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<document id="FF4A20402CFD52FF581E626053E7FF82" ID-CLB-Dataset="8118" ID-DOI="10.5252/z2010n1a2" ID-GBIF-Dataset="4cecd899-5900-4411-8446-0d9552b1b098" ID-ISSN="1638-9387" ID-Zenodo-Dep="4520753" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612808309234" checkinUser="felipe" docAuthor="Randriamanantsoa, Richard, Aberlenc, Henri-Pierre, Ralisoa, Olga Bakoly, Ratnadass, Alain &amp; Vercambre, Bernard" docDate="2010" docId="901F8794FF9A746EFD0DFC5DFCC1FD26" docLanguage="fr" docName="Zoosystema.32.1.19-72.pdf" docOrigin="Zoosystema 32 (1)" docSource="http://www.bioone.org/doi/abs/10.5252/z2010n1a2" docStyle="DocumentStyle:E638A3C99FA728B529662B46F84D75FD.3:Zoosystema.2009-2010.journal_article" docStyleId="E638A3C99FA728B529662B46F84D75FD" docStyleName="Zoosystema.2009-2010.journal_article" docStyleVersion="3" docTitle="Paramorphochelus cornutus" docType="treatment" docVersion="12" lastPageNumber="34" masterDocId="6C26FFECFF947461FFB5FFEDFF94FFE8" masterDocTitle="Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar" masterLastPageNumber="72" masterPageNumber="19" pageNumber="33" updateTime="1698928337295" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EFE46DD831F4E8ACE7BDD08F21A0BA31">Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar</mods:title>
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<mods:namePart id="A0FF91FD3725B141D85C8E3A5F2DFB8F">Randriamanantsoa, Richard</mods:namePart>
<mods:affiliation id="E1BC32C58DF6ACF4A7AC3B7D1E0501DC">Fofifa - URP Scrid, 110 Antsirabe (Madagascar) r _ randriamanantsoa @ yahoo. fr</mods:affiliation>
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<mods:namePart id="F02AC53B19CEF9AB0CD9C1569B873204">Aberlenc, Henri-Pierre</mods:namePart>
<mods:affiliation id="BB8D61981670EC12B799C0DA08A8F87E">Cirad Bios, UMR CBGP (INRA / IRD / Cirad / Montpellier SupAgro), Campus international de Baillarguet, Ta A- 55 / L, F- 34398 Montpellier cedex 5 (France) henri-pierre. aberlenc @ cirad. fr</mods:affiliation>
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<mods:namePart id="2C8BE0A70A90E8B44A8C4F3C0F94737E">Ralisoa, Olga Bakoly</mods:namePart>
<mods:affiliation id="D0AE43838A9D0F49F931DCE08FAE2344">Département dEntomologie, Faculté des Sciences, Université dAntananarivo, Antananarivo 101 (Madagascar)</mods:affiliation>
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<mods:namePart id="56B1A352C3647E4E1732AA48C38B991E">Ratnadass, Alain</mods:namePart>
<mods:affiliation id="BBBF1402337532A186B29164A4A30813">Cirad, UPR HortSys - ICRISAT BP 12404, Niamey (Niger) a. ratnadass @ cgiar. org</mods:affiliation>
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<mods:namePart id="BE61D0A82721FF2BCBF4A879DC18E399">Vercambre, Bernard</mods:namePart>
<mods:affiliation id="2F9DC489C63DB09A3CC8B21D6F3DA081">Cirad, UPR Systèmes de Cultures annuels, avenue Agropolis, F- 34398 Montpellier cedex 5 (France) bernard. vercambre @ cirad. fr</mods:affiliation>
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<mods:title id="0E3681947FFAEFDB375557CABD7FA0B9">Zoosystema</mods:title>
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<mods:date id="8E0AB78C007FF28E7C73002AD4A70D4C">2010</mods:date>
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<treatment id="901F8794FF9A746EFD0DFC5DFCC1FD26" ID-DOI="http://doi.org/10.5281/zenodo.4549538" ID-GBIF-Taxon="177425648" ID-Zenodo-Dep="4549538" LSID="urn:lsid:plazi:treatment:901F8794FF9A746EFD0DFC5DFCC1FD26" httpUri="http://treatment.plazi.org/id/901F8794FF9A746EFD0DFC5DFCC1FD26" lastPageId="15" lastPageNumber="34" pageId="14" pageNumber="33">
<subSubSection id="50AC6509FF9A746FFD0DFC5DFCE1FBA0" pageId="14" pageNumber="33" type="nomenclature">
<paragraph id="18093682FF9A746FFD0DFC5DFB32FC23" blockId="14.[696,1190,944,1035]" box="[696,1190,944,971]" pageId="14" pageNumber="33">
<taxonomicName id="DFB64D01FF9A746FFD0DFC5DFB32FC23" authority="(Nonfried, 1892)" baseAuthorityName="Nonfried" baseAuthorityYear="1892" box="[696,1190,944,971]" class="Insecta" family="Melolonthidae" genus="Paramorphochelus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="33" phylum="Arthropoda" rank="species" species="cornutus">
<emphasis id="2AC2EA90FF9A746FFD0DFC5DFC77FC23" bold="true" box="[696,995,944,971]" italics="true" pageId="14" pageNumber="33">Paramorphochelus cornutus</emphasis>
(Nonfried, 1892)
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</paragraph>
<paragraph id="18093682FF9A746FFD41FC3DFBEDFBE3" blockId="14.[696,1190,944,1035]" pageId="14" pageNumber="33">
(
<figureCitation id="808D2A07FF9A746FFD4EFC3DFCD3FC03" box="[763,839,976,1003]" captionStart="FIG" captionStartId="29.[113,124,1567,1584]" captionTargetBox="[152,1172,222,1545]" captionTargetId="figure-18@29.[148,709,692,1169]" captionTargetPageId="29" captionText="FIG. 2. — Larves de Scarabaeoidea de Madagascar, habitus: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paramorphochelus cornutus (Nonfried,1892); K, Encya sikorai (Brenske,1891); L, Apicencya waterloti (Dewailly, 1950); M, Empecta scutata (Fairmaire,1901); N, Hoplochelus betanimena (Künckel,1887); O, Triodontus nitidulus (Guérin, 1844). Échelles: A-I, K, L, N, 5 mm; J, O, 2 mm; M, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4520757" httpUri="https://zenodo.org/record/4520757/files/figure.png" pageId="14" pageNumber="33">Figs 2J</figureCitation>
;
<figureCitation id="808D2A07FF9A746FFCEDFC3DFCE2FC03" box="[856,886,976,1003]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="14" pageNumber="33">4F</figureCitation>
;
<figureCitation id="808D2A07FF9A746FFC32FC3DFC3FFC03" box="[903,939,976,1003]" captionStart="FIG" captionStartId="33.[113,124,1635,1652]" captionTargetBox="[117,1206,214,1443]" captionTargetId="figure-14@33.[117,1206,214,1443]" captionTargetPageId="33" captionText="FIG. 6. — Larves de Scarabaeoidea de Madagascar: A-F, tête; A, Anochilia bifida (Olivier, 1789); B, Hexodon unicolor (Olivier, 1789); C, sp.D2; D, Paramorphochelus cornutus (Nonfried,1892); E, Empecta scutata (Fairmaire,1901); F, sp.S2; G, H, épipharynx; G, Bricoptis variolosa (Gory &amp; Percheron, 1833); H, Celidota parvula (Janson, 1881). Échelles: A-C, 2 mm; D-F, 1 mm; G, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520767" httpUri="https://zenodo.org/record/4520767/files/figure.png" pageId="14" pageNumber="33">6D</figureCitation>
;
<figureCitation id="808D2A07FF9A746FFC09FC3DFC49FC03" box="[956,989,976,1003]" captionStart="FIG" captionStartId="35.[113,124,1635,1652]" captionTargetBox="[141,1181,217,1589]" captionTargetId="figure-14@35.[141,1181,217,1589]" captionTargetPageId="35" captionText="FIG. 8. — Larves de Scarabaeoidea de Madagascar, épipharynx: A, Heteroconus paradoxus (Endrödi,1968); B, Paranodon coquerelii (Fairmaire, 1871); C, Paramorphochelus cornutus (Nonfried,1892); D, sp. M1; E, Encya sikorai (Brenske,1891); F, Enaria melanictera (Klug, 1833); G, sp. M2. Échelles: A, F, 0,5 mm; B, C, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520775" httpUri="https://zenodo.org/record/4520775/files/figure.png" pageId="14" pageNumber="33">8C</figureCitation>
;
<figureCitation id="808D2A07FF9A746FFC5AFC3DFB8AFC03" box="[1007,1054,976,1003]" captionStart="FIG" captionStartId="37.[113,124,1589,1606]" captionTargetBox="[146,1173,219,1547]" captionTargetId="figure-14@37.[146,1173,218,1547]" captionTargetPageId="37" captionText="FIG. 10. — Larves de Scarabaeoidea de Madagascar,hypopharynx: A, Bricoptis variolosa (Gory &amp; Percheron,1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K, Paramorphochelus cornutus (Nonfried,1892); L, Encya sikorai (Brenske, 1891). Échelles: A, J, 0,2 mm; C, D, F, G, I, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520783" httpUri="https://zenodo.org/record/4520783/files/figure.png" pageId="14" pageNumber="33">10K</figureCitation>
;
<figureCitation id="808D2A07FF9A746FFB85FC3DFBCBFC03" box="[1072,1119,976,1003]" captionStart="FIG" captionStartId="40.[113,124,1612,1629]" captionTargetBox="[116,1208,217,1563]" captionTargetId="figure-21@40.[113,1202,214,1628]" captionTargetPageId="40" captionText="FIG. 13. — Larves de Scarabaeoidea de Madagascar: A-E, labre; A, Anochilia bifida (Olivier, 1789); B, Heteronychus plebeius (Klug, 1833); C, Paramorphochelus cornutus (Nonfried, 1892); D, Enaria melanictera (Klug, 1833); E, Triodontus nitidulus (Guérin, 1844); F-J, Celidota parvula (Janson, 1881), mandibule; F, gauche, face dorsale; G, gauche, face ventrale; H, droite, face dorsale; I, droite, face ventrale; J, aire stridulatoire mandibulaire. Échelles: A, B, D, F-I, 1 mm; C, E, J, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520793" httpUri="https://zenodo.org/record/4520793/files/figure.png" pageId="14" pageNumber="33">13C</figureCitation>
;
<figureCitation id="808D2A07FF9A746FFD51FC1DFD6AFBE3" box="[740,766,1008,1035]" captionStart="FIG" captionStartId="42.[113,124,1589,1606]" captionTargetBox="[135,1177,220,1549]" captionTargetId="figure-21@42.[136,1178,214,1549]" captionTargetPageId="42" captionText="FIG. 15. — Larves de Scarabaeoidea de Madagascar, mandibule: A, E, I, M, Q, U, Y, gauche, face dorsale; B, F, J, N, R, V, Z1, gauche, face ventrale; C, G, K, O, S, W, Z2, droite,face dorsale; D, H, L, P, T, X, Z3, droite,face ventrale; A-D, Paranodon coquerelii (Fairmaire, 1871); E-H, Paramorphochelus cornutus (Nonfried, 1892); I-L, Encya sikorai (Brenske, 1891); M-P, Enaria melanictera (Klug, 1833); Q-T, Apicencya waterloti (Dewailly, 1950); U-X, Empecta scutata (Fairmaire, 1901); Y-Z3, Hoplochelus betanimena (Künckel, 1887). Échelles: A-D, M-P, 1 mm; E-H, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520799" httpUri="https://zenodo.org/record/4520799/files/figure.png" pageId="14" pageNumber="33">15</figureCitation>
E-H; 17B; 18E; 22D, E; 24K-M)
</paragraph>
<paragraph id="18093682FF9A746FFD10FBC2FCE1FBA0" blockId="14.[675,1211,1071,1257]" box="[677,885,1071,1098]" pageId="14" pageNumber="33">MATÉRIEL EXAMINÉ</paragraph>
</subSubSection>
<subSubSection id="50AC6509FF9A746EFD10FBA2FCC1FD26" lastPageId="15" lastPageNumber="34" pageId="14" pageNumber="33" type="description">
<paragraph id="18093682FF9A746FFD10FBA2FC02FB01" blockId="14.[675,1211,1071,1257]" pageId="14" pageNumber="33">
Larves provenant dAndranomanelatra et dIvory dans une parcelle de Riz pluvial.
<typeStatus id="C70D8820FF9A746FFBBDFB82FBA9FB62" box="[1032,1085,1135,1162]" pageId="14" pageNumber="33">Type</typeStatus>
de sol: ferrallitique. Spécimens conservés dans les collections du Fofifa (7 larves), du Cirad-CBGP (8 larves) et du MNHN (4 larves).
</paragraph>
<paragraph id="18093682FF9A746FFD10FAE0FCA1FACE" blockId="14.[677,960,1293,1352]" box="[677,821,1293,1320]" pageId="14" pageNumber="33">DESCRIPTION</paragraph>
<paragraph id="18093682FF9A746FFD10FAC0FC54FAA0" blockId="14.[677,960,1293,1352]" box="[677,960,1325,1352]" pageId="14" pageNumber="33">
Couleur blanche (
<figureCitation id="808D2A07FF9A746FFCD2FAC0FC27FAA0" box="[871,947,1325,1352]" captionStart="FIG" captionStartId="29.[113,124,1567,1584]" captionTargetBox="[152,1172,222,1545]" captionTargetId="figure-18@29.[148,709,692,1169]" captionTargetPageId="29" captionText="FIG. 2. — Larves de Scarabaeoidea de Madagascar, habitus: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paramorphochelus cornutus (Nonfried,1892); K, Encya sikorai (Brenske,1891); L, Apicencya waterloti (Dewailly, 1950); M, Empecta scutata (Fairmaire,1901); N, Hoplochelus betanimena (Künckel,1887); O, Triodontus nitidulus (Guérin, 1844). Échelles: A-I, K, L, N, 5 mm; J, O, 2 mm; M, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4520757" httpUri="https://zenodo.org/record/4520757/files/figure.png" pageId="14" pageNumber="33">Fig. 2J</figureCitation>
).
</paragraph>
<paragraph id="18093682FF9A746FFD10FA81FCADFA6E" blockId="14.[676,1212,1388,1605]" box="[677,825,1388,1414]" pageId="14" pageNumber="33">
<emphasis id="2AC2EA90FF9A746FFD10FA81FCADFA6E" box="[677,825,1388,1414]" italics="true" pageId="14" pageNumber="33">
Tête (
<figureCitation id="808D2A07FF9A746FFD6BFA81FCA5FA6E" box="[734,817,1388,1414]" captionStart="FIG" captionStartId="33.[113,124,1635,1652]" captionTargetBox="[117,1206,214,1443]" captionTargetId="figure-14@33.[117,1206,214,1443]" captionTargetPageId="33" captionText="FIG. 6. — Larves de Scarabaeoidea de Madagascar: A-F, tête; A, Anochilia bifida (Olivier, 1789); B, Hexodon unicolor (Olivier, 1789); C, sp.D2; D, Paramorphochelus cornutus (Nonfried,1892); E, Empecta scutata (Fairmaire,1901); F, sp.S2; G, H, épipharynx; G, Bricoptis variolosa (Gory &amp; Percheron, 1833); H, Celidota parvula (Janson, 1881). Échelles: A-C, 2 mm; D-F, 1 mm; G, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520767" httpUri="https://zenodo.org/record/4520767/files/figure.png" pageId="14" pageNumber="33">Fig. 6D</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="18093682FF9A746FFD10FA61FCB1F9AD" blockId="14.[676,1212,1388,1605]" pageId="14" pageNumber="33">Couleur jaune. 1 soie dorso-épicraniale et 1 soie latéro-externe disposées symétriquement. 1 soie frontale postérieure et 2 soies frontales antérieures disposées symétriquement. Clypéus avec 1 soie latérale externe et 1 soie antérieure à disposition symétrique.</paragraph>
<paragraph id="18093682FF9A746EFD10F984FED9FEBA" blockId="14.[674,1209,1641,1700]" lastBlockId="15.[112,646,216,338]" lastPageId="15" lastPageNumber="34" pageId="14" pageNumber="33">
<emphasis id="2AC2EA90FF9A746FFD10F984FC9AF96B" bold="true" box="[677,782,1641,1667]" pageId="14" pageNumber="33">Antennes.</emphasis>
Les longueurs respectives des articles sont: I = 1; II = 2; III = 1,36; IV = 1,27. Dernier article antennaire avec 4 plages sensorielles: 2 ventrales, 1 minuscule plage apicale et 1 dorsale. 1 longue soie sur le second article. Avant-dernier article avec 1 saillie apico-latérale.
</paragraph>
<paragraph id="18093682FF9B746EFFC4FE9BFDE1FD67" blockId="15.[111,648,374,655]" pageId="15" pageNumber="34">
<emphasis id="2AC2EA90FF9B746EFFC4FE9BFF39FE78" bold="true" box="[113,173,374,400]" pageId="15" pageNumber="34">Labre</emphasis>
(
<figureCitation id="808D2A07FF9B746EFF0CFE9BFE83FE79" box="[185,279,374,401]" captionStart="FIG" captionStartId="40.[113,124,1612,1629]" captionTargetBox="[116,1208,217,1563]" captionTargetId="figure-21@40.[113,1202,214,1628]" captionTargetPageId="40" captionText="FIG. 13. — Larves de Scarabaeoidea de Madagascar: A-E, labre; A, Anochilia bifida (Olivier, 1789); B, Heteronychus plebeius (Klug, 1833); C, Paramorphochelus cornutus (Nonfried, 1892); D, Enaria melanictera (Klug, 1833); E, Triodontus nitidulus (Guérin, 1844); F-J, Celidota parvula (Janson, 1881), mandibule; F, gauche, face dorsale; G, gauche, face ventrale; H, droite, face dorsale; I, droite, face ventrale; J, aire stridulatoire mandibulaire. Échelles: A, B, D, F-I, 1 mm; C, E, J, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520793" httpUri="https://zenodo.org/record/4520793/files/figure.png" pageId="15" pageNumber="34">Fig. 13C</figureCitation>
).Transverse, symétrique, faiblement trilobé. Lobe médian avec 6 soies insérées chacune sur une protubérance pigmentée, les deux protubérances médianes accolées, presque fusionnées. Lobe latéral avec 2 soies externes et 2 soies internes, insérées chacune sur une protubérance pigmentée. 1 soie latéro-antérieure. 1 soie latérale. 2 longues soies labro-internes en avant du milieu. Une rangée transverse irrégulière de 4 à 5 soies postérieures.
</paragraph>
<paragraph id="18093682FF9B746EFFC4FD5EFE62FC84" blockId="15.[111,649,691,876]" pageId="15" pageNumber="34">
<emphasis id="2AC2EA90FF9B746EFFC4FD5EFF7BFD26" bold="true" box="[113,239,691,718]" pageId="15" pageNumber="34">Épipharynx</emphasis>
(
<figureCitation id="808D2A07FF9B746EFF49FD5EFEDAFD26" box="[252,334,691,718]" captionStart="FIG" captionStartId="35.[113,124,1635,1652]" captionTargetBox="[141,1181,217,1589]" captionTargetId="figure-14@35.[141,1181,217,1589]" captionTargetPageId="35" captionText="FIG. 8. — Larves de Scarabaeoidea de Madagascar, épipharynx: A, Heteroconus paradoxus (Endrödi,1968); B, Paranodon coquerelii (Fairmaire, 1871); C, Paramorphochelus cornutus (Nonfried,1892); D, sp. M1; E, Encya sikorai (Brenske,1891); F, Enaria melanictera (Klug, 1833); G, sp. M2. Échelles: A, F, 0,5 mm; B, C, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520775" httpUri="https://zenodo.org/record/4520775/files/figure.png" pageId="15" pageNumber="34">Fig. 8C</figureCitation>
). Haptomère formé par 9 à 11 fortes épines disposées en fer à cheval. Aire pédiale large et glabre. Chaetoparia dissymétriques, les soies convergeant vers laire pédiale. Gymnoparia très larges. Acanthoparia avec 6 à 9 courtes soies lamelliformes incurvées vers lavant.
</paragraph>
<paragraph id="18093682FF9B746EFFC4FC7DFDD7FB62" blockId="15.[111,649,912,1162]" pageId="15" pageNumber="34">
<emphasis id="2AC2EA90FF9B746EFFC4FC7DFF63FC42" bold="true" box="[113,247,912,938]" pageId="15" pageNumber="34">Mandibules</emphasis>
(
<figureCitation id="808D2A07FF9B746EFEBCFC7CFECDFC44" box="[265,345,913,940]" captionStart="FIG" captionStartId="42.[113,124,1589,1606]" captionTargetBox="[135,1177,220,1549]" captionTargetId="figure-21@42.[136,1178,214,1549]" captionTargetPageId="42" captionText="FIG. 15. — Larves de Scarabaeoidea de Madagascar, mandibule: A, E, I, M, Q, U, Y, gauche, face dorsale; B, F, J, N, R, V, Z1, gauche, face ventrale; C, G, K, O, S, W, Z2, droite,face dorsale; D, H, L, P, T, X, Z3, droite,face ventrale; A-D, Paranodon coquerelii (Fairmaire, 1871); E-H, Paramorphochelus cornutus (Nonfried, 1892); I-L, Encya sikorai (Brenske, 1891); M-P, Enaria melanictera (Klug, 1833); Q-T, Apicencya waterloti (Dewailly, 1950); U-X, Empecta scutata (Fairmaire, 1901); Y-Z3, Hoplochelus betanimena (Künckel, 1887). Échelles: A-D, M-P, 1 mm; E-H, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520799" httpUri="https://zenodo.org/record/4520799/files/figure.png" pageId="15" pageNumber="34">Fig. 15</figureCitation>
E-H). Aire scissoriale de la mandibule gauche allongée, se terminant par 3 dents, lapicale et la troisième dents aiguës, la deuxième dent émoussée. Aire scissoriale de la mandibule droite allongée, avec 2 dents émoussées. 7 soies latéro-externes et 1 soie dorso-externe. Il nexiste pas daire stridulatoire sous la forme dune aire striée en vis-à-vis des dents stridulatoires maxillaires.
</paragraph>
<paragraph id="18093682FF9B746EFFC4FB43FDA4FAA0" blockId="15.[111,649,1198,1352]" pageId="15" pageNumber="34">
<emphasis id="2AC2EA90FF9B746EFFC4FB43FF5EFB20" bold="true" box="[113,202,1198,1224]" pageId="15" pageNumber="34">Maxilles</emphasis>
(
<figureCitation id="808D2A07FF9B746EFF63FB43FEA6FB21" box="[214,306,1198,1225]" captionStart="FIG" captionStartId="44.[113,124,1612,1629]" captionTargetBox="[117,1206,214,1559]" captionTargetId="figure-21@44.[117,1206,214,1559]" captionTargetPageId="44" captionText="FIG. 17. — Larves de Scarabaeoidea de Madagascar, maxille, face dorsale: A, Paranodon coquerelii (Fairmaire, 1871); B, Paramorphochelus cornutus (Nonfried, 1892); C, Encya sikorai (Brenske, 1891); D, Enaria melanictera (Klug, 1833); E, Apicencya waterloti (Dewailly, 1950); F, Empecta scutata (Fairmaire, 1901); G, Hoplochelus betanimena (Künckel, 1887); H, Triodontus nitidulus (Guérin, 1844). Échelles: A, 0,5 mm; B, 0,1 mm; D, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520807" httpUri="https://zenodo.org/record/4520807/files/figure.png" pageId="15" pageNumber="34">Fig. 17B</figureCitation>
). Galea et lacinia soudées. Lacinia à uncus avec trois dents inégales et une très forte épine, galea à uncus pointu sclérifié et 6 fortes épines. Sur la face dorsale, 10 à 11 dents stridulatoires, séparées, pointues et inclinées vers lapex.
</paragraph>
<paragraph id="18093682FF9B746EFFC4FA81FE67F9EE" blockId="15.[113,648,1388,1542]" pageId="15" pageNumber="34">
<emphasis id="2AC2EA90FF9B746EFFC4FA81FE85FA6E" bold="true" box="[113,273,1388,1414]" pageId="15" pageNumber="34">Hypopharynx</emphasis>
(
<figureCitation id="808D2A07FF9B746EFE96FA81FE1BFA6F" box="[291,399,1388,1415]" captionStart="FIG" captionStartId="37.[113,124,1589,1606]" captionTargetBox="[146,1173,219,1547]" captionTargetId="figure-14@37.[146,1173,218,1547]" captionTargetPageId="37" captionText="FIG. 10. — Larves de Scarabaeoidea de Madagascar,hypopharynx: A, Bricoptis variolosa (Gory &amp; Percheron,1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K, Paramorphochelus cornutus (Nonfried,1892); L, Encya sikorai (Brenske, 1891). Échelles: A, J, 0,2 mm; C, D, F, G, I, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520783" httpUri="https://zenodo.org/record/4520783/files/figure.png" pageId="15" pageNumber="34">Fig. 10K</figureCitation>
). Glosses avec de très longues soies à lapex et de plus courtes et plus fortes soies à la base. Oncyli transverses, avec une dent sclérifiée postérieure triangulaire et une forte dent antérieure, sclérifiée et dressée.
</paragraph>
<paragraph id="18093682FF9B746EFFC4F9C7FF2DF9AC" blockId="15.[113,649,1578,1700]" box="[113,185,1578,1604]" pageId="15" pageNumber="34">
<emphasis id="2AC2EA90FF9B746EFFC4F9C7FF2DF9AC" box="[113,185,1578,1604]" italics="true" pageId="15" pageNumber="34">Thorax</emphasis>
</paragraph>
<paragraph id="18093682FF9B746EFFC4F9A7FC22FE39" blockId="15.[113,649,1578,1700]" lastBlockId="15.[676,1210,216,465]" pageId="15" pageNumber="34">
<emphasis id="2AC2EA90FF9B746EFFC4F9A7FF22F98C" bold="true" box="[113,182,1610,1636]" pageId="15" pageNumber="34">Pattes</emphasis>
(
<figureCitation id="808D2A07FF9B746EFF7DF9A7FEBBF98D" box="[200,303,1610,1637]" captionStart="FIG" captionStartId="45.[113,124,1635,1652]" captionTargetBox="[117,1206,215,1564]" captionTargetId="figure-14@45.[117,1206,214,1564]" captionTargetPageId="45" captionText="FIG. 18. — Larves de Scarabaeoidea de Madagascar, pattes: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Paramorphochelus cornutus (Nonfried, 1892). Échelles: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520809" httpUri="https://zenodo.org/record/4520809/files/figure.png" pageId="15" pageNumber="34">Fig. 18E</figureCitation>
). Longues, de taille croissante de P1 à P3. Coxa très allongée à apex arrondi, trochanter inséré latéralement sur la coxa, fémur, tibiotarse avec de fortes soies mêlées à de longues soies à lapex. Tibiotarse 3 plus court, plus large et plus arrondi que les deux précédents, sans griffe. Apex du tibiotase 1 avec une griffe sclérifiée large à la base, terminée par une longue pointe aiguë. Apex du tibiotase 2 avec une griffe sclérifiée plus courte que la précédente, large et anguleuse à la base, terminée en pointe.
</paragraph>
<paragraph id="18093682FF9B746EFD10FE18FC93FDE7" blockId="15.[675,1212,501,718]" box="[677,775,501,527]" pageId="15" pageNumber="34">
<emphasis id="2AC2EA90FF9B746EFD10FE18FC93FDE7" box="[677,775,501,527]" italics="true" pageId="15" pageNumber="34">Abdomen</emphasis>
</paragraph>
<paragraph id="18093682FF9B746EFD10FDF8FCC1FD26" blockId="15.[675,1212,501,718]" pageId="15" pageNumber="34">
<emphasis id="2AC2EA90FF9B746EFD10FDF8FD7AFDC7" bold="true" box="[677,750,533,559]" pageId="15" pageNumber="34">Raster</emphasis>
(
<figureCitation id="808D2A07FF9B746EFD4AFDF8FCC2FDD8" box="[767,854,533,560]" captionStart="FIG" captionStartId="31.[113,124,1635,1652]" captionTargetBox="[124,1194,218,1601]" captionTargetId="figure-14@31.[106,1195,214,1601]" captionTargetPageId="31" captionText="FIG. 4. — Larves de Scarabaeoidea de Madagascar,raster: A, sp. D1; B, Hexodon unicolor (Olivier, 1789); C, sp. D2; D, Heteroconus paradoxus (Endrödi, 1968); E, Paranodon coquerelii (Fairmaire, 1871); F, Paramorphochelus cornutus (Nonfried, 1892); G, sp. M1; H, Encya sikorai (Brenske, 1891). Échelles: A-D, H, 2 mm; E-G, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520763" httpUri="https://zenodo.org/record/4520763/files/figure.png" pageId="15" pageNumber="34">Fig. 4F</figureCitation>
). Fente anale en « Y ». Présence tout le long du côté dorsal comme du côté ventral de la fente anale de courtes soies mêlées à dautres plus longues. Teges avec une quarantaine de fortes soies aplaties incurvées à lapex. Barbula de longues soies très denses.
</paragraph>
</subSubSection>
</treatment>
</document>