160 lines
18 KiB
XML
160 lines
18 KiB
XML
<document id="01213290891AE7B40172B2F15DCE95B1" ID-CLB-Dataset="26847" ID-DOI="10.11646/zootaxa.4604.1.1" ID-GBIF-Dataset="b5902a9d-c38b-462f-ad8a-95693c1822dc" ID-ISSN="1175-5326" ID-Zenodo-Dep="2783674" ID-ZooBank="E6D82300-8DC6-4560-A945-8F29D3B3D501" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1557723545002" checkinUser="plazi" docAuthor="Ruta, Rafał" docDate="2019" docId="71140E6CF27D142373B5EF44579A2961" docLanguage="en" docName="zootaxa.4604.1.1.pdf" docOrigin="Zootaxa 4604 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calvariopsis sculptipenne Ruta 2019, comb. nov." docType="treatment" docVersion="12" lastPageNumber="22" masterDocId="8D2D7614F26914367322EA4454252806" masterDocTitle="Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="21" updateTime="1698728494285" updateUser="ExternalLinkService">
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<mods:titleInfo id="73A0A24F50105D37B49228E70580B1DF">
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<mods:title id="B2FD0781F1F844F230FDDC3936602D14">Calvariopsis gen. nov., a new genus of Neotropical Scirtidae (Coleoptera: Scirtoidea)</mods:title>
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<mods:name id="06554E3D165C033FAA771E1EF6C3A17A" type="personal">
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<mods:namePart id="EECA6565A8557BB74A7D9E5D16B01ADF">Ruta, Rafał</mods:namePart>
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<mods:typeOfResource id="EBF810C65A2A79CE2E357CD23596A807">text</mods:typeOfResource>
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<mods:title id="C2FA9A14838C48DDEACD99E2CAA98CBB">Zootaxa</mods:title>
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<mods:part id="E827259DAED86B71F42D218B9589F11E">
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<mods:date id="5CF594F5E2B9E9DB7A9F7684ED00F8D7">2019</mods:date>
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<mods:number id="2AE11C269571C5E56E76FFF2311CFE93">2019-05-13</mods:number>
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<mods:number id="97A55EB136A742AC2448670F9D4BB6AC">4604</mods:number>
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<mods:detail id="2A340AA19CAE8003082BEA2B7852CBC2" type="issue">
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<mods:start id="60458FF3AF8CEF328D9C154D8F348678">1</mods:start>
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<mods:classification id="C1E3947B17D80B9611EA0A224039087F">journal article</mods:classification>
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<mods:identifier id="0E43B644F2179E95B1BAE680E0662E85" type="CLB-Dataset">26847</mods:identifier>
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<mods:identifier id="0A4151BC2E8F163F15B4339122363A55" type="DOI">10.11646/zootaxa.4604.1.1</mods:identifier>
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<mods:identifier id="4F8AD4C224A428C87154256783731E0E" type="GBIF-Dataset">b5902a9d-c38b-462f-ad8a-95693c1822dc</mods:identifier>
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<mods:identifier id="536125B99F75C4D5C99922EAEB12D9F9" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0E6268AE53027C2900BCFC389779EA6A" type="Zenodo-Dep">2783674</mods:identifier>
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</mods:mods>
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<treatment id="71140E6CF27D142373B5EF44579A2961" ID-GBIF-Taxon="157135232" LSID="urn:lsid:plazi:treatment:71140E6CF27D142373B5EF44579A2961" httpUri="http://treatment.plazi.org/id/71140E6CF27D142373B5EF44579A2961" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
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<subSubSection id="B1A7ECF1F27D142273B5EF4456C22D1D" box="[151,743,1280,1307]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="F902BF7AF27D142273B5EF4456C22D1D" blockId="20.[151,743,1280,1343]" box="[151,743,1280,1307]" pageId="20" pageNumber="21">
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<heading id="A24A0816F27D142273B5EF4456C22D1D" bold="true" box="[151,743,1280,1307]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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<emphasis id="CBC96368F27D142273B5EF4456C22D1D" bold="true" box="[151,743,1280,1307]" pageId="20" pageNumber="21">
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<taxonomicName id="3EBDC4F9F27D142273B5EF44567C2D1C" authority="(Pic, 1931)" authorityName="Ruta" authorityYear="2019" baseAuthorityName="Pic" baseAuthorityYear="1931" box="[151,601,1280,1307]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="sculptipenne" status="comb. nov.">
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<emphasis id="CBC96368F27D142273B5EF4455E82D1D" bold="true" box="[151,461,1280,1307]" italics="true" pageId="20" pageNumber="21">Calvariopsis sculptipenne</emphasis>
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(
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<bibRefCitation id="9D2CC28BF27D142272FFEF4456742D1C" author="Pic, M." box="[477,593,1280,1306]" pageId="20" pageNumber="21" pagination="1 - 36" refId="ref23111" refString="Pic, M. (1931) Nouveautes diverses. Melanges exotico-entomologiques, 58, 1 - 36." type="journal article" year="1931">Pic, 1931</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="D0FADE13F27D14227142EF4556C22D1D" box="[608,743,1281,1307]" pageId="20" pageNumber="21" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B1A7ECF1F27D142273B5EF6055B72D39" box="[151,402,1316,1343]" pageId="20" pageNumber="21" type="description">
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<paragraph id="F902BF7AF27D142273B5EF6055B72D39" blockId="20.[151,743,1280,1343]" box="[151,402,1316,1343]" pageId="20" pageNumber="21">
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(
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<figureCitation id="6186A3FFF27D142273BDEF6054D92D38" box="[159,252,1316,1342]" captionStart="FIGURE 1" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[183,1404,196,1722]" captionTargetId="figure@3.[183,1404,196,1722]" captionTargetPageId="3" captionText="FIGURE 1. Calvariopsis gen. nov., dorsal view. A) C. borowieci sp. nov., male; B) C. borowieci sp. nov., female; C) C. four- gassiense sp. nov., male; D) C. panamense sp. nov., male; E) C. peruviana sp. nov., Dept. Cusco, male; F) C. peruviana sp. nov., Callanga, male; G) C. peruviana sp. nov., Dept. Junín, male; H) C. picta sp. nov., Callanga, male; I) C. pittieri sp. nov., male; J) C. pittieri sp. nov., female; K) C. sculptipenne (Pic, 1931), female; L) C. venezuelense sp. nov., male; M–O) C. yanay- acuense sp. nov., males; P) C. yanayacuense sp. nov., female; Q) C. bituberculata sp. nov., male; R) C. brasiliense (Pic, 1916), male; S) C. brasiliense, female; T) C. guyanense sp. nov., male; U) C. kawense sp. nov., male; V) C. nana sp. nov., male; W) C. saopaulense sp. nov., male; X) C. wittmeri sp. nov., male; Y) C. sp., Bolivia, female. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/2783682/files/figure.png" pageId="20" pageNumber="21">Figs 1K</figureCitation>
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,
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<figureCitation id="6186A3FFF27D1422722AEF6055302D38" box="[264,277,1316,1342]" captionStart="FIGURE 7" captionStartId="9.[151,250,1631,1656]" captionTargetBox="[165,1422,181,1592]" captionTargetId="figure@9.[165,1424,181,1592]" captionTargetPageId="9" captionText="FIGURE 7. Calvariopsis gen. nov., group I, females, apical portion of elytra, SEM images of excitators. A) C. borowieci sp. nov.; B) C. sp., Costa Rica; C–D) C. sculptipenne (Pic); E–F) C. yanayacuense sp. nov." httpUri="https://zenodo.org/record/2783718/files/figure.png" pageId="20" pageNumber="21">7</figureCitation>
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C–D, 16A)
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</paragraph>
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</subSubSection>
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<subSubSection id="B1A7ECF1F27D142273B5EF2D56DC2D99" pageId="20" pageNumber="21" type="reference_group">
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<paragraph id="F902BF7AF27D142273B5EF2D562F2D87" blockId="20.[151,761,1384,1440]" box="[151,522,1384,1409]" pageId="20" pageNumber="21">
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<treatmentCitationGroup id="D9AD9854F27D142273B5EF2D562F2D87" box="[151,522,1384,1409]" pageId="20" pageNumber="21">
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<emphasis id="CBC96368F27D142273B5EF2D548D2D86" box="[151,168,1385,1408]" italics="true" pageId="20" pageNumber="21">=</emphasis>
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<taxonomicName id="3EBDC4F9F27D1422738DEF2D562F2D87" authority="Pic, 1931: 5" authorityName="Pic" authorityPageNumber="5" authorityYear="1931" box="[175,522,1384,1409]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="sculptipennis">
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<emphasis id="CBC96368F27D1422738DEF2D55AF2D87" box="[175,394,1385,1409]" italics="true" pageId="20" pageNumber="21">Cyphon sculptipennis</emphasis>
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<treatmentCitation id="781C996BF27D142272B2EF2C562F2D87" author="Pic, M." box="[400,522,1384,1409]" page="5" pageId="20" pageNumber="21" year="1931">
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<bibRefCitation id="9D2CC28BF27D142272B2EF2C55D62D87" author="Pic, M." box="[400,499,1384,1409]" pageId="20" pageNumber="21" pagination="1 - 36" refId="ref23111" refString="Pic, M. (1931) Nouveautes diverses. Melanges exotico-entomologiques, 58, 1 - 36." type="journal article" year="1931">Pic, 1931</bibRefCitation>
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: 5
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</treatmentCitation>
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</taxonomicName>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="F902BF7AF27D142273B5EFCC56DC2D99" blockId="20.[151,761,1384,1440]" box="[151,761,1415,1440]" pageId="20" pageNumber="21">
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<emphasis id="CBC96368F27D142273B5EFCC548D2D99" box="[151,168,1416,1439]" italics="true" pageId="20" pageNumber="21">=</emphasis>
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<taxonomicName id="3EBDC4F9F27D1422738DEFC356192D99" authority="(Pic, 1931)" authorityName="Pic" authorityYear="1931" baseAuthorityName="Pic" baseAuthorityYear="1931" box="[175,572,1415,1440]" class="Insecta" family="Scirtidae" genus="Contacyphon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="sculptipennis">
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<emphasis id="CBC96368F27D1422738DEFC355E72D99" box="[175,450,1415,1439]" italics="true" pageId="20" pageNumber="21">Contacyphon sculptipennis</emphasis>
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(
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<bibRefCitation id="9D2CC28BF27D142272EDEFC356102D99" author="Pic, M." box="[463,565,1415,1440]" pageId="20" pageNumber="21" pagination="1 - 36" refId="ref23111" refString="Pic, M. (1931) Nouveautes diverses. Melanges exotico-entomologiques, 58, 1 - 36." type="journal article" year="1931">Pic, 1931</bibRefCitation>
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)
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</taxonomicName>
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: Zwick
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<emphasis id="CBC96368F27D142271AFEFCC56E52D99" box="[653,704,1415,1439]" italics="true" pageId="20" pageNumber="21">et al.</emphasis>
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2013
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</paragraph>
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</subSubSection>
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||
<subSubSection id="B1A7ECF1F27D142273B5EF8A506C2E0A" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="F902BF7AF27D142273B5EF8A506C2E0A" blockId="20.[151,1437,1485,2016]" pageId="20" pageNumber="21">
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<emphasis id="CBC96368F27D142273B5EF8A55622DEE" bold="true" box="[151,327,1485,1512]" pageId="20" pageNumber="21">Type material.</emphasis>
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<materialsCitation id="49D5B527F27D14227272EF8A506C2E0A" ID-GBIF-Occurrence="2239706645" collectionCode="MNHN" pageId="20" pageNumber="21" specimenCount="2" typeStatus="holotype">
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<typeStatus id="260601D8F27D14227272EF8A559E2DEE" box="[336,443,1486,1512]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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, female (
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<collectionCode id="9FAC27BFF27D14227104EF8A56A62DEE" box="[550,643,1486,1512]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="20" pageNumber="21">MNHN</collectionCode>
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): “Corumba \ Matt Grosso” [printed label]; “sculptipennis \ n sp” [in Pic’s hand]; “
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<typeStatus id="260601D8F27D1422720FEFB655F22E0A" box="[301,471,1522,1548]" pageId="20" pageNumber="21" type="holotype">HOLOTYPUS</typeStatus>
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\
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<taxonomicName id="3EBDC4F9F27D142272CEEFB656FB2E0A" authorityName="Pic" authorityYear="1931" box="[492,734,1522,1548]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="sculptipennis">
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<emphasis id="CBC96368F27D142272CEEFB656FB2E0A" box="[492,734,1522,1548]" italics="true" pageId="20" pageNumber="21">Cyphon sculptipennis</emphasis>
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</taxonomicName>
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\
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<bibRefCitation id="9D2CC28BF27D142271D1EFB6577A2E0A" author="Pic, M." box="[755,863,1522,1548]" pageId="20" pageNumber="21" pagination="1 - 36" refId="ref23111" refString="Pic, M. (1931) Nouveautes diverses. Melanges exotico-entomologiques, 58, 1 - 36." type="journal article" year="1931">Pic, 1931</bibRefCitation>
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” [red label, printed].
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="B1A7ECF1F27D142373E5EC51570328D1" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
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<paragraph id="F902BF7AF27D142273E5EC5157062FBA" blockId="20.[151,1437,1485,2016]" pageId="20" pageNumber="21">
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<emphasis id="CBC96368F27D142273E5EC5155502E36" bold="true" box="[199,373,1557,1584]" pageId="20" pageNumber="21">Redescription.</emphasis>
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Female. Body oblong oval, convex, covered with yellowish suberect setae. Body uniformly dark brown, venter brown. Head very broad, 1.75× wider than width of interocular space, with distinct but fine punctation, punctures small, granulate, separated by ca. 2.0–3.0× diameter of a puncture, eyes moderately large, not protuberant. Pronotum transverse, widest at its base. Disc of pronotum with fine, granulate punctation, punctures separated by 1.5–2.0× diameter of a puncture. Posterior margin of pronotum subtly bisinuate. Anterior angles protruding anteriorly, subtriangular, posterior angles obtuse. Scutellar shield subtriangular, with very fine, granulate punctation. Angle between pronotum and elytra subtly marked in dorsal outline. Elytra oblong, without traces of longitudinal ridges, broadest in the middle of their length. Sides slightly curved in apical 1/2, regularly converging to apex in posterior 1/3. Humeri well marked. Punctation stronger than on pronotum, punctures elongate, distance between punctures ca. 0.7–1.0× diameter of a puncture. Apical portion of elytra depressed in adsutural portion, with two deep cavities on each elytron (
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<figureCitation id="6186A3FFF27D1422713CED3A56402F9E" box="[542,613,1918,1944]" captionStart="FIGURE 7" captionStartId="9.[151,250,1631,1656]" captionTargetBox="[165,1422,181,1592]" captionTargetId="figure@9.[165,1424,181,1592]" captionTargetPageId="9" captionText="FIGURE 7. Calvariopsis gen. nov., group I, females, apical portion of elytra, SEM images of excitators. A) C. borowieci sp. nov.; B) C. sp., Costa Rica; C–D) C. sculptipenne (Pic); E–F) C. yanayacuense sp. nov." httpUri="https://zenodo.org/record/2783718/files/figure.png" pageId="20" pageNumber="21">Figs 7</figureCitation>
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C–D). Sternite VIII elongate, Y-shaped, with single apodeme. Remaining structures of genital tract missing in the studied specimen.
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</paragraph>
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<paragraph id="F902BF7AF27D142273E5ED8255A02FE6" blockId="20.[151,1437,1485,2016]" box="[199,389,1990,2016]" pageId="20" pageNumber="21">Male. Unknown.</paragraph>
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<paragraph id="F902BF7AF27C142373E5EADD570328D1" blockId="21.[151,1437,152,359]" pageId="21" pageNumber="22">
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<emphasis id="CBC96368F27C142373E5EADD562528B5" bold="true" box="[199,512,152,179]" pageId="21" pageNumber="22">Measurements and ratios.</emphasis>
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Female (n = 1) TL
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<quantity id="3E45129FF27C142371C1EADD576E28B5" box="[739,843,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="2.83" pageId="21" pageNumber="22" unit="mm" value="2.83">2.83 mm</quantity>
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, PL
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<quantity id="3E45129FF27C142370A2EADC57C228B5" box="[896,999,152,179]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="21" pageNumber="22" unit="mm" value="0.57">0.57 mm</quantity>
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, PW
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<quantity id="3E45129FF27C14237704EADD50A828B5" box="[1062,1165,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="21" pageNumber="22" unit="mm" value="1.23">1.23 mm</quantity>
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, EL
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<quantity id="3E45129FF27C142377E1EADD510E28B5" box="[1219,1323,153,179]" metricMagnitude="-3" metricUnit="m" metricValue="2.34" pageId="21" pageNumber="22" unit="mm" value="2.34">2.34 mm</quantity>
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, EW
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<quantity id="3E45129FF27C14237649EADD54E028D1" metricMagnitude="-3" metricUnit="m" metricValue="1.83" pageId="21" pageNumber="22" unit="mm" value="1.83">1.83 mm</quantity>
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, TL/EW 1.55, PW/PL 2.16, EL/EW 1.28, EL/PL 4.11.
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</paragraph>
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</subSubSection>
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<subSubSection id="B1A7ECF1F27C142373E5EAA454D22918" pageId="21" pageNumber="22" type="distribution">
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||
<paragraph id="F902BF7AF27C142373E5EAA454D22918" blockId="21.[151,1437,152,359]" pageId="21" pageNumber="22">
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||
<emphasis id="CBC96368F27C142373E5EAA4554428FD" bold="true" box="[199,353,224,251]" pageId="21" pageNumber="22">Distribution.</emphasis>
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Known only from the
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<typeStatus id="260601D8F27C14237149EAA556BE28FD" box="[619,667,225,251]" pageId="21" pageNumber="22">type</typeStatus>
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locality in
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<collectingCountry id="81AAFFEAF27C14237002EAA4574228FC" box="[800,871,224,250]" name="Brazil" pageId="21" pageNumber="22">Brazil</collectingCountry>
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, close to the border with
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<collectingCountry id="81AAFFEAF27C142377ADEAA450C128FC" box="[1167,1252,224,250]" name="Bolivia" pageId="21" pageNumber="22">Bolivia</collectingCountry>
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, in the Pantanal wetland.
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</paragraph>
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</subSubSection>
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<subSubSection id="B1A7ECF1F27C142373E5EB6D579A2961" pageId="21" pageNumber="22" type="discussion">
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||
<paragraph id="F902BF7AF27C142373E5EB6D579A2961" blockId="21.[151,1437,152,359]" pageId="21" pageNumber="22">
|
||
<emphasis id="CBC96368F27C142373E5EB6D551E2945" bold="true" box="[199,315,297,323]" pageId="21" pageNumber="22">Remarks.</emphasis>
|
||
Presence of well developed, foveate excitators in apical portions of elytra, and TL larger than
|
||
<quantity id="3E45129FF27C1423766AEB6D51B82945" box="[1352,1437,297,323]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="21" pageNumber="22" unit="mm" value="2.4">2.4 mm</quantity>
|
||
support inclusion of this species in the group I of
|
||
<taxonomicName id="3EBDC4F9F27C142371E0EB0857742960" authority="Ruta, 2019" authorityName="Ruta" authorityYear="2019" box="[706,849,332,358]" class="Insecta" family="Scirtidae" genus="Calvariopsis" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="CBC96368F27C142371E0EB0857742960" box="[706,849,332,358]" italics="true" pageId="21" pageNumber="22">Calvariopsis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D0FADE13F27C1423707BEB09579A2961" box="[857,959,333,359]" pageId="21" pageNumber="22" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |