treatments-xml/data/71/66/B3/7166B372006C461BA871FA7E82ACD52A.xml
2024-06-21 12:40:05 +02:00

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<document id="EE9BC54DF02B2D88BCFBA6034A634FB9" ID-DOI="10.5281/zenodo.10905322" ID-ISSN="1407-8619" ID-Zenodo-Dep="10905322" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712002783355" checkinUser="felipe" docAuthor="Fanti, Fabrizio &amp; Damgaard, Anders Leth" docDate="2019" docId="7166B372006C461BA871FA7E82ACD52A" docLanguage="en" docName="BalticJColeopterol.19.2.101-125.pdf" docOrigin="Baltic Journal of Coleopterology 19 (2)" docStyle="DocumentStyle:AA156050B09C0A4609AEA89F10CC943C.2:BalticJColeopterol.2012-2021.journal_article" docStyleId="AA156050B09C0A4609AEA89F10CC943C" docStyleName="BalticJColeopterol.2012-2021.journal_article" docStyleVersion="2" docTitle="Sanaungulus fabriciusi Fanti &amp; Damgaard 2019, sp. nov." docType="treatment" docVersion="2" lastPageNumber="113" masterDocId="8D5FCB0A00654617AB77FF818631D03D" masterDocTitle="New soldier beetles (Cantharidae) from Baltic, Burmese and Dominican ambers of the Anders Damgaard amber collection" masterLastPageNumber="125" masterPageNumber="101" pageNumber="110" updateTime="1712003226581" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="69E29224B94F8ED07636DB7C4F3D3338">New soldier beetles (Cantharidae) from Baltic, Burmese and Dominican ambers of the Anders Damgaard amber collection</mods:title>
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<mods:namePart id="AC1715D78FAE92C95693E5BF1ED7B6BA">Fanti, Fabrizio</mods:namePart>
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<mods:namePart id="EBADC60E872344173C617492EB2D6B49">Damgaard, Anders Leth</mods:namePart>
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<mods:title id="4424A6D6DFA75D94C8750773E9735D3F">Baltic Journal of Coleopterology</mods:title>
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<mods:date id="FF9CA2D812E393C80B7A1AE9D40DA7FD">2019</mods:date>
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<mods:number id="A2ACEFCA4453DF0922F0200C4723A1A5">2019-12-31</mods:number>
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<treatment id="7166B372006C461BA871FA7E82ACD52A" ID-DOI="http://doi.org/10.5281/zenodo.10905382" ID-Zenodo-Dep="10905382" LSID="urn:lsid:plazi:treatment:7166B372006C461BA871FA7E82ACD52A" httpUri="http://treatment.plazi.org/id/7166B372006C461BA871FA7E82ACD52A" lastPageId="12" lastPageNumber="113" pageId="9" pageNumber="110">
<subSubSection id="B1D551EF006C461EA871FA7E826BD62A" box="[774,1114,1535,1559]" pageId="9" pageNumber="110" type="nomenclature">
<paragraph id="F9700264006C461EA871FA7E826BD62A" blockId="9.[774,1114,1535,1591]" box="[774,1114,1535,1559]" pageId="9" pageNumber="110">
<heading id="A238B508006C461EA871FA7E826BD62A" bold="true" box="[774,1114,1535,1559]" fontSize="10" level="2" pageId="9" pageNumber="110" reason="4">
<emphasis id="CBBBDE76006C461EA871FA7E826BD62A" bold="true" box="[774,1114,1535,1559]" pageId="9" pageNumber="110">
<taxonomicName id="3ECF79E7006C461EA871FA7E8235D62A" authority="Fanti &amp; Damgaard, 2019" authorityName="Fanti &amp; Damgaard" authorityYear="2019" box="[774,1028,1535,1559]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="fabriciusi" status="sp. nov.">
<emphasis id="CBBBDE76006C461EA871FA7E8235D62A" bold="true" box="[774,1028,1535,1559]" italics="true" pageId="9" pageNumber="110">Sanaungulus fabriciusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D088630D006C461EAF7EFA7E826BD62A" box="[1033,1114,1535,1559]" pageId="9" pageNumber="110" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="B1D551EF006C461EA871F99E85BBD60A" box="[774,906,1567,1591]" pageId="9" pageNumber="110" type="description">
<paragraph id="F9700264006C461EA871F99E85BBD60A" blockId="9.[774,1114,1535,1591]" box="[774,906,1567,1591]" pageId="9" pageNumber="110">
(
<figureCitation id="61F41EE1006C461EA879F99E85B3D60A" box="[782,898,1567,1591]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="10.[303,342,755,779]" captionStartId-1="10.[303,343,1525,1549]" captionTargetBox-0="[446,1220,192,707]" captionTargetBox-1="[447,1216,953,1466]" captionTargetId-0="figure-2@10.[446,1220,192,707]" captionTargetId-1="figure-55@10.[447,1216,953,1466]" captionTargetPageId-0="10" captionTargetPageId-1="10" captionText-0="Fig. 9. Sanaungulus fabriciusi sp. nov. ALDC0541/ALD.Bu.208 in Burmese amber. A: Holotype, lateral view, bar = 500 μm; B: Holotype, lateral view (more detailed), bar = 500 μm; C: Holotype, ventral view, bar = 500 μm; D: Holotype, detail of head, palp, mandible and pronotum, bar = 200 μm." captionText-1="Fig. 10. Sanaungulus fabriciusi sp. nov.ALDC0541/ALD.Bu.208 in Burmese amber.A: Holotype, detail of mandibles and head, bar = 100 μm; B: Holotype, detail of antennae, bar = 100 μm; C: Holotype, detail of pronotum, bar = 100 μm; D: Holotype, detail of coxae, trochanters and femora, bar = 100 μm." figureDoi-0="http://doi.org/10.5281/zenodo.10905346" figureDoi-1="http://doi.org/10.5281/zenodo.10905348" httpUri-0="https://zenodo.org/record/10905346/files/figure.png" httpUri-1="https://zenodo.org/record/10905348/files/figure.png" pageId="9" pageNumber="110">Figs. 9 - 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B1D551EF006C461BA871F9DE8479D16A" lastPageId="12" lastPageNumber="113" pageId="9" pageNumber="110" type="materials_examined">
<paragraph id="F9700264006C461EA871F9DE82BBD6AA" blockId="9.[774,1285,1631,1687]" pageId="9" pageNumber="110">
<materialsCitation id="49A70839006C461EA871F9DE82BBD6AA" collectionCode="ALDC" pageId="9" pageNumber="110" specimenCode="ALDC0541" specimenCount="1" typeStatus="holotype">
<emphasis id="CBBBDE76006C461EA871F9DE8545D64A" bold="true" box="[774,884,1631,1655]" pageId="9" pageNumber="110">
<typeStatus id="2674BCC6006C461EA871F9DE855FD64A" box="[774,878,1631,1655]" pageId="9" pageNumber="110" type="holotype">Holotype</typeStatus>
.
</emphasis>
Probably male, in Burmese amber, accession No.
<specimenCode id="A969AA1F006C461EA8F7F9FE85CED6AA" box="[896,1023,1663,1687]" collectionCode="ALDC" pageId="9" pageNumber="110">ALDC0541</specimenCode>
/ALD.Bu.208
</materialsCitation>
</paragraph>
<caption id="ADB052EC006F461DAA58FD728332D371" ID-DOI="http://doi.org/10.5281/zenodo.10905346" ID-Zenodo-Dep="10905346" httpUri="https://zenodo.org/record/10905346/files/figure.png" pageId="10" pageNumber="111" startId="10.[303,342,755,779]" targetBox="[446,1220,192,707]" targetPageId="10" targetType="figure">
<paragraph id="F9700264006F461DAA58FD728332D371" blockId="10.[303,1359,755,844]" pageId="10" pageNumber="111">
Fig. 9.
<taxonomicName id="3ECF79E7006F461DAA0BFD728449D336" authority="Fanti &amp; Damgaard, 2019" authorityName="Fanti &amp; Damgaard" authorityYear="2019" box="[380,632,755,779]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="fabriciusi" status="sp. nov.">
<emphasis id="CBBBDE76006F461DAA0BFD728449D336" box="[380,632,755,779]" italics="true" pageId="10" pageNumber="111">Sanaungulus fabriciusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D088630D006F461DA908FD7584E2D336" box="[639,723,756,779]" pageId="10" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
ALDC0541/ALD.Bu.208 in Burmese amber. A: Holotype, lateral view, bar = 500 μm; B: Holotype, lateral view (more detailed), bar = 500 μm; C: Holotype, ventral view, bar = 500 μm; D: Holotype, detail of head, palp, mandible and pronotum, bar = 200 μm.
</paragraph>
</caption>
<caption id="ADB052EC006F461DAA58FA7487A4D653" ID-DOI="http://doi.org/10.5281/zenodo.10905348" ID-Zenodo-Dep="10905348" httpUri="https://zenodo.org/record/10905348/files/figure.png" pageId="10" pageNumber="111" startId="10.[303,343,1525,1549]" targetBox="[447,1216,953,1466]" targetPageId="10" targetType="figure">
<paragraph id="F9700264006F461DAA58FA7487A4D653" blockId="10.[303,1358,1525,1646]" pageId="10" pageNumber="111">
Fig. 10.
<taxonomicName id="3ECF79E7006F461DAAFFFA7484B2D630" authority="Fanti &amp; Damgaard, 2019" authorityName="Fanti &amp; Damgaard" authorityYear="2019" box="[392,643,1525,1549]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="fabriciusi" status="sp. nov.">
<emphasis id="CBBBDE76006F461DAAFFFA7484B2D630" box="[392,643,1525,1549]" italics="true" pageId="10" pageNumber="111">Sanaungulus fabriciusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D088630D006F461DA9FFFA7784EAD630" box="[648,731,1526,1549]" pageId="10" pageNumber="111" rank="species">sp. nov.</taxonomicNameLabel>
ALDC0541/ALD.Bu.208 in Burmese amber. A: Holotype, detail of mandibles and head, bar = 100 μm; B: Holotype, detail of antennae, bar = 100 μm; C: Holotype, detail of pronotum, bar = 100 μm; D: Holotype, detail of coxae, trochanters and femora, bar = 100 μm.
</paragraph>
</caption>
<caption id="ADB052EC006E461CAB94FC8F8334D35A" ID-DOI="http://doi.org/10.5281/zenodo.10905352" ID-Zenodo-Dep="10905352" httpUri="https://zenodo.org/record/10905352/files/figure.png" pageId="11" pageNumber="112" startId="11.[227,267,782,806]" targetBox="[347,1152,192,730]" targetPageId="11" targetType="figure">
<paragraph id="F9700264006E461CAB94FC8F8334D35A" blockId="11.[227,1287,782,871]" pageId="11" pageNumber="112">
Fig. 11.
<taxonomicName id="3ECF79E7006E461CAA4AFC8F841AD31B" authority="Fanti &amp; Damgaard, 2019" authorityName="Fanti &amp; Damgaard" authorityYear="2019" box="[317,555,782,806]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="strungei" status="sp. nov.">
<emphasis id="CBBBDE76006E461CAA4AFC8F841AD31B" box="[317,555,782,806]" italics="true" pageId="11" pageNumber="112">Sanaungulus strungei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D088630D006E461CA944FC8E84B6D31B" box="[563,647,783,806]" pageId="11" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
ALDC0535/ALD.Bu.207 in Burmese amber. A: Holotype, dorso-lateral view, bar = 1.0 mm; B: Holotype, detail of pronotum and scutellar shield, bar = 200 μm; C: Holotype, detail of head and pronotum, bar = 200 μm; D: Holotype, detail of elytra, bar = 500 μm.
</paragraph>
</caption>
<caption id="ADB052EC006E461CAB94F9C485C0D6A3" ID-DOI="http://doi.org/10.5281/zenodo.10905354" ID-Zenodo-Dep="10905354" httpUri="https://zenodo.org/record/10905354/files/figure.png" pageId="11" pageNumber="112" startId="11.[227,267,1605,1629]" targetBox="[344,1132,1017,1541]" targetPageId="11" targetType="figure">
<paragraph id="F9700264006E461CAB94F9C485C0D6A3" blockId="11.[227,1285,1605,1694]" pageId="11" pageNumber="112">
Fig. 12.
<taxonomicName id="3ECF79E7006E461CAA48F9C4841AD660" authority="Fanti &amp; Damgaard, 2019" authorityName="Fanti &amp; Damgaard" authorityYear="2019" box="[319,555,1605,1629]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="strungei" status="sp. nov.">
<emphasis id="CBBBDE76006E461CAA48F9C4841AD660" box="[319,555,1605,1629]" italics="true" pageId="11" pageNumber="112">Sanaungulus strungei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D088630D006E461CA944F9C784B6D660" box="[563,647,1606,1629]" pageId="11" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
ALDC0535/ALD.Bu.207 in Burmese amber. A: Holotype, dorsal view, bar = 500 μm; B: Holotype, detail of antenna, bar = 400 μm; C: Holotype, detail of legs, bar = 200 μm; D: Holotype, detail of eye and last maxillary palp, bar = 200 μm.
</paragraph>
</caption>
<paragraph id="F97002640069461BAA58FF3E84C4D0CA" blockId="12.[303,813,191,247]" pageId="12" pageNumber="113">
<emphasis id="CBBBDE760069461BAA58FF3E87F0D0EA" bold="true" box="[303,449,191,215]" pageId="12" pageNumber="113">Type locality.</emphasis>
<collectingCountry id="81D842F40069461BAAB2FF41841ED0E5" box="[453,559,192,216]" name="Myanmar" pageId="12" pageNumber="113">Myanmar</collectingCountry>
:
<collectingRegion id="3B0BCC860069461BA940FF3E8488D0EA" box="[567,697,191,215]" country="Myanmar" name="Kachin" pageId="12" pageNumber="113">Kachin state</collectingRegion>
, Myitkyina District, Tanai Township, Hukawng Valley.
</paragraph>
<paragraph id="F97002640069461BAA58FE9E8479D16A" blockId="12.[303,813,287,343]" pageId="12" pageNumber="113">
<emphasis id="CBBBDE760069461BAA58FE9E87F9D10A" bold="true" box="[303,456,287,311]" pageId="12" pageNumber="113">Type horizon.</emphasis>
Lowermost Cenomanian (98.79 ± 0.62 Ma), mid-Cretaceous.
</paragraph>
</subSubSection>
<subSubSection id="B1D551EF0069461BAA58FEFE8758D24A" pageId="12" pageNumber="113" type="diagnosis">
<paragraph id="F97002640069461BAA58FEFE8758D24A" blockId="12.[303,814,383,631]" pageId="12" pageNumber="113">
<emphasis id="CBBBDE760069461BAA58FEFE841ED1AA" bold="true" box="[303,559,383,407]" pageId="12" pageNumber="113">Differential diagnosis.</emphasis>
This species is characterized by head posteriorly to the eyes strongly rounded, and triangular only at the base. No fossil species of the genus
<taxonomicName id="3ECF79E70069461BA941FE5E84F3D1CA" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[566,706,479,503]" class="Insecta" family="Cantharidae" genus="Sanaungulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="CBBBDE760069461BA941FE5E84F3D1CA" box="[566,706,479,503]" italics="true" pageId="12" pageNumber="113">Sanaungulus</emphasis>
</taxonomicName>
show the antennomeres III-IX with long rami.
<taxonomicName id="3ECF79E70069461BA9D9FE7E851CD22A" authorityName="Fanti, Damgaard &amp; Ellenberger" authorityYear="2018" box="[686,813,511,535]" class="Insecta" family="Cantharidae" genus="Burmomiles" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="CBBBDE760069461BA9D9FE7E851CD22A" box="[686,813,511,535]" italics="true" pageId="12" pageNumber="113">Burmomiles</emphasis>
</taxonomicName>
has the the same number of rami but has longer and wider elytra, the shorter legs and roundish head.
</paragraph>
</subSubSection>
<subSubSection id="B1D551EF0069461BAA58FD1E831AD34A" pageId="12" pageNumber="113" type="description">
<paragraph id="F97002640069461BAA58FD1E853CD32A" blockId="12.[303,813,671,791]" pageId="12" pageNumber="113">
<emphasis id="CBBBDE760069461BAA58FD1E878DD28A" bold="true" box="[303,444,671,695]" pageId="12" pageNumber="113">Description.</emphasis>
Adult, winged, probably male on the basis of the last ventrite vaguely triangular and narrower than last tergite. Testaceous-brown with elytra dark brown. Body length: 4.0 mm.
</paragraph>
<paragraph id="F97002640069461BAA58FCBE831AD34A" blockId="12.[303,814,831,1719]" lastBlockId="12.[851,1362,191,887]" pageId="12" pageNumber="113">Head partially covered by head, anteriorly slightly larger than pronotum and posteriorly narrower, strongly rounded behind the eyes and triangular only at the base, without punctation and pubescence. Eyes wide, rounded, convex, strongly prominent. Mandibles extremely elongate, thin, and with a tooth. Maxillary palpi 4-segmented, palpomeres unequal in length, first palpomere slightly elongate, second robust and elongate, third palpomere of intermediate length between first and second palpomere, last palpomere very elongate and securiform and with apical margin rounded. Labial palpi 3-segmented.Antennae 11- segmented, pectinate, relative short, surpassing the half of the elytra and not reaching the half of abdomen, antennal insertion in proximity the eyes; scape very stout, elongate; pedicel filiform about 3.0 times shorter than scape; antennomere III slightly longer and wider than pedicel, equipped with a short and squadrate antennal process; antennomeres IV-VIII very elongate and thin, each equipped with a short and squadrate antennal process inserted in the ventral-apical part; antennomeres IX-XI shorter and more robust than previous ones, antennomeres X-XI without antennal processes; antennomere XI stout, rounded apically, all antennomeres feebly pubescent. Pronotum wide, anterior margin protruding to the head and strongly bordered, posterior margin slightly sinuous and bordered, lateral margins straight and strongly bordered, corners rounded, surface with shallow punctation. Scutellum triangular-shaped, rounded apically. Elytra as large as pronotum, long, with only apical part of the abdomen uncovered, parallel-sided, rounded at apex; surface strongly rugouse with punctation, and with some striae in faint traces. Posterior wings slightly surpassing the elytra and not reaching the last abdominal segments. Abdominal segments transverse, last ventrite vaguely triangular and narrower than last tergite. Legs extremely long; coxae robust and long; trochanters elongate, rounded apically; femora long, slightly curved, more robust than tibiae; tibiae long, cylindrical, thin, longer than femora; tarsal formula 5-5-5, with the first tarsomere elongate, slightly robust and about 2.5 times longer than second; third tarsomere very short; fourth tarsomere bilobed; fifth elongate, flat, slightly enlarged apically; claws simple.</paragraph>
</subSubSection>
<subSubSection id="B1D551EF0069461BA824FC1E85D2D4AA" pageId="12" pageNumber="113" type="etymology">
<paragraph id="F97002640069461BA824FC1E85EED405" blockId="12.[851,1361,927,1080]" pageId="12" pageNumber="113">
<emphasis id="CBBBDE760069461BA824FC1E85FED38A" bold="true" box="[851,975,927,951]" pageId="12" pageNumber="113">Etymology.</emphasis>
In memory of the Danish natural historian and scientist Johan Christian Fabricius (Tønder,
<date id="8D7124A40069461BA8C1FC61826BD3CA" box="[950,1114,991,1016]" pageId="12" pageNumber="113" value="1745-01-07">7 January 1745</date>
- Kiel,
<date id="8D7124A40069461BAFDDFC5E8375D3CA" box="[1194,1348,991,1015]" pageId="12" pageNumber="113" value="1808-03-03">3 March 1808</date>
), one of the most important entomologists of the 18th century.
</paragraph>
<paragraph id="F97002640069461BA824FBDE85D2D4AA" blockId="12.[851,1360,1119,1175]" pageId="12" pageNumber="113">
<emphasis id="CBBBDE760069461BA824FBDE85C4D44A" bold="true" box="[851,1013,1119,1143]" pageId="12" pageNumber="113">Syninclusions.</emphasis>
Debris, air bubbles, stellate hair, and one mite.
</paragraph>
</subSubSection>
<subSubSection id="B1D551EF0069461BA824FB3E82ACD52A" pageId="12" pageNumber="113" type="discussion">
<paragraph id="F97002640069461BA824FB3E82ACD52A" blockId="12.[851,1360,1215,1304]" pageId="12" pageNumber="113">
<emphasis id="CBBBDE760069461BA824FB3E85F2D4EA" bold="true" box="[851,963,1215,1239]" pageId="12" pageNumber="113">Remarks.</emphasis>
The amber measures 20 x 20 x
<quantity id="3E37AF810069461BAE64FB3E8361D4EA" box="[1299,1360,1215,1239]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="113" unit="mm" value="9.0">9 mm</quantity>
and the matrix is transparent. The inclusion is complete and very well visible.
</paragraph>
</subSubSection>
</treatment>
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