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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1159.99218" ID-Pensoft-Pub="1313-2970-1159-87" ID-Pensoft-UUID="ECCECB868071523DB7628E304850BD62" ID-ZooBank="CD97DD120549412F976508D8DE21605B" ModsDocID="1313-2970-1159-87" checkinTime="1682458941538" checkinUser="pensoft" docAuthor="Okada, Ryohei, Jaitrong, Weeyawat &amp; Wewalka, Guenther" docDate="2023" docId="0E028AD633F156FC96DC5E7C696DE758" docLanguage="en" docName="ZooKeys 1159: 87-119" docOrigin="ZooKeys 1159" docPubDate="2023-04-25" docSource="http://dx.doi.org/10.3897/zookeys.1159.99218" docTitle="Microdytes dimorphus Wewalka 1997" docType="treatment" docVersion="1" id="ECCECB868071523DB7628E304850BD62" lastPageNumber="87" masterDocId="ECCECB868071523DB7628E304850BD62" masterDocTitle="A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae)" masterLastPageNumber="119" masterPageNumber="87" pageNumber="87" updateTime="1682458941538" updateUser="pensoft">
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<mods:title>A review of Microdytes J. Balfour-Browne, 1946 from Thailand, Laos, and Cambodia with descriptions of five new species and new records (Coleoptera, Dytiscidae)</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Okada, Ryohei</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8488-0660</mods:nameIdentifier>
<mods:affiliation>Thailand Natural History Museum, National Science Museum, Technopolis, Pathum Thani, Thailand &amp; Coleopterological Society of Japan, National Museum of Nature and Science, Tsukuba, Japan</mods:affiliation>
<mods:nameIdentifier type="email">wasserinsekt@kub.biglobe.ne.jp</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jaitrong, Weeyawat</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1362-0754</mods:nameIdentifier>
<mods:affiliation>Thailand Natural History Museum, National Science Museum, Technopolis, Pathum Thani, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wewalka, Guenther</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-2994-8001</mods:nameIdentifier>
<mods:affiliation>Starkfriedgasse 16 / 1 / 3, A - 1190 Vienna, Austria</mods:affiliation>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-04-25</mods:number>
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<mods:detail type="volume">
<mods:number>1159</mods:number>
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<mods:start>87</mods:start>
<mods:end>119</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1159.99218</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1159.99218</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1159-87</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:0E028AD633F156FC96DC5E7C696DE758" httpUri="http://treatment.plazi.org/id/0E028AD633F156FC96DC5E7C696DE758" lastPageNumber="87" pageId="0" pageNumber="87">
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<taxonomicName ID-CoL="6RGYQ" LSID="0E028AD6-33F1-56FC-96DC-5E7C696DE758" authority="Wewalka, 1997" authorityName="Wewalka" authorityYear="1997" class="Insecta" family="Dytiscidae" genus="Microdytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microdytes dimorphus" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="dimorphus">Microdytes dimorphus Wewalka, 1997</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="87" type="description">
<paragraph pageId="0" pageNumber="87">
<figureCitation captionStart="Figures 1837" captionStartId="F7" captionText="Figures 18 - 37. Habitus of 18 Microdytes akitai 19 M. balkei 20 M. dimorphus 21 M. feryi (paratype, from Myanmar) 22 M. franzi 23 M. gabrielae 24 M. heineri 25 M. maculatus 26 M. mariannae 27 M. menopausis 28 M. paoloi 29 M. pasiricus 30 M. pederzanii 31 M. rocchii 32 M. schoedli 33 M. schoenmanni (from Myanmar) 34 M. schwendingeri 35 M. shepardi 36 M. shunichii 37 M. zetteli. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures18-37" httpUri="https://binary.pensoft.net/fig/842227" pageId="0" pageNumber="87">Fig. 20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="reference_group">
<paragraph pageId="0" pageNumber="87">
<taxonomicName authorityName="Wewalka" authorityYear="1997" class="Insecta" family="Dytiscidae" genus="Microdytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microdytes dimorphus" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="dimorphus">Microdytes dimorphus</taxonomicName>
Wewalka, 1997: 22;
<bibRefCitation author="Wewalka, G" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="87" pagination="21 - 39" refId="B24" refString="Wewalka, G, 2011. New species, new records and taxonomic updates on Microdytes Balfour-Browne, 1946. Koleopterologische Rundschau 81: 21 - 39" title="New species, new records and taxonomic updates on Microdytes Balfour-Browne, 1946." volume="81" year="2011">Wewalka 2011</bibRefCitation>
: 37;
<bibRefCitation author="Nilsson, AN" journalOrPublisher="Etudes Entomologiques Motschulsky" pageId="0" pageNumber="87" publicationUrl="http://www.norrent.se" refId="B13" refString="Nilsson, AN, Hajek, J, 2023. A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1.I.2023. http://www.norrent.se" title="A world catalogue of the family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2023." url="http://www.norrent.se" year="2023">
Nilsson and
<normalizedToken originalValue="Hájek">Hajek</normalizedToken>
2023
</bibRefCitation>
: 211.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="type locality">
<paragraph pageId="0" pageNumber="87">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="87">Thailand, Nakhon Ratchasima or Nakhon Nayok Province, Khao Yai NP.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="materials_examined">
<paragraph pageId="0" pageNumber="87">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="87">
<materialsCitation collectingDate="2020-12-26" collectorName="R. Okada" country="Thailand" elevation="90" location="Bang Phra St." municipality="Si Racha District" specimenCount="1" specimenCount-male="1" stateProvince="Chonburi Province">
<collectingCountry name="Thailand">Thailand</collectingCountry>
:
<collectingRegion country="Thailand" name="Chon Buri">Chonburi Province</collectingRegion>
.
<specimenCount type="male">1♂</specimenCount>
,
<collectingMunicipality>Si Racha District</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:0E028AD633F156FC96DC5E7C696DE758:1DD4ED1D01DD66A32CAE51130E98353F" country="Thailand" municipality="Si Racha District" name="Bang Phra St." stateProvince="Chonburi Province">Bang Phra St.</location>
269 (alt.
<quantity metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">
<elevation metricMagnitude="1" metricUnit="m" metricValue="9.0" unit="m" value="90.0">90 m</elevation>
</quantity>
),
<collectingDate value="2020-12-26">26.XII.2020</collectingDate>
,
<collectorName>R. Okada</collectorName>
leg. (CRO) (Fig.
<figureCitation captionStart="Figures 1837" captionStartId="F7" captionText="Figures 18 - 37. Habitus of 18 Microdytes akitai 19 M. balkei 20 M. dimorphus 21 M. feryi (paratype, from Myanmar) 22 M. franzi 23 M. gabrielae 24 M. heineri 25 M. maculatus 26 M. mariannae 27 M. menopausis 28 M. paoloi 29 M. pasiricus 30 M. pederzanii 31 M. rocchii 32 M. schoedli 33 M. schoenmanni (from Myanmar) 34 M. schwendingeri 35 M. shepardi 36 M. shunichii 37 M. zetteli. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures18-37" httpUri="https://binary.pensoft.net/fig/842227" pageId="0" pageNumber="87">20</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="comments">
<paragraph pageId="0" pageNumber="87">Comments.</paragraph>
<paragraph pageId="0" pageNumber="87">
<taxonomicName authorityName="Wewalka" authorityYear="1997" class="Insecta" family="Dytiscidae" genus="Microdytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Microdytes dimorphus" order="Coleoptera" pageId="0" pageNumber="87" phylum="Arthropoda" rank="species" species="dimorphus">
<emphasis italics="true" pageId="0" pageNumber="87">Microdytes dimorphus</emphasis>
</taxonomicName>
was described based on a single male specimen from Khao Yai National Park and no additional records are known so far. Therefore, this is the second record of this species.
<bibRefCitation author="Wewalka, G" journalOrPublisher="Koleopterologische Rundschau" pageId="0" pageNumber="87" pagination="13 - 51" refId="B22" refString="Wewalka, G, 1997. Taxonomic revision of Microdytes Balfour-Browne. (Coleoptera: Dytiscidae). Koleopterologische Rundschau 67: 13 - 51" title="Taxonomic revision of Microdytes Balfour-Browne. (Coleoptera: Dytiscidae)." volume="67" year="1997">Wewalka (1997)</bibRefCitation>
suggested
<taxonomicName lsidName="M. dimorphus" pageId="0" pageNumber="87" rank="species" species="dimorphus">
<emphasis italics="true" pageId="0" pageNumber="87">M. dimorphus</emphasis>
</taxonomicName>
is more closely related to
<taxonomicName lsidName="M. menopausis" pageId="0" pageNumber="87" rank="species" species="menopausis">
<emphasis italics="true" pageId="0" pageNumber="87">M. menopausis</emphasis>
</taxonomicName>
Wewalka, 1997 (Fig.
<figureCitation captionStart="Figures 1837" captionStartId="F7" captionText="Figures 18 - 37. Habitus of 18 Microdytes akitai 19 M. balkei 20 M. dimorphus 21 M. feryi (paratype, from Myanmar) 22 M. franzi 23 M. gabrielae 24 M. heineri 25 M. maculatus 26 M. mariannae 27 M. menopausis 28 M. paoloi 29 M. pasiricus 30 M. pederzanii 31 M. rocchii 32 M. schoedli 33 M. schoenmanni (from Myanmar) 34 M. schwendingeri 35 M. shepardi 36 M. shunichii 37 M. zetteli. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures18-37" httpUri="https://binary.pensoft.net/fig/842227" pageId="0" pageNumber="87">27</figureCitation>
) but differs by having coarser punctures on the elytra similar in size, a punctured abdomen, a more produced clypeus, and larger second segment of antennae. The specimen examined in this study corresponds with
<taxonomicName lsidName="M. dimorphus" pageId="0" pageNumber="87" rank="species" species="dimorphus">
<emphasis italics="true" pageId="0" pageNumber="87">M. dimorphus</emphasis>
</taxonomicName>
due to the status of punctures on the elytra and abdomen, although the characteristics of clypeus and antenna are intermediate between
<taxonomicName lsidName="M. dimorphus" pageId="0" pageNumber="87" rank="species" species="dimorphus">
<emphasis italics="true" pageId="0" pageNumber="87">M. dimorphus</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="M. menopausis" pageId="0" pageNumber="87" rank="species" species="menopausis">
<emphasis italics="true" pageId="0" pageNumber="87">M. menopausis</emphasis>
</taxonomicName>
. This species can be distinguished from
<taxonomicName lsidName="M. menopausis" pageId="0" pageNumber="87" rank="species" species="menopausis">
<emphasis italics="true" pageId="0" pageNumber="87">M. menopausis</emphasis>
</taxonomicName>
in having a more slender body shape and indistinct elytral markings at both shoulders and post-median part (Fig.
<figureCitation captionStart="Figures 1837" captionStartId="F7" captionText="Figures 18 - 37. Habitus of 18 Microdytes akitai 19 M. balkei 20 M. dimorphus 21 M. feryi (paratype, from Myanmar) 22 M. franzi 23 M. gabrielae 24 M. heineri 25 M. maculatus 26 M. mariannae 27 M. menopausis 28 M. paoloi 29 M. pasiricus 30 M. pederzanii 31 M. rocchii 32 M. schoedli 33 M. schoenmanni (from Myanmar) 34 M. schwendingeri 35 M. shepardi 36 M. shunichii 37 M. zetteli. Scale bar: 1.0 mm." figureDoi="10.3897/zookeys.1159.99218.figures18-37" httpUri="https://binary.pensoft.net/fig/842227" pageId="0" pageNumber="87">20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="87" type="distribution">
<paragraph pageId="0" pageNumber="87">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="87">Thailand: Khao Yai NP [Nakhon Ratchasima or Nakhon Nayok Province] and Chonburi (first record) Province.</paragraph>
</subSubSection>
</treatment>
</document>