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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1145.96724" ID-Pensoft-Pub="1313-2970-1145-1" ID-Pensoft-UUID="ACBBD138B7375B0D9F63CE792A82F653" ID-ZooBank="830628C276CD4641BFC6144CD775ED6B" ModsDocID="1313-2970-1145-1" checkinTime="1675506069289" checkinUser="pensoft" docAuthor="Hedin, Marshal &amp; Milne, Marc A." docDate="2023" docId="424DB1213E9A5F6B822188C5D82DDA24" docLanguage="en" docName="ZooKeys 1145: 1-130" docOrigin="ZooKeys 1145" docPubDate="2023-02-03" docSource="http://dx.doi.org/10.3897/zookeys.1145.96724" docTitle="Nesticus stygius Gertsch 1984" docType="treatment" docVersion="1" id="ACBBD138B7375B0D9F63CE792A82F653" lastPageNumber="1" masterDocId="ACBBD138B7375B0D9F63CE792A82F653" masterDocTitle="New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains" masterLastPageNumber="130" masterPageNumber="1" pageNumber="1" updateTime="1675506069289" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>New species in old mountains: integrative taxonomy reveals ten new species and extensive short-range endemism in Nesticus spiders (Araneae, Nesticidae) from the southern Appalachian Mountains</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hedin, Marshal</mods:namePart>
<mods:affiliation>Department of Biology, San Diego State University, San Diego, California 92182 - 4614, USA</mods:affiliation>
<mods:nameIdentifier type="email">mhedin@sdsu.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Milne, Marc A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1943-0161</mods:nameIdentifier>
<mods:affiliation>Department of Biology, University of Indianapolis, Indianapolis, Indiana 46227, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1145</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>130</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1145.96724</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1145.96724</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1145-1</mods:identifier>
<mods:identifier type="ZooBank">830628C276CD4641BFC6144CD775ED6B</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:424DB1213E9A5F6B822188C5D82DDA24" httpUri="http://treatment.plazi.org/id/424DB1213E9A5F6B822188C5D82DDA24" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<taxonomicName ID-CoL="6S8W2" ID-ENA="60740" LSID="424DB121-3E9A-5F6B-8221-88C5D82DDA24" authority="Gertsch, 1984" authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus stygius" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stygius">Nesticus stygius Gertsch, 1984</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Graphical overview of terminology used to describe genitalic morphology. Tennessee, Overton Co., Obe Lee Cave, Nesticus stygius, ♂ MCH specimen # 1882 palp ventral (A), dorsal (B) C North Carolina, Haywood Co., near Steestachee Bald overlook, N. silvanus, MCH specimen # 1145, ♂ palp dorsal D North Carolina, Rutherford Co., Moonshiner's Cave, N. brimleyi, MCH 99 _ 014, ♂ paracymbium medial E North Carolina, Clay Co., Big Tuni Creek, MCH 02 _ 171, epigynum ventral F North Carolina, Henderson Co., W of Bat Cave, MCH 07 _ 134, epigynum ventral G North Carolina, Buncombe Co., SW of Cane River Gap, MCH 01 _ 167, epigynum dorsal." figureDoi="10.3897/zookeys.1145.96724.figure2" httpUri="https://binary.pensoft.net/fig/804453" pageId="0" pageNumber="1">Fig. 2A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Gertsch" authorityYear="1984" class="Arachnida" family="Nesticidae" genus="Nesticus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nesticus stygius" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="stygius">Nesticus stygius</taxonomicName>
Gertsch, 1984: 36, figs 170-172;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.904.1.1" author="Hedin, M" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 19" refId="B17" refString="Hedin, M, Dellinger, B, 2005. Descriptions of a new species and previously unknown males of Nesticus (Araneae: Nesticidae) from caves in eastern North America, with comments on species rarity. Zootaxa 904: 1 - 19, DOI: https://doi.org/10.11646/zootaxa.904.1.1" title="Descriptions of a new species and previously unknown males of Nesticus (Araneae: Nesticidae) from caves in eastern North America, with comments on species rarity." url="https://doi.org/10.11646/zootaxa.904.1.1" volume="904" year="2005">Hedin and Dellinger 2005</bibRefCitation>
: 8, figs 11, 12.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
New collections from
<materialsCitation collectingDate="1993-10-11" collectorName="M. Hedin, C. Phillips, Non" country="USA" county="Overton Co." location="Obe Lee Cave" specimenCount="3" specimenCount-female="3" stateProvince="Tennessee" typeStatus="type">
<typeStatus>type</typeStatus>
locality:
<collectingCountry name="United States of America">USA</collectingCountry>
-
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Tennessee">Tennessee</collectingRegion>
,
<collectingCounty>Overton Co.</collectingCounty>
</emphasis>
<specimenCount></specimenCount>
,
<specimenCount type="female">3♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:424DB1213E9A5F6B822188C5D82DDA24:A59421F5ABF3E6EFED9DB8BCB6775BE0" country="USA" county="Overton Co." name="Obe Lee Cave" stateProvince="Tennessee">Obe Lee Cave</location>
,
<locationDeviation location="Monterey">N of Monterey</locationDeviation>
;
<collectingDate value="1993-10-11">11 Oct. 1993</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>C. Phillips</collectorName>
leg.;
<emphasis bold="true" pageId="0" pageNumber="1">
<collectorName>Non</collectorName>
<materialsCitation collectingDate="1991-10-01" collectingDateMax="2017-10-01" collectingDateMin="1991-10-01" collectorName="M. L. Niemiller, K. D. Kendall, M. Hedin, S. O'Kane, K. Crandall, A. Gerber, R. Keith, J. Starrett, S. Thomas, N. Mann, N. S. Gladstone, E. T. Carter, L. Hayter" country="United States of America" county="Overton Co." location="Raven Bluff Cave" municipality="East Water Supply Cave" specimenCount="13" specimenCount-female="11" specimenCount-male="2" stateProvince="Tennessee" typeStatus="type">
<typeStatus>type</typeStatus>
material: -
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCounty>Overton Co.</collectingCounty>
</emphasis>
<specimenCount></specimenCount>
, 1 imm;
<collectingMunicipality>East Water Supply Cave</collectingMunicipality>
, TOV15;
<collectingDate value="2013-07-13">13 Jul. 2013</collectingDate>
;
<collectorName>M.L. Niemiller</collectorName>
,
<collectorName>K.D. Kendall</collectorName>
leg.; MLN 13-036; •
<specimenCount type="female">7♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:424DB1213E9A5F6B822188C5D82DDA24:CCA4082619FB8C4C1B3B48AE9ADE6222" country="United States of America" county="Overton Co." municipality="East Water Supply Cave" name="Raven Bluff Cave" stateProvince="Tennessee">Raven Bluff Cave</location>
,
<locationDeviation location="Allons">NW of Allons</locationDeviation>
;
<collectingDate value="1992-09-26">26 Sep. 1992</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>
S.
<normalizedToken originalValue="OKane">O'Kane</normalizedToken>
</collectorName>
leg.; •
<specimenCount type="male">2♂</specimenCount>
,
<specimenCount></specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:424DB1213E9A5F6B822188C5D82DDA24:B80A5324BAE9C2FC0AA6D578A377A549" country="United States of America" county="Overton Co." municipality="East Water Supply Cave" name="Raven Bluff Cave" stateProvince="Tennessee">Raven Bluff Cave</location>
;
<collectingDate value="1991-10-01">1 Oct. 1991</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>K. Crandall</collectorName>
,
<collectorName>A. Gerber</collectorName>
leg.; •
<specimenCount></specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:424DB1213E9A5F6B822188C5D82DDA24:2CB137917F36DD91F5554FBF47CE721A" country="United States of America" county="Overton Co." municipality="East Water Supply Cave" name="Raven Bluff Cave" stateProvince="Tennessee">Raven Bluff Cave</location>
;
<collectingDate value="2005-08-28">28 Aug. 2005</collectingDate>
;
<collectorName>M. Hedin</collectorName>
,
<collectorName>R. Keith</collectorName>
,
<collectorName>J. Starrett</collectorName>
,
<collectorName>S. Thomas</collectorName>
leg.; MCH 05_101; •
<specimenCount type="female">4♀</specimenCount>
, 1 imm;
<location LSID="urn:lsid:plazi:treatment:424DB1213E9A5F6B822188C5D82DDA24:1DE20B6ECC86E401FFAEA765AF504861" country="United States of America" county="Overton Co." municipality="East Water Supply Cave" name="Raven Bluff Cave" stateProvince="Tennessee">Raven Bluff Cave</location>
, TOV28;
<collectingDate value="2017-09-03">3 Sep. 2017</collectingDate>
;
<collectorName>M.L. Niemiller</collectorName>
,
<collectorName>N. Mann</collectorName>
leg.; MLN 17-008.6; •
<specimenCount></specimenCount>
, 1 imm;
<location LSID="urn:lsid:plazi:treatment:424DB1213E9A5F6B822188C5D82DDA24:BE09CD903C57EEEB8BB4F0152ADFAA20" country="United States of America" county="Overton Co." municipality="East Water Supply Cave" name="Webb Cave" stateProvince="Tennessee">Webb Cave</location>
, TOV39;
<collectingDate value="2017-10-01">1 Oct. 2017</collectingDate>
;
<collectorName>N.S. Gladstone</collectorName>
,
<collectorName>E.T. Carter</collectorName>
,
<collectorName>L. Hayter</collectorName>
leg.; NSG 17-TOV39.17
</materialsCitation>
</emphasis>
</materialsCitation>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Morphological diagnosis as in
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.904.1.1" author="Hedin, M" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 19" refId="B17" refString="Hedin, M, Dellinger, B, 2005. Descriptions of a new species and previously unknown males of Nesticus (Araneae: Nesticidae) from caves in eastern North America, with comments on species rarity. Zootaxa 904: 1 - 19, DOI: https://doi.org/10.11646/zootaxa.904.1.1" title="Descriptions of a new species and previously unknown males of Nesticus (Araneae: Nesticidae) from caves in eastern North America, with comments on species rarity." url="https://doi.org/10.11646/zootaxa.904.1.1" volume="904" year="2005">Hedin and Dellinger (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This highly troglomorphic taxon is only known from a few caves on the western margin of the Cumberland Plateau in Overton County, north-central Tennessee (Fig.
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution of the Nesticus archeri group, including Nesticus archeri, N. pecki, N. jemisinae, N. cressleri, and N. stygius. State boundaries and major cities shown for geographic context. Dashed lines circumscribe the distributions of N. pecki and N. stygius." figureDoi="10.3897/zookeys.1145.96724.figure8" httpUri="https://binary.pensoft.net/fig/804459" pageId="0" pageNumber="1">8</figureCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.904.1.1" author="Hedin, M" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="1 - 19" refId="B17" refString="Hedin, M, Dellinger, B, 2005. Descriptions of a new species and previously unknown males of Nesticus (Araneae: Nesticidae) from caves in eastern North America, with comments on species rarity. Zootaxa 904: 1 - 19, DOI: https://doi.org/10.11646/zootaxa.904.1.1" title="Descriptions of a new species and previously unknown males of Nesticus (Araneae: Nesticidae) from caves in eastern North America, with comments on species rarity." url="https://doi.org/10.11646/zootaxa.904.1.1" volume="904" year="2005">Hedin and Dellinger 2005</bibRefCitation>
: fig. 1).
</paragraph>
</subSubSection>
</treatment>
</document>