254 lines
29 KiB
XML
254 lines
29 KiB
XML
<document id="DF7CAF45464FC8517B53FB5D729D9076" ID-CLB-Dataset="3261" ID-DOI="10.11646/phytotaxa.525.4.2" ID-GBIF-Dataset="086b1f35-7629-458d-84fc-cad50c34e236" ID-ISSN="1179-3163" ID-Zenodo-Dep="5723135" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637741515242" checkinUser="plazi" docAuthor="Jara-Muñoz, Orlando Adolfo, Richardson, James E. & Zabala-Rivera, Juan Carlos" docDate="2021" docId="7D6687F2FFC6FFB4BCA70A16650049F8" docLanguage="en" docName="phytotaxa.525.4.2.pdf" docOrigin="Phytotaxa 525 (4)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Begonia vinagrera Jara & Zabala-Rivera 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="276" masterDocId="815FFF8AFFD6FFA6BC2F0971662D4868" masterDocTitle="Five new species and three new varieties of Begonia section Casparya endemic to Colombia" masterLastPageNumber="280" masterPageNumber="258" pageNumber="274" updateTime="1699469805618" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="E4DF4BC35044D8421CDF14D7FBABDA9F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D8A7DA0F5BF5670FEDEFF0E1A4DD49F5">
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<mods:title id="538DC0C06537F60370573DB34DA461E3">Five new species and three new varieties of Begonia section Casparya endemic to Colombia</mods:title>
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</mods:titleInfo>
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<mods:name id="AEA8C1E56E4512388C26ECB0580CCA9F" type="personal">
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<mods:role id="4148AC35BF6B438AE43CC7489E3C5DE3">
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<mods:roleTerm id="BE1BE8F09293ABE1B1C672A74BA34729">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BF4EE81D356202EF307023887B873C2E">Jara-Muñoz, Orlando Adolfo</mods:namePart>
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<mods:affiliation id="07BA45EAAC9335D53F7B709442E5E1C1">Universidad Nacional de Colombia, Instituto de Ciencias Naturales, Cra. 30 No. 45 - 03, Edificio 425, Bogotá D. C. Colombia; & Jardín Botánico de Bogotá José Celestino Mutis, Bogotá D. C., Colombia</mods:affiliation>
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</mods:name>
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<mods:name id="FD5D4553ABC6B5567E0F4704D7D32830" type="personal">
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<mods:role id="6793E56744F7687CD3780296DCDBC9DA">
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<mods:roleTerm id="89E667CF0B484CEB06230EF172FD1DAF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4A66985D84D50930BE85460C994C1EA8">Richardson, James E.</mods:namePart>
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<mods:nameIdentifier id="30D586CEAFC618D4154974428D53E083" type="ORCID">0000-0001-9014-4865</mods:nameIdentifier>
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<mods:affiliation id="B225AD130ECAC507966E081553BFA94E">Departamento de Biología, Facultad de Ciencias Naturales, Universidad del Rosario, Bogotá D. C., Colombia. https: // orcid. org / 0000 - 0001 - 9014 - 4865 & Tropical Diversity Section, Royal Botanic Garden Edinburgh, United Kingdom.</mods:affiliation>
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</mods:name>
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<mods:name id="6A06ADBA29B4D017752EF8D54CF08E82" type="personal">
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<mods:role id="9A96EF57AEE7951886DA5CACBD7AC3C7">
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<mods:roleTerm id="885A5AF66AC660707CFEFBB97EA7C09B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0F1F6D0F1A5CF126C1836B519F900179">Zabala-Rivera, Juan Carlos</mods:namePart>
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<mods:nameIdentifier id="1A709971B6E4662349A5E17E9B67702D" type="ORCID">0000-0003-2679-5473</mods:nameIdentifier>
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<mods:affiliation id="82AFBC1C37F155D1AED5FE4DEDE96E85">Grupo Interdisciplinario en Ciencias (GICI), Universidad Pedagógica y Tecnológica de Colombia, Tunja, Colombia. https: // orcid. org / 0000 - 0003 - 2679 - 5473</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="2E07F216F25C71BBDF073DA23B2F1586">text</mods:typeOfResource>
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<mods:titleInfo id="66FBD20B265D80BD330F6A0F5E7AB4A6">
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<mods:title id="DED32238948118BB02F95DCE2689E585">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1AEF42C954AF4E5563A739AD3738CD27">
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<mods:date id="7C3A187CD3ABF427DB105E165C6B3EE3">2021</mods:date>
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<mods:detail id="E2941101F3B860F39890B3055A456262" type="pubDate">
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<mods:number id="912303922394F7007C510AC782F5D3F2">2021-11-23</mods:number>
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</mods:detail>
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<mods:detail id="519BBF5BABC1F9D95228FFFE826A90C3" type="volume">
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<mods:number id="EDEF1605EA6AD9430D7769000B721A88">525</mods:number>
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</mods:detail>
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<mods:detail id="C641E2072E7A4BA42604FFAA4DDA6557" type="issue">
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<mods:number id="7F03218EA2B0ED378FD4CA48A19DF1C4">4</mods:number>
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<mods:identifier id="E15F98D0C2C1222337EDF83F114F8042" type="DOI">10.11646/phytotaxa.525.4.2</mods:identifier>
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<mods:identifier id="4174364F0BB88C7A1DA99717701D17B0" type="GBIF-Dataset">086b1f35-7629-458d-84fc-cad50c34e236</mods:identifier>
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<treatment id="7D6687F2FFC6FFB4BCA70A16650049F8" ID-DOI="http://doi.org/10.5281/zenodo.5723251" ID-GBIF-Taxon="190872868" ID-Zenodo-Dep="5723251" LSID="urn:lsid:plazi:treatment:7D6687F2FFC6FFB4BCA70A16650049F8" httpUri="http://treatment.plazi.org/id/7D6687F2FFC6FFB4BCA70A16650049F8" lastPageId="18" lastPageNumber="276" pageId="16" pageNumber="274">
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<subSubSection id="BDD5656FFFC6FFB6BCA70A16655F4BEA" box="[136,882,871,898]" pageId="16" pageNumber="274" type="nomenclature">
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<paragraph id="F57036E4FFC6FFB6BCA70A16655F4BEA" blockId="16.[136,882,871,898]" box="[136,882,871,898]" pageId="16" pageNumber="274">
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<heading id="AE388188FFC6FFB6BCA70A16655F4BEA" box="[136,882,871,898]" fontSize="11" level="2" pageId="16" pageNumber="274" reason="3">
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<taxonomicName id="32CF4D67FFC6FFB6BCA70A1664774BE9" ID-CoL="8TK4H" authority="Jara & Zabala-Rivera" authorityName="Jara & Zabala-Rivera" authorityYear="2021" box="[136,602,871,898]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="16" pageNumber="274" phylum="Tracheophyta" rank="species" species="vinagrera" status="sp. nov.">
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<emphasis id="C7BBEAF6FFC6FFB6BCA70A16674D4BE9" bold="true" box="[136,352,871,897]" italics="true" pageId="16" pageNumber="274">Begonia vinagrera</emphasis>
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Jara & Zabala-Rivera
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</taxonomicName>
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,
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<taxonomicNameLabel id="DC88578DFFC6FFB6BE4A0A1964914BEA" box="[613,700,872,898]" pageId="16" pageNumber="274" rank="species">
|
||
<emphasis id="C7BBEAF6FFC6FFB6BE4A0A1964994BE9" box="[613,692,872,897]" italics="true" pageId="16" pageNumber="274">sp. nov</emphasis>
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||
.
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||
</taxonomicNameLabel>
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||
<figureCitation id="6DF42A61FFC6FFB6BEEC0A16653D4BEA" box="[707,784,871,898]" captionStart="FIGURE 8" captionStartId="17.[136,229,1961,1983]" captionTargetBox="[168,1429,211,1913]" captionTargetId="figure-27@17.[151,1436,190,1937]" captionTargetPageId="17" captionText="FIGURE 8. Begonia vinagrera var. vinagrera. A) Branch with flowers, B) Staminate flower, C) Pistillate flower showing perianth, ovary, and bracteoles, D) Fruit. Based on J. Betancur 5805, J.L. Fernandez-Alonso 6250, and pictures of live plants, by O. Bernal." figureDoi="http://doi.org/10.5281/zenodo.5723153" httpUri="https://zenodo.org/record/5723153/files/figure.png" pageId="16" pageNumber="274">Figs. 8</figureCitation>
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,
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<figureCitation id="6DF42A61FFC6FFB6BF320A1965004BEA" box="[797,813,872,898]" captionStart="FIGURE 9" captionStartId="19.[136,229,1991,2013]" captionTargetBox="[253,1324,203,1954]" captionTargetId="figure-27@19.[162,1426,190,1966]" captionTargetPageId="19" captionText="FIGURE 9. Begonia vinagrera var. pomecensis. A) Branch, B) Staminate flower, C) Pistillate flower, D) Fruit, E) Cross section of the fruit. Based on J.C. Zabala-Rivera et al. 2693 and pictures of live plants, by O. Bernal." figureDoi="http://doi.org/10.5281/zenodo.5723156" httpUri="https://zenodo.org/record/5723156/files/figure.png" pageId="16" pageNumber="274">9</figureCitation>
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, & 10
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="BDD5656FFFC6FFB6BCA70ADC647B4B8F" pageId="16" pageNumber="274" type="materials_examined">
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<paragraph id="F57036E4FFC6FFB6BCA70ADC647B4B8F" blockId="16.[136,1451,940,999]" pageId="16" pageNumber="274">
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<materialsCitation id="45A73CB9FFC6FFB6BCA70ADC647B4B8F" ID-GBIF-Occurrence="3412417345" collectingDate="1994-11-23" collectionCode="J" collectorName="J. Betancur" country="Colombia" elevation="2300" location="Santander" pageId="16" pageNumber="274" specimenCount="1" stateProvince="Santander" typeStatus="holotype">
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<typeStatus id="2A748846FFC6FFB6BCA70ADC66974BAA" box="[136,186,941,962]" pageId="16" pageNumber="274">Type</typeStatus>
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:—
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<collectingCountry id="8DD87674FFC6FFB6BCF90ADC67774BAA" box="[214,346,941,962]" name="Colombia" pageId="16" pageNumber="274">COLOMBIA</collectingCountry>
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.
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<collectingRegion id="370BF806FFC6FFB6BD4A0ADD67E44BAA" box="[357,457,940,962]" country="Colombia" name="Santander" pageId="16" pageNumber="274">Santander</collectingRegion>
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: Municipio Charalá, sitio
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||
<collectorName id="583A5332FFC6FFB6BEE20ADD65754BAA" box="[717,856,940,962]" pageId="16" pageNumber="274">El Bogotacito</collectorName>
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, después
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<collectorName id="583A5332FFC6FFB6BF960ADD626C4BAA" box="[953,1089,940,962]" pageId="16" pageNumber="274">de El Taladro</collectorName>
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, vía
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<collectorName id="583A5332FFC6FFB6B85B0ADD63014BAA" box="[1140,1324,940,962]" pageId="16" pageNumber="274">El Carmen-Virolín</collectorName>
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,
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||
<quantity id="32379B01FFC6FFB6B9160ADC63AA4BAA" box="[1337,1415,941,962]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="16" pageNumber="274" unit="m" value="2300.0">
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||
<elevation id="7EE2D1D7FFC6FFB6B9160ADC63AA4BAA" box="[1337,1415,941,962]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="16" pageNumber="274" unit="m" value="2300.0">2300 m</elevation>
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||
</quantity>
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||
,
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<date id="81711024FFC6FFB6B9BC0ADC67764B8E" pageId="16" pageNumber="274" value="1994-11-23">
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<collectingDate id="9135E9CCFFC6FFB6B9BC0ADC67764B8E" pageId="16" pageNumber="274" value="1994-11-23">23 November 1994</collectingDate>
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||
</date>
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||
,
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||
<emphasis id="C7BBEAF6FFC6FFB6BD4A0AA064244B8E" box="[357,521,977,998]" italics="true" pageId="16" pageNumber="274">
|
||
<collectionCode id="93DEAE21FFC6FFB6BD4A0AA0675E4B8E" box="[357,371,977,998]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="16" pageNumber="274" type="Herbarium">J</collectionCode>
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. Betancur 5805
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</emphasis>
|
||
(
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<collectionCode id="93DEAE21FFC6FFB6BE390AA064664B8F" box="[534,587,977,999]" country="Colombia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15629" lsid="urn:lsid:biocol.org:col:15629" name="Universidad Nacional de Colombia" pageId="16" pageNumber="274" type="Herbarium">COL</collectionCode>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="BDD5656FFFC6FFB6BCA70D7C644C4C07" pageId="16" pageNumber="274" type="diagnosis">
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<paragraph id="F57036E4FFC6FFB6BCA70D7C644C4C07" blockId="16.[136,1452,1037,2086]" pageId="16" pageNumber="274">
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<emphasis id="C7BBEAF6FFC6FFB6BCA70D7C66D64C4F" bold="true" box="[136,251,1037,1063]" pageId="16" pageNumber="274">Diagnosis</emphasis>
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:—
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<taxonomicName id="32CF4D67FFC6FFB6BD300D7F67C24C4F" authorityName="Jara & Zabala-Rivera" authorityYear="2021" box="[287,495,1038,1063]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="16" pageNumber="274" phylum="Tracheophyta" rank="species" species="vinagrera">
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<emphasis id="C7BBEAF6FFC6FFB6BD300D7F67C24C4F" box="[287,495,1038,1063]" italics="true" pageId="16" pageNumber="274">Begonia vinagrera</emphasis>
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</taxonomicName>
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is morphologically similar to
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<taxonomicName id="32CF4D67FFC6FFB6BF160D7F65834C4F" box="[825,942,1038,1063]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="16" pageNumber="274" phylum="Tracheophyta" rank="species" species="cornuta">
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<emphasis id="C7BBEAF6FFC6FFB6BF160D7F65834C4F" box="[825,942,1038,1063]" italics="true" pageId="16" pageNumber="274">B. cornuta</emphasis>
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</taxonomicName>
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but can be easily distinguished by its glabrous leaves (vs. pubescent) and oblong (vs. ovate) leaf laminae, and the inflorescence that is a monochasium with few flowers (vs. dichasial with many flowers).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="BDD5656FFFC6FFB6BC920D0665BE4F39" pageId="16" pageNumber="274" type="description">
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||
<paragraph id="F57036E4FFC6FFB6BC920D0665BE4F39" blockId="16.[136,1452,1037,2086]" pageId="16" pageNumber="274">
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<emphasis id="C7BBEAF6FFC6FFB6BC920D06676A4CF9" bold="true" box="[189,327,1143,1169]" pageId="16" pageNumber="274">Description</emphasis>
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:—Caulescent herb, to
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||
<quantity id="32379B01FFC6FFB6BE7E0D0664534CFA" box="[593,638,1143,1170]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="16" pageNumber="274" unit="m" value="2.0">2 m</quantity>
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high.
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<emphasis id="C7BBEAF6FFC6FFB6BEE60D0664D24CF9" box="[713,767,1143,1169]" italics="true" pageId="16" pageNumber="274">Stem</emphasis>
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||
erect, internodes to
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||
<quantity id="32379B01FFC6FFB6BFC20D0662164CFA" box="[1005,1083,1143,1170]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="16" pageNumber="274" unit="cm" value="7.4">7.4 cm</quantity>
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||
long, to
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||
<quantity id="32379B01FFC6FFB6B8890D0662D34CFA" box="[1190,1278,1143,1170]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="16" pageNumber="274" unit="mm" value="9.6">9.6 mm</quantity>
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thick, reddishgreen, glabrous.
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<emphasis id="C7BBEAF6FFC6FFB6BD6E0DEB67B74CDC" box="[321,410,1178,1204]" italics="true" pageId="16" pageNumber="274">Stipules</emphasis>
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||
sub-persistent or early deciduous, lanceolate, oblong or ovate, 6.6–22.9 ×
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||
<quantity id="32379B01FFC6FFB6B8F90DEB63414CDD" box="[1238,1388,1178,1205]" metricMagnitude="-3" metricUnit="m" metricValue="7.6499999999999995" metricValueMax="11.7" metricValueMin="3.6" pageId="16" pageNumber="274" unit="mm" value="7.6499999999999995" valueMax="11.7" valueMin="3.6">3.6–11.7 mm</quantity>
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||
, apex cuspidate, obtuse or acute, light green or reddish-green, glabrous, margin entire.
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||
<emphasis id="C7BBEAF6FFC6FFB6B80D0DCC625C4CBF" box="[1058,1137,1213,1239]" italics="true" pageId="16" pageNumber="274">Petiole</emphasis>
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||
<quantity id="32379B01FFC6FFB6B8550DCC62D44CB0" box="[1146,1273,1213,1240]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="5.5" metricValueMin="0.9" pageId="16" pageNumber="274" unit="cm" value="3.2" valueMax="5.5" valueMin="0.9">0.9–5.5 cm</quantity>
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||
long, glabrous;
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||
<emphasis id="C7BBEAF6FFC6FFB6BCA70D9066FB4C93" box="[136,214,1249,1275]" italics="true" pageId="16" pageNumber="274">lamina</emphasis>
|
||
of the leaf blade straight to the petiole, coriaceous, strongly asymmetric, oblong or oblong-lanceolate, 3.6−14.9 ×
|
||
<quantity id="32379B01FFC6FFB6BC8F0C75670E4D76" box="[160,291,1284,1311]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" metricValueMax="6.5" metricValueMin="0.9" pageId="16" pageNumber="274" unit="cm" value="3.7" valueMax="6.5" valueMin="0.9">0.9−6.5 cm</quantity>
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||
, base unequal, rounded to acute on one side and auriculate on the other side, margin serrate to double serrate, apex acuminate to long acuminate, glabrous on both sides, venation pinnate, 5–8 veined from the base on each side.
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||
<emphasis id="C7BBEAF6FFC6FFB6BCEB0C3B67774D0C" box="[196,346,1354,1380]" italics="true" pageId="16" pageNumber="274">Inflorescence</emphasis>
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||
a dichasium with 2–4 branches, axillary, erect, bisexual or only staminate, protandrous; peduncle 3.1−8.3 (14.3) cm long, glabrous, bracts early deciduous or sub-persistent, glabrous, light green or reddish-green, ovate or obovate, 4.4–7.1 ×
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||
<quantity id="32379B01FFC6FFB6BD920CE164684DC3" box="[445,581,1424,1451]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.7" metricValueMin="2.6" pageId="16" pageNumber="274" unit="mm" value="3.15" valueMax="3.7" valueMin="2.6">2.6–3.7 mm</quantity>
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||
, margin entire, apex obtuse or cuspidate.
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||
<emphasis id="C7BBEAF6FFC6FFB6B8200CE062F94DC3" box="[1039,1236,1425,1451]" italics="true" pageId="16" pageNumber="274">staminate flowers</emphasis>
|
||
: pedicels glabrous, white to reddish,
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||
<quantity id="32379B01FFC6FFB6BD680CC567F14DA6" box="[327,476,1460,1486]" metricMagnitude="-3" metricUnit="m" metricValue="7.449999999999999" metricValueMax="13.2" metricValueMin="1.7" pageId="16" pageNumber="274" unit="mm" value="7.449999999999999" valueMax="13.2" valueMin="1.7">1.7−13.2 mm</quantity>
|
||
long; tepals 4, spreading, outer 2 glabrous, white, sometimes pinkish variegated, oblong or ovate, 6.3–13.6 ×
|
||
<quantity id="32379B01FFC6FFB6BD410CA664294D9A" box="[366,516,1495,1522]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="10.1" metricValueMin="3.5" pageId="16" pageNumber="274" unit="mm" value="6.8" valueMax="10.1" valueMin="3.5">3.5–10.1 mm</quantity>
|
||
, margin entire, apex rounded, inner 2 glabrous, white, elliptic, oblong-lanceolate, or linear-lanceolate, 4.2–8.7 ×
|
||
<quantity id="32379B01FFC6FFB6BDEC0C8B64604E7D" box="[451,589,1530,1557]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.8" metricValueMin="1.2" pageId="16" pageNumber="274" unit="mm" value="2.0" valueMax="2.8" valueMin="1.2">1.2–2.8 mm</quantity>
|
||
, margin entire, apex rounded or short acuminate; stamens 12–15, projecting, yellow, filaments
|
||
<quantity id="32379B01FFC6FFB6BD750F6F67CB4E50" box="[346,486,1566,1592]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.8" metricValueMin="0.2" pageId="16" pageNumber="274" unit="mm" value="1.0" valueMax="1.8" valueMin="0.2">0.2–1.8 mm</quantity>
|
||
long, on a short column of
|
||
<quantity id="32379B01FFC6FFB6BF6F0F6F65E34E50" box="[832,974,1566,1592]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="16" pageNumber="274" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 mm</quantity>
|
||
, anthers oblong, 1.4–2.2 ×
|
||
<quantity id="32379B01FFC6FFB6B9360F6C638A4E50" box="[1305,1447,1565,1592]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="16" pageNumber="274" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.4–0.6 mm</quantity>
|
||
, connectives scarcely projecting, to
|
||
<quantity id="32379B01FFC6FFB6BE390F30645D4E33" box="[534,624,1601,1627]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="16" pageNumber="274" unit="mm" value="0.4">0.4 mm</quantity>
|
||
.
|
||
<emphasis id="C7BBEAF6FFC6FFB6BE540F3065174E33" box="[635,826,1601,1627]" italics="true" pageId="16" pageNumber="274">Pistillate flowers</emphasis>
|
||
: pedicels glabrous, white to reddish-white, to
|
||
<quantity id="32379B01FFC6FFB6B9680F3063814E33" box="[1351,1452,1601,1627]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="16" pageNumber="274" unit="mm" value="13.7">13.7 mm</quantity>
|
||
long; bracteoles 2, glabrous, reddish, ovate or obovate, 5.4–9.8 ×
|
||
<quantity id="32379B01FFC6FFB6BF730F1565C94E16" box="[860,996,1636,1662]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="7.6" metricValueMin="4.6" pageId="16" pageNumber="274" unit="mm" value="6.1" valueMax="7.6" valueMin="4.6">4.6–7.6 mm</quantity>
|
||
, apex obtuse and 1-setose, margin entire; ovary glabrous, turbinate, 9.7–16 ×
|
||
<quantity id="32379B01FFC6FFB6BE060FF664964ECA" box="[553,699,1671,1698]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" metricValueMax="2.7" metricValueMin="1.18" pageId="16" pageNumber="274" unit="mm" value="19.4" valueMax="27.0" valueMin="11.8">11.8–27 mm</quantity>
|
||
, green, equally 3-horned, horns slightly curved upward, placenta branches simple, bearing ovules on both surfaces; perianth with 5 spreading tepals, glabrous, white, and sometimes red and white variegated, the outer longer, obovate, 6–18.9 ×
|
||
<quantity id="32379B01FFC6FFB6BF240FBC65A14E80" box="[779,908,1741,1768]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" metricValueMax="11.6" metricValueMin="3.0" pageId="16" pageNumber="274" unit="mm" value="7.3" valueMax="11.6" valueMin="3.0">3–11.6 mm</quantity>
|
||
, margin entire, apex obtuse, the smallest elliptic, 5–5.5 ×
|
||
<quantity id="32379B01FFC6FFB6BCC90F8167424F63" box="[230,367,1776,1803]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.4" metricValueMin="2.5" pageId="16" pageNumber="274" unit="mm" value="2.95" valueMax="3.4" valueMin="2.5">2.5–3.4 mm</quantity>
|
||
, margin entire, apex acute, rounded or short acuminate; styles multifid, yellow, fused toward the base,
|
||
<quantity id="32379B01FFC6FFB6BCE70E6567084F46" box="[200,293,1812,1838]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="16" pageNumber="274" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
|
||
long, stigmatic papillae restricted to the apex.
|
||
<emphasis id="C7BBEAF6FFC6FFB6BF1E0E6565A04F46" box="[817,909,1812,1838]" italics="true" pageId="16" pageNumber="274">Capsule</emphasis>
|
||
body and horns shape like those of the ovary, to 21 ×
|
||
<quantity id="32379B01FFC6FFB6BCED0E46673C4F3A" box="[194,273,1847,1874]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="16" pageNumber="274" unit="mm" value="33.0">33 mm</quantity>
|
||
when mature, drying maroon, column
|
||
<quantity id="32379B01FFC6FFB6BEEA0E46657E4F3A" box="[709,851,1847,1874]" metricMagnitude="-2" metricUnit="m" metricValue="1.4449999999999998" metricValueMax="1.89" metricValueMin="1.0" pageId="16" pageNumber="274" unit="mm" value="14.45" valueMax="18.9" valueMin="10.0">10–18.9 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BDD5656FFFC6FFB6BC920E2B65234FF0" pageId="16" pageNumber="274" type="etymology">
|
||
<paragraph id="F57036E4FFC6FFB6BC920E2B65234FF0" blockId="16.[136,1452,1037,2086]" pageId="16" pageNumber="274">
|
||
<emphasis id="C7BBEAF6FFC6FFB6BC920E2B67134F1C" bold="true" box="[189,318,1882,1908]" pageId="16" pageNumber="274">Etymology</emphasis>
|
||
:—The specific epithet refers to the common name of this species in the department of
|
||
<collectingRegion id="370BF806FFC6FFB6B91A0E2B638A4F1C" box="[1333,1447,1882,1908]" country="Colombia" name="Santander" pageId="16" pageNumber="274">Santander</collectingRegion>
|
||
, where it is known as Vinagrera in Spanish (
|
||
<emphasis id="C7BBEAF6FFC6FFB6BE5F0E0C64D34FF0" box="[624,766,1917,1944]" italics="true" pageId="16" pageNumber="274">Agudelo 170</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BDD5656FFFC6FFB4BC920ED1650049F8" lastPageId="18" lastPageNumber="276" pageId="16" pageNumber="274" type="distribution">
|
||
<paragraph id="F57036E4FFC6FFB6BC920ED162ED404D" blockId="16.[136,1452,1037,2086]" pageId="16" pageNumber="274">
|
||
<emphasis id="C7BBEAF6FFC6FFB6BC920ED167C94FD2" bold="true" box="[189,484,1952,1979]" pageId="16" pageNumber="274">Distribution and habitat</emphasis>
|
||
:—The species grows in cloud forest at the border of the departments of
|
||
<collectingRegion id="370BF806FFC6FFB6B9090ED063574FD3" box="[1318,1402,1953,1979]" country="Colombia" name="Boyaca" pageId="16" pageNumber="274">Boyacá</collectingRegion>
|
||
and
|
||
<collectingRegion id="370BF806FFC6FFB6BCA70EB567034FB6" box="[136,302,1988,2014]" country="Colombia" name="Cundinamarca" pageId="16" pageNumber="274">Cundinamarca</collectingRegion>
|
||
, with
|
||
<taxonomicName id="32CF4D67FFC6FFB6BD5A0EB464874FB6" authority="var. vinagrera" authorityName="var. vinagrera" authorityYear="2021" box="[373,682,1989,2014]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="16" pageNumber="274" phylum="Tracheophyta" rank="variety" species="vinagrera" variety="vinagrera">
|
||
<emphasis id="C7BBEAF6FFC6FFB6BD5A0EB464874FB6" box="[373,682,1989,2014]" italics="true" pageId="16" pageNumber="274">B. vinagrera var. vinagrera</emphasis>
|
||
</taxonomicName>
|
||
restricted to the locality of Virolín and surrounded areas, mainly in the department of
|
||
<collectingRegion id="370BF806FFC6FFB6BD780E9667E44069" box="[343,457,2023,2049]" country="Colombia" name="Santander" pageId="16" pageNumber="274">Santander</collectingRegion>
|
||
, between 1700 and
|
||
<quantity id="32379B01FFC6FFB6BE850E96652F406A" box="[682,770,2023,2050]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="274" unit="m" value="2500.0">2500 m</quantity>
|
||
, and
|
||
<taxonomicName id="32CF4D67FFC6FFB6BF100E9962AB4069" authorityName="Jara-Muñoz & Richardson & Zabala-Rivera" authorityYear="2021" box="[831,1158,2024,2049]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="16" pageNumber="274" phylum="Tracheophyta" rank="variety" species="vinagrera" variety="pomecensis">
|
||
<emphasis id="C7BBEAF6FFC6FFB6BF100E9962AB4069" box="[831,1158,2024,2049]" italics="true" pageId="16" pageNumber="274">B. vinagrera var. pomecensis</emphasis>
|
||
</taxonomicName>
|
||
exclusive to the valley of the Pomeca river in
|
||
<collectingRegion id="370BF806FFC6FFB6BD45017A67EF404D" box="[362,450,2059,2085]" country="Colombia" name="Boyaca" pageId="16" pageNumber="274">Boyacá</collectingRegion>
|
||
. More details can be consulted under the description of each variety.
|
||
</paragraph>
|
||
<caption id="A1B0666CFFC7FFB7BCA70ED863064F8B" ID-DOI="http://doi.org/10.5281/zenodo.5723153" ID-Zenodo-Dep="5723153" httpUri="https://zenodo.org/record/5723153/files/figure.png" pageId="17" pageNumber="275" startId="17.[136,229,1961,1983]" targetBox="[168,1429,211,1913]" targetPageId="17">
|
||
<paragraph id="F57036E4FFC7FFB7BCA70ED863064F8B" blockId="17.[136,1452,1961,2019]" pageId="17" pageNumber="275">
|
||
<emphasis id="C7BBEAF6FFC7FFB7BCA70ED866DA4FD6" bold="true" box="[136,247,1961,1983]" pageId="17" pageNumber="275">FIGURE 8</emphasis>
|
||
.
|
||
<taxonomicName id="32CF4D67FFC7FFB7BD2D0EDB646F4FD7" authority="var. vinagrera" authorityName="var. vinagrera" authorityYear="2021" box="[258,578,1962,1983]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="17" pageNumber="275" phylum="Tracheophyta" rank="variety" species="vinagrera" variety="vinagrera">
|
||
<emphasis id="C7BBEAF6FFC7FFB7BD2D0EDB646F4FD7" box="[258,578,1962,1983]" italics="true" pageId="17" pageNumber="275">Begonia vinagrera var. vinagrera</emphasis>
|
||
</taxonomicName>
|
||
. A) Branch with flowers, B) Staminate flower, C) Pistillate flower showing perianth, ovary, and bracteoles, D) Fruit. Based on
|
||
<emphasis id="C7BBEAF6FFC7FFB7BDF80EBF64564F8A" box="[471,635,1997,2019]" italics="true" pageId="17" pageNumber="275">J. Betancur 5805</emphasis>
|
||
,
|
||
<emphasis id="C7BBEAF6FFC7FFB7BEA80EBF65B44F8A" box="[647,921,1997,2019]" italics="true" pageId="17" pageNumber="275">J.L. Fernandez-Alonso 6250</emphasis>
|
||
, and pictures of live plants, by O. Bernal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="F57036E4FFC4FFB4BC9209EF659E494D" blockId="18.[136,1452,158,400]" pageId="18" pageNumber="276">
|
||
<emphasis id="C7BBEAF6FFC4FFB4BC9209EF679A48D0" bold="true" box="[189,439,158,184]" pageId="18" pageNumber="276">Conservation status:</emphasis>
|
||
―According to the criterion B, with AOO =
|
||
<quantity id="32379B01FFC4FFB4BFE409EF623848D0" box="[971,1045,158,184]" metricMagnitude="4" metricUnit="m" metricValue="4.3" pageId="18" pageNumber="276" unit="km" value="43.0">43 km</quantity>
|
||
<superScript id="02BA9BACFFC4FFB4B83A09EF623048C4" attach="left" box="[1045,1053,158,172]" fontSize="6" pageId="18" pageNumber="276">2</superScript>
|
||
, growing in a region not severely fragmented, and partially protected by the Santuario de Flora y Fauna de Iguaque, but known from no more than five locations (criterion B2a), and with an observed decline in the AOO and quality of habitat (criterion B2b(iii)), we conclude that this species is in the category Endangered (EN B2a,b(iii)).
|
||
</paragraph>
|
||
<paragraph id="F57036E4FFC4FFB4BC92085F650049F8" blockId="18.[136,1452,158,400]" pageId="18" pageNumber="276">
|
||
<emphasis id="C7BBEAF6FFC4FFB4BC92085F672D4920" bold="true" box="[189,256,302,328]" pageId="18" pageNumber="276">Notes</emphasis>
|
||
:—Specimens of this species (both varieties) have been frequently misidentified in herbaria as
|
||
<taxonomicName id="32CF4D67FFC4FFB4B961085E66F24904" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="18" pageNumber="276" phylum="Tracheophyta" rank="species" species="cornuta">
|
||
<emphasis id="C7BBEAF6FFC4FFB4B961085E66F24904" italics="true" pageId="18" pageNumber="276">Begonia cornuta</emphasis>
|
||
</taxonomicName>
|
||
; however, both species can be separated as indicated in the diagnosis. A synthetic description of both varieties, including only distinguishing characters, is presented below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |