245 lines
28 KiB
XML
245 lines
28 KiB
XML
<document id="2F990C436F519EF095F9191C2BA5AC34" ID-CLB-Dataset="31262" ID-DOI="http://dx.doi.org/10.3897/zookeys.717.21885" ID-GBIF-Dataset="7140b994-d67e-44a7-a4e1-daa7216c6b46" ID-PMC="PMC5784208" ID-Pensoft-Pub="1313-2970-717-1" ID-PubMed="29391848" ID-ZooBank="C19B43B1B3214CB1B1B2A246CEAC56BC" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-717-1" ModsDocOrigin="ZooKeys 717" ModsDocTitle="Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda)" checkinTime="1512083542653" checkinUser="pensoft" docAuthor="Korshunova, Tatiana, Martynov, Alexander, Bakken, Torkild, Evertsen, Jussi, Fletcher, Karin, Mudianta, I Wayan, Saito, Hiroshi, Lundin, Kennet, Michael Schroedl, & Picton, Bernard" docDate="2017" docId="3FF942D0187EB091DC2D946D99F89B89" docLanguage="en" docName="ZooKeys 717: 1-139" docOrigin="ZooKeys 717" docSource="http://dx.doi.org/10.3897/zookeys.717.21885" docTitle="Flabellinidae Bergh 1889" docType="treatment" docVersion="8" lastPageNumber="40" masterDocId="FFED910FFFF5FFC82005844DFFF1FFA8" masterDocTitle="Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda)" masterLastPageNumber="139" masterPageNumber="1" pageNumber="39" updateTime="1732584470242" updateUser="ExternalLinkService">
|
||
<mods:mods id="FCB07BFDEB660D6DC00B06E59F012836" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="8E6C215F1E534373E7114A481F5BCE43">
|
||
<mods:title id="826131E72AE912C4D0F156A3EBBCD315">Polyphyly of the traditional family Flabellinidae affects a major group of Nudibranchia: aeolidacean taxonomic reassessment with descriptions of several new families, genera, and species (Mollusca, Gastropoda)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="0F7A2C8DE6A5EEB145429DA07C659EB0" type="personal">
|
||
<mods:role id="BBBC6524DDE1D2027A41E77B5350C4A2">
|
||
<mods:roleTerm id="334C9FD33D63B719028AB2258E9F240A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C7D738F308EA80C3C3F54E1B5A94597C">Korshunova, Tatiana</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="13E2D4D9B471E351781DA333B2F731DB" type="personal">
|
||
<mods:role id="79A0BA556C1D76C0F769E2B83E0CBABF">
|
||
<mods:roleTerm id="B953C9F315339C2017AEEF7D6D7AE20C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A75D7B4A587B498437A9EFE67BC553CA">Martynov, Alexander</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="1E7768D7D4E82122A90050F54F3D6A5A" type="personal">
|
||
<mods:role id="A6BEBA43EF7B716E0A817CDCC58B92A6">
|
||
<mods:roleTerm id="EE922C0793A6DD584CBEAAB26D2B7AE1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1E648ACA9DB775048B5978333C90BFF4">Bakken, Torkild</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="450237F696FEF8881D8E71C468B992B6" type="personal">
|
||
<mods:role id="95F5B45BE1C3D7A340CC4811176FFB79">
|
||
<mods:roleTerm id="F7F83CD9713E426F887335FD4B7EF79F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6DB13C72B0214879CE0704CD1396AA37">Evertsen, Jussi</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="3726B631466294CEB5692FAFB6AD3DD5" type="personal">
|
||
<mods:role id="268DD0CE0562CEDDF8EBE30C5310E320">
|
||
<mods:roleTerm id="87845D045F9B0945994DF75A27738CDB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="23CD86E87763F0EFBDF7924387B9A754">Fletcher, Karin</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="650AE1D3FE85A0028A04CC58513D0E74" type="personal">
|
||
<mods:role id="4F2AA59BD6B0119A6AEC94BE8CD4F2FF">
|
||
<mods:roleTerm id="13774338A0FD344B6DE60F70CEDC349D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="48766E06311182DEF944D610D000F3C3">Mudianta, I Wayan</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="67145646269ADBCFDF74D666AB92C3A6" type="personal">
|
||
<mods:role id="09C3189AC62F0974215D04FB2A9AF1B5">
|
||
<mods:roleTerm id="8E85558C88CB0DDBDE45BD7C8501CD5B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="29147B63BF6BCEC6DF7475CA0C78B7F5">Saito, Hiroshi</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="26958BDC16982F55E6445BE746FC0A44" type="personal">
|
||
<mods:role id="208D0E82BBCDFE3A635C9223191689D2">
|
||
<mods:roleTerm id="21096FBE650D6094862C003AC2423DF5">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="462843DC811F8ACCA4031B489817F188">Lundin, Kennet</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="477CC498653DAA07CE4799C09F149372" type="personal">
|
||
<mods:role id="5BF166F96328DA090FADC3CE5A36F591">
|
||
<mods:roleTerm id="6334968850F6E4B13072F3D06CB81EAD">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E1C07A20320B514E11FEE5CCF39A8185">Michael Schroedl,</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="530925F85F0FFB7FAA68A812CEDCF011" type="personal">
|
||
<mods:role id="917EC59AC0CE026968C65A756C840F95">
|
||
<mods:roleTerm id="984317E070606266534C6CB809D96723">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="AEE80E3854FFF482808DB82FC96EDF97">Picton, Bernard</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="23817FC0265EE060A57CA3AA6EE4B13C">text</mods:typeOfResource>
|
||
<mods:relatedItem id="13DAA7FF9081C66351E87C1B0B8548A2" type="host">
|
||
<mods:titleInfo id="6AAC984A145309371DA34FAA6917A14D">
|
||
<mods:title id="221CDA13A67EA70C788CE40D8A8BF247">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="10DC3FC3FC4704280816D4662C6A4DC2">
|
||
<mods:date id="03B448CEEC77E19F7D2B7DFCC91E0254">2017</mods:date>
|
||
<mods:detail id="A9558D1B48827C559569268547852171" type="volume">
|
||
<mods:number id="E2BA5B0F1881799D8B15B1E8245D8C35">717</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="5E7812AF32B6D8108F5850A0997E4CA1" unit="page">
|
||
<mods:start id="01B6CFBEBD9D947B0A95CB2EFFF7CEDE">1</mods:start>
|
||
<mods:end id="85C3A3C949D013F646E672396C76C322">139</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="DD76F881D73BAD87D91B46574F9D0E53">
|
||
<mods:url id="69CDB1D34767DEE294C79D9BB620347E">http://dx.doi.org/10.3897/zookeys.717.21885</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="C1784EA674DAD125182768549ED815BE">journal article</mods:classification>
|
||
<mods:identifier id="D34D963365D2D3AF050C39B5BF23AB36" type="DOI">http://dx.doi.org/10.3897/zookeys.717.21885</mods:identifier>
|
||
<mods:identifier id="77C95743D6D65C5CCBF66433F83DAF1D" type="Pensoft-Pub">1313-2970-717-1</mods:identifier>
|
||
<mods:identifier id="1A1D5DECA7E6EDD48716CCE8B268C01E" type="ZooBank">C19B43B1B3214CB1B1B2A246CEAC56BC</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="3FF942D0187EB091DC2D946D99F89B89" ID-GBIF-Taxon="137581536" LSID="urn:lsid:plazi:treatment:3FF942D0187EB091DC2D946D99F89B89" httpUri="http://treatment.plazi.org/id/3FF942D0187EB091DC2D946D99F89B89" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" scope_class="Gastropoda" scope_order="Nudibranchia" scope_phylum="Mollusca">
|
||
<subSubSection id="1B352A2819D03652D37996DB6DF62B59" pageId="38" pageNumber="39" type="nomenclature">
|
||
<paragraph id="51D5DB44ABF0242AF41D53AC45F79965" pageId="38" pageNumber="39">
|
||
Family
|
||
<taxonomicName id="732766B0A68A9AB747DF5800555FE980" ID-CoL="7NHLN" authority="Bergh, 1889, restricted" class="Gastropoda" family="Flabellinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="family">Flabellinidae Bergh, 1889, restricted</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="736D22CE1F952D45635E87B5AAA3032D" pageId="38" pageNumber="39" type="diagnosis">
|
||
<paragraph id="29B9FF859723BDA3750D800B9DCD9408" pageId="38" pageNumber="39">Diagnosis.</paragraph>
|
||
<paragraph id="7CD0B780DB37E6EF9664DFBE73FA6C73" pageId="38" pageNumber="39">Body commonly narrow. Notal edge discontinuous or fully reduced. Cerata in separate clusters, on elevations or distinct stalks. Rhinophores smooth, annulated or papillated. Anus mixed (pleuroproctic in higher acleioproctic position) or pleuroproctic under the reduced notal edge. Distinct oral glands present, commonly penetrate below anterior cerata. Radula formula 1.1.1. Rachidian teeth usually compressed by adjacent lateral denticles. Lateral teeth with attenuated process basally, usually denticulated, rarely smooth. Number and position of receptaculum seminis variable: two separate ones, or double proximal and single distal, or double distal one, in few cases proximal receptaculum not evident. Vas deferens usually long, with indistinct prostate. External permanent penial collar absent. Penis usually elongated conical, narrow, always internal unarmed.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A3CB750341C5556849D9E2BA3AD7FA76" pageId="38" pageNumber="39" type="genera included">
|
||
<paragraph id="FD045BDD3A9CB76EFD2B5FF32F675A4E" pageId="38" pageNumber="39">Genera included.</paragraph>
|
||
<paragraph id="49F1E0C07680F2DD728398185D07DEC7" pageId="38" pageNumber="39">
|
||
<taxonomicName id="E4DEE089515BA59E09FB4EB24E0D973F" class="Gastropoda" family="Flabellinidae" genus="Calmella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calmella" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">Calmella</taxonomicName>
|
||
Eliot, 1910,
|
||
<taxonomicName id="A909008CA6237AA53890DBBD652F8CB9" class="Gastropoda" family="Flabellinidae" genus="Carronella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carronella" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">Carronella</taxonomicName>
|
||
gen. n.,
|
||
<taxonomicName id="E05FD8371CBA7D7D5DB5CA639D6E25ED" class="Gastropoda" family="Flabellinidae" genus="Coryphellina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coryphellina" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">Coryphellina</taxonomicName>
|
||
<normalizedToken id="D3EC25E7340AE1E19270ACACFCBC7F8E" originalValue="O’Donoghue">O'Donoghue</normalizedToken>
|
||
, 1929,
|
||
<taxonomicName id="1743191364F4BB7ACBDB78AC46FFDF95" class="Gastropoda" family="Flabellinidae" genus="Edmundsella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Edmundsella" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">Edmundsella</taxonomicName>
|
||
gen. n.,
|
||
<taxonomicName id="57FD7AC76EAD5DDCCCB6C38312E584BA" class="Gastropoda" family="Flabellinidae" genus="Flabellina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Flabellina" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">Flabellina</taxonomicName>
|
||
Gray, 1833 in Griffith and Pidgeon, 1833-1834,
|
||
<taxonomicName id="E812E90562C68C4DEFD847DE712D1883" class="Gastropoda" family="Flabellinidae" genus="Paraflabellina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraflabellina" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">Paraflabellina</taxonomicName>
|
||
gen. n.,
|
||
<taxonomicName id="BA9FEE71A11B5C79AE4680E9942A6CF9" class="Gastropoda" family="Piseinotecidae" genus="Piseinotecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Piseinotecus" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="genus">Piseinotecus</taxonomicName>
|
||
(?) Marcus, 1955.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="17B0DC90437E4E8CE318433198A8E6D8" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="remarks">
|
||
<paragraph id="EF9E8C809A6EEA80B338F382D4E51C0B" pageId="38" pageNumber="39">Remarks.</paragraph>
|
||
<paragraph id="F8A130AAA5B5D6C4990D14CE532DEE54" pageId="38" pageNumber="39">
|
||
The name
|
||
<taxonomicName id="48930F3EE65D77D4752F136BABA38D49" class="Gastropoda" family="Flabellinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="family">Flabellinidae</taxonomicName>
|
||
has been used for a long time to encompass all the diversity of aeolidaceans with a triseriate radula and without a clearly defined supplementary (or penial) gland in the male part of reproductive system. However, according to our present molecular phylogenetic analysis (Figs 1, 2), the former
|
||
<taxonomicName id="8FCE455C3FAA1E9E613951E8EB2455ED" class="Gastropoda" family="Flabellinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="family">Flabellinidae</taxonomicName>
|
||
form several distinct clades. Remarkably, some of them appear to be closely related to traditional
|
||
<taxonomicName id="88C0326871A9734E61916BF176E7729F" class="Gastropoda" family="Tergipedidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="family">Tergipedidae</taxonomicName>
|
||
or
|
||
<taxonomicName id="D4C05F4A04EA7E229518A8FC63CB78A3" class="Gastropoda" family="Facelinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="family">Facelinidae</taxonomicName>
|
||
thus making any notion that traditional
|
||
<taxonomicName id="EDE441AECDB3EF370B6D05A41BCC8122" class="Gastropoda" family="Flabellinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="38" pageNumber="39" phylum="Mollusca" rank="family">Flabellinidae</taxonomicName>
|
||
can be saved as a monophyletic unit highly unlikely.
|
||
</paragraph>
|
||
<paragraph id="96DFA0723F781F8E9609FB59E6BE8177" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
|
||
Therefore, following the phylogenetic pattern discovered from the DNA analysis (Fig. 1) and consolidated morphological data (Fig. 2) we consistently utilise
|
||
<pageBreakToken id="6CFBE7E716E2BF810E2B96C9556A7715" pageId="39" pageNumber="40" start="start">a</pageBreakToken>
|
||
differential approach, and propose several family level groups within traditional "
|
||
<taxonomicName id="3FD87508055A5F182E58650358039079" class="Gastropoda" family="Flabellinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="family">Flabellinidae</taxonomicName>
|
||
". The super-cluster of the majority of traditional
|
||
<taxonomicName id="E044C9E85F9C0FC270D200B0214798BF" class="Gastropoda" family="Flabellinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="family">Flabellinidae</taxonomicName>
|
||
holds a central position within this group and is apparently monophyletic (Fig. 2). It is, however, comprised of three clearly defined clades, which roughly correspond to the traditional genera
|
||
<taxonomicName id="C5C015F6C61E0D7AAD190E3ECA3DF7F3" class="Gastropoda" family="Paracoryphellidae" genus="Chlamylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chlamylla" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">Chlamylla</taxonomicName>
|
||
,
|
||
<taxonomicName id="D7EE94EC4FD9E33F0B0D8B04CD558B15" class="Gastropoda" family="Coryphellidae" genus="Coryphella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coryphella" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">Coryphella</taxonomicName>
|
||
, and
|
||
<taxonomicName id="D14A1CC65F584D2644EA8A7B9976A7C8" class="Gastropoda" family="Flabellinidae" genus="Flabellina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Flabellina" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">Flabellina</taxonomicName>
|
||
. However, a decision to maintain these phylogenetic superclusters just at generic levels will considerably obscure the complicated evolutionary patterns discovered from the molecular analysis (Fig. 1) which must be consistently reflected in a classification. Importantly, every large phylogenetic cluster has clear morphological trends. For example, the reinstated family
|
||
<taxonomicName id="ED07F0281431876676612BEBF8BB829B" class="Gastropoda" family="Paracoryphellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="family">Paracoryphellidae</taxonomicName>
|
||
is characterised by the invariable presence of a wide body and continuous notal edge, elongated penis, and presence of complicated external penial collars or external permanent penis in two genera, and also invariably an uncompressed cusp of the rachidian teeth. The reinstated family
|
||
<taxonomicName id="89EA0DCE0181EA6280B3AD125C2B8255" class="Gastropoda" family="Coryphellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="family">Coryphellidae</taxonomicName>
|
||
contains mostly taxa with a narrow body and discontinuous notal edge, but importantly there are several genera not directly related to other coryphellids (e.g.,
|
||
<taxonomicName id="42227CF2A96453B483CA19CF8C72003B" class="Gastropoda" family="Coryphellidae" genus="Borealia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Borealia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">Borealia</taxonomicName>
|
||
,
|
||
<taxonomicName id="F82116B43D6FDC36EC5118CE3F9FF55F" class="Gastropoda" family="Coryphellidae" genus="Himatina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Himatina" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">Himatina</taxonomicName>
|
||
,
|
||
<taxonomicName id="587A4C8A242D5B0996D2552D13D914E5" class="Gastropoda" family="Coryphellidae" genus="Gulenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gulenia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">Gulenia</taxonomicName>
|
||
,
|
||
<taxonomicName id="50B76BA5C39535A687CE052D6F1CD352" class="Gastropoda" family="Coryphellidae" genus="Itaxia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Itaxia" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">Itaxia</taxonomicName>
|
||
, see above) that possess a continuous notal edge, which are all basal to various clades of narrow-bodied coryphellids. Internally most of the genera of the family
|
||
<taxonomicName id="FB35DD5C1086FCE2B8D70DD5C3F3DAD1" class="Gastropoda" family="Coryphellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="family">Coryphellidae</taxonomicName>
|
||
possess a non-compressed cusp of the rachidian teeth, short vas deferens, and broad penis always without an external penial collar. In turn, the considerably restricted family
|
||
<taxonomicName id="7FFDE49A634256F97AAA43A0BB306C81" class="Gastropoda" family="Flabellinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="family">Flabellinidae</taxonomicName>
|
||
s. str. includes taxa which usually have various ceratal elevations, up to highly ramified stalks in the genus
|
||
<taxonomicName id="6AA1584110E5B69BEC474829C2A92EBE" class="Gastropoda" family="Flabellinidae" genus="Flabellina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Flabellina" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="genus">Flabellina</taxonomicName>
|
||
s. str. The family
|
||
<taxonomicName id="F510DA9A49E35EF3071184CE80EFE868" class="Gastropoda" family="Flabellinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="family">Flabellinidae</taxonomicName>
|
||
in the restricted sense also includes such clear morphological trends as an almost invariably compressed cusp of the rachidian teeth, lateral teeth with attenuated bases, commonly large oral glands and usually a relatively long vas deferens and elongated penis (see below).
|
||
</paragraph>
|
||
<paragraph id="C7760DBBBD7F34B1BFE2C261AE0AA933" pageId="39" pageNumber="40">
|
||
The minimum uncorrected p-distances of the mitochondrial barcode marker (COI) between the species of family
|
||
<taxonomicName id="4602F579A3F5B179B24645ABF51E93E5" class="Gastropoda" family="Flabellinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="family">Flabellinidae</taxonomicName>
|
||
are given in Table 2. The minimum uncorrected p-distances between all genera range from 13.1 to 20.2%. The lowest distances occurred between
|
||
<taxonomicName id="8988890334F861B187807510A85C6A2C" class="Gastropoda" family="Flabellinidae" genus="Coryphellina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coryphellina rubrolineata" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="rubrolineata">Coryphellina rubrolineata</taxonomicName>
|
||
and
|
||
<taxonomicName id="7E741684B3CDBDE7EE378D6BD7D4318D" class="Gastropoda" family="Flabellinidae" genus="Edmundsella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Edmundsella pedata" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="pedata">Edmundsella pedata</taxonomicName>
|
||
(13.1%). Such considerable molecular divergence between taxa of the family
|
||
<taxonomicName id="FAEDE30184006A7E366357B4838D3049" class="Gastropoda" family="Flabellinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="family">Flabellinidae</taxonomicName>
|
||
supports the reliability of the multi-genera approach consistently applied in this study.
|
||
</paragraph>
|
||
<caption id="A8034C077508330ADC2BD0204B2367EC" pageId="39" pageNumber="40">
|
||
<paragraph id="41196611CD52EC9252EEE2F21D8B6EDF" pageId="39" pageNumber="40">
|
||
Table 2. Minimum uncorrected p-distances (%) between representatives of the genera of the family
|
||
<taxonomicName id="DF28661381B4E8BCC8C2184EF9C15BF7" class="Gastropoda" family="Flabellinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="family">Flabellinidae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EBF1E2904E017168E091DCE7A1050EC6" pageId="39" pageNumber="40">
|
||
<table id="33A24A107D1C899ADC019449F6613110" pageId="39" pageNumber="40">
|
||
<tr id="D29CD67286D493BA56AEB7A7F63D44B2" pageId="39" pageNumber="40">
|
||
<th id="609B140363EEFAAB911504B89BE5DEDC" colspan="1" pageId="39" pageNumber="40" rowspan="1">
|
||
<taxonomicName id="34CA21B4F75A0B69D227BD2126A9BF6F" class="Gastropoda" family="Flabellinidae" genus="Paraflabellina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraflabellina funeka" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="funeka">Paraflabellina funeka</taxonomicName>
|
||
</th>
|
||
<th id="1D0E0AA66B65AC4B95D21A44F4AA5588" colspan="1" pageId="39" pageNumber="40" rowspan="1">
|
||
<taxonomicName id="565B9C119C708F4F4381A10B5CFA9BC9" class="Gastropoda" family="Flabellinidae" genus="Calmella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calmella cavolini" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="cavolini">Calmella cavolini</taxonomicName>
|
||
</th>
|
||
<th id="57D316AC5F5E1829B85FD3687135E30B" colspan="1" pageId="39" pageNumber="40" rowspan="1">
|
||
<taxonomicName id="E9932846C347F4FDE8DE3DCC50738024" class="Gastropoda" family="Flabellinidae" genus="Carronella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carronella pellucida" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="pellucida">Carronella pellucida</taxonomicName>
|
||
</th>
|
||
<th id="65E96D19DB5A775EE0029D3DDD890592" colspan="1" pageId="39" pageNumber="40" rowspan="1">
|
||
<taxonomicName id="B713C03E7250E9CEB0F076BD702DF2CA" class="Gastropoda" family="Flabellinidae" genus="Flabellina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Flabellina affinis" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="affinis">Flabellina affinis</taxonomicName>
|
||
</th>
|
||
<th id="4F5AF975FF54A079858A0152F474A76A" colspan="1" pageId="39" pageNumber="40" rowspan="1">
|
||
<taxonomicName id="BB2A8F0BF03C74E46A727A432CEB21A7" class="Gastropoda" family="Flabellinidae" genus="Coryphellina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coryphellina rubrolineata" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="rubrolineata">Coryphellina rubrolineata</taxonomicName>
|
||
</th>
|
||
<th id="1A3740A44DE88CEF95A9CD2EB82A512E" colspan="1" pageId="39" pageNumber="40" rowspan="1">
|
||
<taxonomicName id="FFAA33C8556AD51C4649425F25F3F603" class="Gastropoda" family="Flabellinidae" genus="Edmundsella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Edmundsella pedata" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="pedata">Edmundsella pedata</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="C21787630A2647579A37DBE80D0A8B49" pageId="39" pageNumber="40">
|
||
<td id="ED6A7613ED503A84F17978B5FB18E7A3" colspan="1" pageId="39" pageNumber="40" rowspan="1">
|
||
<taxonomicName id="6C1FA21D8706C1D82A925145C2219188" class="Gastropoda" family="Flabellinidae" genus="Paraflabellina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paraflabellina funeka" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="funeka">Paraflabellina funeka</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr id="74FCFD4C3E251380E37762D0DF9F5FCB" pageId="39" pageNumber="40">
|
||
<td id="7C2679B8B28953CFB80569F0A3F8A4CE" colspan="1" pageId="39" pageNumber="40" rowspan="1">
|
||
<taxonomicName id="A58AE14D44DC5F920FA80B1AFE2EA59C" class="Gastropoda" family="Flabellinidae" genus="Calmella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calmella cavolini" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="cavolini">Calmella cavolini</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr id="EEB4ED77D33BE152AA40E2325ED12D8E" pageId="39" pageNumber="40">
|
||
<td id="5235429FA8EA95340043475EAA12FB45" colspan="1" pageId="39" pageNumber="40" rowspan="1">
|
||
<taxonomicName id="567F2ADC32E27952E01E4DB3792C5616" class="Gastropoda" family="Flabellinidae" genus="Carronella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carronella pellucida" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="pellucida">Carronella pellucida</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr id="0BAF35AA64FB5B8D64281B28134A3DB6" pageId="39" pageNumber="40">
|
||
<td id="EDE136E72731530127F36900E56D78BA" colspan="1" pageId="39" pageNumber="40" rowspan="1">
|
||
<taxonomicName id="78CA29CA0771CFE994AB27730830983E" class="Gastropoda" family="Flabellinidae" genus="Flabellina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Flabellina affinis" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="affinis">Flabellina affinis</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr id="EC408521B7D22356C299AB81DD15CC80" pageId="39" pageNumber="40">
|
||
<td id="B8BD1C3494CB14AD6ABD0559288A9018" colspan="1" pageId="39" pageNumber="40" rowspan="1">
|
||
<taxonomicName id="79765B60473B9011850CCE3E5A363F2C" class="Gastropoda" family="Flabellinidae" genus="Coryphellina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coryphellina rubrolineata" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="rubrolineata">Coryphellina rubrolineata</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr id="415D23E2FC6C099C15E130BF20657667" pageId="39" pageNumber="40">
|
||
<td id="3946A88B7C592C01629B4A6A7CD3D8E9" colspan="1" pageId="39" pageNumber="40" rowspan="1">
|
||
<taxonomicName id="5052D29037E6405026D937787538266C" class="Gastropoda" family="Flabellinidae" genus="Edmundsella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Edmundsella pedata" order="Nudibranchia" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="pedata">Edmundsella pedata</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |