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<document id="3240B9038B2E4884D96BBA269FC64B97" ID-CLB-Dataset="26049" ID-DOI="10.11646/zootaxa.4654.1.1" ID-GBIF-Dataset="9ce1d103-9234-4fbe-9544-f1c294bb8783" ID-ISSN="1175-5326" ID-Zenodo-Dep="3365803" ID-ZooBank="CFD82CC0-3110-472E-972B-7ADC0C523A04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565592061838" checkinUser="plazi" docAuthor="Hansen, Thomas" docDate="2019" docId="BF18F633A9A4FF412B9CC22EFBA3F976" docLanguage="en" docName="zootaxa.4654.1.1.pdf" docOrigin="Zootaxa 4654 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Ringicula Deshayes in Deshayes &amp; Milne-Edwards 1838" docType="treatment" docVersion="20" lastPageNumber="165" masterDocId="43218E4BA900FFE52B0BC14DFFA9FFE3" masterDocTitle="Gastropods from the Cretaceous-Palaeogene boundary in Denmark" masterLastPageNumber="196" masterPageNumber="1" pageNumber="165" updateTime="1732572783792" updateUser="ExternalLinkService">
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<mods:title id="EAEA32593B40E1E20E34F94BDCBF5A3A">Gastropods from the Cretaceous-Palaeogene boundary in Denmark</mods:title>
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<mods:namePart id="2AC06C20A4AFC08772568CC65DC09A9E">Hansen, Thomas</mods:namePart>
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<mods:title id="C278F3E89FCDB57185B4D35146CD5E75">Zootaxa</mods:title>
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<mods:date id="427EA6B2D1C50ED5397F1878DC22A506">2019</mods:date>
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<mods:number id="41EBB577DAC953332691B4E1FD92E782">4654</mods:number>
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<treatment id="BF18F633A9A4FF412B9CC22EFBA3F976" ID-GBIF-Taxon="159074635" LSID="urn:lsid:plazi:treatment:BF18F633A9A4FF412B9CC22EFBA3F976" httpUri="http://treatment.plazi.org/id/BF18F633A9A4FF412B9CC22EFBA3F976" lastPageNumber="165" pageId="164" pageNumber="165" scope_family="Ringiculidae">
<subSubSection id="7FAB14AEA9A4FF412B9CC22EFE90FC9D" box="[151,313,867,894]" pageId="164" pageNumber="165" type="nomenclature">
<paragraph id="370E4725A9A4FF412B9CC22EFE90FC9D" blockId="164.[151,313,867,930]" box="[151,313,867,894]" pageId="164" pageNumber="165">
<heading id="6C46F049A9A4FF412B9CC22EFE90FC9D" bold="true" box="[151,313,867,894]" fontSize="11" level="1" pageId="164" pageNumber="165" reason="1">
<emphasis id="05C59B37A9A4FF412B9CC22EFE90FC9D" bold="true" box="[151,313,867,894]" pageId="164" pageNumber="165">
<taxonomicName id="F0B13CA6A9A4FF412B9CC22EFEA7FC9D" ID-CoL="7PFTM" authorityName="Deshayes in Deshayes &amp; Milne-Edwards" authorityYear="1838" box="[151,270,867,894]" class="Gastropoda" family="Ringiculidae" genus="Ringicula" kingdom="Animalia" pageId="164" pageNumber="165" phylum="Mollusca" rank="genus">
<emphasis id="05C59B37A9A4FF412B9CC22EFEA7FC9D" bold="true" box="[151,270,867,894]" italics="true" pageId="164" pageNumber="165">Ringicula</emphasis>
</taxonomicName>
sp.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA9A4FF412B9CC2CAFEB0FC41" box="[151,281,903,930]" pageId="164" pageNumber="165" type="description">
<paragraph id="370E4725A9A4FF412B9CC2CAFEB0FC41" blockId="164.[151,313,867,930]" box="[151,281,903,930]" pageId="164" pageNumber="165">
<figureCitation id="AF8A5BA0A9A4FF412B9CC2CAFF41FC41" box="[151,232,903,930]" captionStart="FIGURE 35" captionStartId="167.[151,250,188,213]" captionTargetId="figure@166.[151,1435,179,1918]" captionTargetPageId="166" captionText="FIGURE 35. AB. Pseudomalaxis sp., Silicone cast of MGUH 33306. Width 4.3 mm. CD. Climacopoma? sp., Silicone cast of MGUH 33307. Width 13.1 and 9.9 mm respectively. EI. Mathilda dania n. sp. E. Silicone cast of holotype MGUH 33308. Width 1.9 mm. F. Silicone cast of paratype MGUH 33309. Width 2.0 mm. G. Silicone cast of paratype MGUH 33310. Width 0.8 mm. H. Silicone cast of paratype MGUH 33311. Width 0.7 mm. I. Silicone cast of paratype MGUH 33312. Width 1.7 mm. JK. Clathrobaculus sp., Silicone cast of MGUH 33313. Width 0.5 mm. LP. Ringicula sp. L. Silicone cast of MGUH 33314. Width 7.0 mm. MN. Silicone cast of MGUH 33315 and a partly hidden second specimen. Width 7.3 and 6.6 mm respectively. OP. Silicone cast of MGUH 33316. Width 5.6, 5.3 mm respectively. QT. Cylichna stevniensis n. sp. Q. Silicone cast of holotype MGUH 33317. Width 1.4 mm. RS. Internal mould of paratype MGUH 33318. Width 1.5 mm. T. Internal mould of paratype MGUH 33319. Width 2.8 mm. UV. Cylichna? sp. A. Cast of MGUH 33320. Width 2.3, 2.4 mm respectively. WY. Cylichna? sp. B. W. Wax cast of MGUH 33321. Width 2.4 mm. XY. Inner mould of MGUH 33322. Width 1.3 mm. ZAE. Roxania sp. Cast of MGUH 33323. Width 3.2, 3.3 mm respectively." figureDoi="http://doi.org/10.5281/zenodo.3442126" httpUri="https://zenodo.org/record/3442126/files/figure.png" pageId="164" pageNumber="165">Figs 35</figureCitation>
LP
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA9A4FF412B9CC282FBA3F976" pageId="164" pageNumber="165" type="materials_examined">
<paragraph id="370E4725A9A4FF412B9CC282FF4CFBED" blockId="164.[151,1437,975,1686]" pageId="164" pageNumber="165">
<emphasis id="05C59B37A9A4FF412B9CC282FEAFFC09" bold="true" box="[151,262,975,1002]" pageId="164" pageNumber="165">Material.</emphasis>
<collectionCode id="51A0DFE0A9A4FF412A05C29DFECDFC09" box="[270,356,976,1002]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="164" pageNumber="165">MGUH</collectionCode>
33314,
<collectionCode id="51A0DFE0A9A4FF412AB4C29DFDBCFC09" box="[447,533,976,1002]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="164" pageNumber="165">MGUH</collectionCode>
33315,
<collectionCode id="51A0DFE0A9A4FF41297AC29DFD6EFC09" box="[625,711,976,1002]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="164" pageNumber="165">MGUH</collectionCode>
33316, and an additional specimen sitting together with
<collectionCode id="51A0DFE0A9A4FF412E4CC29DFA34FC09" box="[1351,1437,976,1002]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="164" pageNumber="165">MGUH</collectionCode>
33315.
</paragraph>
<paragraph id="370E4725A9A4FF412BCCC55AFB79FBD2" blockId="164.[151,1437,975,1686]" box="[199,1232,1047,1074]" pageId="164" pageNumber="165">
<emphasis id="05C59B37A9A4FF412BCCC55AFEF3FBD1" bold="true" box="[199,346,1047,1074]" pageId="164" pageNumber="165">Occurrence.</emphasis>
The lower Danian dead layer exposed in the Dania Quarry, Northern Jutland.
</paragraph>
<paragraph id="370E4725A9A4FF412BCCC576FEB6FB99" blockId="164.[151,1437,975,1686]" pageId="164" pageNumber="165">
<emphasis id="05C59B37A9A4FF412BCCC576FEF0FBB5" bold="true" box="[199,345,1083,1110]" pageId="164" pageNumber="165">Description.</emphasis>
Protoconch apparently conical in outline,
<quantity id="F049EAC0A9A4FF412838C571FC21FBB5" box="[819,904,1084,1110]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="164" pageNumber="165" unit="mm" value="0.8">0.8 mm</quantity>
wide with smooth, convex whorls separated by deep suture.
</paragraph>
<paragraph id="370E4725A9A4FF412BCCC5CEFDF6FA96" blockId="164.[151,1437,975,1686]" pageId="164" pageNumber="165">Teleoconch low spired, rotund with flattened, adpressed spire whorls; whorls nearly four times as wide as high. Last whorl evenly convex all the way to anterior canal, taking up more than 80 % of shell height. Aperture narrow, outer lip strengthened by strong varix reaching from anterior canal to adapical suture of transition between last whorl and penultimate whorl. Outer lip carrying five stronger, elongated teeth on abapical part, succeeded adapically by several more, but very weak teeth fading out completely adapically. Inner lip with moderately wide callus and two sinuses separated by a strong and furrowed ridge. Adapical parietal region carrying strong but low and ridged transverse thickening of the shell.</paragraph>
<paragraph id="370E4725A9A4FF412BCCC432FA3DFA7A" blockId="164.[151,1437,975,1686]" box="[199,1428,1407,1434]" pageId="164" pageNumber="165">Teleoconch sculpture consisting of around 20 moderately weak spiral grooves, which continue onto the varix.</paragraph>
<paragraph id="370E4725A9A4FF412BCCC4E9FAFCFA02" blockId="164.[151,1437,975,1686]" pageId="164" pageNumber="165">
<emphasis id="05C59B37A9A4FF412BCCC4E9FED7FA5D" bold="true" box="[199,382,1444,1470]" pageId="164" pageNumber="165">Measurements.</emphasis>
Most complete specimen
<collectionCode id="51A0DFE0A9A4FF4129AAC4E9FD5EFA5D" box="[673,759,1444,1470]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="164" pageNumber="165">MGUH</collectionCode>
33314 is 6.0 mm wide and more than
<quantity id="F049EAC0A9A4FF412FABC4EEFB5CFA5D" box="[1184,1269,1443,1470]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="164" pageNumber="165" unit="mm" value="6.9">6.9 mm</quantity>
high, but lacking anterior canal and protoconch apex. It consists of the final protoconch whorl and 3 ½ teleoconch whorls.
</paragraph>
<paragraph id="370E4725A9A4FF412BCCC4A1FBA3F976" blockId="164.[151,1437,975,1686]" pageId="164" pageNumber="165">
<emphasis id="05C59B37A9A4FF412BCCC4A1FE92F9E5" bold="true" box="[199,315,1516,1542]" pageId="164" pageNumber="165">Remarks.</emphasis>
This species differs from the Danian
<taxonomicName id="F0B13CA6A9A4FF4129FAC4A1FBF3F9E5" authority="Glibert, 1973" authorityName="Glibert" authorityYear="1973" box="[753,1114,1515,1542]" class="Gastropoda" family="Ringiculidae" genus="Ringicula" kingdom="Animalia" pageId="164" pageNumber="165" phylum="Mollusca" rank="species" species="pinguis">
<emphasis id="05C59B37A9A4FF4129FAC4A1FC10F9E6" box="[753,953,1516,1542]" italics="true" pageId="164" pageNumber="165">Ringicula pinguis</emphasis>
<bibRefCitation id="53203AD4A9A4FF4128C8C4A6FBF3F9E5" author="Glibert, M." box="[963,1114,1515,1542]" pageId="164" pageNumber="165" pagination="1 - 116" refId="ref130648" refString="Glibert, M. (1973) Revision des Gastropoda du Danien et du Montien de la Belgique. 1. Les Gastropoda du Calcaire de Mons. Institut Royal des Sciences Naturelles de Belgique Memoire, 173, 1 - 116, pls. 1 - 11." type="journal article" year="1973">Glibert, 1973</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="4FA607B5A9A4FF412FA8C4A6FAAAF9E6" box="[1187,1283,1515,1541]" name="Belgium" pageId="164" pageNumber="165">Belgium</collectingCountry>
by the much more rotund outline, adpressed whorls and furrowed lower ridge on the inner lip. It differs from the Campanian
<taxonomicName id="F0B13CA6A9A4FF412E8FC75DFDB8F9AD" authority="Kiel &amp; Bandel, 2001 a" authorityName="Kiel &amp; Bandel" authorityYear="2001" class="Gastropoda" family="Ringiculidae" genus="Ringicula" kingdom="Animalia" pageId="164" pageNumber="165" phylum="Mollusca" rank="species" species="abundanta">
<emphasis id="05C59B37A9A4FF412E8FC75DFEA6F9AD" italics="true" pageId="164" pageNumber="165">R. abundanta</emphasis>
<bibRefCitation id="53203AD4A9A4FF412A1CC77EFDB8F9AD" author="Kiel, S. &amp; Bandel, K." box="[279,529,1587,1614]" pageId="164" pageNumber="165" pagination="319 - 334" refId="ref133596" refString="Kiel, S. &amp; Bandel, K. (2001 a) About Heterostropha (Gastropoda) of the Campanian of Torallola, Spain. Journal of the Czech Geological Society, 46, 319 - 334." type="journal article" year="2001">Kiel &amp; Bandel, 2001a</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="F0B13CA6A9A4FF412940C779FC52F9AD" authority="Kiel &amp; Bandel, 2001 a" authorityName="Kiel &amp; Bandel" authorityYear="2001" box="[587,1019,1587,1614]" class="Gastropoda" family="Ringiculidae" genus="Ringicula" kingdom="Animalia" pageId="164" pageNumber="165" phylum="Mollusca" rank="species" species="multidentata">
<emphasis id="05C59B37A9A4FF412940C779FD50F9AD" box="[587,761,1588,1614]" italics="true" pageId="164" pageNumber="165">R. multidentata</emphasis>
<bibRefCitation id="53203AD4A9A4FF41280AC77EFC52F9AD" author="Kiel, S. &amp; Bandel, K." box="[769,1019,1587,1614]" pageId="164" pageNumber="165" pagination="319 - 334" refId="ref133596" refString="Kiel, S. &amp; Bandel, K. (2001 a) About Heterostropha (Gastropoda) of the Campanian of Torallola, Spain. Journal of the Czech Geological Society, 46, 319 - 334." type="journal article" year="2001">Kiel &amp; Bandel, 2001a</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="4FA607B5A9A4FF412F48C77EFB2AF9AE" box="[1091,1155,1587,1613]" name="Spain" pageId="164" pageNumber="165">Spain</collectingCountry>
by the rotund shape and from the latter also by a different tooth pattern on the outer lip. It differs from the Palaeocene
<taxonomicName id="F0B13CA6A9A4FF412FC8C715FF66F975" authority="Roedel, 1937" authorityName="Roedel" authorityYear="1937" class="Gastropoda" family="Ringiculidae" genus="Ringicula" kingdom="Animalia" pageId="164" pageNumber="165" phylum="Mollusca" rank="species" species="erratica">
<emphasis id="05C59B37A9A4FF412FC8C715FA97F992" box="[1219,1342,1624,1649]" italics="true" pageId="164" pageNumber="165">R. erratica</emphasis>
<bibRefCitation id="53203AD4A9A4FF412E4DC71AFF66F975" author="Roedel, H." pageId="164" pageNumber="165" pagination="184 - 222" refId="ref138710" refString="Roedel, H. (1937) Die Fauna der norddeutschen Palaozangeschiebe: Schnecken, Scaphopoden, Brachiopoden und Korallen. Zeitschrift fur Geschiebeforschung, 13, 184 - 222." type="journal article" year="1937">Roedel, 1937</bibRefCitation>
</taxonomicName>
from Northern Europe by its lower spire and more flattened spire whorls.
</paragraph>
</subSubSection>
</treatment>
</document>