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<document id="787B8EB98204885233766F347BD43DE0" ID-CLB-Dataset="8118" ID-DOI="10.5252/z2010n1a2" ID-GBIF-Dataset="4cecd899-5900-4411-8446-0d9552b1b098" ID-ISSN="1638-9387" ID-Zenodo-Dep="4520753" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612808309234" checkinUser="felipe" docAuthor="Randriamanantsoa, Richard, Aberlenc, Henri-Pierre, Ralisoa, Olga Bakoly, Ratnadass, Alain &amp; Vercambre, Bernard" docDate="2010" docId="901F8794FF927466FCB6FC3DFC40FD06" docLanguage="fr" docName="Zoosystema.32.1.19-72.pdf" docOrigin="Zoosystema 32 (1)" docSource="http://www.bioone.org/doi/abs/10.5252/z2010n1a2" docStyle="DocumentStyle:E638A3C99FA728B529662B46F84D75FD.3:Zoosystema.2009-2010.journal_article" docStyleId="E638A3C99FA728B529662B46F84D75FD" docStyleName="Zoosystema.2009-2010.journal_article" docStyleVersion="3" docTitle="Anochilia bifida" docType="treatment" docVersion="11" lastPageNumber="26" masterDocId="6C26FFECFF947461FFB5FFEDFF94FFE8" masterDocTitle="Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar" masterLastPageNumber="72" masterPageNumber="19" pageNumber="25" updateTime="1698928337295" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="47628898341C6FFAFFD68EF977AC5524">Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar</mods:title>
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<mods:namePart id="E28820DF66883636B0F8D5D8E15B91BC">Randriamanantsoa, Richard</mods:namePart>
<mods:affiliation id="673496A2DF8744AA844B4FCBBD524BAC">Fofifa - URP Scrid, 110 Antsirabe (Madagascar) r _ randriamanantsoa @ yahoo. fr</mods:affiliation>
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<mods:namePart id="43FD5E24AE3D5FA4A4CE55449D3937B6">Aberlenc, Henri-Pierre</mods:namePart>
<mods:affiliation id="C897149A48EB9C6A2A0EED7CAC67AF6D">Cirad Bios, UMR CBGP (INRA / IRD / Cirad / Montpellier SupAgro), Campus international de Baillarguet, Ta A- 55 / L, F- 34398 Montpellier cedex 5 (France) henri-pierre. aberlenc @ cirad. fr</mods:affiliation>
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<mods:namePart id="39AF0B4F5B5B865B8DD4B7D9CCD1B4FB">Ralisoa, Olga Bakoly</mods:namePart>
<mods:affiliation id="84F0525A4813B607553E3E9737235293">Département dEntomologie, Faculté des Sciences, Université dAntananarivo, Antananarivo 101 (Madagascar)</mods:affiliation>
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<mods:namePart id="A8697935B4EE386B46B8AA5BEF6EE094">Ratnadass, Alain</mods:namePart>
<mods:affiliation id="B960CAABBBF08D28B4A58DF400E921C6">Cirad, UPR HortSys - ICRISAT BP 12404, Niamey (Niger) a. ratnadass @ cgiar. org</mods:affiliation>
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<mods:namePart id="1F3A211FDBE8CF7FC20D572E1B630F21">Vercambre, Bernard</mods:namePart>
<mods:affiliation id="1E7121FA597EB1D6099DB5CF88F67C2F">Cirad, UPR Systèmes de Cultures annuels, avenue Agropolis, F- 34398 Montpellier cedex 5 (France) bernard. vercambre @ cirad. fr</mods:affiliation>
<mods:nameIdentifier id="9BC5922D906F85A997F43241E09E03A8" type="email">vercambre@cirad.fr</mods:nameIdentifier>
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<mods:title id="935836BBD84ACAA506B804DAEA2A4EDA">Zoosystema</mods:title>
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<mods:date id="330AE89231F360EB0A141B9E9261BDE5">2010</mods:date>
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<treatment id="901F8794FF927466FCB6FC3DFC40FD06" ID-DOI="http://doi.org/10.5281/zenodo.4557661" ID-GBIF-Taxon="177425670" ID-Zenodo-Dep="4557661" LSID="urn:lsid:plazi:treatment:901F8794FF927466FCB6FC3DFC40FD06" httpUri="http://treatment.plazi.org/id/901F8794FF927466FCB6FC3DFC40FD06" lastPageId="7" lastPageNumber="26" pageId="6" pageNumber="25">
<subSubSection id="50AC6509FF927467FCB6FC3DFBCFFC03" box="[771,1115,976,1003]" pageId="6" pageNumber="25" type="nomenclature">
<paragraph id="18093682FF927467FCB6FC3DFBCFFC03" blockId="6.[771,1115,976,1003]" box="[771,1115,976,1003]" pageId="6" pageNumber="25">
<taxonomicName id="DFB64D01FF927467FCB6FC3DFBCFFC03" authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[771,1115,976,1003]" class="Insecta" family="Cetoniidae" genus="Anochilia" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="25" phylum="Arthropoda" rank="species" species="bifida">
<emphasis id="2AC2EA90FF927467FCB6FC3DFC26FC03" bold="true" box="[771,946,976,1003]" italics="true" pageId="6" pageNumber="25">Anochilia bifida</emphasis>
(Olivier, 1789)
</taxonomicName>
</paragraph>
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<subSubSection id="50AC6509FF927467FD44FC1DFBFFFBC2" pageId="6" pageNumber="25" type="description">
<paragraph id="18093682FF927467FD44FC1DFBFFFBC2" blockId="6.[753,1133,1008,1066]" pageId="6" pageNumber="25">
(
<figureCitation id="808D2A07FF927467FD4DFC1DFCD9FBE3" box="[760,845,1008,1035]" captionStart="FIG" captionStartId="29.[113,124,1567,1584]" captionTargetBox="[152,1172,222,1545]" captionTargetId="figure-18@29.[148,709,692,1169]" captionTargetPageId="29" captionText="FIG. 2. — Larves de Scarabaeoidea de Madagascar, habitus: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paramorphochelus cornutus (Nonfried,1892); K, Encya sikorai (Brenske,1891); L, Apicencya waterloti (Dewailly, 1950); M, Empecta scutata (Fairmaire,1901); N, Hoplochelus betanimena (Künckel,1887); O, Triodontus nitidulus (Guérin, 1844). Échelles: A-I, K, L, N, 5 mm; J, O, 2 mm; M, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4520757" httpUri="https://zenodo.org/record/4520757/files/figure.png" pageId="6" pageNumber="25">Figs 2C</figureCitation>
;
<figureCitation id="808D2A07FF927467FCEAFC1DFC17FBE3" box="[863,899,1008,1035]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="6" pageNumber="25">3D</figureCitation>
;
<figureCitation id="808D2A07FF927467FC21FC1DFC27FBE3" box="[916,947,1008,1035]" captionStart="FIG" captionStartId="33.[113,124,1635,1652]" captionTargetBox="[117,1206,214,1443]" captionTargetId="figure-14@33.[117,1206,214,1443]" captionTargetPageId="33" captionText="FIG. 6. — Larves de Scarabaeoidea de Madagascar: A-F, tête; A, Anochilia bifida (Olivier, 1789); B, Hexodon unicolor (Olivier, 1789); C, sp.D2; D, Paramorphochelus cornutus (Nonfried,1892); E, Empecta scutata (Fairmaire,1901); F, sp.S2; G, H, épipharynx; G, Bricoptis variolosa (Gory &amp; Percheron, 1833); H, Celidota parvula (Janson, 1881). Échelles: A-C, 2 mm; D-F, 1 mm; G, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520767" httpUri="https://zenodo.org/record/4520767/files/figure.png" pageId="6" pageNumber="25">6A</figureCitation>
;
<figureCitation id="808D2A07FF927467FC70FC1DFC70FBE3" box="[965,996,1008,1035]" captionStart="FIG" captionStartId="34.[113,124,1635,1652]" captionTargetBox="[155,1168,220,1591]" captionTargetId="figure-21@34.[155,1168,220,1591]" captionTargetPageId="34" captionText="FIG. 7. — Larves de Scarabaeoidea de Madagascar,épipharynx: A, Anochilia bifida (Olivier,1789); B, Euryomia argentea (Olivier,1789); C, Heteronychus arator rugifrons (Fairmaire, 1871); D, Heteronychus bituberculatus (Kolbe, 1900); E, F, sp. D1; G, Hexodon unicolor (Olivier, 1789); H, sp. D2. Échelles: A, B, D, E, G, 0,5 mm; C, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520771" httpUri="https://zenodo.org/record/4520771/files/figure.png" pageId="6" pageNumber="25">7A</figureCitation>
;
<figureCitation id="808D2A07FF927467FC43FC1DFBB2FBE3" box="[1014,1062,1008,1035]" captionStart="FIG" captionStartId="37.[113,124,1589,1606]" captionTargetBox="[146,1173,219,1547]" captionTargetId="figure-14@37.[146,1173,218,1547]" captionTargetPageId="37" captionText="FIG. 10. — Larves de Scarabaeoidea de Madagascar,hypopharynx: A, Bricoptis variolosa (Gory &amp; Percheron,1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K, Paramorphochelus cornutus (Nonfried,1892); L, Encya sikorai (Brenske, 1891). Échelles: A, J, 0,2 mm; C, D, F, G, I, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520783" httpUri="https://zenodo.org/record/4520783/files/figure.png" pageId="6" pageNumber="25">10C</figureCitation>
;
<figureCitation id="808D2A07FF927467FB82FC1DFBF6FBE3" box="[1079,1122,1008,1035]" captionStart="FIG" captionStartId="39.[113,124,1567,1584]" captionTargetBox="[118,1207,216,1486]" captionTargetId="figure-14@39.[118,1207,216,1486]" captionTargetPageId="39" captionText="FIG. 12. — Larves de Scarabaeoidea de Madagascar: A-E, hypopharynx; A, Enaria melanictera (Klug, 1833); B, Apicencya waterloti (Dewailly, 1950); C, Empecta scutata (Fairmaire, 1901); D, Hoplochelus betanimena (Künckel, 1887); E, Triodontus nitidulus (Guérin, 1844); F-O, antenne; F, Anochilia bifida (Olivier, 1789); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K-M, Encya sikorai (Brenske,1891); L, M, plages sensorielles de larticle apical; N, Apicencya waterloti (Dewailly, 1950); O, Empecta scutata (Fairmaire, 1901). Échelles: A, F, G, I, J, O, 0,5 mm; E, L, M, 0,2 mm; H, K, N, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520787" httpUri="https://zenodo.org/record/4520787/files/figure.png" pageId="6" pageNumber="25">12F</figureCitation>
;
<figureCitation id="808D2A07FF927467FD41FBE2FCB5FBC2" box="[756,801,1039,1066]" captionStart="FIG" captionStartId="40.[113,124,1612,1629]" captionTargetBox="[116,1208,217,1563]" captionTargetId="figure-21@40.[113,1202,214,1628]" captionTargetPageId="40" captionText="FIG. 13. — Larves de Scarabaeoidea de Madagascar: A-E, labre; A, Anochilia bifida (Olivier, 1789); B, Heteronychus plebeius (Klug, 1833); C, Paramorphochelus cornutus (Nonfried, 1892); D, Enaria melanictera (Klug, 1833); E, Triodontus nitidulus (Guérin, 1844); F-J, Celidota parvula (Janson, 1881), mandibule; F, gauche, face dorsale; G, gauche, face ventrale; H, droite, face dorsale; I, droite, face ventrale; J, aire stridulatoire mandibulaire. Échelles: A, B, D, F-I, 1 mm; C, E, J, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520793" httpUri="https://zenodo.org/record/4520793/files/figure.png" pageId="6" pageNumber="25">13A</figureCitation>
;
<figureCitation id="808D2A07FF927467FC86FBE2FCD9FBC2" box="[819,845,1039,1066]" captionStart="FIG" captionStartId="41.[113,124,1589,1606]" captionTargetBox="[161,1157,219,1557]" captionTargetId="figure-14@41.[157,1165,214,1560]" captionTargetPageId="41" captionText="FIG. 14. — Larves de Scarabaeoidea de Madagascar,mandibule: A, D, H, L, P, T, X, gauche,face dorsale; B, E, I, M, Q, U, Y, gauche, face ventrale; C, F, J, N, R, V, Z1, droite, face dorsale; G, K, O, S, W, Z2, droite, face ventrale; A-C, Bricoptis variolosa (Gory &amp; Percheron, 1833); D-G, Anochilia bifida (Olivier, 1789); H-K, Euryomia argentea (Olivier, 1789); L-O, Heteronychus arator rugifrons (Fairmaire,1871); P-S, Heteronychus bituberculatus (Kolbe,1900); T-W, Hexodon unicolor (Olivier,1789); X-Z2, Heteroconus paradoxus (Endrödi, 1968). Échelles: A-C, H-O, 0,5 mm; D-G, P-Z2, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520795" httpUri="https://zenodo.org/record/4520795/files/figure.png" pageId="6" pageNumber="25">14</figureCitation>
D-G; 16G; 18C; 23E-G)
</paragraph>
</subSubSection>
<subSubSection id="50AC6509FF927467FD10FBA2FB39FB01" pageId="6" pageNumber="25" type="materials_examined">
<paragraph id="18093682FF927467FD10FBA2FB39FB01" blockId="6.[677,1211,1103,1257]" pageId="6" pageNumber="25">
MATÉRIEL EXAMINÉ Larves provenant dIbity. Lespèce a été collectée dans du fumier de Bovins (utilisé pour fertiliser les rizières). Spécimens conservés dans les collections du Fofifa (
<specimenCount id="0EB0FD0BFF927467FCADFB23FCF4FB01" box="[792,864,1230,1257]" pageId="6" pageNumber="25" type="larva">1 larve</specimenCount>
) et du Cirad-CBGP (2 larves).
</paragraph>
</subSubSection>
<subSubSection id="50AC6509FF927466FD10FAE0FC40FD06" lastPageId="7" lastPageNumber="26" pageId="6" pageNumber="25" type="description">
<paragraph id="18093682FF927467FD10FAE0FCA1FACE" blockId="6.[677,1011,1293,1352]" box="[677,821,1293,1320]" pageId="6" pageNumber="25">DESCRIPTION</paragraph>
<paragraph id="18093682FF927467FD10FAC0FC67FAA0" blockId="6.[677,1011,1293,1352]" box="[677,1011,1325,1352]" pageId="6" pageNumber="25">
Couleur blanc ivoire (
<figureCitation id="808D2A07FF927467FC25FAC0FC72FAA0" box="[912,998,1325,1352]" captionStart="FIG" captionStartId="29.[113,124,1567,1584]" captionTargetBox="[152,1172,222,1545]" captionTargetId="figure-18@29.[148,709,692,1169]" captionTargetPageId="29" captionText="FIG. 2. — Larves de Scarabaeoidea de Madagascar, habitus: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paramorphochelus cornutus (Nonfried,1892); K, Encya sikorai (Brenske,1891); L, Apicencya waterloti (Dewailly, 1950); M, Empecta scutata (Fairmaire,1901); N, Hoplochelus betanimena (Künckel,1887); O, Triodontus nitidulus (Guérin, 1844). Échelles: A-I, K, L, N, 5 mm; J, O, 2 mm; M, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4520757" httpUri="https://zenodo.org/record/4520757/files/figure.png" pageId="6" pageNumber="25">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="18093682FF927467FD10FA81FCA1FA6E" blockId="6.[677,1209,1388,1510]" box="[677,821,1388,1414]" pageId="6" pageNumber="25">
<emphasis id="2AC2EA90FF927467FD10FA81FCA1FA6E" box="[677,821,1388,1414]" italics="true" pageId="6" pageNumber="25">
Tête (
<figureCitation id="808D2A07FF927467FD6BFA81FCBAFA6E" box="[734,814,1388,1414]" captionStart="FIG" captionStartId="33.[113,124,1635,1652]" captionTargetBox="[117,1206,214,1443]" captionTargetId="figure-14@33.[117,1206,214,1443]" captionTargetPageId="33" captionText="FIG. 6. — Larves de Scarabaeoidea de Madagascar: A-F, tête; A, Anochilia bifida (Olivier, 1789); B, Hexodon unicolor (Olivier, 1789); C, sp.D2; D, Paramorphochelus cornutus (Nonfried,1892); E, Empecta scutata (Fairmaire,1901); F, sp.S2; G, H, épipharynx; G, Bricoptis variolosa (Gory &amp; Percheron, 1833); H, Celidota parvula (Janson, 1881). Échelles: A-C, 2 mm; D-F, 1 mm; G, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520767" httpUri="https://zenodo.org/record/4520767/files/figure.png" pageId="6" pageNumber="25">Fig. 6A</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="18093682FF927467FD10FA61FBC4FA0E" blockId="6.[677,1209,1388,1510]" pageId="6" pageNumber="25">Couleur brunâtre. Pas docelle. 1 soie dorso-épicraniale et 5 soies épicraniales latéro-externes. Pas de soie frontale. 1 soie dans langle frontal.</paragraph>
<paragraph id="18093682FF927467FD10F9E7FC3DF94C" blockId="6.[674,1212,1546,1700]" pageId="6" pageNumber="25">
<emphasis id="2AC2EA90FF927467FD10F9E7FC99F9CC" bold="true" box="[677,781,1546,1572]" pageId="6" pageNumber="25">Antennes</emphasis>
(
<figureCitation id="808D2A07FF927467FCAEF9E7FCEFF9CD" box="[795,891,1546,1573]" captionStart="FIG" captionStartId="39.[113,124,1567,1584]" captionTargetBox="[118,1207,216,1486]" captionTargetId="figure-14@39.[118,1207,216,1486]" captionTargetPageId="39" captionText="FIG. 12. — Larves de Scarabaeoidea de Madagascar: A-E, hypopharynx; A, Enaria melanictera (Klug, 1833); B, Apicencya waterloti (Dewailly, 1950); C, Empecta scutata (Fairmaire, 1901); D, Hoplochelus betanimena (Künckel, 1887); E, Triodontus nitidulus (Guérin, 1844); F-O, antenne; F, Anochilia bifida (Olivier, 1789); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K-M, Encya sikorai (Brenske,1891); L, M, plages sensorielles de larticle apical; N, Apicencya waterloti (Dewailly, 1950); O, Empecta scutata (Fairmaire, 1901). Échelles: A, F, G, I, J, O, 0,5 mm; E, L, M, 0,2 mm; H, K, N, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520787" httpUri="https://zenodo.org/record/4520787/files/figure.png" pageId="6" pageNumber="25">Fig. 12F</figureCitation>
). Article I un peu moins long que les II et III pris ensemble. Une aire sensorielle apico-latérale sur lavant-dernier article, 8 plages sensorielles sur larticle apical: une petite aire apicale, 4 ventrales et 3 dorsales.
</paragraph>
<paragraph id="18093682FF937466FFC4FF35FF4AFE39" blockId="7.[113,649,216,465]" pageId="7" pageNumber="26">
<emphasis id="2AC2EA90FF937466FFC4FF35FF24FF1A" bold="true" box="[113,176,216,242]" pageId="7" pageNumber="26">Labre</emphasis>
(
<figureCitation id="808D2A07FF937466FF0AFF35FEB6FF1B" box="[191,290,216,243]" captionStart="FIG" captionStartId="40.[113,124,1612,1629]" captionTargetBox="[116,1208,217,1563]" captionTargetId="figure-21@40.[113,1202,214,1628]" captionTargetPageId="40" captionText="FIG. 13. — Larves de Scarabaeoidea de Madagascar: A-E, labre; A, Anochilia bifida (Olivier, 1789); B, Heteronychus plebeius (Klug, 1833); C, Paramorphochelus cornutus (Nonfried, 1892); D, Enaria melanictera (Klug, 1833); E, Triodontus nitidulus (Guérin, 1844); F-J, Celidota parvula (Janson, 1881), mandibule; F, gauche, face dorsale; G, gauche, face ventrale; H, droite, face dorsale; I, droite, face ventrale; J, aire stridulatoire mandibulaire. Échelles: A, B, D, F-I, 1 mm; C, E, J, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520793" httpUri="https://zenodo.org/record/4520793/files/figure.png" pageId="7" pageNumber="26">Fig. 13A</figureCitation>
). Transverse, trilobé, symétrique. Lobe médian avec 4 fortes soies externes et 4 soies internes. Lobe latéral avec, davant en arrière, 4 soies antérieures externes et 3 longues soies + 1 très longue soie antérieures internes, et 1 longue soie latérale. 2 longues soies labro-internes. 10 à 12 courtes soies postérieures (en 2 groupes de 5 à 6 soies).
</paragraph>
<paragraph id="18093682FF937466FFC4FE18FEF5FCA5" blockId="7.[111,649,501,845]" pageId="7" pageNumber="26">
<emphasis id="2AC2EA90FF937466FFC4FE18FF65FDF8" bold="true" box="[113,241,501,528]" pageId="7" pageNumber="26">Épipharynx</emphasis>
(
<figureCitation id="808D2A07FF937466FF4BFE18FEC5FDF8" box="[254,337,501,528]" captionStart="FIG" captionStartId="34.[113,124,1635,1652]" captionTargetBox="[155,1168,220,1591]" captionTargetId="figure-21@34.[155,1168,220,1591]" captionTargetPageId="34" captionText="FIG. 7. — Larves de Scarabaeoidea de Madagascar,épipharynx: A, Anochilia bifida (Olivier,1789); B, Euryomia argentea (Olivier,1789); C, Heteronychus arator rugifrons (Fairmaire, 1871); D, Heteronychus bituberculatus (Kolbe, 1900); E, F, sp. D1; G, Hexodon unicolor (Olivier, 1789); H, sp. D2. Échelles: A, B, D, E, G, 0,5 mm; C, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520771" httpUri="https://zenodo.org/record/4520771/files/figure.png" pageId="7" pageNumber="26">Fig. 7A</figureCitation>
). Haptomère avec une rangée de 9 à 13 épines formant un peigne. Aire pédiale glabre. Chaetoparia dissymétriques, avec 2 rangées concentriques de soies convergeant vers laire pédiale du côté gauche, les plus internes plus fortes, et avec de nombreuses soies épineuses non alignées, devenant plus petites vers lintérieur du côté droit, avec de fines soies du côté externe. Acanthoparia formés de 8 à 11 soies de taille croissante vers lavant. Clithra bien marqués, la concavité étant légèrement anguleuse.
</paragraph>
<paragraph id="18093682FF937466FFC4FC9CFF7BFB21" blockId="7.[111,648,881,1225]" pageId="7" pageNumber="26">
<emphasis id="2AC2EA90FF937466FFC4FC9CFF68FC63" bold="true" box="[113,252,881,907]" pageId="7" pageNumber="26">Mandibules</emphasis>
(
<figureCitation id="808D2A07FF937466FEBAFC9CFEF0FC64" box="[271,356,881,908]" captionStart="FIG" captionStartId="41.[113,124,1589,1606]" captionTargetBox="[161,1157,219,1557]" captionTargetId="figure-14@41.[157,1165,214,1560]" captionTargetPageId="41" captionText="FIG. 14. — Larves de Scarabaeoidea de Madagascar,mandibule: A, D, H, L, P, T, X, gauche,face dorsale; B, E, I, M, Q, U, Y, gauche, face ventrale; C, F, J, N, R, V, Z1, droite, face dorsale; G, K, O, S, W, Z2, droite, face ventrale; A-C, Bricoptis variolosa (Gory &amp; Percheron, 1833); D-G, Anochilia bifida (Olivier, 1789); H-K, Euryomia argentea (Olivier, 1789); L-O, Heteronychus arator rugifrons (Fairmaire,1871); P-S, Heteronychus bituberculatus (Kolbe,1900); T-W, Hexodon unicolor (Olivier,1789); X-Z2, Heteroconus paradoxus (Endrödi, 1968). Échelles: A-C, H-O, 0,5 mm; D-G, P-Z2, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520795" httpUri="https://zenodo.org/record/4520795/files/figure.png" pageId="7" pageNumber="26">Fig. 14</figureCitation>
D-G). Aire scissoriale de la mandibule gauche avec 4 dents développées à bord arrondi. Aire scissoriale de la mandibule droite avec 3 dents, la première très développée et pointue. Mandibule gauche avec 3 longues + 4 courtes soies baso-latérales et 2 soies dorso-externes. Mandibule droite avec 2 longues + 6 courtes soies baso-latérales et 2 soies dorso-externes. Sur la face ventrale des 2 mandibules, présence dune aire stridulatoire composée de 21 à 24 bourrelets transverses.
</paragraph>
<paragraph id="18093682FF937466FFC4FB00FE2AFA4F" blockId="7.[111,648,1261,1447]" pageId="7" pageNumber="26">
<emphasis id="2AC2EA90FF937466FFC4FB00FF5DFAEF" bold="true" box="[113,201,1261,1287]" pageId="7" pageNumber="26">Maxilles</emphasis>
(
<figureCitation id="808D2A07FF937466FF63FB00FEA0FAE0" box="[214,308,1261,1288]" captionStart="FIG" captionStartId="43.[113,124,1567,1584]" captionTargetBox="[152,1180,217,1546]" captionTargetId="figure-14@43.[151,1175,219,1547]" captionTargetPageId="43" captionText="FIG. 16. — Larves de Scarabaeoidea de Madagascar: A-D, mandibule, Triodontus nitidulus (Guérin, 1844); A, gauche, face dorsale; B, gauche, face ventrale; C, droite, face dorsale; D, droite, face ventrale; E-O, maxille, face dorsale; E, Bricoptis variolosa (Gory &amp; Percheron, 1833); F, Celidota parvula (Janson, 1881); G, Anochilia bifida (Olivier, 1789); H, Euryomia argentea (Olivier, 1789); I, J, Heteronychus arator rugifrons (Fairmaire, 1871); J, dents stridulatoires; K, L, Heteronychus bituberculatus (Kolbe,1900); L, dents stridulatoires; M, Heteronychus plebeius (Klug,1833); N, Hexodon unicolor (Olivier,1789); O, Heteroconus paradoxus (Endrödi,1968). Échelles: A-D, N, O, 0,5 mm; E, F, H, I, K, 0,4 mm; G, 1 mm; J, 0,01 mm; L, 0,02 mm." figureDoi="http://doi.org/10.5281/zenodo.4520805" httpUri="https://zenodo.org/record/4520805/files/figure.png" pageId="7" pageNumber="26">Fig. 16G</figureCitation>
). Galea et lacinia soudées. Lacinia à uncus avec 2 dents sclérifiées (1 grande et 1 petite), soudées à leur base. Galea avec uncus sclérifié. Sur la face dorsale, présence dune aire stridulatoire composée de 6 à 8 + 1 dents pointues, coniques, espacées, penchées vers lavant.
</paragraph>
<paragraph id="18093682FF937466FFC4FA26FE9AF94C" blockId="7.[113,648,1483,1700]" pageId="7" pageNumber="26">
<emphasis id="2AC2EA90FF937466FFC4FA26FE9BFA0D" bold="true" box="[113,271,1483,1509]" pageId="7" pageNumber="26">Hypopharynx</emphasis>
(
<figureCitation id="808D2A07FF937466FE94FA26FE1FFA0E" box="[289,395,1483,1510]" captionStart="FIG" captionStartId="37.[113,124,1589,1606]" captionTargetBox="[146,1173,219,1547]" captionTargetId="figure-14@37.[146,1173,218,1547]" captionTargetPageId="37" captionText="FIG. 10. — Larves de Scarabaeoidea de Madagascar,hypopharynx: A, Bricoptis variolosa (Gory &amp; Percheron,1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K, Paramorphochelus cornutus (Nonfried,1892); L, Encya sikorai (Brenske, 1891). Échelles: A, J, 0,2 mm; C, D, F, G, I, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520783" httpUri="https://zenodo.org/record/4520783/files/figure.png" pageId="7" pageNumber="26">Fig. 10C</figureCitation>
). Partie antérieure des glosses avec 2 rangées de 3 longues soies, les plus longues à lapex, les plus courtes à la base. Une touffe de soies latérales à lapex. En avant de loncylus, de courtes et fortes soies sont disposées en «U». Oncylus transverse, fortement sclérifié, avec 2 lobes saillants.
</paragraph>
<paragraph id="18093682FF937466FD10FF35FD79FF1A" blockId="7.[675,1210,216,401]" box="[677,749,216,242]" pageId="7" pageNumber="26">
<emphasis id="2AC2EA90FF937466FD10FF35FD79FF1A" box="[677,749,216,242]" italics="true" pageId="7" pageNumber="26">Thorax</emphasis>
</paragraph>
<paragraph id="18093682FF937466FD10FF15FCA4FE79" blockId="7.[675,1210,216,401]" pageId="7" pageNumber="26">
<emphasis id="2AC2EA90FF937466FD10FF15FD73FEFA" bold="true" box="[677,743,248,274]" pageId="7" pageNumber="26">Pattes</emphasis>
(
<figureCitation id="808D2A07FF937466FD41FF1AFCC2FEFA" box="[756,854,247,274]" captionStart="FIG" captionStartId="45.[113,124,1635,1652]" captionTargetBox="[117,1206,215,1564]" captionTargetId="figure-14@45.[117,1206,214,1564]" captionTargetPageId="45" captionText="FIG. 18. — Larves de Scarabaeoidea de Madagascar, pattes: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Paramorphochelus cornutus (Nonfried, 1892). Échelles: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520809" httpUri="https://zenodo.org/record/4520809/files/figure.png" pageId="7" pageNumber="26">Fig. 18C</figureCitation>
). Courtes, de longueur croissante de P1 à P3, toutes de structure analogue: coxa, trochanter, fémur tibia et tarse différenciés. Tarse court, fortement pigmenté, apex sclérifié avec une courte griffe.
</paragraph>
<paragraph id="18093682FF937466FD10FE58FC93FE27" blockId="7.[675,1210,437,750]" box="[677,775,437,463]" pageId="7" pageNumber="26">
<emphasis id="2AC2EA90FF937466FD10FE58FC93FE27" box="[677,775,437,463]" italics="true" pageId="7" pageNumber="26">Abdomen</emphasis>
</paragraph>
<paragraph id="18093682FF937466FD10FE38FC40FD06" blockId="7.[675,1210,437,750]" pageId="7" pageNumber="26">
<emphasis id="2AC2EA90FF937466FD10FE38FD7DFE07" bold="true" box="[677,745,469,495]" pageId="7" pageNumber="26">Raster</emphasis>
(
<figureCitation id="808D2A07FF937466FD40FE38FCDDFE18" box="[757,841,469,496]" captionStart="FIG" captionStartId="30.[113,124,1612,1629]" captionTargetBox="[139,1184,217,1560]" captionTargetId="figure-21@30.[139,1184,217,1561]" captionTargetPageId="30" captionText="FIG. 3. — Larves de Scarabaeoidea de Madagascar, raster: A, Bricoptis variolosa (Gory &amp; Percheron, 1833); B, Celidota parvula (Janson, 1881); C, sp. C1; D, Anochilia bifida (Olivier, 1789); E, Euryomia argentea (Olivier, 1789); F, Heteronychus arator rugifrons (Fairmaire, 1871); G, Heteronychus bituberculatus (Kolbe, 1900); H, Heteronychus plebeius (Klug, 1833). Échelles: A-C, G, H, 1 mm; D, 0,5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520761" httpUri="https://zenodo.org/record/4520761/files/figure.png" pageId="7" pageNumber="26">Fig. 3D</figureCitation>
). Palidia formés de 2 rangées subparallèles, très légèrement incurvées et divergeant vers larrière, constituées de 11 à 14 épines inclinées vers le septula et vers larrière. Septula glabre. De chaque côté de la partie antérieure des palidia, 2 très longues soies, la plus interne en arrière, la plus externe en avant. Teges formés de soies courtes inclinées vers larrière. Fente anale transverse, faiblement arquée, proche de lapex abdominal.
</paragraph>
</subSubSection>
</treatment>
</document>