E26CB1E494BFAB9FD8BFA6D" ID-CoL="7DXPH" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" box="[264,553,1398,1424]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E26CB1E494BFAB9FD8BFA6D" bold="true" box="[264,553,1398,1424]" italics="true" pageId="1" pageNumber="2">Urophonius somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E26CB1E4A72FAB8FDCDFA6C" box="[561,623,1399,1425]" pageId="1" pageNumber="2" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35983E90E26CB1E494BFA50FECAFA44" box="[264,360,1439,1465]" pageId="1" pageNumber="2" type="description">
<paragraph id="8BFCD0620E26CB1E494BFA50FECAFA44" blockId="1.[264,623,1398,1465]" box="[264,360,1439,1465]" pageId="1" pageNumber="2">
<figureCitation id="1378CCE70E26CB1E494BFA50FECAFA44" box="[264,360,1439,1465]" captionStart="FIGURES 1­8" captionStartId="2.[264,383,1693,1717]" captionTargetBox="[296,1291,350,1665]" captionTargetId="figure-105@2.[268,1320,329,1682]" captionTargetPageId="2" captionText="FIGURES 1­8. Urophonius somuncura n. sp. 1­7: holotype female (CDA), 8: generalised from several individuals. 1, metasomal segment V and telson, lateral view; 2, metasomal segment V, ventral view; 3­4, sternite V and metasomal segments I­II, ventral view, 3, morphology (ventral macrosetae as black dots), 4, pigment pattern (macrosetae as open circles); 5­6, right pedipalp chela, 5, ventromedial view, 6, lateral view; 7, right pedipalp femur, retrolateral view (indicated trichobothrium e and macroseta M1). 8: pigment pattern on tergite V. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5014524" httpUri="https://zenodo.org/record/5014524/files/figure.png" pageId="1" pageNumber="2">Figs.1­8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35983E90E26CB1E494BFA28FD34F9E3" pageId="1" pageNumber="2" type="reference_group">
<paragraph id="8BFCD0620E26CB1E494BFA28FD34F9E3" blockId="1.[264,1323,1511,1566]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="AB53F74C0E26CB1E494BFA28FD34F9E3" pageId="1" pageNumber="2">
<taxonomicName id="4C43ABE10E26CB1E494BFA28FDD9FA03" box="[264,635,1511,1534]" class="Arachnida" family="Bothriuridae" genus="Urophonius" kingdom="Animalia" order="Scorpiones" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9370C700E26CB1E494BFA28FE24FA03" box="[264,390,1511,1534]" italics="true" pageId="1" pageNumber="2">Urophonius</emphasis>
sp.
<treatmentCitation id="0AE2F6730E26CB1E49FBFA28FDD9FA03" author="Maury, E. A." box="[440,635,1511,1534]" page="67" pageId="1" pageNumber="2" year="1979">
<bibRefCitation id="EFD2AD930E26CB1E49FBFA28FDD9FA03" author="Maury, E. A." box="[440,635,1511,1534]" pageId="1" pageNumber="2" pagination="57 - 68" refId="ref5947" refString="Maury, E. A. (1979 a) Escorpiofauna patagonica. II. Urophonius granulatus Pocock 1898 (Bothriuridae). Physis, Secc. C, 38 (94), 57 - 68." type="journal article" year="1979">Maury, 1979a: 67</bibRefCitation>
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AE2F6730E26CB1E4AC9FA28FCE4FA03" author="Acosta, L. E." box="[650,838,1511,1534]" page="28" pageId="1" pageNumber="2" year="1986">
<bibRefCitation id="EFD2AD930E26CB1E4AC9FA28FCE4FA03" author="Acosta, L. E." box="[650,838,1511,1534]" pageId="1" pageNumber="2" pagination="28" refId="ref5538" refString="Acosta, L. E. (1986) Escorpiofauna de la Meseta de Somuncura (provincia de Rio Negro, Argentina). IIIras Jornadas Cientificas de la Sociedad de Biologia de Cordoba, p. 28." type="book chapter" year="1986">Acosta, 1986: 28</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0AE2F6730E26CB1E4B17FA28FC15FA03" author="Acosta, L. E." box="[852,951,1511,1534]" page="27" pageId="1" pageNumber="2" year="1988">
<bibRefCitation id="EFD2AD930E26CB1E4B17FA28FC15FA03" author="Acosta, L. E." box="[852,951,1511,1534]" pageId="1" pageNumber="2" pagination="23 - 33" refId="ref5577" refString="Acosta, L. E. (1988) Contribucion al conocimiento taxonomico del genero Urophonius Pocock, 1893 (Scorpiones, Bothriuridae). The Journal of Arachnology, 16 (1), 23 - 33." type="journal article" year="1988">1988: 27</bibRefCitation>
</treatmentCitation>
, 32;
<treatmentCitation id="0AE2F6730E26CB1E4BACFA28FA85FA03" author="Acosta, L. &amp; Maury, E. A." box="[1007,1319,1511,1534]" page="554" pageId="1" pageNumber="2" year="1998">
<bibRefCitation id="EFD2AD930E26CB1E4BACFA28FA85FA03" author="Acosta, L. &amp; Maury, E. A." box="[1007,1319,1511,1534]" pageId="1" pageNumber="2" pagination="545 - 559" refId="ref5616" refString="Acosta, L. E &amp; Maury, E. A. (1998) Scorpiones. In: Morrone, J. J. &amp; Coscaron, S. (Eds.) Biodiversidad de Artropodos argentinos. Una perspectiva biotaxonomica. Editorial Sur, La Plata. pp. 545 - 559." type="book chapter" year="1998">Acosta &amp; Maury, 1998: 554</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AE2F6730E26CB1E497BF9C8FD30F9E3" author="Ojanguren Affilastro, A. A." box="[312,658,1543,1566]" page="185" pageId="1" pageNumber="2" year="2002">
<bibRefCitation id="EFD2AD930E26CB1E497BF9C8FD30F9E3" author="Ojanguren Affilastro, A. A." box="[312,658,1543,1566]" pageId="1" pageNumber="2" pagination="181 - 186" refId="ref6017" refString="Ojanguren Affilastro, A. A. (2002) Nuevos aportes al conocimiento del genero Urophonius Pocock, 1893 (Scorpiones, Bothriuridae). Revista Iberica de Aracnologia, 6, 181 - 186." type="journal article" year="2002">Ojanguren Affilastro, 2002: 185</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35983E90E26CB1C494BF981FD93FE2B" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BFCD0620E26CB1E494BF981FC5BF8F4" blockId="1.[264,1324,1614,1801]" pageId="1" pageNumber="2">
<emphasis id="B9370C700E26CB1E494BF981FE3CF995" bold="true" box="[264,414,1614,1640]" pageId="1" pageNumber="2">Type series:</emphasis>
<materialsCitation id="3B2BDA3F0E26CB1E49EBF980FC5BF8F4" ID-GBIF-Occurrence="3320658302" collectingDate="1985-12-07" collectingDateMax="1985-12-07" collectingDateMin="1968-12-18" collectionCode="CDA" collectorName="L. Acosta" country="Argentina" county="Argentina" location="Laguna Pelada" municipality="Laguna Pelada" pageId="1" pageNumber="2" specimenCount="1" specimenCount-female="11" specimenCount-juvenile="7" stateProvince="Province of Rio Negro" typeStatus="holotype">
<typeStatus id="54F86EC00E26CB1E49EBF980FDB7F994" box="[424,533,1615,1641]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
<specimenCount id="9D451BEB0E26CB1E4A5CF980FDCDF994" box="[543,623,1615,1641]" pageId="1" pageNumber="2" type="female">female</specimenCount>
(
<collectionCode id="ED5248A70E26CB1E4AC2F980FD63F994" box="[641,705,1615,1641]" country="Canada" httpUri="http://grbio.org/cool/89m4-4i9r" name="Canadian Department of Agriculture" pageId="1" pageNumber="2">CDA</collectionCode>
000.151), 9 female and 7 juvenile paratypes (CDA 000.152), 2 female paratypes (FML), 2 female paratypes (IADIZA), 9 female and 4 juvenile paratypes (LEA 000.232):
<collectingCounty id="629DA8EE0E26CB1E4A39F950FD51F944" box="[634,755,1695,1721]" pageId="1" pageNumber="2">Argentina</collectingCounty>
.
<collectingRegion id="49871E800E26CB1E4ABCF950FBB2F944" box="[767,1040,1695,1721]" pageId="1" pageNumber="2">Province of Río Negro</collectingRegion>
. Meseta de Somuncurá,
<location id="8E9C86B90E26CB1E494BF908FE1BF91C" LSID="urn:lsid:plazi:treatment:03EA61740E26CB15494BFAB9FCD6FD13:8E9C86B90E26CB1E494BF908FE1BF91C" box="[264,441,1735,1761]" country="Argentina" county="Argentina" municipality="Laguna Pelada" name="Laguna Pelada" pageId="1" pageNumber="2" stateProvince="Province of Rio Negro">Laguna Pelada</location>
,
<date id="FFFDF6A20E26CB1E4985F908FD3EF91C" box="[454,668,1735,1761]" pageId="1" pageNumber="2" value="1985-12-07">
<collectingDate id="EFB90F4A0E26CB1E4985F908FD3EF91C" box="[454,668,1735,1761]" pageId="1" pageNumber="2" value="1985-12-07">7 December 1985</collectingDate>
</date>
,
<collectorName id="26B6B5B40E26CB1E4AEAF908FCBEF91C" box="[681,796,1735,1761]" pageId="1" pageNumber="2">L. Acosta</collectorName>
leg. — 1 paratype female (MACN), Meseta de Somuncurá, Laguna Chara,
<date id="FFFDF6A20E26CB1E4A37F920FCF4F8F4" box="[628,854,1775,1801]" pageId="1" pageNumber="2" value="1968-12-18">18 December 1968</date>
, J.M. Cei leg.
</materialsCitation>
</paragraph>
<caption id="DF3C80EA0E25CB1D494BF952FBE5F88D" ID-DOI="http://doi.org/10.5281/zenodo.5014524" ID-Zenodo-Dep="5014524" httpUri="https://zenodo.org/record/5014524/files/figure.png" pageId="2" pageNumber="3" startId="2.[264,383,1693,1717]" targetBox="[296,1291,350,1665]" targetPageId="2">
<paragraph id="8BFCD0620E25CB1D494BF952FBE5F88D" blockId="2.[264,1324,1693,1905]" pageId="2" pageNumber="3">
<emphasis id="B9370C700E25CB1D494BF952FE11F948" bold="true" box="[264,435,1693,1717]" pageId="2" pageNumber="3">FIGURES 1­8.</emphasis>
<taxonomicName id="4C43ABE10E25CB1D49FEF952FD1EF949" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" box="[445,700,1693,1716]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E25CB1D49FEF952FD1EF949" box="[445,700,1693,1716]" italics="true" pageId="2" pageNumber="3">Urophonius somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E25CB1D4A85F952FCA0F949" box="[710,770,1693,1716]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
1­7: holotype female (CDA), 8: generalised from several individuals. 1, metasomal segment V and telson, lateral view; 2, metasomal segment V, ventral view; 3­4, sternite V and metasomal segments I­II, ventral view, 3, morphology (ventral macrosetae as black dots), 4, pigment pattern (macrosetae as open circles); 5­6, right pedipalp chela, 5, ventromedial view, 6, lateral view; 7, right pedipalp femur, retrolateral view (indicated trichobothrium
<emphasis id="B9370C700E25CB1D4902F895FEEFF88C" box="[321,333,1882,1905]" italics="true" pageId="2" pageNumber="3">e</emphasis>
and macroseta M1). 8: pigment pattern on tergite V. Scale lines: 1 mm.
</paragraph>
</caption>
<paragraph id="8BFCD0620E24CB1C497BFED4FD93FE2B" blockId="3.[264,1324,283,1830]" pageId="3" pageNumber="4">
<emphasis id="B9370C700E24CB1C497BFED4FD69FEC8" bold="true" box="[312,715,283,309]" pageId="3" pageNumber="4">Additional material examined:</emphasis>
<materialsCitation id="3B2BDA3F0E24CB1C4A9FFED3FB27FE7B" ID-GBIF-Occurrence="3320658301" collectingDate="1985-01-12" collectingDateMax="1985-01-14" collectingDateMin="1985-01-12" collectorName="L. Acosta" country="Argentina" county="Laguna Pelada" location="Province of Rio Negro" municipality="Meseta de Somuncura" pageId="3" pageNumber="4" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1">
<collectingCountry id="F35490F20E24CB1C4A9FFED3FCF7FECB" box="[732,853,284,310]" name="Argentina" pageId="3" pageNumber="4">Argentina</collectingCountry>
.
<location id="8E9C86B90E24CB1C4B2AFED3FB37FECB" LSID="urn:lsid:plazi:treatment:03EA61740E26CB15494BFAB9FCD6FD13:8E9C86B90E24CB1C4B2AFED3FB37FECB" box="[873,1173,284,310]" country="Argentina" county="Laguna Pelada" municipality="Meseta de Somuncura" name="Province of Rio Negro" pageId="3" pageNumber="4">Province of Río Negro</location>
:
<collectingMunicipality id="6B984A180E24CB1C4CEAFED3FE32FEA3" pageId="3" pageNumber="4">Meseta de Somuncurá</collectingMunicipality>
,
<collectingCounty id="629DA8EE0E24CB1C49DFFE8BFDEEFEA3" box="[412,588,324,350]" pageId="3" pageNumber="4">Laguna Pelada</collectingCounty>
,
<date id="FFFDF6A20E24CB1C4A1BFE8BFCBEFEA3" box="[600,796,324,350]" pageId="3" pageNumber="4" value="1985-01-12">
<collectingDate id="EFB90F4A0E24CB1C4A1BFE8BFCBEFEA3" box="[600,796,324,350]" pageId="3" pageNumber="4" value="1985-01-12">12 January 1985</collectingDate>
</date>
,
<collectorName id="26B6B5B40E24CB1C4B6BFE8BFC38FEA3" box="[808,922,324,350]" pageId="3" pageNumber="4">L. Acosta</collectorName>
leg.,
<specimenCount id="9D451BEB0E24CB1C4B9FFE8BFBE7FEA3" box="[988,1093,324,350]" pageId="3" pageNumber="4" type="female">1 female</specimenCount>
,
<specimenCount id="9D451BEB0E24CB1C4C13FE8BFB65FEA3" box="[1104,1223,324,350]" pageId="3" pageNumber="4" type="juvenile">1 juvenile</specimenCount>
, under a stone, daytime (
<collectionCode id="ED5248A70E24CB1C4982FEA3FE5EFE7B" box="[449,508,364,390]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14547" lsid="urn:lsid:biocol.org:col:14547" name="University of Lethbridge" pageId="3" pageNumber="4" type="Herbarium">LEA</collectionCode>
000.233); same loc., 13/
<date id="FFFDF6A20E24CB1C4B5DFEA3FC43FE7B" box="[798,993,364,390]" pageId="3" pageNumber="4" value="1985-01-14">
<collectingDate id="EFB90F4A0E24CB1C4B5DFEA3FC43FE7B" box="[798,993,364,390]" pageId="3" pageNumber="4" value="1985-01-14">14 January 1985</collectingDate>
</date>
,
<collectorName id="26B6B5B40E24CB1C4BAEFEA3FBC3FE7B" box="[1005,1121,364,390]" pageId="3" pageNumber="4">L. Acosta</collectorName>
, G
</materialsCitation>
.
<materialsCitation id="3B2BDA3F0E24CB1C4CD2FEA3FD8FFE2B" ID-GBIF-Occurrence="3320658303" collectingDate="1968-12-17" collectorName="J. M. Cei" country="Indonesia" location="Cerro Corona" pageId="3" pageNumber="4" specimenCount="13" specimenCount-female="8" specimenCount-juvenile="5">
<collectingCountry id="F35490F20E24CB1C4CD2FEA3FB78FE7B" box="[1169,1242,364,390]" name="Indonesia" pageId="3" pageNumber="4">Flores</collectingCountry>
leg.,
<specimenCount id="9D451BEB0E24CB1C4D58FEA3FEC4FE53" pageId="3" pageNumber="4" type="female">8 females</specimenCount>
,
<specimenCount id="9D451BEB0E24CB1C4937FE5BFE54FE53" box="[372,502,404,430]" pageId="3" pageNumber="4" type="juvenile">4 juveniles</specimenCount>
(U.V.) (
<collectionCode id="ED5248A70E24CB1C4A18FE5BFD34FE53" box="[603,662,404,430]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14547" lsid="urn:lsid:biocol.org:col:14547" name="University of Lethbridge" pageId="3" pageNumber="4" type="Herbarium">LEA</collectionCode>
000.234);
<location id="8E9C86B90E24CB1C4B59FE5BFC62FE53" LSID="urn:lsid:plazi:treatment:03EA61740E26CB15494BFAB9FCD6FD13:8E9C86B90E24CB1C4B59FE5BFC62FE53" box="[794,960,404,430]" country="Indonesia" name="Cerro Corona" pageId="3" pageNumber="4">Cerro Corona</location>
,
<date id="FFFDF6A20E24CB1C4B8CFE5BFB17FE53" box="[975,1205,404,430]" pageId="3" pageNumber="4" value="1968-12-17">
<collectingDate id="EFB90F4A0E24CB1C4B8CFE5BFB17FE53" box="[975,1205,404,430]" pageId="3" pageNumber="4" value="1968-12-17">17 December 1968</collectingDate>
</date>
,
<collectorName id="26B6B5B40E24CB1C4C87FE5BFA88FE53" box="[1220,1322,404,430]" pageId="3" pageNumber="4">J.M. Cei</collectorName>
leg.,
<specimenCount id="9D451BEB0E24CB1C4901FE73FE14FE2B" box="[322,438,444,470]" pageId="3" pageNumber="4" type="juvenile">1 juvenile</specimenCount>
(
<collectionCode id="ED5248A70E24CB1C4985FE73FD87FE2B" box="[454,549,444,470]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="3" pageNumber="4" type="Museum">MACN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35983E90E24CB1C497BFE2CFA8EFE03" box="[312,1324,483,510]" pageId="3" pageNumber="4" type="etymology">
<paragraph id="8BFCD0620E24CB1C497BFE2CFA8EFE03" blockId="3.[264,1324,283,1830]" box="[312,1324,483,510]" pageId="3" pageNumber="4">
<emphasis id="B9370C700E24CB1C497BFE2CFE6AFE00" bold="true" box="[312,456,483,509]" pageId="3" pageNumber="4">Etymology:</emphasis>
The species is named after the Meseta de Somuncurá; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C35983E90E24CB1C497BFDC4FA8AFC4B" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="8BFCD0620E24CB1C497BFDC4FA8AFC4B" blockId="3.[264,1324,283,1830]" pageId="3" pageNumber="4">
<emphasis id="B9370C700E24CB1C497BFDC4FE18FDD8" bold="true" box="[312,442,523,549]" pageId="3" pageNumber="4">Diagnosis:</emphasis>
Member of the '
<taxonomicName id="4C43ABE10E24CB1C4A3FFDC3FD59FDD8" authorityName="Pocock" authorityYear="1898" box="[636,763,524,549]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E24CB1C4A3FFDC3FD59FDD8" box="[636,763,524,549]" italics="true" pageId="3" pageNumber="4">granulatus</emphasis>
</taxonomicName>
species group' (
<bibRefCitation id="EFD2AD930E24CB1C4BF5FDC3FBEDFDDB" author="Acosta, L. E." box="[950,1103,524,550]" pageId="3" pageNumber="4" pagination="23 - 33" refId="ref5577" refString="Acosta, L. E. (1988) Contribucion al conocimiento taxonomico del genero Urophonius Pocock, 1893 (Scorpiones, Bothriuridae). The Journal of Arachnology, 16 (1), 23 - 33." type="journal article" year="1988">Acosta 1988</bibRefCitation>
), as defined by the ventral pigment pattern of metasoma (ventrolateral and axial stripes, not joining in any segment), the number of ventral macrosetae on metasomal segments I­II (2+2, 3+3 respectively), and by the position of trichobothrium
<emphasis id="B9370C700E24CB1C4B72FD4BFC9CFD60" box="[817,830,644,669]" italics="true" pageId="3" pageNumber="4">e</emphasis>
of femur (close to macroseta M1); since males of the new species are unknown, it remains to be determined if features of the lobe region of the hemispermatophore fit into the group definition. Ventral surface of sternite V and metasomal segment I granulous, only VL carinae discernible. Insertion of ventral setae on metasomal segments I­II forming a hemicircular tegumentary border. Carapace pigment almost continuous from the eye mound to the anterior border. Lateral pigment stripe on metasomal segments I­IV not joining the ventrolateral pigment. The nearest relatives of
<taxonomicName id="4C43ABE10E24CB1C494BFC53FE0CFC48" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" box="[264,430,924,949]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E24CB1C494BFC53FE0CFC48" box="[264,430,924,949]" italics="true" pageId="3" pageNumber="4">U. somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E24CB1C49F5FC53FE56FC4B" box="[438,500,924,950]" pageId="3" pageNumber="4" rank="species">n. sp.</taxonomicNameLabel>
are
<taxonomicName id="4C43ABE10E24CB1C4A64FC53FD6BFC48" authorityName="Pocock" authorityYear="1898" box="[551,713,924,949]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E24CB1C4A64FC53FD6BFC48" box="[551,713,924,949]" italics="true" pageId="3" pageNumber="4">U. granulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C43ABE10E24CB1C4B41FC53FC47FC48" authorityName="Cekalovic" authorityYear="1981" box="[770,997,924,949]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tregualemuensis">
<emphasis id="B9370C700E24CB1C4B41FC53FC47FC48" box="[770,997,924,949]" italics="true" pageId="3" pageNumber="4">U. tregualemuensis</emphasis>
</taxonomicName>
(see 'Comparisons' below).
</paragraph>
</subSubSection>
<subSubSection id="C35983E90E24CB15497BFC0CFCD6FD13" lastPageId="10" lastPageNumber="11" pageId="3" pageNumber="4" type="description">
<paragraph id="8BFCD0620E24CB1C497BFC0CFD4FFB83" blockId="3.[264,1324,283,1830]" pageId="3" pageNumber="4">
<emphasis id="B9370C700E24CB1C497BFC0CFE70FC20" bold="true" box="[312,466,963,989]" pageId="3" pageNumber="4">Description:</emphasis>
Small to medium sized bothriurids; total length (adult females) up to
<quantity id="4CBB7D870E24CB1C4D4DFC0BFE97FBFB" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="4" unit="mm" value="32.0">32 mm</quantity>
(measurements of
<typeStatus id="54F86EC00E24CB1C4A58FC23FD24FBFB" box="[539,646,1004,1030]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
:
<tableCitation id="C6C1E5D90E24CB1C4AD0FC23FD48FBFB" box="[659,746,1004,1030]" captionStart="TABLE 1" captionStartId="4.[264,352,285,309]" captionTargetBox="[272,1208,348,1100]" captionTargetPageId="4" captionText="TABLE 1. Measurements (mm) of the holotype female of Urophonius somuncura n. sp." httpUri="http://table.plazi.org/id/DF3C80EA0E23CB1B494BFED2FB14FECB" pageId="3" pageNumber="4" tableUuid="DF3C80EA0E23CB1B494BFED2FB14FECB">Table 1</tableCitation>
). Males unknown. General colour straw yellow, slightly more orange towards the caudal end of the metasoma; pedipalps and legs lighter, pectines whitish yellow; irregular pigment on carapace, mesosomal tergites, metasoma, pedipalps and legs, very faint in pectines.
</paragraph>
<paragraph id="8BFCD0620E24CB1C497BFB43FCFDFAE3" blockId="3.[264,1324,283,1830]" pageId="3" pageNumber="4">
<emphasis id="B9370C700E24CB1C497BFB43FDAAFB58" box="[312,520,1164,1189]" italics="true" pageId="3" pageNumber="4">Pigment patterns.</emphasis>
Carapace. With a dense spot between the eyes, the pigment extends, less densely, up to the anterior border, except for some small clear areas; posterior half with larger depigmented areas, the pigment forms several diagonal stripes and an enlarged triangular spot on each side of the posterior border.
</paragraph>
<paragraph id="8BFCD0620E24CB1C497BFAE3FD16F8DB" blockId="3.[264,1324,283,1830]" pageId="3" pageNumber="4">
Tergites. With pigment very similar to
<taxonomicName id="4C43ABE10E24CB1C4B40FAE3FC05FAB8" authorityName="Pocock" authorityYear="1898" box="[771,935,1324,1349]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E24CB1C4B40FAE3FC05FAB8" box="[771,935,1324,1349]" italics="true" pageId="3" pageNumber="4">U. granulatus</emphasis>
</taxonomicName>
: general appearance of four longitudinal dark stripes intercalated with five more or less depigmented areas on tergites I­ VI; the median clear area is approximately diamond­shaped on the posterior half of tergites III­VI; paramedian pigment consisting in a darker basal portion, an inclined or bracket­shaped middle part, and a fainter medial­anterior portion, often joining medially the opposite paramedian stripe; each paramedian stripe is aligned to a well­defined spot on the pretergite; the paralateral pigment, with an irregular medial edge, is continued in the pretergite, too. Tergite VII: border of the pretergite with a dark line, only medially interrupted; on the tergite proper, two subtrapezoidal dark spots with open areas inside, reaching the posterior border but separated from the anterior and lateral borders, and from each other (clear median line, except for faint anterior projections connecting medially). Sternites I­V depigmented, except for the lateral edges on sternites II­V, and three small dots on the caudal border of the last sternite.
</paragraph>
<caption id="DF3C80EA0E23CB1B494BFED2FB14FECB" ID-Table-UUID="DF3C80EA0E23CB1B494BFED2FB14FECB" box="[264,1206,284,310]" httpUri="http://table.plazi.org/id/DF3C80EA0E23CB1B494BFED2FB14FECB" pageId="4" pageNumber="5" startId="4.[264,352,285,309]" targetBox="[272,1208,348,1100]" targetIsTable="true" targetPageId="4">
<paragraph id="8BFCD0620E23CB1B494BFED2FB14FECB" blockId="4.[264,1206,284,310]" box="[264,1206,284,310]" pageId="4" pageNumber="5">
<emphasis id="B9370C700E23CB1B494BFED2FED6FEC8" bold="true" box="[264,372,285,309]" pageId="4" pageNumber="5">TABLE 1</emphasis>
. Measurements (mm) of the holotype female of
<taxonomicName id="4C43ABE10E23CB1B4B39FED1FBD7FEC8" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" box="[890,1141,286,309]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E23CB1B4B39FED1FBD7FEC8" box="[890,1141,286,309]" italics="true" pageId="4" pageNumber="5">Urophonius somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E23CB1B4C3FFED1FB14FECB" box="[1148,1206,284,310]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BFCD0620E23CB1B4953FE93FBDDFBB1" pageId="4" pageNumber="5">
<table id="F94322C20E2334E04953FE93FB1AFBB1" box="[272,1208,348,1100]" gridcols="2" gridrows="17" pageId="4" pageNumber="5">
<tr id="3573D2200E2334E04953FE93FB1AFE8E" box="[272,1208,348,371]" gridrow="0" pageId="4" pageNumber="5">
<th id="76A2BB5C0E2334E04953FE93FD50FE8E" box="[272,754,348,371]" gridcol="0" gridrow="0" pageId="4" pageNumber="5">Body length</th>
<th id="76A2BB5C0E2334E04C6BFE93FB1AFE8E" box="[1064,1208,348,371]" gridcol="1" gridrow="0" pageId="4" pageNumber="5">31.8</th>
</tr>
<tr id="3573D2200E2334E04953FE45FB1AFE5C" box="[272,1208,394,417]" gridrow="1" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FE45FD50FE5C" box="[272,754,394,417]" gridcol="0" gridrow="1" pageId="4" pageNumber="5">Carapace length</td>
<td id="76A2BB5C0E2334E04C6BFE45FB1AFE5C" box="[1064,1208,394,417]" gridcol="1" gridrow="1" pageId="4" pageNumber="5">4.1</td>
</tr>
<tr id="3573D2200E2334E04953FE77FB1AFE32" box="[272,1208,440,463]" gridrow="2" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FE77FD50FE32" box="[272,754,440,463]" gridcol="0" gridrow="2" pageId="4" pageNumber="5">anterior / posterior width</td>
<td id="76A2BB5C0E2334E04C6BFE77FB1AFE32" box="[1064,1208,440,463]" gridcol="1" gridrow="2" pageId="4" pageNumber="5">2.8 / 4.4</td>
</tr>
<tr id="3573D2200E2334E04953FE2AFB1AFE01" box="[272,1208,485,508]" gridrow="3" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FE2AFD50FE01" box="[272,754,485,508]" gridcol="0" gridrow="3" pageId="4" pageNumber="5">Mesosoma length</td>
<td id="76A2BB5C0E2334E04C6BFE2AFB1AFE01" box="[1064,1208,485,508]" gridcol="1" gridrow="3" pageId="4" pageNumber="5">8.9</td>
</tr>
<tr id="3573D2200E2334E04953FDDDFB1AFDD4" box="[272,1208,530,553]" gridrow="4" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FDDDFD50FDD4" box="[272,754,530,553]" gridcol="0" gridrow="4" pageId="4" pageNumber="5">Metasoma length</td>
<td id="76A2BB5C0E2334E04C6BFDDDFB1AFDD4" box="[1064,1208,530,553]" gridcol="1" gridrow="4" pageId="4" pageNumber="5">18.8</td>
</tr>
<tr id="3573D2200E2334E04953FD8FFB1AFDAA" box="[272,1208,576,599]" gridrow="5" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FD8FFD50FDAA" box="[272,754,576,599]" gridcol="0" gridrow="5" pageId="4" pageNumber="5">Metasomal segment I length / width</td>
<td id="76A2BB5C0E2334E04C6BFD8FFB1AFDAA" box="[1064,1208,576,599]" gridcol="1" gridrow="5" pageId="4" pageNumber="5">1.9 / 2.5</td>
</tr>
<tr id="3573D2200E2334E04953FDA1FB1AFD78" box="[272,1208,622,645]" gridrow="6" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FDA1FD50FD78" box="[272,754,622,645]" gridcol="0" gridrow="6" pageId="4" pageNumber="5">Metasomal segment II length / width</td>
<td id="76A2BB5C0E2334E04C6BFDA1FB1AFD78" box="[1064,1208,622,645]" gridcol="1" gridrow="6" pageId="4" pageNumber="5">2.2 / 2.3</td>
</tr>
<tr id="3573D2200E2334E04953FD54FB1AFD4F" box="[272,1208,667,690]" gridrow="7" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FD54FD50FD4F" box="[272,754,667,690]" gridcol="0" gridrow="7" pageId="4" pageNumber="5">Metasomal segment III length / width</td>
<td id="76A2BB5C0E2334E04C6BFD54FB1AFD4F" box="[1064,1208,667,690]" gridcol="1" gridrow="7" pageId="4" pageNumber="5">2.3 / 2.2</td>
</tr>
<tr id="3573D2200E2334E04953FD07FB1AFD22" box="[272,1208,712,735]" gridrow="8" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FD07FD50FD22" box="[272,754,712,735]" gridcol="0" gridrow="8" pageId="4" pageNumber="5">Metasomal segment IV length / width</td>
<td id="76A2BB5C0E2334E04C6BFD07FB1AFD22" box="[1064,1208,712,735]" gridcol="1" gridrow="8" pageId="4" pageNumber="5">2.9 / 2.2</td>
</tr>
<tr id="3573D2200E2334E04953FD39FB1AFCF0" box="[272,1208,758,781]" gridrow="9" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FD39FD50FCF0" box="[272,754,758,781]" gridcol="0" gridrow="9" pageId="4" pageNumber="5">Metasomal segment V length / width / height</td>
<td id="76A2BB5C0E2334E04C6BFD39FB1AFCF0" box="[1064,1208,758,781]" gridcol="1" gridrow="9" pageId="4" pageNumber="5">4.3 / 2.1 / 1.9</td>
</tr>
<tr id="3573D2200E2334E04953FCECFB1AFCC7" box="[272,1208,803,826]" gridrow="10" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FCECFD50FCC7" box="[272,754,803,826]" gridcol="0" gridrow="10" pageId="4" pageNumber="5">Telson length / width / height</td>
<td id="76A2BB5C0E2334E04C6BFCECFB1AFCC7" box="[1064,1208,803,826]" gridcol="1" gridrow="10" pageId="4" pageNumber="5">5.2 / 1.9 / 1.5</td>
</tr>
<tr id="3573D2200E2334E04953FC9EFB1AFC95" box="[272,1208,849,872]" gridrow="11" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FC9EFD50FC95" box="[272,754,849,872]" gridcol="0" gridrow="11" pageId="4" pageNumber="5">Aculeus length</td>
<td id="76A2BB5C0E2334E04C6BFC9EFB1AFC95" box="[1064,1208,849,872]" gridcol="1" gridrow="11" pageId="4" pageNumber="5">1.3</td>
</tr>
<tr id="3573D2200E2334E04953FCB0FB1AFC6B" box="[272,1208,895,918]" gridrow="12" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FCB0FD50FC6B" box="[272,754,895,918]" gridcol="0" gridrow="12" pageId="4" pageNumber="5">Pedipalp total length</td>
<td id="76A2BB5C0E2334E04C6BFCB0FB1AFC6B" box="[1064,1208,895,918]" gridcol="1" gridrow="12" pageId="4" pageNumber="5">13.4</td>
</tr>
<tr id="3573D2200E2334E04953FC63FB1AFC3E" box="[272,1208,940,963]" gridrow="13" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FC63FD50FC3E" box="[272,754,940,963]" gridcol="0" gridrow="13" pageId="4" pageNumber="5">Femur length / width</td>
<td id="76A2BB5C0E2334E04C6BFC63FB1AFC3E" box="[1064,1208,940,963]" gridcol="1" gridrow="13" pageId="4" pageNumber="5">3.5 / 1.0</td>
</tr>
<tr id="3573D2200E2334E04953FC16FB1AFC0D" box="[272,1208,985,1008]" gridrow="14" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FC16FD50FC0D" box="[272,754,985,1008]" gridcol="0" gridrow="14" pageId="4" pageNumber="5">Patella length / width</td>
<td id="76A2BB5C0E2334E04C6BFC16FB1AFC0D" box="[1064,1208,985,1008]" gridcol="1" gridrow="14" pageId="4" pageNumber="5">3.8 / 1.3</td>
</tr>
<tr id="3573D2200E2334E04953FBC8FB1AFBE3" box="[272,1208,1031,1054]" gridrow="15" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FBC8FD50FBE3" box="[272,754,1031,1054]" gridcol="0" gridrow="15" pageId="4" pageNumber="5">Chela length / width / height</td>
<td id="76A2BB5C0E2334E04C6BFBC8FB1AFBE3" box="[1064,1208,1031,1054]" gridcol="1" gridrow="15" pageId="4" pageNumber="5">6.1 / 1.2 / 1.3</td>
</tr>
<tr id="3573D2200E2334E04953FBFAFB1AFBB1" box="[272,1208,1077,1100]" gridrow="16" pageId="4" pageNumber="5">
<td id="76A2BB5C0E2334E04953FBFAFD50FBB1" box="[272,754,1077,1100]" gridcol="0" gridrow="16" pageId="4" pageNumber="5">Movable finger length</td>
<td id="76A2BB5C0E2334E04C6BFBFAFB1AFBB1" box="[1064,1208,1077,1100]" gridcol="1" gridrow="16" pageId="4" pageNumber="5">4.0</td>
</tr>
</table>
</paragraph>
<paragraph id="8BFCD0620E23CB1B497BFB48FE20F88C" blockId="4.[264,1324,1159,1905]" pageId="4" pageNumber="5">
Metasoma. Segments I­IV: a dark line along the DL carinae, in some specimens dashed; on segments I­III this line reaches the posterior border, diverging from there laterally and almost touching the lateral pigment; the dorsal surface has paired spots on segments I­III (in a few specimens joining to form a single median spot), as diagonal stripes on segment I, as shield­like spots on segments II­III, in all cases touching the DL lines in the middle; segment IV only has tenuous longitudinal lines on the dorsal face; lateral side of segments I­IV with a well­defined dark stripe, with irregular borders, it remains independent both from the dorsal and the ventral pigment; ventral face with well­defined ventrolateral and axial lines, which do not coalesce all along the metasoma; segment I is variable concerning the ventral pigment, in many specimens very faint or almost vanishing; this variation ranges from 'axial line present', bordered by vestigial paramedian 'ramifications' originating in the ventrolateral stripes (as in the
<typeStatus id="54F86EC00E23CB1B4B9BF9F0FBE1F9A4" box="[984,1091,1599,1625]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
:
<figureCitation id="1378CCE70E23CB1B4C10F9F0FB3FF9A4" box="[1107,1181,1599,1625]" captionStart="FIGURES 1­8" captionStartId="2.[264,383,1693,1717]" captionTargetBox="[296,1291,350,1665]" captionTargetId="figure-105@2.[268,1320,329,1682]" captionTargetPageId="2" captionText="FIGURES 1­8. Urophonius somuncura n. sp. 1­7: holotype female (CDA), 8: generalised from several individuals. 1, metasomal segment V and telson, lateral view; 2, metasomal segment V, ventral view; 3­4, sternite V and metasomal segments I­II, ventral view, 3, morphology (ventral macrosetae as black dots), 4, pigment pattern (macrosetae as open circles); 5­6, right pedipalp chela, 5, ventromedial view, 6, lateral view; 7, right pedipalp femur, retrolateral view (indicated trichobothrium e and macroseta M1). 8: pigment pattern on tergite V. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5014524" httpUri="https://zenodo.org/record/5014524/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
) to 'ventral surface nearly completely depigmented' (see Variability); the axial line can also be slightly interrupted shortly before the posterior border on segment II, less frequently on segment III. Segment V: ventral axial line complete, lateroventral stripes widening posteriorly (the area has no dense pigment but a rather 'dotted' appearance); on the caudal end the pigment expands to the lateral side (joining there an irregular lateral stripe), and to the dorsal side, forming a dorsolateral stripe. Telson with brown pigment on the ventral side, dorsal side yellowish.
</paragraph>
<paragraph id="8BFCD0620E22CB1A497BFED3FB5CFE7B" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">Pedipalps. Pro and retrodorsal borders of femur with irregular pigment; patella with similar pigmentation, plus a dark stripe on the retrolateral surface; chela with longitudinal, irregular stripes; legs irregularly spotted, being more remarkable on femur and patella.</paragraph>
<paragraph id="8BFCD0620E22CB1A497BFE5BFE52FA6B" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">
<emphasis id="B9370C700E22CB1A497BFE5BFE65FE50" box="[312,455,404,429]" italics="true" pageId="5" pageNumber="6">Morphology</emphasis>
. Carapace, tergites I­VI and sternites I­IV smooth; tergite VII with granulous carinae on the posterior half; sternite V with a median­posterior granular area. Metasomal carinae on segments I­IV: DL complete, denticulate and conspicuous; LSM also complete, formed by small granules roughly aligned; the surface between DL and LSM with scattered small granules; LIM less defined, granulous, forming an irregular row on segments I­II, but almost disappearing in a less granulous sector on segments III­IV; VL less developed, yet well defined on segments I­II, they become tenuous on segments III­ IV; on segment I they delimit a ventral area, irregularly granulous, which represents the undefined VSM; the granulous ventral area is less accentuated on segment II, almost disappearing on III­IV; 2+2 ventral macrosetae on segment I, 3+3 on segments II­III; on segments I­II, the insertion of each macroseta is anteriorly bordered by a slight tegumentary hemicircle, often formed by an arch of granules. Segment V: DL restricted to the basal quarter, with large, blunt granules; LM represented by an inconspicuous 'stripe' of tiny granules, which joins caudally to the ventral granulation; ventral side densely granulous; VL and VM carinae complete, they are normally somewhat hidden by the general granulation, but can still be recognised by their larger granules (especially on the posterior half) and because they are on slightly elevated ridges; VM widening posteriorly into a triangular granulous area; the ventral granulation leaves only smooth basal sectors on each side of VM. Telson irregularly granulous on the ventral side, dorsally smooth. Pedipalps: femur slender, with granulous pro­ and retrodorsal borders, trichobothrium
<emphasis id="B9370C700E22CB1A4C69FB43FB95FB58" box="[1066,1079,1164,1189]" italics="true" pageId="5" pageNumber="6">e</emphasis>
situated close below the macroseta M1 or a little more basally; chela slender, with relatively long fingers; cutting edge of fingers with one single row of granules, and 5+5 accesory granules; like in females of other
<taxonomicName id="4C43ABE10E22CB1A4994FACBFD61FAE3" box="[471,707,1284,1310]" class="Arachnida" family="Bothriuridae" genus="Urophonius" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9370C700E22CB1A4994FACBFDC0FAE0" box="[471,610,1284,1309]" italics="true" pageId="5" pageNumber="6">Urophonius</emphasis>
species
</taxonomicName>
, the medial side of the chela shows a small (vestigial?) tubercle near
<emphasis id="B9370C700E22CB1A49B8FAE3FDB0FAB8" box="[507,530,1324,1349]" italics="true" pageId="5" pageNumber="6">ib</emphasis>
. Trichobothriotaxy as for the genus. Tarsal spine count:
<typeStatus id="54F86EC00E22CB1A4C87FAE3FA89FABB" box="[1220,1323,1324,1350]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
with III 5­6 / 5­6, IV 6­7 / 5­6 (see Variability). Pectinal teeth 13 to 16,
<typeStatus id="54F86EC00E22CB1A4C1AFA9BFB62FA93" box="[1113,1216,1364,1390]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
with 16­ 15 (see Variability).
</paragraph>
<paragraph id="8BFCD0620E22CB1A497BFA6BFB65F9CB" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">
<emphasis id="B9370C700E22CB1A497BFA6BFE16FA40" box="[312,436,1444,1469]" italics="true" pageId="5" pageNumber="6">Variability</emphasis>
: Tarsal spine formula, based on
<specimenCount id="9D451BEB0E22CB1A4B06FA6BFC48FA43" box="[837,1002,1444,1470]" pageId="5" pageNumber="6" type="generic">50 specimens</specimenCount>
, with some tarsi lost (in a pair, the first number refers to the retrolateral row:
<bibRefCitation id="EFD2AD930E22CB1A4B23FA03FC57FA1B" author="Maury, E. A." box="[864,1013,1484,1510]" pageId="5" pageNumber="6" pagination="143 - 160" refId="ref5903" refString="Maury, E. A. (1977) Comentarios sobre dos especies de escorpiones del genero Urophonius (Bothriuridae). Revista del Museo Argentino de Ciencias Naturales ' Bernardino Rivadavia', Entomologia, 5 (7), 143 - 160." type="journal article" year="1977">Maury 1977</bibRefCitation>
). Frequencies on T III (
<emphasis id="B9370C700E22CB1A4D48FA03FAB8FA18" box="[1291,1306,1484,1509]" italics="true" pageId="5" pageNumber="6">n</emphasis>
= 97 tarsi): 4­4 spines (2 tarsi), 3­5 (2), 4­5 (15), 4­6 (3), 5­5 (39), 5­6 (35), 5­7 (1). Frequencies on T IV (
<emphasis id="B9370C700E22CB1A49E8F9D3FE18F9C8" box="[427,442,1564,1589]" italics="true" pageId="5" pageNumber="6">n</emphasis>
= 94): 4­5 (1), 4­6 (2), 5­5 (4), 5­6 (38), 6­6 (15), 6­7 (32), 5­7 (2).
</paragraph>
<paragraph id="8BFCD0620E22CB1A497BF98BFD35F97B" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">
Number of pectinal teeth. Frequencies in adult females (
<emphasis id="B9370C700E22CB1A4B9DF98BFC4FF9A0" box="[990,1005,1604,1629]" italics="true" pageId="5" pageNumber="6">n</emphasis>
= 66 pectines): 13 teeth (2 pectines), 14 (20), 15 (35), 16 (9).
</paragraph>
<paragraph id="8BFCD0620E22CB1A497BF95BFE3FF8B3" blockId="5.[264,1324,284,1870]" pageId="5" pageNumber="6">
Ventral pigment on metasomal segment I (
<emphasis id="B9370C700E22CB1A4B69F95BFC9BF950" box="[810,825,1684,1709]" italics="true" pageId="5" pageNumber="6">n</emphasis>
=
<specimenCount id="9D451BEB0E22CB1A4B12F95BFC54F953" box="[849,1014,1684,1710]" pageId="5" pageNumber="6" type="generic">50 specimens</specimenCount>
): with evident paramedian spots, median line well developed (
<specimenCount id="9D451BEB0E22CB1A4AE9F973FCE3F92B" box="[682,833,1724,1750]" pageId="5" pageNumber="6" type="generic">8 specimens</specimenCount>
); with just small traces of the paramedian spots, median line may be more or less reduced (18); no trace of the paramedial spots, median line either well developed or reduced (15); all parts of ventral pigment absent or vestigial (8).
</paragraph>
<paragraph id="8BFCD0620E21CB19497BFED4FB1DFD8B" blockId="6.[264,1324,283,1630]" pageId="6" pageNumber="7">
<emphasis id="B9370C700E21CB19497BFED4FE45FEC8" bold="true" box="[312,487,283,309]" pageId="6" pageNumber="7">Comparisons:</emphasis>
<taxonomicName id="4C43ABE10E21CB1949B3FED3FD3AFEC8" authorityName="Acosta" authorityYear="2003" box="[496,664,284,309]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="somuncura">
<emphasis id="B9370C700E21CB1949B3FED3FD3AFEC8" box="[496,664,284,309]" italics="true" pageId="6" pageNumber="7">U. somuncura</emphasis>
</taxonomicName>
differs from its nearest relatives,
<taxonomicName id="4C43ABE10E21CB194C6EFED3FB73FEC8" authorityName="Pocock" authorityYear="1898" box="[1069,1233,284,309]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E21CB194C6EFED3FB73FEC8" box="[1069,1233,284,309]" italics="true" pageId="6" pageNumber="7">U. granulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C43ABE10E21CB194D4DFED3FE65FEA0" authorityName="Cekalovic" authorityYear="1981" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tregualemuensis">
<emphasis id="B9370C700E21CB194D4DFED3FE65FEA0" italics="true" pageId="6" pageNumber="7">U. tregualemuensis</emphasis>
</taxonomicName>
, by the following characters: VL and VSM carinae on metasomal segment I well developed and granulous in
<taxonomicName id="4C43ABE10E21CB194ADFFEA3FC9CFE78" authorityName="Pocock" authorityYear="1898" box="[668,830,364,389]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E21CB194ADFFEA3FC9CFE78" box="[668,830,364,389]" italics="true" pageId="6" pageNumber="7">U. granulatus</emphasis>
</taxonomicName>
, forming conspicuous tegumentary elevations (
<bibRefCitation id="EFD2AD930E21CB194913FE5BFE53FE53" author="Maury, E. A." box="[336,497,404,430]" pageId="6" pageNumber="7" pagination="57 - 68" refId="ref5947" refString="Maury, E. A. (1979 a) Escorpiofauna patagonica. II. Urophonius granulatus Pocock 1898 (Bothriuridae). Physis, Secc. C, 38 (94), 57 - 68." type="journal article" year="1979">Maury 1979a</bibRefCitation>
,
<bibRefCitation id="EFD2AD930E21CB1949BFFE5BFD36FE53" author="Acosta, L. E." box="[508,660,404,430]" pageId="6" pageNumber="7" pagination="23 - 33" refId="ref5577" refString="Acosta, L. E. (1988) Contribucion al conocimiento taxonomico del genero Urophonius Pocock, 1893 (Scorpiones, Bothriuridae). The Journal of Arachnology, 16 (1), 23 - 33." type="journal article" year="1988">Acosta 1988</bibRefCitation>
); in
<taxonomicName id="4C43ABE10E21CB194A84FE5BFC0BFE50" authorityName="Cekalovic" authorityYear="1981" box="[711,937,404,429]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tregualemuensis">
<emphasis id="B9370C700E21CB194A84FE5BFC0BFE50" box="[711,937,404,429]" italics="true" pageId="6" pageNumber="7">U. tregualemuensis</emphasis>
</taxonomicName>
the VSM carinae are low but still recognizable as an irregular row of pearl shaped granules, spreading also in a granulous area towards VL; in
<taxonomicName id="4C43ABE10E21CB194A45FE2BFD58FE00" authority=", VSM" authorityName="VSM" box="[518,762,484,510]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="somuncura">
<emphasis id="B9370C700E21CB194A45FE2BFD12FE00" box="[518,688,484,509]" italics="true" pageId="6" pageNumber="7">U. somuncura</emphasis>
, VSM
</taxonomicName>
carinae no longer exist, but instead are completely dispersed in a flat ventral area, with small scattered granules. Similarly, ventral carinae are well developed on the posterior half of sternite V of
<taxonomicName id="4C43ABE10E21CB194B9CFDFBFB21FDB0" authorityName="Pocock" authorityYear="1898" box="[991,1155,564,589]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E21CB194B9CFDFBFB21FDB0" box="[991,1155,564,589]" italics="true" pageId="6" pageNumber="7">U. granulatus</emphasis>
</taxonomicName>
, but not in
<taxonomicName id="4C43ABE10E21CB194D4DFDFBFE65FD88" authorityName="Cekalovic" authorityYear="1981" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tregualemuensis">
<emphasis id="B9370C700E21CB194D4DFDFBFE65FD88" italics="true" pageId="6" pageNumber="7">U. tregualemuensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C43ABE10E21CB194A43FD93FD04FD88" box="[512,678,604,629]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="somuncura">
<emphasis id="B9370C700E21CB194A43FD93FD04FD88" box="[512,678,604,629]" italics="true" pageId="6" pageNumber="7">U. somuncura</emphasis>
</taxonomicName>
, just represented by a posterior granular area.
</paragraph>
<paragraph id="8BFCD0620E21CB19497BFD4BFCB8FCC3" blockId="6.[264,1324,283,1630]" pageId="6" pageNumber="7">
For both
<taxonomicName id="4C43ABE10E21CB1949E7FD4BFD25FD60" authorityName="Cekalovic" authorityYear="1981" box="[420,647,644,669]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tregualemuensis">
<emphasis id="B9370C700E21CB1949E7FD4BFD25FD60" box="[420,647,644,669]" italics="true" pageId="6" pageNumber="7">U. tregualemuensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C43ABE10E21CB194A83FD4BFCC4FD60" box="[704,870,644,669]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="somuncura">
<emphasis id="B9370C700E21CB194A83FD4BFCC4FD60" box="[704,870,644,669]" italics="true" pageId="6" pageNumber="7">U. somuncura</emphasis>
</taxonomicName>
the insertion of the ventral submedian macrosetae on metasomal segments I­II determines the formation of a hemicircular tegumentary border around each seta (see
<bibRefCitation id="EFD2AD930E21CB194A88FD1BFCC6FD13" author="Acosta, L. E." box="[715,868,724,750]" pageId="6" pageNumber="7" pagination="23 - 33" refId="ref5577" refString="Acosta, L. E. (1988) Contribucion al conocimiento taxonomico del genero Urophonius Pocock, 1893 (Scorpiones, Bothriuridae). The Journal of Arachnology, 16 (1), 23 - 33." type="journal article" year="1988">Acosta 1988</bibRefCitation>
: fig. 3); these borders are conspicuous in the Chilean species, and much less noticeable in
<taxonomicName id="4C43ABE10E21CB194BC4FD33FB93FCE8" box="[903,1073,764,789]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="somuncura">
<emphasis id="B9370C700E21CB194BC4FD33FB93FCE8" box="[903,1073,764,789]" italics="true" pageId="6" pageNumber="7">U. somuncura</emphasis>
</taxonomicName>
. Ventral setae of
<taxonomicName id="4C43ABE10E21CB194D4DFD33FE25FCC0" authorityName="Pocock" authorityYear="1898" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E21CB194D4DFD33FE25FCC0" italics="true" pageId="6" pageNumber="7">U. granulatus</emphasis>
</taxonomicName>
do not form tegumentary borders.
</paragraph>
<paragraph id="8BFCD0620E21CB19497BFC83FC55FC23" blockId="6.[264,1324,283,1630]" pageId="6" pageNumber="7">
The position of trichobothrium
<emphasis id="B9370C700E21CB194AE8FC83FD1AFC98" box="[683,696,844,869]" italics="true" pageId="6" pageNumber="7">e</emphasis>
of femur shows a subtle variation among the species in the group (
<bibRefCitation id="EFD2AD930E21CB1949E5FCBBFDE0FC73" author="Acosta, L. E." box="[422,578,884,910]" pageId="6" pageNumber="7" pagination="23 - 33" refId="ref5577" refString="Acosta, L. E. (1988) Contribucion al conocimiento taxonomico del genero Urophonius Pocock, 1893 (Scorpiones, Bothriuridae). The Journal of Arachnology, 16 (1), 23 - 33." type="journal article" year="1988">Acosta 1988</bibRefCitation>
): in
<taxonomicName id="4C43ABE10E21CB194A37FCBBFCBAFC70" authorityName="Pocock" authorityYear="1898" box="[628,792,884,909]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E21CB194A37FCBBFCBAFC70" box="[628,792,884,909]" italics="true" pageId="6" pageNumber="7">U. granulatus</emphasis>
</taxonomicName>
it is more distal than M1; in most specimens of
<taxonomicName id="4C43ABE10E21CB194969FC53FDACFC48" authorityName="Cekalovic" authorityYear="1981" box="[298,526,924,949]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tregualemuensis">
<emphasis id="B9370C700E21CB194969FC53FDACFC48" box="[298,526,924,949]" italics="true" pageId="6" pageNumber="7">U. tregualemuensis</emphasis>
</taxonomicName>
it is clearly more basal, while in
<taxonomicName id="4C43ABE10E21CB194BE1FC53FBEBFC48" box="[930,1097,924,949]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="somuncura">
<emphasis id="B9370C700E21CB194BE1FC53FBEBFC48" box="[930,1097,924,949]" italics="true" pageId="6" pageNumber="7">U. somuncura</emphasis>
</taxonomicName>
it lies close below the previously mentioned macroseta or just a little more basally.
</paragraph>
<paragraph id="8BFCD0620E21CB19497BFC23FDE2FBAB" blockId="6.[264,1324,283,1630]" pageId="6" pageNumber="7">
Like
<taxonomicName id="4C43ABE10E21CB194936FC23FDBBFBF8" authorityName="Pocock" authorityYear="1898" box="[373,537,1004,1029]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E21CB194936FC23FDBBFBF8" box="[373,537,1004,1029]" italics="true" pageId="6" pageNumber="7">U. granulatus</emphasis>
</taxonomicName>
, the metasomal segment I of
<taxonomicName id="4C43ABE10E21CB194B37FC23FBB9FBF8" box="[884,1051,1004,1029]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="somuncura">
<emphasis id="B9370C700E21CB194B37FC23FBB9FBF8" box="[884,1051,1004,1029]" italics="true" pageId="6" pageNumber="7">U. somuncura</emphasis>
</taxonomicName>
always bears 2+2 ventral macrosetae; in
<taxonomicName id="4C43ABE10E21CB1949B5FBDBFD7FFBD0" authorityName="Cekalovic" authorityYear="1981" box="[502,733,1044,1069]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tregualemuensis">
<emphasis id="B9370C700E21CB1949B5FBDBFD7FFBD0" box="[502,733,1044,1069]" italics="true" pageId="6" pageNumber="7">U. tregualemuensis</emphasis>
</taxonomicName>
some variability exists (either 2+2, 2+3 or 3+3 macrosetae;
<bibRefCitation id="EFD2AD930E21CB1949D8FBF3FD91FBAB" author="Acosta, L. E." box="[411,563,1084,1110]" pageId="6" pageNumber="7" pagination="23 - 33" refId="ref5577" refString="Acosta, L. E. (1988) Contribucion al conocimiento taxonomico del genero Urophonius Pocock, 1893 (Scorpiones, Bothriuridae). The Journal of Arachnology, 16 (1), 23 - 33." type="journal article" year="1988">Acosta 1988</bibRefCitation>
).
</paragraph>
<paragraph id="8BFCD0620E21CB19497BFBABFC5FFB33" blockId="6.[264,1324,283,1630]" pageId="6" pageNumber="7">
Both in
<taxonomicName id="4C43ABE10E21CB1949D4FBABFD98FB80" authorityName="Pocock" authorityYear="1898" box="[407,570,1124,1149]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E21CB1949D4FBABFD98FB80" box="[407,570,1124,1149]" italics="true" pageId="6" pageNumber="7">U. granulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C43ABE10E21CB194A30FBABFCB8FB80" box="[627,794,1124,1149]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="somuncura">
<emphasis id="B9370C700E21CB194A30FBABFCB8FB80" box="[627,794,1124,1149]" italics="true" pageId="6" pageNumber="7">U. somuncura</emphasis>
</taxonomicName>
the lateral pigment stripe on metasomal segments I­IV does not join the ventrolateral pigment; in
<taxonomicName id="4C43ABE10E21CB194BC2FB43FBC6FB58" authorityName="Cekalovic" authorityYear="1981" box="[897,1124,1164,1189]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tregualemuensis">
<emphasis id="B9370C700E21CB194BC2FB43FBC6FB58" box="[897,1124,1164,1189]" italics="true" pageId="6" pageNumber="7">U. tregualemuensis</emphasis>
</taxonomicName>
an irregular reticle fills the area between those stripes, thereby connecting them.
</paragraph>
<paragraph id="8BFCD0620E21CB19497BFB13FEF9FA6B" blockId="6.[264,1324,283,1630]" pageId="6" pageNumber="7">
In
<taxonomicName id="4C43ABE10E21CB19491AFB13FDA2FB08" box="[345,512,1244,1269]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="somuncura">
<emphasis id="B9370C700E21CB19491AFB13FDA2FB08" box="[345,512,1244,1269]" italics="true" pageId="6" pageNumber="7">U. somuncura</emphasis>
</taxonomicName>
, the carapace pigment covers almost continuously the area from the eye mound to the front edge, while in
<taxonomicName id="4C43ABE10E21CB194A87FACBFCC5FAE0" authorityName="Pocock" authorityYear="1898" box="[708,871,1284,1309]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E21CB194A87FACBFCC5FAE0" box="[708,871,1284,1309]" italics="true" pageId="6" pageNumber="7">U. granulatus</emphasis>
</taxonomicName>
a transversal clear sector separates the ocular and frontal spots; the pattern in
<taxonomicName id="4C43ABE10E21CB194AA2FAE3FC65FAB8" authorityName="Cekalovic" authorityYear="1981" box="[737,967,1324,1349]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tregualemuensis">
<emphasis id="B9370C700E21CB194AA2FAE3FC65FAB8" box="[737,967,1324,1349]" italics="true" pageId="6" pageNumber="7">U. tregualemuensis</emphasis>
</taxonomicName>
is distinct, since the pigment leaves a large triangular area clear on the anterior border, or at most with a tiny median dot or line.
</paragraph>
<paragraph id="8BFCD0620E21CB19497BFA6BFE50F9F3" blockId="6.[264,1324,283,1630]" pageId="6" pageNumber="7">
Some specimens of
<taxonomicName id="4C43ABE10E21CB194A6BFA6BFD6DFA40" box="[552,719,1444,1469]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="somuncura">
<emphasis id="B9370C700E21CB194A6BFA6BFD6DFA40" box="[552,719,1444,1469]" italics="true" pageId="6" pageNumber="7">U. somuncura</emphasis>
</taxonomicName>
show vestigial paramedian pigment on the ventral side of metasomal segment I (see
<figureCitation id="1378CCE70E21CB194AD0FA03FD7BFA1B" box="[659,729,1484,1510]" captionStart="FIGURES 1­8" captionStartId="2.[264,383,1693,1717]" captionTargetBox="[296,1291,350,1665]" captionTargetId="figure-105@2.[268,1320,329,1682]" captionTargetPageId="2" captionText="FIGURES 1­8. Urophonius somuncura n. sp. 1­7: holotype female (CDA), 8: generalised from several individuals. 1, metasomal segment V and telson, lateral view; 2, metasomal segment V, ventral view; 3­4, sternite V and metasomal segments I­II, ventral view, 3, morphology (ventral macrosetae as black dots), 4, pigment pattern (macrosetae as open circles); 5­6, right pedipalp chela, 5, ventromedial view, 6, lateral view; 7, right pedipalp femur, retrolateral view (indicated trichobothrium e and macroseta M1). 8: pigment pattern on tergite V. Scale lines: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5014524" httpUri="https://zenodo.org/record/5014524/files/figure.png" pageId="6" pageNumber="7">Fig. 4</figureCitation>
), a condition never seen in both
<taxonomicName id="4C43ABE10E21CB194C15FA03FB5BFA18" authorityName="Pocock" authorityYear="1898" box="[1110,1273,1484,1509]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E21CB194C15FA03FB5BFA18" box="[1110,1273,1484,1509]" italics="true" pageId="6" pageNumber="7">U. granulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C43ABE10E21CB19494BFA3BFE49F9F0" authorityName="Cekalovic" authorityYear="1981" box="[264,491,1524,1549]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tregualemuensis">
<emphasis id="B9370C700E21CB19494BFA3BFE49F9F0" box="[264,491,1524,1549]" italics="true" pageId="6" pageNumber="7">U. tregualemuensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFCD0620E21CB19497BF9D3FD86F9A3" blockId="6.[264,1324,283,1630]" pageId="6" pageNumber="7">
To insert the new species in the genus key provided by
<bibRefCitation id="EFD2AD930E21CB194B80F9D3FBCFF9CB" author="Acosta, L. E." box="[963,1133,1564,1590]" pageId="6" pageNumber="7" pagination="23 - 33" refId="ref5577" refString="Acosta, L. E. (1988) Contribucion al conocimiento taxonomico del genero Urophonius Pocock, 1893 (Scorpiones, Bothriuridae). The Journal of Arachnology, 16 (1), 23 - 33." type="journal article" year="1988">Acosta (1988)</bibRefCitation>
, couple 5 has to be modified, as follows:
</paragraph>
<paragraph id="8BFCD0620E21CB19494BF95BFA88F90D" blockId="6.[264,1323,1684,1880]" pageId="6" pageNumber="7">
5. VSM carinae on metasomal segment I and sternite V well defined, on tegumentary elevations. Insertion of ventral setae (segments I­II) does not form tegumentary borders. Trichobothrium
<emphasis id="B9370C700E21CB194D5CF978FA89F933" box="[1311,1323,1719,1742]" italics="true" pageId="6" pageNumber="7">e</emphasis>
more distal than M1
<emphasis id="B9370C700E21CB194A58F916FA88F90D" box="[539,1322,1753,1776]" italics="true" pageId="6" pageNumber="7">
...............................................................................................
<taxonomicName id="4C43ABE10E21CB194CD8F916FA88F90D" authorityName="Pocock" authorityYear="1898" box="[1179,1322,1753,1776]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="granulatus">U. granulatus</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BFCD0620E21CB18494BF933FA89FE1D" blockId="6.[264,1323,1684,1880]" lastBlockId="7.[264,1323,284,654]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
­ VSM carinae on metasomal segment I and sternite V dispersed on a granulous area. Insertion of ventral setae (segments I­II) forms anteriorly a tegumentary hemicircle. Trichobothrium
<emphasis id="B9370C700E21CB194CA1F8D0FB4CF8CB" box="[1250,1262,1823,1846]" italics="true" pageId="6" pageNumber="7">e</emphasis>
close below or more basal than M1...................................................................................................... 6 6. Carapace with an anterior clear triangle free of pigment, with just a tiny spot or line. Mesosomal tergites I­VI with a single median clear stripe. Lateral pigment stripe on metasomal segments I­IV connected with the ventrolateral stripe through a dense reticle. Ventral surface of metasomal segment I with pearl­shaped granulation. Ventral setae of segment I can be either 2+2, 2+3 or 3+3; hemicircular tegumentary elevations around the setae well developed............
<emphasis id="B9370C700E20CB18497AFE06FA89FE1D" box="[313,1323,457,480]" italics="true" pageId="7" pageNumber="8">
........................................................................................................................
<taxonomicName id="4C43ABE10E20CB184C1AFE06FA89FE1D" box="[1113,1323,457,480]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subSpecies" species="treg" subSpecies="ualemuensis">U. treg ualemuensis</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BFCD0620E20CB18494BFE23FA89FD73" blockId="7.[264,1323,284,654]" pageId="7" pageNumber="8">
­ Carapace with pigment filling the preocular area up to the anterior border. Mesosomal tergites I­VI with one median and two paralateral clear stripes. Lateral pigment stripe on metasomal segments I­IV does not join the ventrolateral stripe. Ventral surface of metasomal segment I with small, scattered granulation. Always 2+2 ventral setae on segment I; hemicircular tegumentary elevations around the setae feeble
<emphasis id="B9370C700E20CB184B48FDB8FB4BFD73" box="[779,1257,631,654]" italics="true" pageId="7" pageNumber="8">
.................................................
<taxonomicName id="4C43ABE10E20CB184C14FDB8FB4BFD73" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" box="[1111,1257,631,654]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">U. somuncura</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A204B10B0E20CB184CB1FDB8FA89FD73" box="[1266,1323,631,654]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFCD0620E20CB18497BFD29FC62FA2C" blockId="7.[264,1324,742,1888]" pageId="7" pageNumber="8">
<emphasis id="B9370C700E20CB18497BFD29FD05FCFD" bold="true" box="[312,679,742,768]" pageId="7" pageNumber="8">Habitat and natural history:</emphasis>
<taxonomicName id="4C43ABE10E20CB184AF1FD28FC69FCFD" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" box="[690,971,743,768]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E20CB184AF1FD28FC69FCFD" box="[690,971,743,768]" italics="true" pageId="7" pageNumber="8">Urophonius somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E20CB184B96FD28FBB5FCFC" box="[981,1047,743,769]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
has been hitherto collected only on the higher plains of the Meseta de Somuncurá. According to the detailed descriptions by
<bibRefCitation id="EFD2AD930E20CB184988FCF8FDEFFCAC" author="Cei, J. M." box="[459,589,823,849]" pageId="7" pageNumber="8" pagination="257 - 271" refId="ref5714" refString="Cei, J. M. (1969) La meseta basaltica de Somuncura, Rio Negro. Herpetofauna endemica y sus peculiares equilibrios biocenoticos. Physis, 28 (77), 257 - 271." type="journal article" year="1969">Cei (1969)</bibRefCitation>
and
<bibRefCitation id="EFD2AD930E20CB184AD3FCF8FCC7FCAC" author="Ruiz Leal, A." box="[656,869,823,849]" pageId="7" pageNumber="8" pagination="89 - 118" refId="ref6054" refString="Ruiz Leal, A. (1972) Los confines boreal y austral de las Provincias Patagonica y Central respectivamente. Boletin de la Sociedad Argentina de Botanica, 13 (supl.), 89 - 118." type="journal article" year="1972">Ruiz Leal (1972)</bibRefCitation>
, well­defined altitudinal belts can be identified in this tableland, which are easily recognizable along the road that leads from the locality of Valcheta (
<quantity id="4CBB7D870E20CB184A66FC48FDD2FC5C" box="[549,624,903,929]" metricMagnitude="2" metricUnit="m" metricValue="1.92" pageId="7" pageNumber="8" unit="m" value="192.0">192 m</quantity>
a.s.l.,
<geoCoordinate id="EE77B6A50E20CB184AFFFC48FCBAFC5C" box="[700,792,903,929]" degrees="40" direction="south" minutes="39" orientation="latitude" pageId="7" pageNumber="8" precision="925" value="-40.65">40°39'S</geoCoordinate>
<geoCoordinate id="EE77B6A50E20CB184B5CFC48FC2AFC5C" box="[799,904,903,929]" degrees="66" direction="west" minutes="09" orientation="longitude" pageId="7" pageNumber="8" precision="925" value="-66.15">66°09'W</geoCoordinate>
) up to the Meseta (
<figureCitation id="1378CCE70E20CB184C2BFC48FB0DFC5C" box="[1128,1199,903,929]" captionStart="FIGURE 9" captionStartId="8.[264,368,1315,1339]" captionTargetBox="[277,1311,574,1285]" captionTargetId="figure-292@8.[272,1315,569,1288]" captionTargetPageId="8" captionText="FIGURE 9. Location of the Meseta de Somuncurá (S) in northern Patagonia, Argentina. The type locality is indicated with a star, the Cerro Corona with a black triangle. Dark grey: areas above 900 m; adjacent solid line: 600 m altitude. Dotted line near Valcheta: the 'Arroyo Valcheta' stream. Dashed line: administrative boundary. Inset: location of the represented area in South America." figureDoi="http://doi.org/10.5281/zenodo.5014526" httpUri="https://zenodo.org/record/5014526/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
). Between 200 and
<quantity id="4CBB7D870E20CB184936FC60FE66FC34" box="[373,452,943,969]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="8" unit="m" value="500.0">500 m</quantity>
the typical Monte physiognomy, dominated by
<taxonomicName id="4C43ABE10E20CB184C51FC60FB3CFC34" box="[1042,1182,943,969]" class="Magnoliopsida" family="Zygophyllaceae" genus="Larrea" kingdom="Plantae" order="Zygophyllales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9370C700E20CB184C51FC60FBC0FC35" box="[1042,1122,943,968]" italics="true" pageId="7" pageNumber="8">Larrea</emphasis>
spp.
</taxonomicName>
('jarilla') is observed. On the initial spurs of the tableland some Patagonian elements appear, gradually replacing Monte elements as the ascent progresses; it is thus an ecotonal belt (~
<quantity id="4CBB7D870E20CB184C84FC30FE80FBBC" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="7" pageNumber="8" unit="m" value="700.0" valueMax="900.0" valueMin="500.0">500­900 m</quantity>
), which was denoted by
<bibRefCitation id="EFD2AD930E20CB184A1DFBE8FD43FBBC" author="Cei, J. M." box="[606,737,1063,1089]" pageId="7" pageNumber="8" pagination="257 - 271" refId="ref5714" refString="Cei, J. M. (1969) La meseta basaltica de Somuncura, Rio Negro. Herpetofauna endemica y sus peculiares equilibrios biocenoticos. Physis, 28 (77), 257 - 271." type="journal article" year="1969">Cei (1969)</bibRefCitation>
as 'belt of the ravines' or 'belt of the filtrating streams'. From
<quantity id="4CBB7D870E20CB184982FB80FDACFB94" box="[449,526,1103,1129]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="8" unit="m" value="900.0">900 m</quantity>
upwards the environment has a definite Patagonian character.
<bibRefCitation id="EFD2AD930E20CB184D41FB80FEF4FB6C" author="Cei, J. M." pageId="7" pageNumber="8" pagination="257 - 271" refId="ref5714" refString="Cei, J. M. (1969) La meseta basaltica de Somuncura, Rio Negro. Herpetofauna endemica y sus peculiares equilibrios biocenoticos. Physis, 28 (77), 257 - 271." type="journal article" year="1969">Cei (1969)</bibRefCitation>
named the sector between 900 and
<quantity id="4CBB7D870E20CB184B57FBB8FCD2FB6C" box="[788,880,1143,1169]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="8" unit="m" value="1400.0">1400 m</quantity>
as 'belt of the plains' or 'belt of the argillaceous ponds'. Indeed, the lowest part of this belt (
<quantity id="4CBB7D870E20CB184BE6FB50FB9AFB44" box="[933,1080,1183,1209]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="7" pageNumber="8" unit="m" value="1050.0" valueMax="1200.0" valueMin="900.0">900­1200 m</quantity>
) still shows a rough and undulated landscape, with extensive shrub­sized vegetation, especially formed by the hemispheric bush known as 'neneo' (
<taxonomicName id="4C43ABE10E20CB184AF0FB20FC30FAF5" box="[691,914,1263,1288]" class="Magnoliopsida" family="Apiaceae" genus="Mulinum" kingdom="Plantae" order="Apiales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="spinosum">
<emphasis id="B9370C700E20CB184AF0FB20FC30FAF5" box="[691,914,1263,1288]" italics="true" pageId="7" pageNumber="8">Mulinum spinosum</emphasis>
</taxonomicName>
); near the
<quantity id="4CBB7D870E20CB184C4CFB20FBC5FAF4" box="[1039,1127,1263,1289]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="8" unit="m" value="1200.0">1200 m</quantity>
altitude the conspicuous silverly plants of
<taxonomicName id="4C43ABE10E20CB184A00FAD8FC92FACD" box="[579,816,1303,1328]" class="Magnoliopsida" family="Asteraceae" genus="Senecio" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="filaginoides">
<emphasis id="B9370C700E20CB184A00FAD8FC92FACD" box="[579,816,1303,1328]" italics="true" pageId="7" pageNumber="8">Senecio filaginoides</emphasis>
</taxonomicName>
also appear. The scorpion species
<taxonomicName id="4C43ABE10E20CB184C84FAD8FD32FAA4" authority="Kraepelin, 1894" authorityName="Kraepelin" authorityYear="1894" class="Arachnida" family="Bothriuridae" genus="Bothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="B9370C700E20CB184C84FAD8FE62FAA5" italics="true" pageId="7" pageNumber="8">Bothriurus burmeisteri</emphasis>
Kraepelin, 1894
</taxonomicName>
, widespread in the Monte region (
<bibRefCitation id="EFD2AD930E20CB184C78FAF0FB47FAA4" author="Maury, E. A." box="[1083,1253,1343,1369]" pageId="7" pageNumber="8" pagination="703 - 719" refId="ref5985" refString="Maury, E. A. (1979 b) Apuntes para una zoogeografia de la escorpiofauna argentina. Acta zoologica Lilloana, 35 (2): 703 - 719." type="journal article" year="1979">Maury 1979b</bibRefCitation>
), was found to reach both the ecotonal sector (
<quantity id="4CBB7D870E20CB184AB3FAA8FCD4FA7C" box="[752,886,1383,1409]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="7" pageNumber="8" unit="m" value="700.0" valueMax="900.0" valueMin="500.0">500­900 m</quantity>
) and the lower portion of the plains belt (
<bibRefCitation id="EFD2AD930E20CB184900FA40FE7FFA54" author="Acosta, L. E." box="[323,477,1423,1449]" pageId="7" pageNumber="8" pagination="28" refId="ref5538" refString="Acosta, L. E. (1986) Escorpiofauna de la Meseta de Somuncura (provincia de Rio Negro, Argentina). IIIras Jornadas Cientificas de la Sociedad de Biologia de Cordoba, p. 28." type="book chapter" year="1986">Acosta 1986</bibRefCitation>
); in January, many specimens of
<taxonomicName id="4C43ABE10E20CB184B23FA40FBAAFA55" authorityName="Kraepelin" authorityYear="1894" box="[864,1032,1423,1448]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="burmeisteri">
<emphasis id="B9370C700E20CB184B23FA40FBAAFA55" box="[864,1032,1423,1448]" italics="true" pageId="7" pageNumber="8">B. burmeisteri</emphasis>
</taxonomicName>
were collected at Puesto La Cortadera (ca.
<quantity id="4CBB7D870E20CB18499FFA78FD9AFA2C" box="[476,568,1463,1489]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="m" value="1000.0">1000 m</quantity>
), including a couple in courtship.
</paragraph>
<paragraph id="8BFCD0620E20CB17497BFA10FC93FE03" blockId="7.[264,1324,742,1888]" lastBlockId="8.[264,1324,284,510]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
Above, on the plains proper, the vegetation turns to a grassland dominated by the 'coirón' (
<taxonomicName id="4C43ABE10E20CB184935F9C8FE47F9DC" box="[374,485,1543,1569]" class="Liliopsida" family="Poaceae" genus="Poa" kingdom="Plantae" order="Poales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9370C700E20CB184935F9C8FE07F9DD" box="[374,421,1543,1568]" italics="true" pageId="7" pageNumber="8">Poa</emphasis>
spp.
</taxonomicName>
,
<taxonomicName id="4C43ABE10E20CB1849BBF9C8FDD7F9DC" box="[504,629,1543,1569]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9370C700E20CB1849BBF9C8FD96F9DD" box="[504,564,1543,1568]" italics="true" pageId="7" pageNumber="8">Stipa</emphasis>
spp.
</taxonomicName>
,
<taxonomicName id="4C43ABE10E20CB184ACBF9C8FC84F9DC" box="[648,806,1543,1569]" class="Liliopsida" family="Poaceae" genus="Festuca" kingdom="Plantae" order="Poales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9370C700E20CB184ACBF9C8FD47F9DD" box="[648,741,1543,1568]" italics="true" pageId="7" pageNumber="8">Festuca</emphasis>
spp.
</taxonomicName>
), together with numerous cushion­shaped plants, like the rock­hard
<taxonomicName id="4C43ABE10E20CB184A02F9E0FD72F9B5" box="[577,720,1582,1608]" class="Magnoliopsida" family="Apiaceae" genus="Azorella" kingdom="Plantae" order="Apiales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9370C700E20CB184A02F9E0FD01F9B5" box="[577,675,1583,1608]" italics="true" pageId="7" pageNumber="8">Azorella</emphasis>
sp.
</taxonomicName>
('leña de piedra' = 'stone firewood'). These upper plains are only slightly undulating and have a rigorous climate, with wide daily and seasonal variations in temperature and continuous winds, predominantly from the SW. They contain several depressions or 'ponds' of centripetal drainage, whose borders are partially surrounded by basaltic cliffs of diverse size. The water content of these ponds originates from the sporadic rainfalls, as well as from summer thawing, and may persist for several weeks, even for years in the larger ones, thanks to their argillaceous bottom.
<taxonomicName id="4C43ABE10E20CB184CE3F8D0FE28F89D" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E20CB184CE3F8D0FE28F89D" italics="true" pageId="7" pageNumber="8">Urophonius somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E20CB1849D1F889FE70F89D" box="[402,466,1862,1888]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
was collected in this belt, together with
<taxonomicName id="4C43ABE10E20CB184BECF888FAA8F89D" box="[943,1290,1862,1888]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9370C700E20CB184BECF888FB8CF89D" box="[943,1070,1863,1888]" italics="true" pageId="7" pageNumber="8">Bothriurus</emphasis>
sp. (
<bibRefCitation id="EFD2AD930E20CB184C24F889FAA3F89D" author="Acosta, L. E." box="[1127,1281,1862,1888]" pageId="7" pageNumber="8" pagination="28" refId="ref5538" refString="Acosta, L. E. (1986) Escorpiofauna de la Meseta de Somuncura (provincia de Rio Negro, Argentina). IIIras Jornadas Cientificas de la Sociedad de Biologia de Cordoba, p. 28." type="book chapter" year="1986">Acosta 1986</bibRefCitation>
)
</taxonomicName>
. A few volcanic peaks elevate above the plains level, especially in the SE portion of the Meseta, such as the Cerro Corona (ca.
<quantity id="4CBB7D870E2FCB174A9AFE8BFC95FEA3" box="[729,823,324,350]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="9" unit="m" value="1700.0">1700 m</quantity>
), the highest elevation in Somuncurá;
<bibRefCitation id="EFD2AD930E2FCB174D40FE8BFEF4FE7B" author="Cei, J. M." pageId="8" pageNumber="9" pagination="257 - 271" refId="ref5714" refString="Cei, J. M. (1969) La meseta basaltica de Somuncura, Rio Negro. Herpetofauna endemica y sus peculiares equilibrios biocenoticos. Physis, 28 (77), 257 - 271." type="journal article" year="1969">Cei (1969)</bibRefCitation>
distinguishes here the 'belt of the volcanic peaks' (1400­ca.
<quantity id="4CBB7D870E2FCB174C66FEA3FB2AFE7B" box="[1061,1160,364,390]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="9" unit="m" value="1700.0">1700 m</quantity>
). Most specimens studied of
<taxonomicName id="4C43ABE10E2FCB174991FE5BFDD9FE50" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" box="[466,635,404,429]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E2FCB174991FE5BFDD9FE50" box="[466,635,404,429]" italics="true" pageId="8" pageNumber="9">U. somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E2FCB174AC6FE5BFD65FE53" box="[645,711,404,430]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
come from the
<typeStatus id="54F86EC00E2FCB174BCEFE5BFC1DFE53" box="[909,959,404,430]" pageId="8" pageNumber="9">type</typeStatus>
locality (near the pond called 'Laguna Pelada'), while material already examined by
<bibRefCitation id="EFD2AD930E2FCB174BD1FE73FBE6FE2B" author="Maury, E. A." box="[914,1092,444,470]" pageId="8" pageNumber="9" pagination="57 - 68" refId="ref5947" refString="Maury, E. A. (1979 a) Escorpiofauna patagonica. II. Urophonius granulatus Pocock 1898 (Bothriuridae). Physis, Secc. C, 38 (94), 57 - 68." type="journal article" year="1979">Maury (1979a)</bibRefCitation>
was collected near Laguna Chara and at the base of Cerro Corona.
</paragraph>
<caption id="DF3C80EA0E2FCB17494BFAECFB5CFA5F" ID-DOI="http://doi.org/10.5281/zenodo.5014526" ID-Zenodo-Dep="5014526" httpUri="https://zenodo.org/record/5014526/files/figure.png" pageId="8" pageNumber="9" startId="8.[264,368,1315,1339]" targetBox="[277,1311,574,1285]" targetPageId="8">
<paragraph id="8BFCD0620E2FCB17494BFAECFB5CFA5F" blockId="8.[264,1325,1315,1442]" pageId="8" pageNumber="9">
<emphasis id="B9370C700E2FCB17494BFAECFE29FAC6" bold="true" box="[264,395,1315,1339]" pageId="8" pageNumber="9">FIGURE 9.</emphasis>
Location of the Meseta de Somuncurá (S) in northern Patagonia, Argentina. The type locality is indicated with a star, the Cerro Corona with a black triangle. Dark grey: areas above 900 m; adjacent solid line: 600 m altitude. Dotted line near Valcheta: the 'Arroyo Valcheta' stream. Dashed line: administrative boundary. Inset: location of the represented area in South America.
</paragraph>
</caption>
<paragraph id="8BFCD0620E2FCB16497BFA19FE34FB0B" blockId="8.[264,1324,1494,1880]" lastBlockId="9.[264,1324,284,1870]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
The east­faced cliffs that surround the Laguna Pelada offer some protection against the WSW winds, and this allows the development of quite dense shrub vegetation (
<figureCitation id="1378CCE70E2FCB174C87FA31FABCF9E5" box="[1220,1310,1534,1560]" captionStart="FIGURE 10" captionStartId="10.[264,367,1690,1714]" captionTargetBox="[377,1205,809,1668]" captionTargetId="figure-262@10.[477,1111,809,1241]" captionTargetPageId="10" captionText="FIGURE 10. Above: west side of Laguna Pelada, type locality of Urophonius somuncura n. sp.; the collecting site is located around the cliffs and rocky outcrops, with scrub vegetation on the right. Below: Schematic representation of a profile of the type locality (not to scale); locations where scorpions were found are indicated by numbers (1-5). In January, U. somuncura n. sp. was collected only at location number 2, and Bothriurus sp. at locations 1 and 3-5. In December, U. somuncura n. sp. was detected in all 1-5 spots." figureDoi="http://doi.org/10.5281/zenodo.5014528" httpUri="https://zenodo.org/record/5014528/files/figure.png" pageId="8" pageNumber="9">Fig. 10</figureCitation>
). Amongst the blocks of rock, tortuous and almost hanging saxicolous shrubs (
<taxonomicName id="4C43ABE10E2FCB174CDDF9E9FA87F9BD" box="[1182,1317,1574,1600]" class="Magnoliopsida" family="Anacardiaceae" genus="Schinus" kingdom="Plantae" order="Sapindales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9370C700E2FCB174CDDF9E9FB5BF9C2" box="[1182,1273,1574,1599]" italics="true" pageId="8" pageNumber="9">Schinus</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="4C43ABE10E2FCB17494BF981FE3BF995" box="[264,409,1614,1640]" class="Magnoliopsida" family="Berberidaceae" genus="Berberis" kingdom="Plantae" order="Ranunculales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9370C700E2FCB17494BF981FECFF99A" box="[264,365,1614,1639]" italics="true" pageId="8" pageNumber="9">Berberis</emphasis>
sp.
</taxonomicName>
) develop, forming together with the numerous and irregular crevices many potential shelters for faunal elements. Other pools were examined (Lagunas Valerio,
<collectingCountry id="F35490F20E2FCB174CAEF9B9FEE3F945" name="Paraguay" pageId="8" pageNumber="9">Paraguay</collectingCountry>
Chico,
<collectingCountry id="F35490F20E2FCB1749DEF951FDA8F945" box="[413,522,1694,1720]" name="Paraguay" pageId="8" pageNumber="9">Paraguay</collectingCountry>
Grande), but only Laguna Pelada had a cliff protected enough, with such dense shrub vegetation. Samples were obtained during separate summers, in
<date id="FFFDF6A20E2FCB174C92F909FE57F8F5" pageId="8" pageNumber="9" value="1985-01" valueMax="1985-12-31" valueMin="1985-01-00">January and December 1985</date>
, in the same site around Laguna Pelada. Diurnal search was very difficult (specimens are suspected to hide in the deep crevices), but with U.V. light, scorpions were easily detected during the night. In December,
<specimenCount id="9D451BEB0E2FCB174B32F8F1FBB3F8A5" box="[881,1041,1854,1880]" pageId="8" pageNumber="9" type="generic">34 specimens</specimenCount>
(females and juveniles) of
<taxonomicName id="4C43ABE10E2ECB164964FED3FE6FFEC8" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" box="[295,461,284,309]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E2ECB164964FED3FE6FFEC8" box="[295,461,284,309]" italics="true" pageId="9" pageNumber="10">U. somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E2ECB164997FED3FDB0FECB" box="[468,530,284,310]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
were caught, together with a single juvenile
<taxonomicName id="4C43ABE10E2ECB164C5DFED3FB67FECB" box="[1054,1221,284,310]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9370C700E2ECB164C5DFED3FB3FFEC8" box="[1054,1181,284,309]" italics="true" pageId="9" pageNumber="10">Bothriurus</emphasis>
sp.
</taxonomicName>
In January, the sampling yielded
<specimenCount id="9D451BEB0E2ECB164A77FE8BFD70FEA3" box="[564,722,324,350]" pageId="9" pageNumber="10" type="generic">14 specimens</specimenCount>
of
<taxonomicName id="4C43ABE10E2ECB164ABBFE8BFC3FFEA0" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" box="[760,925,324,349]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E2ECB164ABBFE8BFC3FFEA0" box="[760,925,324,349]" italics="true" pageId="9" pageNumber="10">U. somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E2ECB164BE7FE8BFC40FEA3" box="[932,994,324,350]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
(females and juveniles) and
<specimenCount id="9D451BEB0E2ECB16494BFEA3FE09FE7B" box="[264,427,364,390]" pageId="9" pageNumber="10" type="generic">16 specimens</specimenCount>
of
<taxonomicName id="4C43ABE10E2ECB16499FFEA3FD2AFE7B" box="[476,648,364,390]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9370C700E2ECB16499FFEA3FDF9FE78" box="[476,603,364,389]" italics="true" pageId="9" pageNumber="10">Bothriurus</emphasis>
sp.
</taxonomicName>
(males, females, couples in courtship). No male of
<taxonomicName id="4C43ABE10E2ECB164D4DFEA3FE28FE50" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E2ECB164D4DFEA3FE28FE50" italics="true" pageId="9" pageNumber="10">U. somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E2ECB1649D4FE5BFE7EFE53" box="[407,476,404,430]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
was found during the mentioned months. To interpret these results, observations made by
<bibRefCitation id="EFD2AD930E2ECB164A4DFE73FD1CFE2B" author="Maury, E. A." box="[526,702,444,470]" pageId="9" pageNumber="10" pagination="57 - 68" refId="ref5947" refString="Maury, E. A. (1979 a) Escorpiofauna patagonica. II. Urophonius granulatus Pocock 1898 (Bothriuridae). Physis, Secc. C, 38 (94), 57 - 68." type="journal article" year="1979">Maury (1979a)</bibRefCitation>
on the close relative
<taxonomicName id="4C43ABE10E2ECB164BF6FE73FBFAFE28" authorityName="Pocock" authorityYear="1898" box="[949,1112,444,469]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E2ECB164BF6FE73FBFAFE28" box="[949,1112,444,469]" italics="true" pageId="9" pageNumber="10">U. granulatus</emphasis>
</taxonomicName>
could be extrapolated here. In southern Patagonia, males of
<taxonomicName id="4C43ABE10E2ECB164B4CFE2BFC11FE00" authorityName="Pocock" authorityYear="1898" box="[783,947,484,509]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E2ECB164B4CFE2BFC11FE00" box="[783,947,484,509]" italics="true" pageId="9" pageNumber="10">U. granulatus</emphasis>
</taxonomicName>
are most frequent at the beginning of the surface activity period (October­November), and become very rare in JanuaryFebruary, when almost only females and juveniles are collected.
<taxonomicName id="4C43ABE10E2ECB164C6CFDFBFB78FDB0" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" box="[1071,1242,564,589]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E2ECB164C6CFDFBFB78FDB0" box="[1071,1242,564,589]" italics="true" pageId="9" pageNumber="10">U. somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E2ECB164CA4FDFBFA89FDB3" box="[1255,1323,564,590]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
might have a similar pattern of surface activity, with females showing a decreasing activity in December­January, as revealed in the samples. For their part, males and females of
<taxonomicName id="4C43ABE10E2ECB16494BFD63FE12FD3B" box="[264,432,684,710]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9370C700E2ECB16494BFD63FE25FD38" box="[264,391,684,709]" italics="true" pageId="9" pageNumber="10">Bothriurus</emphasis>
sp.
</taxonomicName>
seem to appear almost simultaneously by the turn of the year. These results may be consistent with slightly asynchronous surface activities of the two scorpion species that are sympatric in the pond area. In January, when the proportion of these species is approximately similar,
<taxonomicName id="4C43ABE10E2ECB164A59FCEBFD61FCC3" box="[538,707,804,830]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9370C700E2ECB164A59FCEBFD3BFCC0" box="[538,665,804,829]" italics="true" pageId="9" pageNumber="10">Bothriurus</emphasis>
sp.
</taxonomicName>
was found in more diverse locations within the site (
<figureCitation id="1378CCE70E2ECB164952FC83FECAFC9B" box="[273,360,844,870]" captionStart="FIGURE 10" captionStartId="10.[264,367,1690,1714]" captionTargetBox="[377,1205,809,1668]" captionTargetId="figure-262@10.[477,1111,809,1241]" captionTargetPageId="10" captionText="FIGURE 10. Above: west side of Laguna Pelada, type locality of Urophonius somuncura n. sp.; the collecting site is located around the cliffs and rocky outcrops, with scrub vegetation on the right. Below: Schematic representation of a profile of the type locality (not to scale); locations where scorpions were found are indicated by numbers (1-5). In January, U. somuncura n. sp. was collected only at location number 2, and Bothriurus sp. at locations 1 and 3-5. In December, U. somuncura n. sp. was detected in all 1-5 spots." figureDoi="http://doi.org/10.5281/zenodo.5014528" httpUri="https://zenodo.org/record/5014528/files/figure.png" pageId="9" pageNumber="10">Fig. 10</figureCitation>
, below): on the upper side of the cliff, which is exposed to the winds (either near the base of 'coiron' plants or under the tangle of shrub twigs); at the base of the rocky wall, and even more than
<quantity id="4CBB7D870E2ECB164A43FC53FDEFFC4B" box="[512,589,924,950]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="10" unit="m" value="500.0">500 m</quantity>
away from the pond, under a stone by the road; in very few occasions,
<taxonomicName id="4C43ABE10E2ECB1649CCFC0BFDACFC20" authorityName="Peters" authorityYear="1861" box="[399,526,964,989]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B9370C700E2ECB1649CCFC0BFDACFC20" box="[399,526,964,989]" italics="true" pageId="9" pageNumber="10">Bothriurus</emphasis>
</taxonomicName>
specimens were detected climbing on the vertical wall. In that month all
<taxonomicName id="4C43ABE10E2ECB1649C4FC23FD93FBF8" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" box="[391,561,1004,1029]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E2ECB1649C4FC23FD93FBF8" box="[391,561,1004,1029]" italics="true" pageId="9" pageNumber="10">U. somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E2ECB164A7FFC23FDDDFBFB" box="[572,639,1004,1030]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
were on the rocky wall, normally half hidden by small irregularities of the rock, none being found on the exposed areas. By contrast, in December (i.e. when
<taxonomicName id="4C43ABE10E2ECB1649CBFBF3FD90FBAB" box="[392,562,1084,1110]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9370C700E2ECB1649CBFBF3FDA5FBA8" box="[392,519,1084,1109]" italics="true" pageId="9" pageNumber="10">Bothriurus</emphasis>
sp.
</taxonomicName>
was apparently still not active), specimens of
<taxonomicName id="4C43ABE10E2ECB164C20FBF3FAA9FBA8" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" box="[1123,1291,1084,1109]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E2ECB164C20FBF3FAA9FBA8" box="[1123,1291,1084,1109]" italics="true" pageId="9" pageNumber="10">U. somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E2ECB164D56FBF3FE8BFB83" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
were seen in all previously mentioned locations (upper exposed area, base, and vertical wall). Thus, it is also possible that occupying a differential 'microhabitat' might help
<taxonomicName id="4C43ABE10E2ECB164D4DFB43FE28FB30" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E2ECB164D4DFB43FE28FB30" italics="true" pageId="9" pageNumber="10">U. somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E2ECB1649D1FB7BFE70FB33" box="[402,466,1204,1230]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
to avoid predation by
<taxonomicName id="4C43ABE10E2ECB164A9EFB7BFC27FB33" box="[733,901,1204,1230]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9370C700E2ECB164A9EFB7BFCFEFB30" box="[733,860,1204,1229]" italics="true" pageId="9" pageNumber="10">Bothriurus</emphasis>
sp.
</taxonomicName>
(of larger size) during this overlapping period.
</paragraph>
<paragraph id="8BFCD0620E2ECB15497BFACCFCD6FD13" blockId="9.[264,1324,284,1870]" lastBlockId="10.[264,1335,284,750]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="B9370C700E2ECB16497BFACCFD2BFAE0" bold="true" box="[312,649,1283,1309]" pageId="9" pageNumber="10">Zoogeographic comments:</emphasis>
As mentioned above, the Meseta de Somuncurá was deemed to be like a Patagonian 'peninsula' within the Monte Province (
<bibRefCitation id="EFD2AD930E2ECB164C24FAE3FA84FABB" author="Ruiz Leal, A." box="[1127,1318,1324,1350]" pageId="9" pageNumber="10" pagination="89 - 118" refId="ref6054" refString="Ruiz Leal, A. (1972) Los confines boreal y austral de las Provincias Patagonica y Central respectivamente. Boletin de la Sociedad Argentina de Botanica, 13 (supl.), 89 - 118." type="journal article" year="1972">Ruiz Leal 1972</bibRefCitation>
;
<bibRefCitation id="EFD2AD930E2ECB16494BFA9BFD8CFA93" author="Cabrera, A. L. &amp; Willink, A." box="[264,558,1364,1390]" pageId="9" pageNumber="10" pagination="1 - 117" refId="ref5671" refString="Cabrera, A. L. &amp; Willink, A. (1973) Biogeografia de America Latina. Coleccion Monografias Cientificas O. E. A., serie Biologia, nº 13, pp. I-VI, 1 - 117." type="book chapter" year="1973">Cabrera &amp; Willink 1973</bibRefCitation>
). Regarding its scorpiofauna, however, it should be best considered a distributional island (
<bibRefCitation id="EFD2AD930E2ECB164A26FAB3FC20FA6B" author="Acosta, L. &amp; Maury, E. A." box="[613,898,1404,1430]" pageId="9" pageNumber="10" pagination="545 - 559" refId="ref5616" refString="Acosta, L. E &amp; Maury, E. A. (1998) Scorpiones. In: Morrone, J. J. &amp; Coscaron, S. (Eds.) Biodiversidad de Artropodos argentinos. Una perspectiva biotaxonomica. Editorial Sur, La Plata. pp. 545 - 559." type="book chapter" year="1998">Acosta &amp; Maury 1998</bibRefCitation>
). The insular condition is actually restricted to the 'coiron'­fields plateau, since, as has been seen, below
<quantity id="4CBB7D870E2ECB164C02FA6BFB39FA43" box="[1089,1179,1444,1470]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="10" unit="m" value="1200.0">1200 m</quantity>
only Monte scorpions have been found. One of the relatives of
<taxonomicName id="4C43ABE10E2ECB164B34FA03FB82FA18" authority="Acosta, 2003" authorityName="Acosta" authorityYear="2003" box="[887,1056,1484,1509]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="somuncura" status="sp. nov.">
<emphasis id="B9370C700E2ECB164B34FA03FB82FA18" box="[887,1056,1484,1509]" italics="true" pageId="9" pageNumber="10">U. somuncura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A204B10B0E2ECB164C69FA03FBCFFA1B" box="[1066,1133,1484,1510]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
,
<emphasis id="B9370C700E2ECB164C3DFA03FA88FA18" box="[1150,1322,1484,1509]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="4C43ABE10E2ECB164C3DFA03FA84FA18" authorityName="Pocock" authorityYear="1898" box="[1150,1318,1484,1509]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="granulatus">U. granulatus</taxonomicName>
,
</emphasis>
inhabits the southern end of the Patagonian steppe, mainly in the Argentinean province of Santa Cruz and adjacent sites in south
<collectingCountry id="F35490F20E2ECB164A95F9D3FCB4F9CB" box="[726,790,1564,1590]" name="Chile" pageId="9" pageNumber="10">Chile</collectingCountry>
(
<bibRefCitation id="EFD2AD930E2ECB164B65F9D3FC69F9CB" author="Maury, E. A." box="[806,971,1564,1590]" pageId="9" pageNumber="10" pagination="57 - 68" refId="ref5947" refString="Maury, E. A. (1979 a) Escorpiofauna patagonica. II. Urophonius granulatus Pocock 1898 (Bothriuridae). Physis, Secc. C, 38 (94), 57 - 68." type="journal article" year="1979">Maury 1979a</bibRefCitation>
); the northernmost locality of this species is Los Altares, province of Chubut (
<geoCoordinate id="EE77B6A50E2ECB164B0DF98BFC09F9A3" box="[846,939,1604,1630]" degrees="43" direction="south" minutes="53" orientation="latitude" pageId="9" pageNumber="10" precision="925" value="-43.883335">43°53'S</geoCoordinate>
<geoCoordinate id="EE77B6A50E2ECB164BF6F98BFBBCF9A3" box="[949,1054,1604,1630]" degrees="68" direction="west" minutes="24" orientation="longitude" pageId="9" pageNumber="10" precision="925" value="-68.4">68°24'W</geoCoordinate>
; see
<bibRefCitation id="EFD2AD930E2ECB164C1FF98BFB54F9A3" author="Acosta, L. E." box="[1116,1270,1604,1630]" pageId="9" pageNumber="10" pagination="23 - 33" refId="ref5577" refString="Acosta, L. E. (1988) Contribucion al conocimiento taxonomico del genero Urophonius Pocock, 1893 (Scorpiones, Bothriuridae). The Journal of Arachnology, 16 (1), 23 - 33." type="journal article" year="1988">Acosta 1988</bibRefCitation>
: fig. 15), which lies around
<quantity id="4CBB7D870E2ECB164A57F9A3FDC9F97B" box="[532,619,1644,1670]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="km" value="300.0">300 km</quantity>
south of the Meseta de Somuncurá. Except for this northern record, the known range of
<taxonomicName id="4C43ABE10E2ECB164A12F95BFD57F950" authorityName="Pocock" authorityYear="1898" box="[593,757,1684,1709]" class="Arachnida" family="Bothriuridae" genus="Urophonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B9370C700E2ECB164A12F95BFD57F950" box="[593,757,1684,1709]" italics="true" pageId="9" pageNumber="10">U. granulatus</emphasis>
</taxonomicName>
corresponds well with the 'southern Patagonian area' as defined by
<bibRefCitation id="EFD2AD930E2ECB1649B4F973FDDEF92B" author="Cei, J. M." box="[503,636,1724,1750]" pageId="9" pageNumber="10" pagination="309 - 339" refId="ref5751" refString="Cei, J. M. (1979) The Patagonian herpetofauna. In: Duellman, W. E. (Ed.) The South American herpetofauna: its origin, evolution, and dispersal. Monograph of the Museum of Natural History, The University of Kansas, Nr 7, pp. 309 - 339." type="journal article" year="1979">Cei (1979)</bibRefCitation>
; the dominant vegetation of this area consists of 'coirón' grasslands, similar to the upper plains in Somuncurá, so it can be stated that these two closely related species occupy comparable biotopes. The northern Patagonian area resembles more the 'neneo' sub­belt, that is, the lower portion of the Somuncurá plains. Concerning
<taxonomicName id="4C43ABE10E2DCB154976FED3FE7CFECB" box="[309,478,284,310]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9370C700E2DCB154976FED3FE16FEC8" box="[309,436,284,309]" italics="true" pageId="10" pageNumber="11">Bothriurus</emphasis>
sp.
</taxonomicName>
, its nearest relatives are the members of the '
<taxonomicName id="4C43ABE10E2DCB154BABFED3FBD5FEC8" authorityName="Maury" authorityYear="1968" box="[1000,1143,284,309]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="patagonicus">
<emphasis id="B9370C700E2DCB154BABFED3FBD5FEC8" box="[1000,1143,284,309]" italics="true" pageId="10" pageNumber="11">patagonicus</emphasis>
</taxonomicName>
species group', which include the nominal species
<taxonomicName id="4C43ABE10E2DCB154AFBFE8BFBACFEA3" authority="Maury 1968" authorityName="Maury" authorityYear="1968" box="[696,1038,324,350]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="patagonicus">
<emphasis id="B9370C700E2DCB154AFBFE8BFCCEFEA0" box="[696,876,324,349]" italics="true" pageId="10" pageNumber="11">B. patagonicus</emphasis>
<bibRefCitation id="EFD2AD930E2DCB154B34FE8BFBACFEA3" author="Maury, E. A." box="[887,1038,324,350]" pageId="10" pageNumber="11" pagination="149 - 164" refId="ref5846" refString="Maury, E. A. (1968) Aportes al conocimiento de los escorpiones de la Republica Argentina. II. Algunas consideraciones sobre el genero Bothriurus en la Patagonia y Tierra del Fuego con la descripcion de una nueva especie (Bothriuridae). Physis, 28 (76), 149 - 164." type="journal article" year="1968">Maury 1968</bibRefCitation>
</taxonomicName>
, and some undescribed entities as well (
<bibRefCitation id="EFD2AD930E2DCB154989FEA3FDC2FE7B" author="Maury, E. A." box="[458,608,364,390]" pageId="10" pageNumber="11" pagination="149 - 164" refId="ref5846" refString="Maury, E. A. (1968) Aportes al conocimiento de los escorpiones de la Republica Argentina. II. Algunas consideraciones sobre el genero Bothriurus en la Patagonia y Tierra del Fuego con la descripcion de una nueva especie (Bothriuridae). Physis, 28 (76), 149 - 164." type="journal article" year="1968">Maury 1968</bibRefCitation>
, Mattoni &amp; Acosta, in prep.). The distribution of the
<taxonomicName id="4C43ABE10E2DCB154C9DFEA3FEF6FE50" authorityName="Maury" authorityYear="1968" class="Arachnida" family="Bothriuridae" genus="Bothriurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="patagonicus">
<emphasis id="B9370C700E2DCB154C9DFEA3FEF6FE50" italics="true" pageId="10" pageNumber="11">patagonicus</emphasis>
</taxonomicName>
group ranging from a narrow strip near the Andes in the province of
<collectingRegion id="49871E800E2DCB154CDFFE5BFAB2FE53" box="[1180,1296,404,430]" country="Argentina" name="Neuquen" pageId="10" pageNumber="11">Neuquén</collectingRegion>
, and widening into a broader area southwards, in the province of
<collectingRegion id="49871E800E2DCB154C7DFE73FB67FE2B" box="[1086,1221,444,470]" country="Argentina" name="Santa Cruz" pageId="10" pageNumber="11">Santa Cruz</collectingRegion>
(
<bibRefCitation id="EFD2AD930E2DCB154C98FE73FEF1FE03" author="Maury, E. A." pageId="10" pageNumber="11" pagination="703 - 719" refId="ref5985" refString="Maury, E. A. (1979 b) Apuntes para una zoogeografia de la escorpiofauna argentina. Acta zoologica Lilloana, 35 (2): 703 - 719." type="journal article" year="1979">Maury 1979b</bibRefCitation>
) fits well into the 'Patagonian scorpiogeographical area' as defined by
<bibRefCitation id="EFD2AD930E2DCB154CE5FE2BFE13FDDB" author="Acosta, L. &amp; Maury, E. A." pageId="10" pageNumber="11" pagination="545 - 559" refId="ref5616" refString="Acosta, L. E &amp; Maury, E. A. (1998) Scorpiones. In: Morrone, J. J. &amp; Coscaron, S. (Eds.) Biodiversidad de Artropodos argentinos. Una perspectiva biotaxonomica. Editorial Sur, La Plata. pp. 545 - 559." type="book chapter" year="1998">Acosta &amp; Maury (1998)</bibRefCitation>
; only
<taxonomicName id="4C43ABE10E2DCB1549BFFDC3FD07FDDB" box="[508,677,524,550]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9370C700E2DCB1549BFFDC3FDD9FDD8" box="[508,635,524,549]" italics="true" pageId="10" pageNumber="11">Bothriurus</emphasis>
sp.
</taxonomicName>
from Somuncurá shows a disjunct pattern. It is worth noting that the Somuncuran herpetofauna (lizards and amphibians) contains six species that are widespread in Patagonia, as well as three species and two subspecies that are endemic to the Meseta (
<bibRefCitation id="EFD2AD930E2DCB154A77FD4BFD08FD63" author="Cei, J. M." box="[564,682,644,670]" pageId="10" pageNumber="11" pagination="309 - 339" refId="ref5751" refString="Cei, J. M. (1979) The Patagonian herpetofauna. In: Duellman, W. E. (Ed.) The South American herpetofauna: its origin, evolution, and dispersal. Monograph of the Museum of Natural History, The University of Kansas, Nr 7, pp. 309 - 339." type="journal article" year="1979">Cei 1979</bibRefCitation>
). The endemics
<taxonomicName id="4C43ABE10E2DCB154B3BFD4BFA88FD63" authority="(Cei, 1969)" baseAuthorityName="Cei" baseAuthorityYear="1969" box="[888,1322,644,670]" class="Amphibia" family="Ceratophryidae" genus="Atelognathus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="reverberii">
<emphasis id="B9370C700E2DCB154B3BFD4BFB36FD60" box="[888,1172,644,669]" italics="true" pageId="10" pageNumber="11">Atelognathus reverberii</emphasis>
(
<bibRefCitation id="EFD2AD930E2DCB154CEAFD4BFA80FD63" author="Cei, J. M." box="[1193,1314,644,670]" pageId="10" pageNumber="11" pagination="257 - 271" refId="ref5714" refString="Cei, J. M. (1969) La meseta basaltica de Somuncura, Rio Negro. Herpetofauna endemica y sus peculiares equilibrios biocenoticos. Physis, 28 (77), 257 - 271." type="journal article" year="1969">Cei, 1969</bibRefCitation>
)
</taxonomicName>
(
<taxonomicName id="4C43ABE10E2DCB154953FD63FE6DFD3B" box="[272,463,684,710]" class="Amphibia" family="Leptodactylidae" kingdom="Animalia" order="Anura" pageId="10" pageNumber="11" phylum="Chordata" rank="family">Leptodactylidae</taxonomicName>
Telmatobiinae) and
<taxonomicName id="4C43ABE10E2DCB154A97FD63FE0CFD13" authority="Donoso Barros &amp; Cei, 1971" authorityName="Donoso Barros &amp; Cei" authorityYear="1971" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="ruizleali">
<emphasis id="B9370C700E2DCB154A97FD63FC1FFD38" box="[724,957,684,709]" italics="true" pageId="10" pageNumber="11">Liolaemus ruizleali</emphasis>
Donoso Barros &amp; Cei, 1971 (Liolaemidae)
</taxonomicName>
were found above
<quantity id="4CBB7D870E2DCB154ACCFD1BFD4AFD13" box="[655,744,724,750]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="m" value="1000.0">1000 m</quantity>
(
<bibRefCitation id="EFD2AD930E2DCB154AB4FD1BFCCAFD13" author="Cei, J. M." box="[759,872,724,750]" pageId="10" pageNumber="11" pagination="309 - 339" refId="ref5751" refString="Cei, J. M. (1979) The Patagonian herpetofauna. In: Duellman, W. E. (Ed.) The South American herpetofauna: its origin, evolution, and dispersal. Monograph of the Museum of Natural History, The University of Kansas, Nr 7, pp. 309 - 339." type="journal article" year="1979">Cei 1979</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink