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<document id="E0669960656AC44EBBB85B5B9D3E1368" ID-CLB-Dataset="21602" ID-DOI="10.5852/ejt.2020.674" ID-GBIF-Dataset="a4358cde-3ef0-4e08-9142-06ca894eefa4" ID-ISSN="2118-9773" ID-Zenodo-Dep="3908375" ID-ZooBank="DF72EEF8-9C29-4A73-89FC-228392709427" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593098001284" checkinUser="valdenar" docAuthor="Pholyotha, Arthit, Sutcharit, Chirasak, Thach, Phanara, Chhuoy, Samol, Ngor, Peng Bun &amp; Panha, Somsak" docDate="2020" docId="CE49BC756A26FFE3FE6CFCFFBA3F3998" docLanguage="en" docName="ejt-674_pholyotha_sutchart_thach.pdf.imf" docOrigin="European Journal of Taxonomy 674" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sarika lactoconcha Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha 2020, sp. nov." docType="treatment" docUuid="0E48F6FB-6775-4470-BF85-345C3D43112A" docUuidSource="ZooBank" docVersion="6" lastPageNumber="10" masterDocId="3270C40D6A21FFE9FF92FFC0BE7C3817" masterDocTitle="Land snail genus Sarika Godwin-Austen, 1907 (Eupulmonata: Ariophantidae) from Cambodia, with description of three new species" masterLastPageNumber="21" masterPageNumber="1" pageNumber="8" updateTime="1698842924734" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F3EDB2D231BC2632B39C8C001F71B7D2">Land snail genus Sarika Godwin-Austen, 1907 (Eupulmonata: Ariophantidae) from Cambodia, with description of three new species</mods:title>
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<mods:namePart id="926D7A322E78569AFE68D7B95BA3422C">Pholyotha, Arthit</mods:namePart>
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<mods:affiliation id="64A2CD2D79DCD4F357D841D1FE62F9E8">Biological Sciences Program, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Email: arthitpolyotha @ gmail. com &amp; urn: lsid: zoobank. org: author: 6 CC 9 B 5 FE- 6586 - 4132 - 8289 - 102 DC 14 D 3844</mods:affiliation>
<mods:nameIdentifier id="75BDBA81BE2DE7B336BEE7FCC063A380" type="email">arthitpolyotha@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D2BC32E4A28636F6DB739BDA8ED3DA8B">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="1D59C2BFE0DDA3728DBD91457873EB77">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Email: jirasak 4 @ yahoo. com &amp; urn: lsid: zoobank. org: author: C 2 E 2 FA 6 B-A 3 F 9 - 4 F 33 - B 447 - B 59 B 1 BD 322 D 4</mods:affiliation>
<mods:nameIdentifier id="F211B8AF3D4F48C3174AA2BBFC18B16D" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="F67C24AE806172CC8864A33DA5D6FED0">Thach, Phanara</mods:namePart>
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<mods:affiliation id="992F6EA762527B9CA6F767B97A05CDD1">Inland Fisheries Research and Development Institute (IFReDI), Fisheries Administration Phnom Penh, Cambodia. &amp; Email: thachphanara @ yahoo. com &amp; urn: lsid: zoobank. org: author: CA 9 CDCEC-E 5 BB- 452 A- 9 C 6 C- 18 F 3280 AAC 02</mods:affiliation>
<mods:nameIdentifier id="E9E38819E1611628092211091C1F0DCA" type="email">thachphanara@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="F7A1616A1D4D25D9E01E6F6ABF1ED1D5">Chhuoy, Samol</mods:namePart>
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<mods:affiliation id="641DC3B7AE0F2480EAEEAE582B24FCFF">Inland Fisheries Research and Development Institute (IFReDI), Fisheries Administration Phnom Penh, Cambodia. &amp; Email: chhuoysamol @ yahoo. com &amp; urn: lsid: zoobank. org: author: 2674 B 4 EE- 7 B 30 - 4285 - 8543 - A 46 FD 1 E 5 B 53 E</mods:affiliation>
<mods:nameIdentifier id="16D88CA3C580057C2BC42E24C46D834C" type="email">chhuoysamol@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="761E037999E50DEF6B2C66F5C0BD58AB">Ngor, Peng Bun</mods:namePart>
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<mods:affiliation id="0357CFE890A541CE987929235D2B0B91">Inland Fisheries Research and Development Institute (IFReDI), Fisheries Administration Phnom Penh, Cambodia. &amp; Wonders of the Mekong Project, c / o IFReDI, # 186, Norodom Blvd., Phnom Penh, Cambodia. &amp; Email: pengbun. ngor @ gmail. com &amp; urn: lsid: zoobank. org: author: 77 BC 565 D- 7067 - 46 C 6 - 8744 - A 0 C 50 A 6 E 2 B 3 F</mods:affiliation>
<mods:nameIdentifier id="7ADC4B54FD0B14C81D29988B21CF8A6A" type="email">pengbun.ngor@gmail.com</mods:nameIdentifier>
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<mods:namePart id="91C5DCA7825A86BDA653D4298D115F65">Panha, Somsak</mods:namePart>
<mods:affiliation id="D4246E886F3913EA128C10E0E110BFA0">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79 &amp; Corresponding author: somsak. pan @ chula. ac. th</mods:affiliation>
<mods:nameIdentifier id="526E240920D8FD3322AA252E0F84F4E2" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:title id="44B1F48C2F49082574886910FE0C863A">European Journal of Taxonomy</mods:title>
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<mods:date id="36FB2439EC3B1022E21A0BE55B551A4A">2020</mods:date>
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<mods:number id="CB00D79C1827A04F767A6FE0083FED05">2020-06-25</mods:number>
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<mods:number id="2922AA6881E73968A56993A2A9713EF5">674</mods:number>
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<mods:start id="A4AD792791B8C18E54BC2B10B4141091">1</mods:start>
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<mods:classification id="1B04DD45E3EBC90AE3BC57EA183DE87B">journal article</mods:classification>
<mods:identifier id="94127855F64D34311E38B9F309D09095" type="CLB-Dataset">21602</mods:identifier>
<mods:identifier id="AD7CFFC01F5DC0520473FF4E9063DB37" type="DOI">10.5852/ejt.2020.674</mods:identifier>
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<treatment id="CE49BC756A26FFE3FE6CFCFFBA3F3998" ID-GBIF-Taxon="164743218" LSID="urn:lsid:zoobank.org:act:0E48F6FB-6775-4470-BF85-345C3D43112A" httpUri="http://treatment.plazi.org/id/CE49BC756A26FFE3FE6CFCFFBA3F3998" lastPageId="10" lastPageNumber="10" pageId="7" pageNumber="8">
<subSubSection id="0EFA5EE86A26FFEEFE6CFCFFBA4A3B4D" box="[510,1078,831,858]" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="465F0D636A26FFEEFE6CFCFFBA4A3B4D" blockId="7.[376,1213,831,928]" box="[510,1078,831,858]" pageId="7" pageNumber="8">
<heading id="1D17BA0F6A26FFEEFE6CFCFFBA4A3B4D" box="[510,1078,831,858]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<taxonomicName id="81E076E06A26FFEEFE6CFCFFBDA83B4D" ID-CoL="7T3FL" authority="Pholyotha &amp; Panha" authorityName="Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha" authorityYear="2020" box="[510,980,831,858]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="lactoconcha" status="sp. nov.">
<emphasis id="7494D1716A26FFEEFE6CFCFFBC943B4D" bold="true" box="[510,744,831,858]" italics="true" pageId="7" pageNumber="8">Sarika lactoconcha</emphasis>
Pholyotha &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="6FA76C0A6A26FFEEFC49FC80BA4A3B4D" box="[987,1078,832,858]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0EFA5EE86A26FFEEFEEAFCA3BDE03BB7" pageId="7" pageNumber="8" type="description">
<paragraph id="465F0D636A26FFEEFEEAFCA3BAC13B69" blockId="7.[376,1213,831,928]" box="[376,1213,867,894]" pageId="7" pageNumber="8">
<uri id="327101616A26FFEEFEEAFCA3BAC13B69" box="[376,1213,867,894]" pageId="7" pageNumber="8">
urn:lsid:zoobank.org:act:
<uuid id="324637B66A26FFEEFD04FCA3BAC13B69" box="[662,1213,867,894]" pageId="7" pageNumber="8">0E48F6FB-6775-4470-BF85-345C3D43112A</uuid>
</uri>
</paragraph>
<paragraph id="465F0D636A26FFEEFD0AFC46BDE03BB7" blockId="7.[376,1213,831,928]" box="[664,924,902,928]" pageId="7" pageNumber="8">
<tableCitation id="0B6238D86A26FFEEFD0AFC46BC933BB7" box="[664,751,902,928]" captionStart="Table 1" captionStartId="2.[189,258,268,294]" captionTargetPageId="2" captionText="Table 1. Taxa attributed to Macrochlamys and Sarika from mainland Southeast Asia, as classified by their maximum shell width and maximum whorl numbers. Data are taken from the original descriptions and additional references. Taxa in bold are confirmed in generic placement by genital characters. 1 Godwin- Austen (1907) and Blanford &amp; Godwin-Austen (1908); 2 Laidlaw (1933); 3 Solem (1966); 4 Maneevong (2000); 5 Sutcharit &amp; Panha (2008): 6 Pholyotha et al. (2018); 7 Pholyotha et al. (2020); 8 This study." httpUri="http://table.plazi.org/id/129F5DEB6A23FFEBFF2FFECCBB3839A5" pageId="7" pageNumber="8" tableUuid="129F5DEB6A23FFEBFF2FFECCBB3839A5">Table 1</tableCitation>
;
<figureCitation id="DEDB11E66A26FFEEFD6CFC46BD3A3BB7" box="[766,838,902,928]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1398,969,1888]" captionTargetId="figure@5.[189,1398,969,1888]" captionTargetPageId="5" captionText="Fig. 2. Shells of Sarika spp. A. Sarika resplendens (Philippi, 1846) (NHMUK 1903.7.1.112). B. S. bocourti (Morelet, 1875) (CUMZ 7900). C. S. lactoconcha Pholyotha &amp; Panha sp. nov., holotype (CUMZ 7901). D. S. lactoconcha Pholyotha &amp; Panha sp. nov., paratype (CUMZ 7902)." figureDoi="http://doi.org/10.5281/zenodo.3908379" httpUri="https://zenodo.org/record/3908379/files/figure.png" pageId="7" pageNumber="8">Figs 2</figureCitation>
C–D, 3
</paragraph>
</subSubSection>
<subSubSection id="0EFA5EE86A26FFEEFF2FFC07BA2D3C41" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="465F0D636A26FFEEFF2FFC07BF4A3BF6" blockId="7.[189,310,967,993]" box="[189,310,967,993]" pageId="7" pageNumber="8">
<heading id="1D17BA0F6A26FFEEFF2FFC07BF4A3BF6" bold="true" box="[189,310,967,993]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="7494D1716A26FFEEFF2FFC07BF4A3BF6" bold="true" box="[189,310,967,993]" pageId="7" pageNumber="8">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="465F0D636A26FFEEFF2FFC36BA2D3C41" blockId="7.[189,1399,1014,1111]" pageId="7" pageNumber="8">
<taxonomicName id="81E076E06A26FFEEFF2FFC36BFDF3C07" authority="Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha, 2020" authorityName="Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha" authorityYear="2020" box="[189,419,1014,1040]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="lactoconcha" status="sp. nov.">
<emphasis id="7494D1716A26FFEEFF2FFC36BFDF3C07" box="[189,419,1014,1040]" italics="true" pageId="7" pageNumber="8">Sarika lactoconcha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6FA76C0A6A26FFEEFE3CFC37BC773C06" box="[430,523,1015,1041]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by a large, depressed and whitish shell, rounded on the periphery. Animal with dark grey body and five mantle lobes. Genitalia with rather long epiphallic caecum and triangular prism-shaped penial pilasters on inner penial sculpture.
</paragraph>
</subSubSection>
<subSubSection id="0EFA5EE86A26FFEEFF2FFBBEBAC73CFD" pageId="7" pageNumber="8" type="etymology">
<paragraph id="465F0D636A26FFEEFF2FFBBEBF383C8F" blockId="7.[189,324,1150,1176]" box="[189,324,1150,1176]" pageId="7" pageNumber="8">
<heading id="1D17BA0F6A26FFEEFF2FFBBEBF383C8F" bold="true" box="[189,324,1150,1176]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="7494D1716A26FFEEFF2FFBBEBF383C8F" bold="true" box="[189,324,1150,1176]" pageId="7" pageNumber="8">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="465F0D636A26FFEEFF2FFB6CBAC73CFD" blockId="7.[189,1399,1196,1258]" pageId="7" pageNumber="8">
The specific epithet ‘
<taxonomicName id="81E076E06A26FFEEFE21FB6CBC3E3CD1" authorityName="Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha" authorityYear="2020" box="[435,578,1196,1222]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="lactoconcha">
<emphasis id="7494D1716A26FFEEFE21FB6CBC3E3CD1" box="[435,578,1196,1222]" italics="true" pageId="7" pageNumber="8">lactoconcha</emphasis>
</taxonomicName>
’ is derived from the Latin word ‘
<emphasis id="7494D1716A26FFEEFC54FB6CBA7A3CD1" box="[966,1030,1196,1222]" italics="true" pageId="7" pageNumber="8">lactis</emphasis>
’ meaning ‘milk, milky’ and the Latin word ‘
<emphasis id="7494D1716A26FFEEFEC2FB10BFD93CFD" box="[336,421,1232,1258]" italics="true" pageId="7" pageNumber="8">concha</emphasis>
’ meaning ’snail’ or ’shell’. It refers to the milky colour of the shell.
</paragraph>
</subSubSection>
<subSubSection id="0EFA5EE86A26FFEEFF2FFAD1BA4D3F15" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="465F0D636A26FFEEFF2FFAD1BFD63D3C" blockId="7.[189,426,1297,1323]" box="[189,426,1297,1323]" pageId="7" pageNumber="8">
<heading id="1D17BA0F6A26FFEEFF2FFAD1BFD63D3C" bold="true" box="[189,426,1297,1323]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="7494D1716A26FFEEFF2FFAD1BFD63D3C" bold="true" box="[189,426,1297,1323]" pageId="7" pageNumber="8">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="465F0D636A26FFEEFF78FA80BF203D4D" blockId="7.[189,1398,1344,1475]" box="[234,348,1344,1370]" pageId="7" pageNumber="8">
<emphasis id="7494D1716A26FFEEFF78FA80BF203D4D" bold="true" box="[234,348,1344,1370]" pageId="7" pageNumber="8">
<typeStatus id="995BB3C16A26FFEEFF78FA80BF203D4D" box="[234,348,1344,1370]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="465F0D636A26FFEEFF2FFAA3BF273DD4" blockId="7.[189,1398,1344,1475]" pageId="7" pageNumber="8">
<materialsCitation id="F688073E6A26FFEEFF2FFAA3BF273DD4" ID-GBIF-Occurrence="2643240306" collectionCode="CUMZ" country="Cambodia" county="Tuek Chhou District" latitude="10.642722" location="Tuek Chhou District" longLatPrecision="1" longitude="104.268974" municipality="Phnom Chhngok Cave Temple" pageId="7" pageNumber="8" specimenCode="CUMZ 7901" specimenCount="1" stateProvince="Kampot Province" typeStatus="holotype">
<collectingCountry id="3EF74DF36A26FFEEFF2FFAA3BF273D6A" box="[189,347,1379,1405]" name="Cambodia" pageId="7" pageNumber="8">CAMBODIA</collectingCountry>
•
<specimenCount id="50E6C6EA6A26FFEEFEEFFAA3BFB23D6A" box="[381,462,1379,1405]" pageId="7" pageNumber="8">1 shell</specimenCount>
(width
<quantity id="8118A0866A26FFEEFDA0FAA3BCDE3D6A" box="[562,674,1379,1405]" metricMagnitude="-2" metricUnit="m" metricValue="2.36" pageId="7" pageNumber="8" unit="mm" value="23.6">23.6 mm</quantity>
, height
<quantity id="8118A0866A26FFEEFC95FAA3BD053D6A" box="[775,889,1379,1405]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="7" pageNumber="8" unit="mm" value="12.1">12.1 mm</quantity>
);
<collectingRegion id="8424C3816A26FFEEFC03FAA3BA1A3D6A" box="[913,1126,1379,1405]" country="Cambodia" name="Kampot" pageId="7" pageNumber="8">Kampot Province</collectingRegion>
,
<collectingCounty id="AF3E75EF6A26FFEEFBE4FAA3BB0E3D6A" box="[1142,1394,1379,1405]" pageId="7" pageNumber="8">Tuek Chhou District</collectingCounty>
,
<locationDeviation id="57ED14416A26FFEEFF2FFA46BD603DB7" box="[189,796,1414,1441]" pageId="7" pageNumber="8">limestone outcrops at Phnom Chhngok Cave Temple</locationDeviation>
, locality code C046;
<geoCoordinate id="23D46BA46A26FFEEFB98FA46BACF3DB6" box="[1034,1203,1412,1441]" degrees="10" direction="north" minutes="38" orientation="latitude" pageId="7" pageNumber="8" precision="1" seconds="33.8" value="10.642722">10°38′33.8″ N</geoCoordinate>
,
<geoCoordinate id="23D46BA46A26FFEEFB2EFA46BB0D3DB6" box="[1212,1393,1412,1441]" degrees="104" direction="east" minutes="16" orientation="longitude" pageId="7" pageNumber="8" precision="1" seconds="08.3" value="104.268974">104°16′08.3″ E</geoCoordinate>
;
<specimenCode id="1646A5186A26FFEEFF2FFA69BF2B3DD4" box="[189,343,1449,1475]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="7" pageNumber="8">CUMZ 7901</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="465F0D636A26FFEEFF78FA2BBF1B3E12" blockId="7.[189,1399,1515,1646]" box="[234,359,1515,1541]" pageId="7" pageNumber="8">
<emphasis id="7494D1716A26FFEEFF78FA2BBF1B3E12" bold="true" box="[234,359,1515,1541]" pageId="7" pageNumber="8">
<typeStatus id="995BB3C16A26FFEEFF78FA2BBF1B3E12" box="[234,359,1515,1541]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="465F0D636A26FFEEFF2FF9CEBE823E79" blockId="7.[189,1399,1515,1646]" pageId="7" pageNumber="8">
<materialsCitation id="F688073E6A26FFEEFF2FF9CEBB0B3E3F" ID-GBIF-Occurrence="2643240309" box="[189,1399,1550,1576]" collectionCode="CUMZ" country="Cambodia" county="Tuek Chhou District" latitude="10.642722" location="CAMBODIA" longLatPrecision="1" longitude="104.268974" municipality="Phnom Chhngok Cave Temple" pageId="7" pageNumber="8" specimenCode="CUMZ 7902" specimenCount="4" stateProvince="Kampot" typeStatus="paratype">
<collectingCountry id="3EF74DF36A26FFEEFF2FF9CEBF273E3F" box="[189,347,1550,1576]" name="Cambodia" pageId="7" pageNumber="8">CAMBODIA</collectingCountry>
•
<specimenCount id="50E6C6EA6A26FFEEFEFDF9CEBFC13E3F" box="[367,445,1550,1576]" pageId="7" pageNumber="8">1 shell</specimenCount>
,
<specimenCount id="50E6C6EA6A26FFEEFE55F9CEBCD13E3F" box="[455,685,1550,1576]" pageId="7" pageNumber="8">3 alcohol-preserved</specimenCount>
specimens; same collection data as for holotype;
<specimenCode id="1646A5186A26FFEEFB70F9CEBB0B3E3F" box="[1250,1399,1550,1576]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="7" pageNumber="8">CUMZ 7902</specimenCode>
</materialsCitation>
•
<materialsCitation id="F688073E6A26FFEEFF42F9F1BD473E5B" ID-GBIF-Occurrence="2643240316" box="[208,827,1585,1612]" collectionCode="NHM" country="Cambodia" county="Tuek Chhou District" latitude="10.642722" location="Phnom Chhngok Cave" longLatPrecision="1" longitude="104.268974" municipality="Phnom Chhngok Cave Temple" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Kampot" typeStatus="paratype">
<specimenCount id="50E6C6EA6A26FFEEFF42F9F1BF5E3E5C" box="[208,290,1585,1611]" pageId="7" pageNumber="8">1 shell</specimenCount>
; same collection data as for preceding;
<collectionCode id="20F195A66A26FFEEFD65F9F2BD473E5B" box="[759,827,1586,1612]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="7" pageNumber="8">NHM</collectionCode>
</materialsCitation>
•
<materialsCitation id="F688073E6A26FFEEFCC5F9F1BE863E79" ID-GBIF-Occurrence="2643240314" collectionCode="ZRC" country="Cambodia" county="Tuek Chhou District" latitude="10.642722" location="Phnom Chhngok Cave" longLatPrecision="1" longitude="104.268974" municipality="Phnom Chhngok Cave Temple" pageId="7" pageNumber="8" specimenCount="1" stateProvince="Kampot" typeStatus="paratype">
<specimenCount id="50E6C6EA6A26FFEEFCC5F9F1BDD53E5C" box="[855,937,1585,1611]" pageId="7" pageNumber="8">1 shell</specimenCount>
; same collection data as for preceding;
<collectionCode id="20F195A66A26FFEEFF2FF994BE863E79" box="[189,250,1620,1646]" collectionName="Singapore, National University of Singapore, Raffles Museum of Biodiversity Research, Zoological Reference Collection" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="8">ZRC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="465F0D636A26FFEEFF2FF955BF053EB8" blockId="7.[189,377,1685,1711]" box="[189,377,1685,1711]" pageId="7" pageNumber="8">
<heading id="1D17BA0F6A26FFEEFF2FF955BF053EB8" bold="true" box="[189,377,1685,1711]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="7494D1716A26FFEEFF2FF955BF053EB8" bold="true" box="[189,377,1685,1711]" pageId="7" pageNumber="8">Other material</emphasis>
</heading>
</paragraph>
<paragraph id="465F0D636A26FFEEFF2FF904BA4D3F15" blockId="7.[189,1398,1732,1794]" pageId="7" pageNumber="8">
<materialsCitation id="F688073E6A26FFEEFF2FF904BA513F15" ID-GBIF-Occurrence="2643240303" collectionCode="CUMZ" country="Cambodia" latitude="10.616556" location="Tuek Chhou District" longLatPrecision="1" longitude="104.24383" municipality="Tuek Chhou District" pageId="7" pageNumber="8" specimenCode="CUMZ 7903" specimenCount="2" stateProvince="Kampot Province">
<collectingCountry id="3EF74DF36A26FFEEFF2FF904BF273EC9" box="[189,347,1732,1758]" name="Cambodia" pageId="7" pageNumber="8">CAMBODIA</collectingCountry>
•
<specimenCount id="50E6C6EA6A26FFEEFEE8F904BFA63EC9" box="[378,474,1732,1758]" pageId="7" pageNumber="8">2 shells</specimenCount>
;
<collectingRegion id="8424C3816A26FFEEFE7BF905BCC13EC9" box="[489,701,1732,1759]" country="Cambodia" name="Kampot" pageId="7" pageNumber="8">Kampot Province</collectingRegion>
,
<collectingMunicipality id="A63B97196A26FFEEFD5EF904BDB93EC9" box="[716,965,1732,1758]" pageId="7" pageNumber="8">Tuek Chhou District</collectingMunicipality>
,
<locationDeviation id="57ED14416A26FFEEFC46F904BF633F15" pageId="7" pageNumber="8">limestone outcrops at Phnom Kbal Romeas</locationDeviation>
, locality code C045;
<geoCoordinate id="23D46BA46A26FFEEFD85F926BCBF3F15" box="[535,707,1766,1794]" degrees="10" direction="north" minutes="36" orientation="latitude" pageId="7" pageNumber="8" precision="1" seconds="59.6" value="10.616556">10°36′59.6″ N</geoCoordinate>
,
<geoCoordinate id="23D46BA46A26FFEEFD5DF926BDFA3F15" box="[719,902,1766,1794]" degrees="104" direction="east" minutes="14" orientation="longitude" pageId="7" pageNumber="8" precision="1" seconds="37.8" value="104.24383">104°14′37.8″ E</geoCoordinate>
;
<specimenCode id="1646A5186A26FFEEFC01F928BA513F15" box="[915,1069,1768,1794]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="7" pageNumber="8">CUMZ 7903</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0EFA5EE86A26FFE0FF2FF8E9BAA43CDA" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="description">
<paragraph id="465F0D636A26FFEEFF2FF8E9BF323F54" blockId="7.[189,334,1833,1859]" box="[189,334,1833,1859]" pageId="7" pageNumber="8">
<heading id="1D17BA0F6A26FFEEFF2FF8E9BF323F54" bold="true" box="[189,334,1833,1859]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="7494D1716A26FFEEFF2FF8E9BF323F54" bold="true" box="[189,334,1833,1859]" pageId="7" pageNumber="8">Description</emphasis>
</heading>
</paragraph>
<paragraph id="465F0D636A26FFE0FF2FF898BB13395F" blockId="7.[189,1399,1879,2012]" lastBlockId="9.[189,1398,267,329]" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
SHELL (
<figureCitation id="DEDB11E66A26FFEEFE86F897BF203F65" box="[276,348,1879,1906]" captionStart="Fig" captionStartId="5.[189,232,1930,1956]" captionTargetBox="[189,1398,969,1888]" captionTargetId="figure@5.[189,1398,969,1888]" captionTargetPageId="5" captionText="Fig. 2. Shells of Sarika spp. A. Sarika resplendens (Philippi, 1846) (NHMUK 1903.7.1.112). B. S. bocourti (Morelet, 1875) (CUMZ 7900). C. S. lactoconcha Pholyotha &amp; Panha sp. nov., holotype (CUMZ 7901). D. S. lactoconcha Pholyotha &amp; Panha sp. nov., paratype (CUMZ 7902)." figureDoi="http://doi.org/10.5281/zenodo.3908379" httpUri="https://zenodo.org/record/3908379/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
C–D). Depressed, large (shell width up to
<quantity id="8118A0866A26FFEEFCCBF898BDBA3F65" box="[857,966,1880,1906]" metricMagnitude="-2" metricUnit="m" metricValue="2.37" pageId="7" pageNumber="8" unit="mm" value="23.7">23.7 mm</quantity>
, shell height up to
<quantity id="8118A0866A26FFEEFB3FF898BB603F65" box="[1197,1308,1880,1906]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="7" pageNumber="8" unit="mm" value="12.1">12.1 mm</quantity>
), rather thin, semitransparent. Shell surface generally smooth, glossy; more glossy below periphery; shell colour monochrome pale milky or whitish colour. 6–6½ whorls, increasing regularly, slightly convex, separated by rather shallow and wide suture. Spire slightly elevated; last whorl broadly rounded. Aperture oval- lunate shaped, open obliquely. Peristome simple, very slightly thickened, not expanded. Parietal callus thin, transparent; columellar margin simple, slightly reflected near umbilicus. Umbilicus narrow, deep.
</paragraph>
<caption id="129F5DEB6A29FFE1FF2FF8A7BD053FFC" ID-DOI="http://doi.org/10.5281/zenodo.3908381" ID-Zenodo-Dep="3908381" httpUri="https://zenodo.org/record/3908381/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1895,1921]" targetBox="[189,1398,374,1853]" targetPageId="8">
<paragraph id="465F0D636A29FFE1FF2FF8A7BD053FFC" blockId="8.[189,1399,1895,2027]" pageId="8" pageNumber="9">
<emphasis id="7494D1716A29FFE1FF2FF8A7BF6C3F96" bold="true" box="[189,272,1895,1921]" pageId="8" pageNumber="9">Fig. 3.</emphasis>
<taxonomicName id="81E076E06A29FFE1FE8EF8A7BD7C3F96" authority="Pholyotha &amp; Panha" authorityName="Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha" authorityYear="2020" box="[284,768,1895,1921]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="lactoconcha" status="sp. nov.">
<emphasis id="7494D1716A29FFE1FE8EF8A7BC783F96" box="[284,516,1895,1921]" italics="true" pageId="8" pageNumber="9">Sarika lactoconcha</emphasis>
Pholyotha &amp; Panha
</taxonomicName>
<taxonomicNameLabel id="6FA76C0A6A29FFE1FC9EF8A8BD123F95" box="[780,878,1896,1922]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (CUMZ 7902).
<emphasis id="7494D1716A29FFE1FB26F8A7BAB53F96" bold="true" box="[1204,1225,1895,1921]" pageId="8" pageNumber="9">A</emphasis>
. Living snail.
<emphasis id="7494D1716A29FFE1FF2FF84ABEAD3FB3" bold="true" box="[189,209,1930,1956]" pageId="8" pageNumber="9">B</emphasis>
. General view of genital system.
<emphasis id="7494D1716A29FFE1FDCDF84ABC083FB3" bold="true" box="[607,628,1930,1956]" pageId="8" pageNumber="9">C</emphasis>
. Internal structure of penis.
<emphasis id="7494D1716A29FFE1FC51F84ABDA43FB3" bold="true" box="[963,984,1930,1956]" pageId="8" pageNumber="9">D</emphasis>
. Representative SEM image of the radula. Central tooth indicated by ‘C’; yellow arrow heads indicate lateral teeth with the transition to marginal teeth; white arrow heads indicate the end of penis.
</paragraph>
</caption>
<paragraph id="465F0D636A28FFE0FF2FFEB0BC3E3A2D" blockId="9.[189,1399,368,570]" pageId="9" pageNumber="10">
EXTERNAL FEATURES (
<figureCitation id="DEDB11E66A28FFE0FE23FEB0BC6E399D" box="[433,530,368,394]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[189,1398,374,1853]" captionTargetId="figure@8.[189,1398,374,1853]" captionTargetPageId="8" captionText="Fig. 3. Sarika lactoconcha Pholyotha &amp; Panha sp. nov., paratype (CUMZ 7902). A. Living snail. B. General view of genital system. C. Internal structure of penis. D. Representative SEM image of the radula. Central tooth indicated by ‘C’; yellow arrow heads indicate lateral teeth with the transition to marginal teeth; white arrow heads indicate the end of penis." figureDoi="http://doi.org/10.5281/zenodo.3908381" httpUri="https://zenodo.org/record/3908381/files/figure.png" pageId="9" pageNumber="10">Fig. 3A</figureCitation>
). Living animals have dark grey body; dorsally with darker colour than below, foot sole. Caudal foss present; caudal horn raised, rather large. Mantle and lung scattered with dark pigment or spots or blotches. Mantle edge well-developed, dark grey to paler in colour, with three dorsal lobes, two shell lobes (see
<bibRefCitation id="227170926A28FFE0FDDCFE19BD2D39E3" author="Pholyotha A. &amp; Sutcharit C. &amp; Panha S." box="[590,849,473,500]" pageId="9" pageNumber="10" pagination="763 - 781" refId="ref9515" refString="Pholyotha A., Sutcharit C. &amp; Panha S. 2018. The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae). Raffles Bulletin of Zoology 66: 763 - 781." type="journal article" year="2018">
Pholyotha
<emphasis id="7494D1716A28FFE0FD5FFE1ABD7539E4" box="[717,777,474,500]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
2018
</bibRefCitation>
: fig. 1). Dorsal lobes large, broad; right dorsal lobe larger than both anterior and posterior left dorsal lobes. Shell lobes large, long; right shell lobe larger, longer than left shell lobe.
</paragraph>
<paragraph id="465F0D636A28FFE0FF2FFDA2BDD33BCC" blockId="9.[189,1399,609,988]" pageId="9" pageNumber="10">
GENITALIA (
<figureCitation id="DEDB11E66A28FFE0FED0FDA1BFF43A6B" box="[322,392,609,636]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[189,1398,374,1853]" captionTargetId="figure@8.[189,1398,374,1853]" captionTargetPageId="8" captionText="Fig. 3. Sarika lactoconcha Pholyotha &amp; Panha sp. nov., paratype (CUMZ 7902). A. Living snail. B. General view of genital system. C. Internal structure of penis. D. Representative SEM image of the radula. Central tooth indicated by ‘C’; yellow arrow heads indicate lateral teeth with the transition to marginal teeth; white arrow heads indicate the end of penis." figureDoi="http://doi.org/10.5281/zenodo.3908381" httpUri="https://zenodo.org/record/3908381/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
B–C). Atrium (at) enlarged, very short. Penis (p) cylindrical, with thin penial sheath covering proximal penis. Inner sculpture of penis very finely folded to nearly smooth surface close to atrium then gradually transforming to triangular prism (rhombic base, acute angle on top) penial pilaster (pp) near epiphallus. Epiphallus (e) cylindrical, elongate, about two times penis length. Epiphallic caecum (ec) similar diameter as epiphallus, straight, located near middle of epiphallus. Penial retractor muscle (prm) thin, attached at tip of epiphallic caecum. Flagellum (fl) slender, rather long, nearly 1.5 times epiphallus length. Vas deferens (vd) thin tube. Vagina (v) cylindrical, about half of penis length. Dart apparatus (da) cylindrical, rather large and long, joined to atrium at vagina and penis junction. Gametolytic sac (gs) elongate bulbous; gametolytic duct (gd) enlarged cylindrical. Free oviduct (fo) enlarged cylindrical, approximately two times vagina length, proximally encircled with brownish tissue. Oviduct large lobules; prostate gland running alongside oviduct.
</paragraph>
<paragraph id="465F0D636A28FFE0FF2FFBC3BAA43CDA" blockId="9.[189,1399,1027,1229]" pageId="9" pageNumber="10">
RADULA (
<figureCitation id="DEDB11E66A28FFE0FEB8FBC3BFF63C0A" box="[298,394,1027,1053]" captionStart="Fig" captionStartId="8.[189,232,1895,1921]" captionTargetBox="[189,1398,374,1853]" captionTargetId="figure@8.[189,1398,374,1853]" captionTargetPageId="8" captionText="Fig. 3. Sarika lactoconcha Pholyotha &amp; Panha sp. nov., paratype (CUMZ 7902). A. Living snail. B. General view of genital system. C. Internal structure of penis. D. Representative SEM image of the radula. Central tooth indicated by ‘C’; yellow arrow heads indicate lateral teeth with the transition to marginal teeth; white arrow heads indicate the end of penis." figureDoi="http://doi.org/10.5281/zenodo.3908381" httpUri="https://zenodo.org/record/3908381/files/figure.png" pageId="9" pageNumber="10">Fig. 3D</figureCitation>
). Teeth arranged in wide-angle U-shape with half row formula: 1–(15–16)–65 teeth. Central tooth symmetrical tricuspid with pointed cusps; mesocone large lanceolate, dull cusp; ectocones very small. Lateral teeth asymmetrical tricuspid; mesocone large with pointed cusp; endocone very small, located near tip; ectocone larger than endocone, located in middle of tooth. Marginal teeth starting around tooth numbers 15–16, obliquely bicuspid; endocone large, pointed cusp; ectocone small. Outermost teeth gradually reduced in size from elongate, narrow teeth to very short teeth.
</paragraph>
</subSubSection>
<subSubSection id="0EFA5EE86A28FFE0FF2FFB34BF483DDD" pageId="9" pageNumber="10" type="distribution">
<paragraph id="465F0D636A28FFE0FF2FFB34BF2A3D19" blockId="9.[189,342,1268,1294]" box="[189,342,1268,1294]" pageId="9" pageNumber="10">
<heading id="1D17BA0F6A28FFE0FF2FFB34BF2A3D19" bold="true" box="[189,342,1268,1294]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="7494D1716A28FFE0FF2FFB34BF2A3D19" bold="true" box="[189,342,1268,1294]" pageId="9" pageNumber="10">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="465F0D636A28FFE0FF2FFAE3BF483DDD" blockId="9.[189,1399,1315,1482]" pageId="9" pageNumber="10">
This new species is currently known from Phnom Chhngok Cave (the
<typeStatus id="995BB3C16A28FFE0FB84FAE3BA353D2A" box="[1046,1097,1315,1341]" pageId="9" pageNumber="10">type</typeStatus>
locality), an undisturbed small karst outcrop, which is about
<quantity id="8118A0866A28FFE0FDF2FA86BCE73D77" box="[608,667,1350,1376]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="km" value="1.0">1 km</quantity>
long and surrounded by rice fields. Another locality is Phnom Kbal Romeas, a small limestone outcrop. This outcrop (
<figureCitation id="DEDB11E66A28FFE0FCC3FAA9BDEA3D94" box="[849,918,1385,1411]" captionStart="Fig" captionStartId="3.[189,232,1965,1991]" captionTargetBox="[189,1398,1446,1923]" captionTargetId="figure@3.[189,1398,1446,1923]" captionTargetPageId="3" captionText="Fig. 1. Geographic distribution of Sarika spp. in Cambodia based on new specimens examined herein. (modified from GinkgoMaps-project http://www.ginkgomaps.com)." figureDoi="http://doi.org/10.5281/zenodo.3908377" httpUri="https://zenodo.org/record/3908377/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
) is situated about
<quantity id="8118A0866A28FFE0FBE3FAA9BAD03D94" box="[1137,1196,1385,1411]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="10" unit="km" value="4.0">4 km</quantity>
southwest of the
<typeStatus id="995BB3C16A28FFE0FF2FFA4DBE8C3DB0" box="[189,240,1421,1447]" pageId="9" pageNumber="10">type</typeStatus>
locality, surrounded by housing areas and an inactive old quarry, and now has become a tourist attraction.
</paragraph>
</subSubSection>
<subSubSection id="0EFA5EE86A28FFE3FF2FFA31BA3F3998" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="discussion">
<paragraph id="465F0D636A28FFE0FF2FFA31BF533E1C" blockId="9.[189,303,1521,1547]" box="[189,303,1521,1547]" pageId="9" pageNumber="10">
<heading id="1D17BA0F6A28FFE0FF2FFA31BF533E1C" bold="true" box="[189,303,1521,1547]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="7494D1716A28FFE0FF2FFA31BF533E1C" bold="true" box="[189,303,1521,1547]" pageId="9" pageNumber="10">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="465F0D636A28FFE0FF2FF9E0BFDD3F27" blockId="9.[189,1399,1567,1840]" pageId="9" pageNumber="10">
<taxonomicName id="81E076E06A28FFE0FF2FF9E0BFDE3E2D" authority="Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha, 2020" authorityName="Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha" authorityYear="2020" box="[189,418,1568,1594]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="lactoconcha" status="sp. nov.">
<emphasis id="7494D1716A28FFE0FF2FF9E0BFDE3E2D" box="[189,418,1568,1594]" italics="true" pageId="9" pageNumber="10">Sarika lactoconcha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6FA76C0A6A28FFE0FE39F9E0BC743E2D" box="[427,520,1568,1594]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="81E076E06A28FFE0FD3AF9E0BD643E2D" baseAuthorityName="Crosse &amp; Fischer" baseAuthorityYear="1863" box="[680,792,1568,1594]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="benoiti">
<emphasis id="7494D1716A28FFE0FD3AF9E0BD643E2D" box="[680,792,1568,1594]" italics="true" pageId="9" pageNumber="10">S. benoiti</emphasis>
</taxonomicName>
,
<taxonomicName id="81E076E06A28FFE0FCBBF9E0BDD73E2D" baseAuthorityName="Morelet" baseAuthorityYear="1875" box="[809,939,1568,1594]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="bocourti">
<emphasis id="7494D1716A28FFE0FCBBF9E0BDD73E2D" box="[809,939,1568,1594]" italics="true" pageId="9" pageNumber="10">S. bocourti</emphasis>
</taxonomicName>
and
<taxonomicName id="81E076E06A28FFE0FC7AF9E0BAEE3E2D" baseAuthorityName="Philippi" baseAuthorityYear="1846" box="[1000,1170,1568,1594]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="resplendens">
<emphasis id="7494D1716A28FFE0FC7AF9E0BAEE3E2D" box="[1000,1170,1568,1594]" italics="true" pageId="9" pageNumber="10">S. resplendens</emphasis>
</taxonomicName>
with whitish shell, while the three latter species have brownish shells. Moreover,
<taxonomicName id="81E076E06A28FFE0FC00F983BA423E4A" authority="Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha, 2020" authorityName="Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha" authorityYear="2020" box="[914,1086,1603,1629]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="lactoconcha" status="sp. nov.">
<emphasis id="7494D1716A28FFE0FC00F983BA423E4A" box="[914,1086,1603,1629]" italics="true" pageId="9" pageNumber="10">S. lactoconcha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6FA76C0A6A28FFE0FBD7F983BAE33E4A" box="[1093,1183,1603,1629]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
has a rounded last whorl, while
<taxonomicName id="81E076E06A28FFE0FECCF9A6BF9E3E97" baseAuthorityName="Morelet" baseAuthorityYear="1875" box="[350,482,1638,1664]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="bocourti">
<emphasis id="7494D1716A28FFE0FECCF9A6BF9E3E97" box="[350,482,1638,1664]" italics="true" pageId="9" pageNumber="10">S. bocourti</emphasis>
</taxonomicName>
has an obtusely angulated last whorl. Unfortunately, no genital information of
<taxonomicName id="81E076E06A28FFE0FF76F94ABFEF3EB4" baseAuthorityName="Philippi" baseAuthorityYear="1846" box="[228,403,1673,1700]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="resplendens">
<emphasis id="7494D1716A28FFE0FF76F94ABFEF3EB4" box="[228,403,1673,1700]" italics="true" pageId="9" pageNumber="10">S. resplendens</emphasis>
</taxonomicName>
from
<collectingCountry id="3EF74DF36A28FFE0FE7BF949BC1E3EB4" box="[489,610,1673,1699]" name="Cambodia" pageId="9" pageNumber="10">Cambodia</collectingCountry>
is available for further comparison. However, the genitalia of
<taxonomicName id="81E076E06A28FFE0FF2FF96DBF1A3ED1" baseAuthorityName="Philippi" baseAuthorityYear="1846" box="[189,358,1708,1735]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="resplendens">
<emphasis id="7494D1716A28FFE0FF2FF96DBF1A3ED1" box="[189,358,1708,1735]" italics="true" pageId="9" pageNumber="10">S. resplendens</emphasis>
</taxonomicName>
from
<collectingCountry id="3EF74DF36A28FFE0FE22F96DBC5E3ED0" box="[432,546,1709,1735]" name="Myanmar" pageId="9" pageNumber="10">Myanmar</collectingCountry>
possess a large penial retractor muscle with about the same diameter as the epiphallic caecum, and short flagellum compared with penis (see Godwin-Austen 1907: 179–181, pl. 111, figs 3, 3a for comparison), while this new species has a small and thin penial retractor muscle and long flagellum.
</paragraph>
<paragraph id="465F0D636A28FFE3FF2FF897BA3F3998" blockId="9.[189,1399,1879,2012]" lastBlockId="10.[189,1398,267,399]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Compared to species of
<emphasis id="7494D1716A28FFE0FE75F897BC963F66" box="[487,746,1879,1906]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="81E076E06A28FFE0FE75F897BC483F66" authorityName="Godwin-Austen" authorityYear="1907" box="[487,564,1879,1905]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Sarika</taxonomicName>
-
<taxonomicName id="81E076E06A28FFE0FDAFF897BC963F66" authorityName="Gray" authorityYear="1847" box="[573,746,1879,1905]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">Macrochlamys</taxonomicName>
</emphasis>
having a whitish shell,
<taxonomicName id="81E076E06A28FFE0FB81F898BABF3F66" authority="Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha, 2020" authorityName="Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha" authorityYear="2020" box="[1043,1219,1879,1906]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="lactoconcha" status="sp. nov.">
<emphasis id="7494D1716A28FFE0FB81F898BABF3F66" box="[1043,1219,1879,1906]" italics="true" pageId="9" pageNumber="10">S. lactoconcha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6FA76C0A6A28FFE0FB5DF898BB513F65" box="[1231,1325,1880,1906]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
has a wider last whorl (loosely coiled), apertural lip simple and genitalia without penial verge. Meanwhile,
<taxonomicName id="81E076E06A28FFE0FF2FF85EBF853FAE" authority="(Benson, 1860)" baseAuthorityName="Benson" baseAuthorityYear="1860" box="[189,505,1950,1977]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="consepta">
<emphasis id="7494D1716A28FFE0FF2FF85EBF3C3FAF" box="[189,320,1950,1976]" italics="true" pageId="9" pageNumber="10">S. consepta</emphasis>
(Benson, 1860)
</taxonomicName>
from
<collectingCountry id="3EF74DF36A28FFE0FDAFF85EBCD33FAF" box="[573,687,1950,1976]" name="Myanmar" pageId="9" pageNumber="10">Myanmar</collectingCountry>
has a narrow last whorl (densely coiled), apertural lip slightly expanded and rather thickened inside, and genitalia with large penial verge (see
<bibRefCitation id="227170926A28FFE0FBEAF801BB0B3FCC" author="Pholyotha A. &amp; Sutcharit C. &amp; Tongkerd P. &amp; Lin A. &amp; Panha S." box="[1144,1399,1985,2011]" pageId="9" pageNumber="10" pagination="183 - 204" refId="ref9561" refString="Pholyotha A., Sutcharit C., Tongkerd P., Lin A. &amp; Panha S. 2020. Taxonomic revision of the land snail genera Macrochlamys Gray, 1847 and Sarika Godwin-Austen, 1907 (Eupulmonata: Ariophantidae) from Southeastern Myanmar, with descriptions of three new species. Molluscan Research 40: 183 - 204. https: // doi. org / 10.1080 / 13235818.2020.1723041" type="journal article" year="2020">
Pholyotha
<emphasis id="7494D1716A28FFE0FB65F802BB4F3FCC" box="[1271,1331,1985,2011]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
2020
</bibRefCitation>
for comparison). Another species,
<taxonomicName id="81E076E06A2BFFE3FDC2FECBBA193932" authority="Vermeulen et al., 2019" authorityName="Vermeulen" authorityYear="2019" box="[592,1125,267,294]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="psyche">
<emphasis id="7494D1716A2BFFE3FDC2FECBBD283932" box="[592,852,267,293]" italics="true" pageId="10" pageNumber="11">Macrochlamys psyche</emphasis>
<bibRefCitation id="227170926A2BFFE3FCC9FECBBA193932" author="Vermeulen J. J. &amp; Luu H. T. &amp; Theary K. &amp; Anker K." box="[859,1125,267,294]" pageId="10" pageNumber="11" pagination="7 - 41" refId="ref9881" refString="Vermeulen J. J., Luu H. T., Theary K. &amp; Anker K. 2019. New species of land snails (Mollusca: Gastropoda: Caenogastropoda and Pulmonata) of the Mekong Delta Limestone Hills (Cambodia, Vietnam). Folia Malacologica 27: 7 - 41. https: // doi. org / 10.12657 / folmal. 027.001" type="journal article" year="2019">
Vermeulen
<emphasis id="7494D1716A2BFFE3FC73FECCBA673932" box="[993,1051,267,293]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
, 2019
</bibRefCitation>
</taxonomicName>
was recently described from southern
<collectingCountry id="3EF74DF36A2BFFE3FEF8FEEEBFAD395F" box="[362,465,302,328]" name="Vietnam" pageId="10" pageNumber="11">Vietnam</collectingCountry>
. This
<taxonomicName id="81E076E06A2BFFE3FD8AFEEFBCB9395E" authorityName="Gray" authorityYear="1847" box="[536,709,303,329]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="7494D1716A2BFFE3FD8AFEEFBCB9395E" box="[536,709,303,329]" italics="true" pageId="10" pageNumber="11">Macrochlamys</emphasis>
</taxonomicName>
species has a flattened to slightly concaved spire, whereas
<taxonomicName id="81E076E06A2BFFE3FF2FFE92BF10397B" authority="Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha, 2020" authorityName="Pholyotha &amp; Sutcharit &amp; Thach &amp; Chhuoy &amp; Ngor &amp; Panha" authorityYear="2020" box="[189,364,338,364]" class="Gastropoda" family="Ariophantidae" genus="Sarika" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="lactoconcha" status="sp. nov.">
<emphasis id="7494D1716A2BFFE3FF2FFE92BF10397B" box="[189,364,338,364]" italics="true" pageId="10" pageNumber="11">S. lactoconcha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6FA76C0A6A2BFFE3FEE7FE92BFAE397B" box="[373,466,338,364]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
has an elevated spire. Unfortunately, the genital anatomy of
<taxonomicName id="81E076E06A2BFFE3FB23FE93BB56397B" box="[1201,1322,338,364]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="psyche">
<emphasis id="7494D1716A2BFFE3FB23FE93BB56397B" box="[1201,1322,338,364]" italics="true" pageId="10" pageNumber="11">M. psyche</emphasis>
</taxonomicName>
is not available for comparison, therefore, its generic placement is still provisional.
</paragraph>
</subSubSection>
</treatment>
</document>
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