507 lines
71 KiB
XML
507 lines
71 KiB
XML
<document id="76AFFE5A4C17418646ACA9098A655B9A" ID-CLB-Dataset="22241" ID-DOI="10.5852/ejt.2013.60" ID-GBIF-Dataset="a44ff2c0-9279-428d-9ed3-840c6a061dcc" ID-ISSN="2118-9773" ID-Zenodo-Dep="3824956" ID-ZooBank="67D17AEB-B622-4184-9265-D3D2D6DA500A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589220985040" checkinUser="carolina" docAuthor="Kok, Philippe J. R." docDate="2013" docId="C76DAB35307EC714FDB340FEB455FE8A" docLanguage="en" docName="ejt-2013-60.pdf.imf" docOrigin="European Journal of Taxonomy 60" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pristimantis imthurni Kok 2013, sp. nov." docType="treatment" docUuid="9AFC18C4-23A0-4C20-96EB-480F0F512791" docUuidSource="ZooBank" docVersion="10" lastPageNumber="21" masterDocId="3B54D34D3070C700FFD54378B17EFFC3" masterDocTitle="Two new charismatic Pristimantis species (Anura: Craugastoridae) from the tepuis of “ The Lost World ” (Pantepui region, South America)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="15" updateTime="1698839205156" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
|
||
<mods:mods id="F54F5FE5D396C3700F849467B3983F75" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="193AB308A285E560C37D69D9BBC8C6E2">
|
||
<mods:title id="19CCF95AADF0ABA17EC310DB4BA5AFB5">Two new charismatic Pristimantis species (Anura: Craugastoridae) from the tepuis of “ The Lost World ” (Pantepui region, South America)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="417EA90A000D5AA5DFF6F8346284528C" type="personal">
|
||
<mods:role id="F8E0ABCFCA61FC031DCF7F1512D54F8E">
|
||
<mods:roleTerm id="569686706C09C3493D7ED7A8C86F1140">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4311D7C80784BAD34C306D1AC0DEADD2">Kok, Philippe J. R.</mods:namePart>
|
||
<mods:affiliation id="8039A64186A542FE9387634F94E3B54A">Department of Vertebrates, Royal Belgian Institute of Natural Sciences, rue Vautier 29, B- 1000 Brussels, Belgium E-mail: philippe. kok @ naturalsciences. be & Amphibian Evolution Lab, Department of Biology, Vrije Universiteit Brussel, Pleinlaan 2, B- 1050 Brussels, Belgium E-mail: philippe. kok @ vub. ac. be urn: lsid: zoobank. org: author: 62 BA 0 CB 3 - FB 94 - 407 F- 90 A 1 - 06 AB 6401 B 911</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="0E4E032B56A718683DF59A9A91B90961">text</mods:typeOfResource>
|
||
<mods:relatedItem id="78A5637DB97AC80AC532BE5027E9F212" type="host">
|
||
<mods:titleInfo id="8E1DADC244C1B264D24B490AE4BD4012">
|
||
<mods:title id="EA5A65610A230109EC0715D8428874C0">European Journal of Taxonomy</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="7E288B7BEFDD3C406D9809CB1B6F9549">
|
||
<mods:date id="F6AAC5F2F3C0808DC4C89D9BB15384FE">2013</mods:date>
|
||
<mods:detail id="7C111D7C6DFBAE46E43993450D0B812A" type="pubDate">
|
||
<mods:number id="C2202BC3EC6F4FA72B4DEFDEEE259EDE">2013-10-22</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="CDA6E2EAEA5F5BE8C2CD6613A3B7F977" type="issue">
|
||
<mods:number id="23B8300D7BBB7054927DB9FAEE1596EB">60</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="505F9A82FE5F927A81124B2F7870E50C" unit="page">
|
||
<mods:start id="D3ECA9FF06FE10C9A555DE6F909E1955">1</mods:start>
|
||
<mods:end id="8F3C4DE80A98A1596C5AF41F82AEAAFE">24</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="B894B85AB8B170A57E0C757464F59548">journal article</mods:classification>
|
||
<mods:identifier id="3BE71FC995A16C6C5694462C30B50E66" type="CLB-Dataset">22241</mods:identifier>
|
||
<mods:identifier id="2A9CA24827BCB90D203D550CBA9BE560" type="DOI">10.5852/ejt.2013.60</mods:identifier>
|
||
<mods:identifier id="883A15B5F52DEA0E56E719957D9FC7C9" type="GBIF-Dataset">a44ff2c0-9279-428d-9ed3-840c6a061dcc</mods:identifier>
|
||
<mods:identifier id="528BE882A3B048A3233209A0321B4648" type="ISSN">2118-9773</mods:identifier>
|
||
<mods:identifier id="1D0F56C3C6A22D5D28E375D0DA148D27" type="Zenodo-Dep">3824956</mods:identifier>
|
||
<mods:identifier id="F8E21FB44AF93466B93D681A43076C7D" type="ZooBank">67D17AEB-B622-4184-9265-D3D2D6DA500A</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="C76DAB35307EC714FDB340FEB455FE8A" ID-DOI="http://doi.org/10.5281/zenodo.3843693" ID-GBIF-Taxon="164044983" ID-Zenodo-Dep="3843693" LSID="urn:lsid:zoobank.org:act:9AFC18C4-23A0-4C20-96EB-480F0F512791" httpUri="http://treatment.plazi.org/id/C76DAB35307EC714FDB340FEB455FE8A" lastPageId="20" lastPageNumber="21" pageId="14" pageNumber="15">
|
||
<subSubSection id="07DE49A8307EC70EFDB340FEB2B0FC63" box="[614,974,901,928]" pageId="14" pageNumber="15" type="nomenclature">
|
||
<paragraph id="4F7B1A23307EC70EFDB340FEB2B0FC63" blockId="14.[372,1215,901,999]" box="[614,974,901,928]" pageId="14" pageNumber="15">
|
||
<heading id="1433AD4F307EC70EFDB340FEB2B0FC63" box="[614,974,901,928]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="15" reason="2">
|
||
<taxonomicName id="88C461A0307EC70EFDB340FEB212FC63" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[614,876,901,928]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="imthurni" status="sp. nov.">
|
||
<emphasis id="7DB0C631307EC70EFDB340FEB212FC63" bold="true" box="[614,876,901,928]" italics="true" pageId="14" pageNumber="15">Pristimantis imthurni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A307EC70EFCA640FEB2B0FC63" box="[883,974,902,928]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="07DE49A8307EC70EFD7A40D1B5C1FC24" pageId="14" pageNumber="15" type="description">
|
||
<paragraph id="4F7B1A23307EC70EFD7A40D1B2FAFC00" blockId="14.[372,1215,901,999]" box="[687,900,937,963]" pageId="14" pageNumber="15">
|
||
<figureCitation id="D7FF06A6307EC70EFD7A40D1B25FFC00" box="[687,801,937,963]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="15.[189,232,1965,1991]" captionStartId-1="16.[189,232,1965,1991]" captionStartId-2="17.[189,232,1966,1992]" captionTargetBox-0="[189,1398,856,1923]" captionTargetBox-1="[375,1184,1149,1920]" captionTargetBox-2="[218,1361,1393,1894]" captionTargetId-0="figure@15.[189,1398,856,1923]" captionTargetId-1="figure@16.[348,1231,1133,1935]" captionTargetId-2="figure@17.[191,1400,1380,1924]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="Fig. 8. Pristimantis imthurni sp. nov. in life. A. Dorsolateral view of the ♂ holotype (IRSNB 4165). B. Ventral view of the same specimen. Photographs by the author." captionText-1="Fig. 9. Pristimantis imthurni sp. nov. Ventral view of left hand and left foot of the ♂ holotype (IRSNB 4165). Photographs by the author." captionText-2="Fig. 10. Pristimantis imthurni sp. nov. in preservative. A. Dorsol view of the ♂ holotype (IRSNB 4165). B. Ventral view of the same specimen. Photographs by the author." figureDoi-0="http://doi.org/10.5281/zenodo.3824972" figureDoi-1="http://doi.org/10.5281/zenodo.3824974" figureDoi-2="http://doi.org/10.5281/zenodo.3824976" httpUri-0="https://zenodo.org/record/3824972/files/figure.png" httpUri-1="https://zenodo.org/record/3824974/files/figure.png" httpUri-2="https://zenodo.org/record/3824976/files/figure.png" pageId="14" pageNumber="15">Figs 8-10</figureCitation>
|
||
,
|
||
<tableCitation id="02462F98307EC70EFCF840D1B2FAFC00" box="[813,900,937,963]" captionStart="Table 1" captionStartId="9.[190,259,268,294]" captionTargetId="graphics@9.[189,1398,361,1247]" captionTargetPageId="9" captionText="Table 1. Sex, measurements (in mm), and HL/SVL ratio of type specimens of Pristimantis jamescameroni sp. nov. and Pristimantis imthurni sp. nov. Abbreviations are defined in the text." httpUri="http://table.plazi.org/id/1BBB4AAB3079C709FF6B4274B519FE8A" pageId="14" pageNumber="15" tableUuid="1BBB4AAB3079C709FF6B4274B519FE8A">Table 1</tableCitation>
|
||
</paragraph>
|
||
<paragraph id="4F7B1A23307EC70EFEA140B4B5C1FC24" blockId="14.[372,1215,901,999]" box="[372,1215,972,999]" pageId="14" pageNumber="15">
|
||
<uri id="3B551621307EC70EFEA140B4B5C1FC24" box="[372,1215,972,999]" pageId="14" pageNumber="15">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="3B6220F6307EC70EFD4140B4B5C1FC24" box="[660,1215,972,999]" pageId="14" pageNumber="15">9AFC18C4-23A0-4C20-96EB-480F0F512791</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="07DE49A8307EC70EFF684776B50DFBEB" box="[189,1139,1037,1064]" pageId="14" pageNumber="15" type="reference_group">
|
||
<paragraph id="4F7B1A23307EC70EFF684776B50DFBEB" blockId="14.[189,1139,1037,1064]" box="[189,1139,1037,1064]" pageId="14" pageNumber="15">
|
||
<treatmentCitationGroup id="6FD43D0D307EC70EFF684776B50DFBEB" box="[189,1139,1037,1064]" pageId="14" pageNumber="15">
|
||
<taxonomicName id="88C461A0307EC70EFF684776B0EFFBEB" ID-CoL="632GS" authorityName="Dumeril & Bibron" authorityYear="1841" box="[189,401,1038,1064]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="7DB0C631307EC70EFF684776B0EFFBEB" box="[189,401,1038,1064]" italics="true" pageId="14" pageNumber="15">Eleutherodactylus</emphasis>
|
||
</taxonomicName>
|
||
sp. H –
|
||
<treatmentCitation id="CE653C32307EC70EFE214776B2F3FBEB" author="McDiarmid R. W. & Donnelly M. A." box="[500,909,1037,1064]" page="514" pageId="14" pageNumber="15" year="2005">
|
||
<bibRefCitation id="2B5567D2307EC70EFE214776B2F3FBEB" author="McDiarmid & Donnelly" box="[500,909,1037,1064]" firstAuthor="McDiarmid" pageId="14" pageNumber="15" pagination="461 - 560" refId="ref11218" refString="McDiarmid R. W. & Donnelly M. A. 2005. The herpetofauna of the Guayana Highlands: amphibians and reptiles of the Lost World. In: Donnelly M. A., Crother B. I., Guyer C., Wake M. H. & White M. E. (eds) Ecology and Evolution in the Tropics: A Herpetological Perspective: 461 - 560. University of Chicago Press, Chicago." type="book chapter" year="2005">McDiarmid & Donnelly 2005: 514</bibRefCitation>
|
||
</treatmentCitation>
|
||
[table 18A.1], 521.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="07DE49A8307EC70EFF684737B3F8F85D" pageId="14" pageNumber="15" type="diagnosis">
|
||
<paragraph id="4F7B1A23307EC70EFF684737B095FBAA" blockId="14.[189,491,1103,1129]" box="[189,491,1103,1129]" pageId="14" pageNumber="15">
|
||
<heading id="1433AD4F307EC70EFF684737B095FBAA" bold="true" box="[189,491,1103,1129]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="7DB0C631307EC70EFF684737B095FBAA" bold="true" box="[189,491,1103,1129]" pageId="14" pageNumber="15">Definition and diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="4F7B1A23307EC70EFF684706B3F8F85D" blockId="14.[189,1399,1150,1951]" pageId="14" pageNumber="15">
|
||
A small species of the genus
|
||
<taxonomicName id="88C461A0307EC70EFDC14707B3DDFB5B" box="[532,675,1151,1176]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="genus">
|
||
<emphasis id="7DB0C631307EC70EFDC14707B3DDFB5B" box="[532,675,1151,1176]" italics="true" pageId="14" pageNumber="15">Pristimantis</emphasis>
|
||
</taxonomicName>
|
||
currently not assigned to any species group, but morphologically most similar to species of the polyphyletic
|
||
<emphasis id="7DB0C631307EC70EFD2B47DAB2F4FB78" box="[766,906,1186,1211]" italics="true" pageId="14" pageNumber="15">unistrigatus</emphasis>
|
||
species group (
|
||
<emphasis id="7DB0C631307EC70EFB9347DAB5F9FB78" box="[1094,1159,1186,1211]" italics="true" pageId="14" pageNumber="15">sensu</emphasis>
|
||
<bibRefCitation id="2B5567D2307EC70EFB4547D9B40CFB78" author="Hedges" box="[1168,1394,1185,1211]" etAl="et al." firstAuthor="Hedges" pageId="14" pageNumber="15" pagination="1 - 182" refId="ref10373" refString="Hedges S. B., Duellman W. E. & Heinicke M. P. 2008. New World direct-developing frogs (Anura: Terrarana): Molecular phylogeny, classification, biogeography, and conservation. Zootaxa 1737: 1 - 182." type="journal article" year="2008">
|
||
Hedges
|
||
<emphasis id="7DB0C631307EC70EFB2547DAB45BFB78" box="[1264,1317,1185,1211]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
, see also discussion in
|
||
<bibRefCitation id="2B5567D2307EC70EFE1247BCB3FCFB1D" author="Kok" box="[455,642,1220,1246]" etAl="et al." firstAuthor="Kok" pageId="14" pageNumber="15" pagination="1 - 19" refId="ref10912" refString="Kok P. J. R, Means D. B. & Bossuyt F. 2011. A new highland species of Pristimantis Jimenez de la Espada, 1871 (Anura: Strabomantidae) from the Pantepui region, northern South America. Zootaxa 2934: 1 - 19." type="journal article" year="2011">
|
||
Kok
|
||
<emphasis id="7DB0C631307EC70EFDD747BDB341FB1D" box="[514,575,1220,1246]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
) mainly characterized in having Finger I shorter than II, Toe V longer than III, extending to the distal edge of the distal subarticular tubercle of Toe IV when toes are adpressed, and by the absence of cranial crests and the presence of vomerine teeth. The new species is characterized by the following unique combination of characters: (1) body small, male
|
||
<quantity id="883CB7C6307EC70EFADF4656B409FA8B" box="[1290,1399,1326,1352]" metricMagnitude="-2" metricUnit="m" metricValue="2.29" pageId="14" pageNumber="15" unit="mm" value="22.9">22.9 mm</quantity>
|
||
SVL; (2) dorsal skin smooth, belly skin coarsely areolate; (3) tympanum absent (tympanic membrane not differentiated and tympanic annulus not visible externally); (4) tiny pharyngeal ostia present; (5) snout broadly rounded in dorsal view and in profile,
|
||
<emphasis id="7DB0C631307EC70EFCEB46EFB57BFA72" box="[830,1029,1431,1457]" italics="true" pageId="14" pageNumber="15">canthus rostralis</emphasis>
|
||
nearly straight in dorsal view, rounded in cross section, loreal region concave, flaring slightly at upper lip; (6) upper eyelid smooth with one barely distinct enlarged tubercle; (7) choanae very small, round, dentigerous processes of vomers inconspicuous, ovoid, posterior and medial to choanae, each bearing 3-4 teeth; (8) absence of vocal slits in male; (9) tongue cordiform; (10) two large, broadly connected, mostly unpigmented nuptial pads on each thumb in male; (11) Finger I shorter than II; (12) fingers with lateral fringes, best developed preaxially on Fingers II-III; (13) ulnar tubercles absent; (14) tarsal tubercles absent, one inconspicuous calcar tubercle present; (15) inner metatarsal tubercle oval, about four times the size of the round, projecting outer metatarsal tubercle; (16) Toe V longer than III, extending to the distal edge of the distal tubercle on Toe IV when toes are adpressed; (17) toes with lateral fringes, best developed preaxially on Toes III-IV, webbing basal between Toes IV-V; (18) in life dorsal background colouration is vivid yellow with a chocolate brown middorsal stripe and chocolate brown flanks, ventral colouration is translucent brown with a few white spots and flecks; (19) in preservative dorsal background colouration is pale yellow with a light brown middorsal stripe and light brown flanks, ventral colouration is light grey with a few white spots and flecks.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="07DE49A8307FC70FFF684273B5AFFDC7" pageId="15" pageNumber="16" type="etymology">
|
||
<paragraph id="4F7B1A23307FC70FFF684273B03AFEE6" blockId="15.[189,324,267,293]" box="[189,324,267,293]" pageId="15" pageNumber="16">
|
||
<heading id="1433AD4F307FC70FFF684273B03AFEE6" bold="true" box="[189,324,267,293]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="7DB0C631307FC70FFF684273B03AFEE6" bold="true" box="[189,324,267,293]" pageId="15" pageNumber="16">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="4F7B1A23307FC70FFF684241B5AFFDC7" blockId="15.[189,1399,313,516]" pageId="15" pageNumber="16">
|
||
The specific epithet is a noun in the genitive case, honouring Sir Everard F. im Thurn (
|
||
<date id="3B7A3CE3307FC70FFB304241B417FE97" box="[1253,1385,313,340]" pageId="15" pageNumber="16" value="1852" valueMax="1932">1852-1932</date>
|
||
), British colonial official, author, explorer, botanist, and photographer. Everard im Thurn was the first to climb a major tepui (Mount
|
||
<collectingRegion id="8D00D4C1307FC70FFDDE42F8B30EFE59" box="[523,624,384,410]" country="Brazil" name="Roraima" pageId="15" pageNumber="16">Roraima</collectingRegion>
|
||
in
|
||
<date id="3B7A3CE3307FC70FFD4242F8B22BFE59" box="[663,853,384,410]" pageId="15" pageNumber="16" value="1884-12">December 1884</date>
|
||
), along with British surveyor Harry Perkins, a Pomeroon Amerindian named Gabriel, and five other unnamed Amerindians (
|
||
<bibRefCitation id="2B5567D2307FC70FFBEA42DBB59DFE7E" author="Dalziell" box="[1087,1251,419,445]" firstAuthor="Dalziell" pageId="15" pageNumber="16" pagination="97 - 116" refId="ref10135" refString="Dalziell R. 2007. Everard im Thurn in British Guiana and the western Pacific. In: Hulme P. & McDougall R. (eds.) Writing, Travel and Empire: 97 - 116. I. B. Tauris & Co. Ltd, London." type="book chapter" year="2007">Dalziell 2007</bibRefCitation>
|
||
). Im Thurn’s expedition on
|
||
<collectingRegion id="8D00D4C1307FC70FFEB642BEB0B6FE23" box="[355,456,454,480]" country="Brazil" name="Roraima" pageId="15" pageNumber="16">Roraima</collectingRegion>
|
||
and his numerous discoveries were partly eclipsed by the popular novel that they inspired: “The Lost World” by Sir Arthur Conan Doyle (see Introduction;
|
||
<bibRefCitation id="2B5567D2307FC70FFBF44291B5BBFDC7" author="Dalziell" box="[1057,1221,489,516]" firstAuthor="Dalziell" pageId="15" pageNumber="16" pagination="97 - 116" refId="ref10135" refString="Dalziell R. 2007. Everard im Thurn in British Guiana and the western Pacific. In: Hulme P. & McDougall R. (eds.) Writing, Travel and Empire: 97 - 116. I. B. Tauris & Co. Ltd, London." type="book chapter" year="2007">Dalziell 2007</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="07DE49A8307FC70FFF684153B409FD78" pageId="15" pageNumber="16" type="materials_examined">
|
||
<paragraph id="4F7B1A23307FC70FFF684153B0D4FD86" blockId="15.[189,426,555,581]" box="[189,426,555,581]" pageId="15" pageNumber="16">
|
||
<heading id="1433AD4F307FC70FFF684153B0D4FD86" bold="true" box="[189,426,555,581]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="7DB0C631307FC70FFF684153B0D4FD86" bold="true" box="[189,426,555,581]" pageId="15" pageNumber="16">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<materialsCitation id="FFAC107E307FC70FFF3F4122B409FD78" ID-GBIF-Occurrence="2612049302" collectingDate="2012-06-20" collectorName="Philippe J. R. Kok" country="Venezuela" elevation="2471" latitude="5.7691665" location="Ptaritepui" longLatPrecision="21" longitude="-61.817223" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Bolivar state" typeStatus="holotype">
|
||
<paragraph id="4F7B1A23307FC70FFF3F4122B022FDB7" blockId="15.[189,1399,602,699]" box="[234,348,602,628]" pageId="15" pageNumber="16">
|
||
<emphasis id="7DB0C631307FC70FFF3F4122B022FDB7" bold="true" box="[234,348,602,628]" pageId="15" pageNumber="16">
|
||
<typeStatus id="907FA481307FC70FFF3F4122B022FDB7" box="[234,348,602,628]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="4F7B1A23307FC70FFF684105B409FD78" blockId="15.[189,1399,602,699]" pageId="15" pageNumber="16">
|
||
<collectingCountry id="37D35AB3307FC70FFF684105B00EFD54" box="[189,368,637,663]" name="Venezuela" pageId="15" pageNumber="16">VENEZUELA</collectingCountry>
|
||
: an
|
||
<specimenCount id="59C2D1AA307FC70FFE4A4105B364FD54" box="[415,538,637,663]" pageId="15" pageNumber="16" type="adult">adult male</specimenCount>
|
||
collected by
|
||
<collectorName id="E2317FF5307FC70FFD6D4105B2EAFD54" box="[696,916,637,663]" pageId="15" pageNumber="16">Philippe J. R. Kok</collectorName>
|
||
,
|
||
<date id="3B7A3CE3307FC70FFC744105B547FD54" box="[929,1081,637,663]" pageId="15" pageNumber="16" value="2012-06-20">
|
||
<collectingDate id="2B3EC50B307FC70FFC744105B547FD54" box="[929,1081,637,663]" pageId="15" pageNumber="16" value="2012-06-20">20 Jun. 2012</collectingDate>
|
||
</date>
|
||
at 20h37, summit of
|
||
<location id="4A1B4CF8307FC70FFAE04105B185FD79" LSID="urn:lsid:plazi:treatment:C76DAB35307EC714FDB340FEB455FE8A:4A1B4CF8307FC70FFAE04105B185FD79" country="Venezuela" latitude="5.7691665" longLatPrecision="21" longitude="-61.817223" name="Ptaritepui" pageId="15" pageNumber="16" stateProvince="Bolivar state">Ptaritepui</location>
|
||
,
|
||
<collectingRegion id="8D00D4C1307FC70FFED041D8B0E3FD79" box="[261,413,672,698]" country="Venezuela" name="Bolivar" pageId="15" pageNumber="16">Bolívar state</collectingRegion>
|
||
,
|
||
<geoCoordinate id="2AF07CE4307FC70FFE7D41D8B34CFD79" box="[424,562,672,699]" degrees="5" direction="north" minutes="46" orientation="latitude" pageId="15" pageNumber="16" precision="15" seconds="09" value="5.7691665">5°46’09” N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2AF07CE4307FC70FFDE841D8B3A7FD79" box="[573,729,672,698]" degrees="61" direction="west" minutes="49" orientation="longitude" pageId="15" pageNumber="16" precision="15" seconds="02" value="-61.817223">61°49’02” W</geoCoordinate>
|
||
,
|
||
<quantity id="883CB7C6307FC70FFD3141D8B242FD79" box="[740,828,672,698]" metricMagnitude="3" metricUnit="m" metricValue="2.471" pageId="15" pageNumber="16" unit="m" value="2471.0">
|
||
<elevation id="C4E9FD10307FC70FFD3141D8B242FD79" box="[740,828,672,698]" metricMagnitude="3" metricUnit="m" metricValue="2.471" pageId="15" pageNumber="16" unit="m" value="2471.0">2471 m</elevation>
|
||
</quantity>
|
||
elevation, IRSNB 4165 (field number PK 3671).
|
||
</paragraph>
|
||
</materialsCitation>
|
||
</subSubSection>
|
||
<caption id="1BBB4AAB307FC70FFF6844D5B2BCF829" ID-DOI="http://doi.org/10.5281/zenodo.3824972" ID-Zenodo-Dep="3824972" httpUri="https://zenodo.org/record/3824972/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,232,1965,1991]" targetBox="[189,1398,856,1923]" targetPageId="15">
|
||
<paragraph id="4F7B1A23307FC70FFF6844D5B2BCF829" blockId="15.[189,1399,1965,2027]" pageId="15" pageNumber="16">
|
||
<emphasis id="7DB0C631307FC70FFF6844D5B073F804" bold="true" box="[189,269,1965,1991]" pageId="15" pageNumber="16">Fig. 8.</emphasis>
|
||
<taxonomicName id="88C461A0307FC70FFEC244D6B36BF804" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[279,533,1965,1991]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="imthurni" status="sp. nov.">
|
||
<emphasis id="7DB0C631307FC70FFEC244D6B36BF804" box="[279,533,1965,1991]" italics="true" pageId="15" pageNumber="16">Pristimantis imthurni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A307FC70FFDCB44D6B305F80B" box="[542,635,1966,1992]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in life.
|
||
<emphasis id="7DB0C631307FC70FFD0844D5B38CF804" bold="true" box="[733,754,1965,1991]" pageId="15" pageNumber="16">A</emphasis>
|
||
. Dorsolateral view of the ♂ holotype (IRSNB 4165).
|
||
<emphasis id="7DB0C631307FC70FFF6844A9B1AFF828" bold="true" box="[189,209,2001,2027]" pageId="15" pageNumber="16">B</emphasis>
|
||
. Ventral view of the same specimen. Photographs by the author.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="07DE49A83060C710FF684273B470FBEB" pageId="16" pageNumber="17" type="discussion">
|
||
<paragraph id="4F7B1A233060C710FF684273B23BFEE6" blockId="16.[189,837,267,293]" box="[189,837,267,293]" pageId="16" pageNumber="17">
|
||
<heading id="1433AD4F3060C710FF684273B23BFEE6" bold="true" box="[189,837,267,293]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
|
||
<emphasis id="7DB0C6313060C710FF684273B23BFEE6" bold="true" box="[189,837,267,293]" pageId="16" pageNumber="17">Morphological comparisons with congeneric species</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="4F7B1A233060C710FF684242B39FFEB4" blockId="16.[189,1398,313,375]" pageId="16" pageNumber="17">
|
||
For the same reasons mentioned above, comparisons with congeners only focus on the 21
|
||
<taxonomicName id="88C461A03060C710FB324242B408FE90" box="[1255,1398,314,339]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
|
||
<emphasis id="7DB0C6313060C710FB324242B408FE90" box="[1255,1398,314,339]" italics="true" pageId="16" pageNumber="17">Pristimantis</emphasis>
|
||
</taxonomicName>
|
||
species known to occur in the Pantepui region.
|
||
</paragraph>
|
||
<paragraph id="4F7B1A233060C710FF6842E7B27EFDAA" blockId="16.[189,1399,414,617]" pageId="16" pageNumber="17">
|
||
<taxonomicName id="88C461A03060C710FF6842E7B0C4FE7B" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[189,442,414,440]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="imthurni" status="sp. nov.">
|
||
<emphasis id="7DB0C6313060C710FF6842E7B0C4FE7B" box="[189,442,414,440]" italics="true" pageId="16" pageNumber="17">Pristimantis imthurni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A3060C710FE1142E7B35EFE7A" box="[452,544,415,441]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is most easily distinguished from
|
||
<taxonomicName id="88C461A03060C710FC6842E7B53EFE7B" authorityName="Rojas-Runjaic, Salerno, Senaris & Pauly" authorityYear="2013" box="[957,1088,414,440]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="abakapa">
|
||
<emphasis id="7DB0C6313060C710FC6842E7B53EFE7B" box="[957,1088,414,440]" italics="true" pageId="16" pageNumber="17">P. abakapa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FB8542E7B47CFE7B" authorityName="Kok, Means & Bossuyt" authorityYear="2011" box="[1104,1282,415,440]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="aureoventris">
|
||
<emphasis id="7DB0C6313060C710FB8542E7B47CFE7B" box="[1104,1282,415,440]" italics="true" pageId="16" pageNumber="17">P. aureoventris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FAC642E7B411FE7B" baseAuthorityName="Myers & Donnelly" baseAuthorityYear="1997" box="[1299,1391,415,440]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="avius">
|
||
<emphasis id="7DB0C6313060C710FAC642E7B411FE7B" box="[1299,1391,415,440]" italics="true" pageId="16" pageNumber="17">P. avius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FF6842BAB035FE18" baseAuthorityName="Myers & Donnelly" baseAuthorityYear="1996" box="[189,331,450,475]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="cantitans">
|
||
<emphasis id="7DB0C6313060C710FF6842BAB035FE18" box="[189,331,450,475]" italics="true" pageId="16" pageNumber="17">P. cantitans</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FE8A42BAB349FE18" authorityName="Means & Savage" authorityYear="2007" box="[351,567,449,475]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="dendrobatoides">
|
||
<emphasis id="7DB0C6313060C710FE8A42BAB349FE18" box="[351,567,449,475]" italics="true" pageId="16" pageNumber="17">P. dendrobatoides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FD9E42BAB250FE18" baseAuthorityName="Schluter & Rodder" baseAuthorityYear="2007" box="[587,814,450,475]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="guaiquinimensis">
|
||
<emphasis id="7DB0C6313060C710FD9E42BAB250FE18" box="[587,814,450,475]" italics="true" pageId="16" pageNumber="17">P. guaiquinimensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FC9642BAB28AFE18" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[835,1012,450,475]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="marmoratus">
|
||
<emphasis id="7DB0C6313060C710FC9642BAB28AFE18" box="[835,1012,450,475]" italics="true" pageId="16" pageNumber="17">P. marmoratus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FBDC42BAB5DBFE18" baseAuthorityName="Myers & Donnelly" baseAuthorityYear="1997" box="[1033,1189,450,475]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="memorans">
|
||
<emphasis id="7DB0C6313060C710FBDC42BAB5DBFE18" box="[1033,1189,450,475]" italics="true" pageId="16" pageNumber="17">P. memorans</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FB6F42BAB432FE18" baseAuthorityName="Myers & Donnelly" baseAuthorityYear="1996" box="[1210,1356,450,475]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="pruinatus">
|
||
<emphasis id="7DB0C6313060C710FB6F42BAB432FE18" box="[1210,1356,450,475]" italics="true" pageId="16" pageNumber="17">P. pruinatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FAB442BAB049FE3D" baseAuthorityName="Rivero" baseAuthorityYear="1968" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="pulvinatus">
|
||
<emphasis id="7DB0C6313060C710FAB442BAB049FE3D" italics="true" pageId="16" pageNumber="17">P. pulvinatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FE9D429DB096FE3D" authorityName="Means & Savage" authorityYear="2007" box="[328,488,484,510]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="saltissimus">
|
||
<emphasis id="7DB0C6313060C710FE9D429DB096FE3D" box="[328,488,484,510]" italics="true" pageId="16" pageNumber="17">P. saltissimus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FE2D429DB3CDFE3D" box="[504,691,485,510]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="sarisarinama">
|
||
<emphasis id="7DB0C6313060C710FE2D429DB3CDFE3D" box="[504,691,485,510]" italics="true" pageId="16" pageNumber="17">P. sarisarinama</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FD11429DB250FE3D" baseAuthorityName="Melin" baseAuthorityYear="1941" box="[708,814,484,510]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="vilarsi">
|
||
<emphasis id="7DB0C6313060C710FD11429DB250FE3D" box="[708,814,484,510]" italics="true" pageId="16" pageNumber="17">P. vilarsi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FCEA429DB28AFE3D" authorityName="Rodder & Jungfer" authorityYear="2008" box="[831,1012,485,510]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="yuruaniensis">
|
||
<emphasis id="7DB0C6313060C710FCEA429DB28AFE3D" box="[831,1012,485,510]" italics="true" pageId="16" pageNumber="17">P. yuruaniensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="88C461A03060C710FBED429DB5A7FE3D" baseAuthorityName="Lynch & Hoogmoed" baseAuthorityYear="1977" box="[1080,1241,484,510]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="zeuctotylus">
|
||
<emphasis id="7DB0C6313060C710FBED429DB5A7FE3D" box="[1080,1241,484,510]" italics="true" pageId="16" pageNumber="17">P. zeuctotylus</emphasis>
|
||
</taxonomicName>
|
||
by lacking a differentiated tympanic membrane and an external tympanic annulus (at least one of these structures is detectable in the latter 15 species). It further differs from
|
||
<taxonomicName id="88C461A03060C710FC884154B2BBFD86" baseAuthorityName="Melin" baseAuthorityYear="1941" box="[861,965,555,581]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="vilarsi">
|
||
<emphasis id="7DB0C6313060C710FC884154B2BBFD86" box="[861,965,555,581]" italics="true" pageId="16" pageNumber="17">P. vilarsi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="88C461A03060C710FC2B4154B5E2FD86" baseAuthorityName="Lynch & Hoogmoed" baseAuthorityYear="1977" box="[1022,1180,555,581]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="zeuctotylus">
|
||
<emphasis id="7DB0C6313060C710FC2B4154B5E2FD86" box="[1022,1180,555,581]" italics="true" pageId="16" pageNumber="17">P. zeuctotylus</emphasis>
|
||
</taxonomicName>
|
||
in having Finger I <II (Finger I> II in
|
||
<taxonomicName id="88C461A03060C710FE7A4137B369FDAB" baseAuthorityName="Melin" baseAuthorityYear="1941" box="[431,535,590,616]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="vilarsi">
|
||
<emphasis id="7DB0C6313060C710FE7A4137B369FDAB" box="[431,535,590,616]" italics="true" pageId="16" pageNumber="17">P. vilarsi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="88C461A03060C710FD854137B391FDAB" baseAuthorityName="Lynch & Hoogmoed" baseAuthorityYear="1977" box="[592,751,590,616]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="zeuctotylus">
|
||
<emphasis id="7DB0C6313060C710FD854137B391FDAB" box="[592,751,590,616]" italics="true" pageId="16" pageNumber="17">P. zeuctotylus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="4F7B1A233060C710FF6841E8B335FD32" blockId="16.[189,1398,655,753]" pageId="16" pageNumber="17">
|
||
<taxonomicName id="88C461A03060C710FF6841E8B0C6FD6A" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[189,440,655,681]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="imthurni" status="sp. nov.">
|
||
<emphasis id="7DB0C6313060C710FF6841E8B0C6FD6A" box="[189,440,655,681]" italics="true" pageId="16" pageNumber="17">Pristimantis imthurni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A3060C710FE1541E8B364FD69" box="[448,538,656,682]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is immediately distinguished from
|
||
<taxonomicName id="88C461A03060C710FC6D41E8B52DFD6A" baseAuthorityName="Myers & Donnelly" baseAuthorityYear="2008" box="[952,1107,656,681]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="auricarens">
|
||
<emphasis id="7DB0C6313060C710FC6D41E8B52DFD6A" box="[952,1107,656,681]" italics="true" pageId="16" pageNumber="17">P. auricarens</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FBB441E8B5C1FD6A" authorityName="Means & Savage" authorityYear="2007" box="[1121,1215,656,681]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="jester">
|
||
<emphasis id="7DB0C6313060C710FBB441E8B5C1FD6A" box="[1121,1215,656,681]" italics="true" pageId="16" pageNumber="17">P. jester</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FB1841E8B40EFD6A" baseAuthorityName="Fuentes-Ramos & Barrio-Amoros" baseAuthorityYear="2004" box="[1229,1392,655,681]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="marahuaka">
|
||
<emphasis id="7DB0C6313060C710FB1841E8B40EFD6A" box="[1229,1392,655,681]" italics="true" pageId="16" pageNumber="17">P. marahuaka</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="88C461A03060C710FF6841CCB02AFD0E" authorityName="Barrio-Amoros, Mesa, Brewer-Carias & McDiarmid" authorityYear="2010" box="[189,340,691,717]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="muchimuk">
|
||
<emphasis id="7DB0C6313060C710FF6841CCB02AFD0E" box="[189,340,691,717]" italics="true" pageId="16" pageNumber="17">P. muchimuk</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="88C461A03060C710FE4041CCB361FD0E" baseAuthorityName="Myers & Donnelly" baseAuthorityYear="1996" box="[405,543,692,717]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="yaviensis">
|
||
<emphasis id="7DB0C6313060C710FE4041CCB361FD0E" box="[405,543,692,717]" italics="true" pageId="16" pageNumber="17">P. yaviensis</emphasis>
|
||
</taxonomicName>
|
||
in having lateral fringes on most fingers and toes (absent or limited to a weak keel in the latter 5 species).
|
||
</paragraph>
|
||
<paragraph id="4F7B1A233060C710FF684060B470FBEB" blockId="16.[189,1399,791,1064]" pageId="16" pageNumber="17">
|
||
By lacking a differentiated tympanic membrane and an external tympanic annulus, and in having lateral fringes on fingers and toes,
|
||
<taxonomicName id="88C461A03060C710FDB04043B21AFC96" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[613,868,827,853]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="imthurni" status="sp. nov.">
|
||
<emphasis id="7DB0C6313060C710FDB04043B21AFC96" box="[613,868,827,853]" italics="true" pageId="16" pageNumber="17">Pristimantis imthurni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A3060C710FCBB4043B2B2FC96" box="[878,972,827,853]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is morphologically most similar to
|
||
<taxonomicName id="88C461A03060C710FF684027B0F3FCBB" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[189,397,863,888]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="jamescameroni" status="sp. nov.">
|
||
<emphasis id="7DB0C6313060C710FF684027B0F3FCBB" box="[189,397,863,888]" italics="true" pageId="16" pageNumber="17">P. jamescameroni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A3060C710FE434026B08BFCBB" box="[406,501,862,888]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, described earlier in this paper. These two species are mainly distinguished based on distinct dorsal and ventral colour patterns (compare
|
||
<figureCitation id="D7FF06A63060C710FC4C40F9B2A1FC58" box="[921,991,897,923]" captionStart="Fig" captionStartId="5.[189,232,1858,1884]" captionTargetBox="[189,1398,1017,1816]" captionTargetId="figure@5.[189,1398,1017,1816]" captionTargetPageId="5" captionText="Fig. 2. Pristimantis jamescameroni sp. nov. Intraspecific variation in dorsal and ventral colour pattern in living specimens. A. Dorsolateral view of the ♂ holotype (IRSNB 4160). B. Ventral view of the same specimen. C. Dorsolateral view of a ♀ paratype (IRSNB 4163). D. Ventral view of the same specimen. E. Dorsolateral view of a ♀ paratype (IRSNB 4162). F. Ventral view of the same specimen. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3824960" httpUri="https://zenodo.org/record/3824960/files/figure.png" pageId="16" pageNumber="17">Fig. 2</figureCitation>
|
||
with
|
||
<figureCitation id="D7FF06A63060C710FBF140F9B515FC58" box="[1060,1131,897,923]" captionStart="Fig" captionStartId="15.[189,232,1965,1991]" captionTargetBox="[189,1398,856,1923]" captionTargetId="figure@15.[189,1398,856,1923]" captionTargetPageId="15" captionText="Fig. 8. Pristimantis imthurni sp. nov. in life. A. Dorsolateral view of the ♂ holotype (IRSNB 4165). B. Ventral view of the same specimen. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3824972" httpUri="https://zenodo.org/record/3824972/files/figure.png" pageId="16" pageNumber="17">Fig. 8</figureCitation>
|
||
), skin texture (smooth in
|
||
<taxonomicName id="88C461A03060C710FF0940DDB021FC7D" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[220,351,932,958]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="imthurni" status="sp. nov.">
|
||
<emphasis id="7DB0C6313060C710FF0940DDB021FC7D" box="[220,351,932,958]" italics="true" pageId="16" pageNumber="17">P. imthurni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A3060C710FEB240DDB0BDFC7C" box="[359,451,933,959]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="7DB0C6313060C710FE1F40DDB094FC7D" box="[458,490,933,958]" italics="true" pageId="16" pageNumber="17">vs.</emphasis>
|
||
shagreen in
|
||
<taxonomicName id="88C461A03060C710FD5740DDB22CFC7D" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[642,850,933,958]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="jamescameroni" status="sp. nov.">
|
||
<emphasis id="7DB0C6313060C710FD5740DDB22CFC7D" box="[642,850,933,958]" italics="true" pageId="16" pageNumber="17">P. jamescameroni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A3060C710FC8F40DDB2C7FC7C" box="[858,953,933,959]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), condition of the supratympanic fold (inconspicuous in
|
||
<taxonomicName id="88C461A03060C710FE4440B0B36FFC22" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[401,529,967,993]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="imthurni" status="sp. nov.">
|
||
<emphasis id="7DB0C6313060C710FE4440B0B36FFC22" box="[401,529,967,993]" italics="true" pageId="16" pageNumber="17">P. imthurni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A3060C710FDC340B0B311FC21" box="[534,623,968,994]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="7DB0C6313060C710FDA140B0B3EAFC22" box="[628,660,968,993]" italics="true" pageId="16" pageNumber="17">vs.</emphasis>
|
||
conspicuous in
|
||
<taxonomicName id="88C461A03060C710FC9940B0B567FC22" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[844,1049,968,993]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="jamescameroni" status="sp. nov.">
|
||
<emphasis id="7DB0C6313060C710FC9940B0B567FC22" box="[844,1049,968,993]" italics="true" pageId="16" pageNumber="17">P. jamescameroni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A3060C710FBCA40B0B505FC21" box="[1055,1147,968,994]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), and head proportion (shorter, with a more rounded snout in
|
||
<taxonomicName id="88C461A03060C710FD4C4093B260FBC6" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[665,798,1003,1029]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="imthurni" status="sp. nov.">
|
||
<emphasis id="7DB0C6313060C710FD4C4093B260FBC6" box="[665,798,1003,1029]" italics="true" pageId="16" pageNumber="17">P. imthurni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A3060C710FCFD4093B2F7FBC6" box="[808,905,1003,1029]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). Besides being morphologically reliably diagnosable, the two species are currently geographically isolated (isolated lineage segments).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="1BBB4AAB3060C710FF6844D5B335F828" ID-DOI="http://doi.org/10.5281/zenodo.3824974" ID-Zenodo-Dep="3824974" httpUri="https://zenodo.org/record/3824974/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,232,1965,1991]" targetBox="[375,1184,1149,1920]" targetPageId="16">
|
||
<paragraph id="4F7B1A233060C710FF6844D5B335F828" blockId="16.[189,1398,1965,2027]" pageId="16" pageNumber="17">
|
||
<emphasis id="7DB0C6313060C710FF6844D5B075F804" bold="true" box="[189,267,1965,1991]" pageId="16" pageNumber="17">Fig. 9.</emphasis>
|
||
<taxonomicName id="88C461A03060C710FEC644D6B371F80B" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[275,527,1966,1992]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="imthurni" status="sp. nov.">
|
||
<emphasis id="7DB0C6313060C710FEC644D6B371F80B" box="[275,527,1966,1992]" italics="true" pageId="16" pageNumber="17">Pristimantis imthurni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A3060C710FDC244D6B30CF80B" box="[535,626,1966,1992]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Ventral view of left hand and left foot of the ♂ holotype (IRSNB 4165). Photographs by the author.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="07DE49A83061C712FF6E4273B0BAFA3D" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
|
||
<paragraph id="4F7B1A233061C711FF6E4273B370FEE6" blockId="17.[187,526,267,293]" box="[187,526,267,293]" pageId="17" pageNumber="18">
|
||
<heading id="1433AD4F3061C711FF6E4273B370FEE6" bold="true" box="[187,526,267,293]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
|
||
<emphasis id="7DB0C6313061C711FF6E4273B370FEE6" bold="true" box="[187,526,267,293]" pageId="17" pageNumber="18">
|
||
Description of the
|
||
<typeStatus id="907FA4813061C711FE774273B370FEE6" box="[418,526,267,293]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="4F7B1A233061C711FF6E4242B22CFC83" blockId="17.[187,1399,313,833]" pageId="17" pageNumber="18">
|
||
An adult male
|
||
<quantity id="883CB7C63061C711FEBC4242B0AFFE97" box="[361,465,314,340]" metricMagnitude="-2" metricUnit="m" metricValue="2.29" pageId="17" pageNumber="18" unit="mm" value="22.9">22.9 mm</quantity>
|
||
SVL (
|
||
<figureCitation id="D7FF06A63061C711FDCB4241B3EFFE97" box="[542,657,313,340]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="15.[189,232,1965,1991]" captionStartId-1="16.[189,232,1965,1991]" captionStartId-2="17.[189,232,1966,1992]" captionTargetBox-0="[189,1398,856,1923]" captionTargetBox-1="[375,1184,1149,1920]" captionTargetBox-2="[218,1361,1393,1894]" captionTargetId-0="figure@15.[189,1398,856,1923]" captionTargetId-1="figure@16.[348,1231,1133,1935]" captionTargetId-2="figure@17.[191,1400,1380,1924]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="Fig. 8. Pristimantis imthurni sp. nov. in life. A. Dorsolateral view of the ♂ holotype (IRSNB 4165). B. Ventral view of the same specimen. Photographs by the author." captionText-1="Fig. 9. Pristimantis imthurni sp. nov. Ventral view of left hand and left foot of the ♂ holotype (IRSNB 4165). Photographs by the author." captionText-2="Fig. 10. Pristimantis imthurni sp. nov. in preservative. A. Dorsol view of the ♂ holotype (IRSNB 4165). B. Ventral view of the same specimen. Photographs by the author." figureDoi-0="http://doi.org/10.5281/zenodo.3824972" figureDoi-1="http://doi.org/10.5281/zenodo.3824974" figureDoi-2="http://doi.org/10.5281/zenodo.3824976" httpUri-0="https://zenodo.org/record/3824972/files/figure.png" httpUri-1="https://zenodo.org/record/3824974/files/figure.png" httpUri-2="https://zenodo.org/record/3824976/files/figure.png" pageId="17" pageNumber="18">Figs 8-10</figureCitation>
|
||
, see
|
||
<tableCitation id="02462F983061C711FD1F4241B25FFE97" box="[714,801,313,340]" captionStart="Table 1" captionStartId="9.[190,259,268,294]" captionTargetId="graphics@9.[189,1398,361,1247]" captionTargetPageId="9" captionText="Table 1. Sex, measurements (in mm), and HL/SVL ratio of type specimens of Pristimantis jamescameroni sp. nov. and Pristimantis imthurni sp. nov. Abbreviations are defined in the text." httpUri="http://table.plazi.org/id/1BBB4AAB3079C709FF6B4274B519FE8A" pageId="17" pageNumber="18" tableUuid="1BBB4AAB3079C709FF6B4274B519FE8A">Table 1</tableCitation>
|
||
for measurements), in very good condition, except a large scar under the right thigh where a piece of muscle was removed prior to preservation. Head slightly longer than wide (HW 97.7% of HL), wider than body; HW 37.1% of SVL; HL 38.0% of SVL; cranial crests absent. Snout longer than eye length (SL 112.1% of EL), broadly rounded in dorsal view and in profile;
|
||
<emphasis id="7DB0C6313061C711FEBB42BEB34AFE23" box="[366,564,454,480]" italics="true" pageId="17" pageNumber="18">canthus rostralis</emphasis>
|
||
nearly straight in dorsal view, rounded in cross section, loreal region concave, flaring slightly at upper lip; eye-naris distance shorter than eye length (EN 72.7% of EL). Nares slightly protuberant, directed posterolaterally, visible in frontal and dorsal views. Widest upper eyelid width narrower than interorbital distance (upper eyelid width 81.5% of IO), upper eyelid smooth with one barely visible enlarged tubercle on each eyelid. Tympanum absent (tympanic membrane not differentiated and tympanic annulus not visible externally); tiny pharyngeal ostia present. Supratympanic fold inconspicuous in life, barely visible and fragmented in preservative, slightly arched, apparently originating at posterior corner of eye, failing to reach shoulder; a few low and small post-rictal tubercles. Choanae very small, round; dentigerous processes of vomers inconspicuous, ovoid, posterior and medial to choanae, each bearing 3-4 teeth. Tongue cordiform, slightly wider than long, rounded posteriorly, posterior one-third free. Vocal slits and vocal sac absent.
|
||
</paragraph>
|
||
<paragraph id="4F7B1A233061C711FF6E4010B0B0FBCD" blockId="17.[187,1399,872,1038]" pageId="17" pageNumber="18">Dorsal skin smooth, including on head; middorsal raphe barely detectable in preservative, well visible in life; no dermal folds or ridges visible on dorsal surface; flanks slightly granular. Throat and upper chest surfaces smooth; no thoracic fold detectable; belly skin coarsely areolate; no discoidal fold detectable; posteroventral thigh and cloacal region coarsely areolate; cloacal sheath absent, two distinctly enlarged tubercles below cloaca.</paragraph>
|
||
<paragraph id="4F7B1A233061C712FF6E474EB40AFE4C" blockId="17.[187,1399,1078,1316]" lastBlockId="18.[189,1398,267,399]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
|
||
Hand length 27.1% of SVL. Finger I 88.6% of II. Relative length of fingers III> IV> II> I; adpressed Finger I fails to reach proximal edge of digital pad of Finger II; adpressed Finger IV reaches proximal edge of digital pad of Finger III. Two large, broadly connected, non-spinous, mostly unpigmented (translucent when wet, white when dry) nuptial pads on each thumb, extending from the preaxial surface of the thenar tubercle and invading most of it, along the dorsal and the preaxial surface of the thumb. Lateral fringes on all fingers, best developed preaxially on Fingers II-III (
|
||
<figureCitation id="D7FF06A63061C711FBDB479EB52BFAC3" box="[1038,1109,1254,1280]" captionStart="Fig" captionStartId="16.[189,232,1965,1991]" captionTargetBox="[375,1184,1149,1920]" captionTargetId="figure@16.[348,1231,1133,1935]" captionTargetPageId="16" captionText="Fig. 9. Pristimantis imthurni sp. nov. Ventral view of left hand and left foot of the ♂ holotype (IRSNB 4165). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3824974" httpUri="https://zenodo.org/record/3824974/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
|
||
). Finger discs expanded, elliptical, broader than long, circumferential groove conspicuous, distal edge of disc rounded; disc of Finger III 1.8 times wider that the distal end of the adjacent phalanx. Palmar tubercle large, well defined, fully pigmented, deeply bifid; thenar tubercle large, protuberant, ovoid; supernumerary tubercles few, of variable sizes, slightly protuberant; subarticular tubercles large, round and protuberant, one each on FI and FII, two each on FIII and FIV. Ulnar tubercles absent; presence of a small antebrachial tubercle.
|
||
</paragraph>
|
||
<caption id="1BBB4AAB3061C711FF6844D6B2BCF828" ID-DOI="http://doi.org/10.5281/zenodo.3824976" ID-Zenodo-Dep="3824976" httpUri="https://zenodo.org/record/3824976/files/figure.png" pageId="17" pageNumber="18" startId="17.[189,232,1966,1992]" targetBox="[218,1361,1393,1894]" targetPageId="17">
|
||
<paragraph id="4F7B1A233061C711FF6844D6B2BCF828" blockId="17.[189,1400,1966,2028]" pageId="17" pageNumber="18">
|
||
<emphasis id="7DB0C6313061C711FF6844D6B066F80B" bold="true" box="[189,280,1966,1992]" pageId="17" pageNumber="18">Fig. 10.</emphasis>
|
||
<taxonomicName id="88C461A03061C711FECB44D7B367F80B" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[286,537,1966,1992]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="imthurni" status="sp. nov.">
|
||
<emphasis id="7DB0C6313061C711FECB44D7B367F80B" box="[286,537,1966,1992]" italics="true" pageId="17" pageNumber="18">Pristimantis imthurni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A3061C711FDF544D7B304F80B" box="[544,634,1966,1993]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in preservative.
|
||
<emphasis id="7DB0C6313061C711FCEE44D6B22EF80B" bold="true" box="[827,848,1966,1992]" pageId="17" pageNumber="18">A</emphasis>
|
||
. Dorsol view of the ♂ holotype (IRSNB 4165).
|
||
<emphasis id="7DB0C6313061C711FF6844AAB1AFF82F" bold="true" box="[189,209,2002,2028]" pageId="17" pageNumber="18">B</emphasis>
|
||
. Ventral view of the same specimen. Photographs by the author.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4F7B1A233062C712FF6842CEB017FCF3" blockId="18.[189,1399,438,817]" pageId="18" pageNumber="19">
|
||
Hind limbs moderate in length, heels slightly overlap when held at right angles to sagittal plane; TIL 50.2% of SVL; FL 40.6% of SVL. Relative length of Toes IV> V> III> II> I; tip of Toe V extends to the distal edge of the distal subarticular tubercle on Toe IV; tip of Toe III extends to the distal edge of the penultimate subarticular tubercle on Toe IV. Lateral fringes on all toes, best developed preaxially on Toes III-IV (
|
||
<figureCitation id="D7FF06A63062C712FE29413BB339FD9E" box="[508,583,579,605]" captionStart="Fig" captionStartId="16.[189,232,1965,1991]" captionTargetBox="[375,1184,1149,1920]" captionTargetId="figure@16.[348,1231,1133,1935]" captionTargetPageId="16" captionText="Fig. 9. Pristimantis imthurni sp. nov. Ventral view of left hand and left foot of the ♂ holotype (IRSNB 4165). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3824974" httpUri="https://zenodo.org/record/3824974/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
|
||
); webbing basal between Toe IV-V. Toe discs slightly larger in size than finger discs, WTD/WFD = 1.08; toe discs expanded, elliptical, broader than long, circumferential groove conspicuous, distal edge of disc rounded. Inner metatarsal tubercle elongate, oval, about four times the size of the projecting, round outer metatarsal tubercle; subarticular tubercles round, large and protuberant; supernumerary plantar tubercles small, low and round, increasing in size distally. Single, inconspicuous, very small, round calcar tubercle; no outer tarsal tubercles detectable; inner tarsal fold not detectable.
|
||
</paragraph>
|
||
<paragraph id="4F7B1A233062C712FF684020B3FCFCB1" blockId="18.[189,642,856,882]" box="[189,642,856,882]" pageId="18" pageNumber="19">
|
||
<emphasis id="7DB0C6313062C712FF684020B08BFCB1" bold="true" box="[189,501,856,882]" pageId="18" pageNumber="19">
|
||
Colour of
|
||
<typeStatus id="907FA4813062C712FEE94020B0D6FCB1" box="[316,424,856,882]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
|
||
in life
|
||
</emphasis>
|
||
(see
|
||
<figureCitation id="D7FF06A63062C712FDE64020B306FCB1" box="[563,632,856,882]" captionStart="Fig" captionStartId="15.[189,232,1965,1991]" captionTargetBox="[189,1398,856,1923]" captionTargetId="figure@15.[189,1398,856,1923]" captionTargetPageId="15" captionText="Fig. 8. Pristimantis imthurni sp. nov. in life. A. Dorsolateral view of the ♂ holotype (IRSNB 4165). B. Ventral view of the same specimen. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3824972" httpUri="https://zenodo.org/record/3824972/files/figure.png" pageId="18" pageNumber="19">Fig. 8</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="4F7B1A233062C712FF6840FEB56CFB1E" blockId="18.[189,1399,902,1246]" pageId="18" pageNumber="19">Dorsal background colouration vivid yellow, top of head greyish brown; chocolate brown middorsal stripe and chocolate brown band on flanks; ill-defined, irregular reddish orange dorsolateral line; a few reddish orange blotches on the posterior part of the dorsum; a few reddish orange minute speckles on flanks; brown facemask; enlarged tubercle on eyelid whitish. Lower flanks yellow; groins, anterior thighs, and armpits brown. Arms and legs greyish brown with a few ill-defined white and dark yellow blotches. Ventral surfaces translucent greyish brown with a few irregular white spots and flecks; posterior surface of thighs and cloacal area brown. Upper face of hands and feet greyish brown with a few white and yellow markings; upper face of tip of fingers white; upper face of tip of Toes I-III white, brown on Toes IV-V. Palms, soles, undersurface of fingers and toes brown; discs greyish brown. Iris silver with dark brown venation and an ill-defined broad horizontal brownish stripe.</paragraph>
|
||
<paragraph id="4F7B1A233062C712FF68467CB274FADC" blockId="18.[189,778,1284,1311]" box="[189,778,1284,1311]" pageId="18" pageNumber="19">
|
||
<emphasis id="7DB0C6313062C712FF68467CB311FADD" bold="true" box="[189,623,1284,1311]" pageId="18" pageNumber="19">
|
||
Colour of
|
||
<typeStatus id="907FA4813062C712FEE9467DB0D0FADC" box="[316,430,1285,1311]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
|
||
in preservative
|
||
</emphasis>
|
||
(see
|
||
<figureCitation id="D7FF06A63062C712FD78467DB27FFADC" box="[685,769,1285,1311]" captionStart="Fig" captionStartId="17.[189,232,1966,1992]" captionTargetBox="[218,1361,1393,1894]" captionTargetId="figure@17.[191,1400,1380,1924]" captionTargetPageId="17" captionText="Fig. 10. Pristimantis imthurni sp. nov. in preservative. A. Dorsol view of the ♂ holotype (IRSNB 4165). B. Ventral view of the same specimen. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3824976" httpUri="https://zenodo.org/record/3824976/files/figure.png" pageId="18" pageNumber="19">Fig. 10</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="4F7B1A233062C712FF68464BB0BAFA3D" blockId="18.[189,1399,1331,1534]" pageId="18" pageNumber="19">
|
||
After 13 months in 70% ethanol (
|
||
<date id="3B7A3CE33062C712FD9E464BB3B8FA8E" box="[587,710,1331,1357]" pageId="18" pageNumber="19" value="2013-07">July 2013</date>
|
||
). Vivid yellow faded to pale yellow. Chocolate brown and brown faded to light brown; top of head is dark brown; ill-defined white and dark yellow blotches on arms and legs are slightly more conspicuous and form irregular, ill-defined transverse bands; dorsal pattern turned generally more conspicuous. Ventral colouration is light grey with a few white spots and flecks. Granules in the cloacal area, as well as subarticular and supernumerary tubercles became dark brown. Discs are grey.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="07DE49A83062C712FF68455CB180F883" pageId="18" pageNumber="19" type="distribution">
|
||
<paragraph id="4F7B1A233062C712FF68455CB08FF9FC" blockId="18.[189,497,1572,1599]" box="[189,497,1572,1599]" pageId="18" pageNumber="19">
|
||
<heading id="1433AD4F3062C712FF68455CB08FF9FC" bold="true" box="[189,497,1572,1599]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
|
||
<emphasis id="7DB0C6313062C712FF68455CB08FF9FC" bold="true" box="[189,497,1572,1599]" pageId="18" pageNumber="19">Distribution and ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="4F7B1A233062C712FF68452CB180F883" blockId="18.[189,1399,1619,1856]" pageId="18" pageNumber="19">
|
||
<taxonomicName id="88C461A03062C712FF68452CB0C7F9AE" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[189,441,1619,1645]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="imthurni" status="sp. nov.">
|
||
<emphasis id="7DB0C6313062C712FF68452CB0C7F9AE" box="[189,441,1619,1645]" italics="true" pageId="18" pageNumber="19">Pristimantis imthurni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A3062C712FE14452CB362F9AD" box="[449,540,1620,1646]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is only known from the summit of Ptari-tepui (
|
||
<figureCitation id="D7FF06A63062C712FB9E452BB5D4F9AE" box="[1099,1194,1619,1645]" captionStart="Fig" captionStartId="1.[189,232,1786,1812]" captionTargetBox="[276,1316,265,1733]" captionTargetId="figure@1.[276,1316,265,1733]" captionTargetPageId="1" captionText="Fig. 1. A. Map of the Pantepui region, located in the western part of the Guiana Shield. The white rectangle corresponds to the area enlarged in (B). B. Distribution of known tepui summit Pristimantis species occurring east of the Rio Caroní in Venezuela and in western Guyana: P. aureoventris (Wei- Assipu-tepui and upper slopes of Mount Roraima), P. yuruaniensis (Yuruaní-tepui), P. muchimuk (Churí-tepui), P. abakapa (Abakapá-tepui), P. auricarens and P. pulvinatus (Auyán-tepui); (1) Apradatepui, type locality of Pristimantis jamescameroni sp. nov. (area enlarged in Fig. 6A); (2) Ptari-tepui, type locality of Pristimantis imthurni sp. nov. (area enlarged in Fig. 6B)." figureDoi="http://doi.org/10.5281/zenodo.3824958" httpUri="https://zenodo.org/record/3824958/files/figure.png" pageId="18" pageNumber="19">Figs 1B</figureCitation>
|
||
,
|
||
<figureCitation id="D7FF06A63062C712FB6D452BB5A3F9AE" box="[1208,1245,1619,1645]" captionStart="Fig" captionStartId="11.[189,232,1896,1922]" captionTargetBox="[249,1344,268,1851]" captionTargetId="figure@11.[249,1344,268,1851]" captionTargetPageId="11" captionText="Fig. 6. A. Satellite map of the Aprada massif, showing the location of Aprada-tepui and neighbouring smaller table mountains. Yellow star indicates the type locality of Pristimantis jamescameroni. B. Satellite map of the Ptari massif, showing the location of Ptari-tepui and neighbouring mountains. Yellow star indicates the type locality of Pristimantis imthurni. Satellite maps © Google Maps." figureDoi="http://doi.org/10.5281/zenodo.3824968" httpUri="https://zenodo.org/record/3824968/files/figure.png" pageId="18" pageNumber="19">6B</figureCitation>
|
||
,
|
||
<figureCitation id="D7FF06A63062C712FB3C452BB479F9AE" box="[1257,1287,1619,1645]" captionStart="Fig" captionStartId="19.[189,232,1964,1990]" captionTargetBox="[189,1398,299,1923]" captionTargetId="figure@19.[189,1398,299,1923]" captionTargetPageId="19" captionText="Fig. 11. A. Ptari-tepui, photographed from the surrounding savannah looking to the NE (21 Jun. 2012). B. Summit of Ptari-tepui (21 Jun. 2012). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3824978" httpUri="https://zenodo.org/record/3824978/files/figure.png" pageId="18" pageNumber="19">11</figureCitation>
|
||
),
|
||
<collectingRegion id="8D00D4C13062C712FACA452BB188F952" country="Venezuela" name="Bolivar" pageId="18" pageNumber="19">Bolívar state</collectingRegion>
|
||
,
|
||
<collectingCountry id="37D35AB33062C712FED1450EB0FEF953" box="[260,384,1654,1680]" name="Venezuela" pageId="18" pageNumber="19">Venezuela</collectingCountry>
|
||
, where it occurs at
|
||
<quantity id="883CB7C63062C712FDA5450EB3B2F952" box="[624,716,1654,1681]" metricMagnitude="3" metricUnit="m" metricValue="2.471" pageId="18" pageNumber="19" unit="m" value="2471.0">2471 m</quantity>
|
||
elevation. Ptari-tepui lies in the Ptari massif, in the Los Testigos subdistrict (
|
||
<bibRefCitation id="2B5567D23062C712FE7A45E1B26DF977" author="McDiarmid & Donnelly" box="[431,787,1689,1716]" firstAuthor="McDiarmid" pageId="18" pageNumber="19" pagination="461 - 560" refId="ref11218" refString="McDiarmid R. W. & Donnelly M. A. 2005. The herpetofauna of the Guayana Highlands: amphibians and reptiles of the Lost World. In: Donnelly M. A., Crother B. I., Guyer C., Wake M. H. & White M. E. (eds) Ecology and Evolution in the Tropics: A Herpetological Perspective: 461 - 560. University of Chicago Press, Chicago." type="book chapter" year="2005">McDiarmid & Donnelly 2005</bibRefCitation>
|
||
). It is located
|
||
<emphasis id="7DB0C6313062C712FC6045E2B2A6F970" box="[949,984,1690,1715]" italics="true" pageId="18" pageNumber="19">ca.</emphasis>
|
||
<quantity id="883CB7C63062C712FC0A45E2B557F970" box="[991,1065,1689,1716]" metricMagnitude="4" metricUnit="m" metricValue="4.7" pageId="18" pageNumber="19" unit="km" value="47.0">47 km</quantity>
|
||
NE of the Chimantá massif, and
|
||
<emphasis id="7DB0C6313062C712FF3B45C6B06FF914" box="[238,273,1726,1751]" italics="true" pageId="18" pageNumber="19">ca.</emphasis>
|
||
<quantity id="883CB7C63062C712FEC245C5B01EF914" box="[279,352,1725,1751]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="18" pageNumber="19" unit="km" value="22.0">22 km</quantity>
|
||
SE of Kamarkawarai-tepui (airline). According to the GPS, Ptari-tepui reaches a maximal elevation of about
|
||
<quantity id="883CB7C63062C712FE434598B091F939" box="[406,495,1760,1786]" metricMagnitude="3" metricUnit="m" metricValue="2.471" pageId="18" pageNumber="19" unit="m" value="2471.0">2471 m</quantity>
|
||
above sea level. The summit area of Ptari-tepui is
|
||
<emphasis id="7DB0C6313062C712FBEE4599B520F939" box="[1083,1118,1761,1786]" italics="true" pageId="18" pageNumber="19">ca.</emphasis>
|
||
<quantity id="883CB7C63062C712FBB04598B5E1F939" box="[1125,1183,1760,1786]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="km" value="1.0">1 km</quantity>
|
||
<superScript id="B8B1B76B3062C712FB7545A6B5D7F92F" attach="left" box="[1184,1193,1758,1772]" fontSize="6" pageId="18" pageNumber="19">2</superScript>
|
||
and is dominated by flat open rock surfaces and small islands of low meadow-like tepui vegetation (
|
||
<bibRefCitation id="2B5567D23062C712FB73447BB447F8DD" author="Huber" box="[1190,1337,1795,1822]" firstAuthor="Huber" pageId="18" pageNumber="19" pagination="1 - 61" refId="ref10419" refString="Huber O. 1995. Geographical and physical features. In: Berry P. E., Holst B. K. & Yatskievych K. (eds) Flora of the Venezuelan Guayana. Volume 1. Introduction: 1 - 61. Missouri Botanical Garden Press, St Louis." type="book chapter" year="1995">Huber 1995</bibRefCitation>
|
||
;
|
||
<figureCitation id="D7FF06A63062C712FA9D447BB18EF883" captionStart="Fig" captionStartId="19.[189,232,1964,1990]" captionTargetBox="[189,1398,299,1923]" captionTargetId="figure@19.[189,1398,299,1923]" captionTargetPageId="19" captionText="Fig. 11. A. Ptari-tepui, photographed from the surrounding savannah looking to the NE (21 Jun. 2012). B. Summit of Ptari-tepui (21 Jun. 2012). Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3824978" httpUri="https://zenodo.org/record/3824978/files/figure.png" pageId="18" pageNumber="19">Fig. 11B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="07DE49A83062C714FF684410B455FE8A" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="discussion">
|
||
<paragraph id="4F7B1A233062C712FF684410B409F80A" blockId="18.[189,1399,1896,1993]" pageId="18" pageNumber="19">
|
||
The male
|
||
<typeStatus id="907FA4813062C712FEFA4410B0E8F841" box="[303,406,1896,1922]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
|
||
– and single specimen collected – was found at night while calling partly concealed in the muddy soil in a small vegetation patch. A few other males were heard calling very sporadically from other small vegetation islands by day or night, but could not be located.
|
||
<taxonomicName id="88C461A03062C712FBC244D7B46DF80B" authority="Kok, 2013" authorityName="Kok" authorityYear="2013" box="[1047,1299,1966,1992]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="imthurni" status="sp. nov.">
|
||
<emphasis id="7DB0C6313062C712FBC244D7B46DF80B" box="[1047,1299,1966,1992]" italics="true" pageId="18" pageNumber="19">Pristimantis imthurni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="66837B4A3062C712FACE44D7B409F80A" box="[1307,1399,1967,1993]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<caption id="1BBB4AAB3063C713FF6844D4B2A4F82A" ID-DOI="http://doi.org/10.5281/zenodo.3824978" ID-Zenodo-Dep="3824978" httpUri="https://zenodo.org/record/3824978/files/figure.png" pageId="19" pageNumber="20" startId="19.[189,232,1964,1990]" targetBox="[189,1398,299,1923]" targetPageId="19">
|
||
<paragraph id="4F7B1A233063C713FF6844D4B2A4F82A" blockId="19.[189,1399,1964,2026]" pageId="19" pageNumber="20">
|
||
<emphasis id="7DB0C6313063C713FF6844D4B04DF805" bold="true" box="[189,307,1964,1990]" pageId="19" pageNumber="20">Fig. 11. A</emphasis>
|
||
. Ptari-tepui, photographed from the surrounding savannah looking to the NE (21 Jun. 2012).
|
||
<emphasis id="7DB0C6313063C713FF6844A8B1AFF829" bold="true" box="[189,209,2000,2026]" pageId="19" pageNumber="20">B</emphasis>
|
||
. Summit of Ptari-tepui (21 Jun. 2012). Photographs by the author.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="4F7B1A233064C714FF684273B455FE8A" blockId="20.[189,1399,267,329]" pageId="20" pageNumber="21">
|
||
is not abundant. The only other anuran reported from the summit of Ptari-tepui is
|
||
<taxonomicName id="88C461A03064C714FB414273B3F4FE8A" authority="Ayarzaguena, Senaris & Gorzula, 1993" authorityName="Ayarzaguena, Senaris & Gorzula" authorityYear="1993" class="Amphibia" family="Hylidae" genus="Tepuihyla" kingdom="Animalia" order="Anura" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="rimarum">
|
||
<emphasis id="7DB0C6313064C714FB414273B409FEE6" box="[1172,1399,267,293]" italics="true" pageId="20" pageNumber="21">Tepuihyla rimarum</emphasis>
|
||
Ayarzagüena, Señaris & Gorzula, 1993
|
||
</taxonomicName>
|
||
, which apparently also occurs in low density (pers. obs.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |