treatments-xml/data/30/09/D4/3009D414FFFDFFB439B6FB05FA0DA0B5.xml

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<mods:title id="D9B9F0E2D51F56860B2DA6A78CE22AE2">The genus Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) in the Colombian Andean Mountains</mods:title>
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<mods:namePart id="142521A24364B1730C550650BD928F21">Vallejo, Luís Fernando</mods:namePart>
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<mods:date id="409A828366ABB630FCA771AABB3026FB">2013</mods:date>
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<emphasis id="8AD4B910FFFDFFB639B6FB05FC56A74C" bold="true" box="[151,967,1268,1294]" pageId="4" pageNumber="105">
Key to the identification of male 
<taxonomicName id="7FA01E81FFFDFFB63B0EFB05FD59A74C" box="[559,712,1268,1294]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="8AD4B910FFFDFFB63B0EFB05FD59A74C" bold="true" box="[559,712,1268,1294]" italics="true" pageId="4" pageNumber="105">Phyllophaga</emphasis>
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species in 
<collectingCountry id="C0B72592FFFDFFB63A6CFB05FC56A74C" box="[845,967,1268,1294]" name="Colombia" pageId="4" pageNumber="105">Colombia</collectingCountry>
</emphasis>
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<paragraph id="B81F6502FFFDFFB639B6FAE4FDE8A76D" blockId="4.[151,967,1268,1327]" box="[151,633,1301,1327]" pageId="4" pageNumber="105">(modified from Morón and Vallejo 2007)</paragraph>
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<paragraph id="B81F6502FFFDFFB639B6FAA7FA0DA729" blockId="4.[151,1437,1365,2033]" box="[151,1436,1365,1387]" pageId="4" pageNumber="105">
<keyLead id="0351C637FFFDFFB639B6FAA7FA0DA729" box="[151,1436,1365,1387]" pageId="4" pageNumber="105">
1. Tarsal claws cleft or bifid (
<figureCitation id="209B7987FFFDFFB638C2FAA7FD88A729" box="[483,537,1366,1387]" captionStart="FIGURES 1 – 2" captionStartId="6.[151,264,1918,1941]" captionTargetBox="[381,1199,1516,1851]" captionTargetId="figure@6.[340,1247,1397,1897]" captionTargetPageId="6" captionText="FIGURES 1 – 2. Phyllophaga luridipennis diagnostic structures (1) Male protarsal claw. (2) Parameres, distal view. Scale: 1 mm." httpUri="https://zenodo.org/record/218999/files/figure.png" pageId="4" pageNumber="105">Fig. 1</figureCitation>
)................................................ 
<taxonomicName id="7FA01E81FFFDFFB63D42FAA4FAD9A729" authority="(Phytalus)" baseAuthorityName="Phytalus" box="[1123,1352,1365,1387]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="8AD4B910FFFDFFB63D42FAA4FB4FA729" box="[1123,1246,1365,1387]" italics="true" pageId="4" pageNumber="105">Phyllophaga</emphasis>
(
<emphasis id="8AD4B910FFFDFFB63DCDFAA4FAD1A729" box="[1260,1344,1365,1387]" italics="true" pageId="4" pageNumber="105">Phytalus</emphasis>
)
</taxonomicName>
..... 2
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFDFFB639B6FA85FA0DA7CB" blockId="4.[151,1437,1365,2033]" box="[151,1436,1395,1417]" pageId="4" pageNumber="105">
<keyLead id="0351C637FFFDFFB639B6FA85FA0DA7CB" box="[151,1436,1395,1417]" pageId="4" pageNumber="105">
- Tarsal claws clearly dentate (
<figureCitation id="209B7987FFFDFFB638D6FA85FDCFA7CB" box="[503,606,1396,1417]" captionStart-0="FIGURES 3 – 10" captionStart-1="FIGURES 11 – 12" captionStart-2="FIGURES 13 – 21" captionStartId-0="7.[151,264,1930,1953]" captionStartId-1="9.[151,264,1610,1633]" captionStartId-2="10.[151,264,1926,1949]" captionTargetBox-0="[171,1369,565,1895]" captionTargetBox-1="[226,1312,1059,1540]" captionTargetBox-2="[210,1329,201,1873]" captionTargetId-0="figure@7.[171,1415,565,1921]" captionTargetId-1="figure@9.[226,1360,1059,1589]" captionTargetId-2="figure@10.[153,1433,147,1924]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionTargetPageId-2="10" captionText-0="FIGURES 3 – 10. Phyllophaga obsoleta diagnostic structures. Dorsal view of pronotum (3), left antenna (4), pro, meso, and metalegs (5), right protarsal claw (6) dorsal view of abdomen apex (7), frontal view (8), lateral view of male genital capsule (9), transversal view (10). Scale: 1 mm." captionText-1="FIGURES 11 – 12. Phyllophaga chiriquina diagnostic structures, male genital capsule, lateral view (11) parameres, distal view (12). Scale: 1 mm." captionText-2="FIGURES 13 – 21. Phyllophaga impressipyga diagnostic structures, pronotum, dorsal view (13), right antenna, lateral view (14), right metatibia, lateral view (15), left anterior claw, lateral view (16), abdomen apex, posterior view (17), frontal view (18) male genital capsule, lateral view (19), frontal view (20), dorsal view (21). Scale: 1 mm." httpUri-0="https://zenodo.org/record/219000/files/figure.png" httpUri-1="https://zenodo.org/record/219001/files/figure.png" httpUri-2="https://zenodo.org/record/219002/files/figure.png" pageId="4" pageNumber="105">Figs. 5–16</figureCitation>
)...................................... 
<taxonomicName id="7FA01E81FFFDFFB63D1CFA82FAD9A7CB" box="[1085,1352,1395,1417]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="8AD4B910FFFDFFB63D1CFA82FB29A7CB" box="[1085,1208,1395,1417]" italics="true" pageId="4" pageNumber="105">Phyllophaga</emphasis>
(
<emphasis id="8AD4B910FFFDFFB63DE4FA82FAD1A7CB" box="[1221,1344,1395,1417]" italics="true" pageId="4" pageNumber="105">Phyllophaga</emphasis>
)
</taxonomicName>
..... 3
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<paragraph id="B81F6502FFFDFFB639B6FA60FA0DA7A3" blockId="4.[151,1437,1365,2033]" pageId="4" pageNumber="105">
<keyLead id="0351C637FFFDFFB639B6FA60FA0DA7A3" pageId="4" pageNumber="105">
2. Inner spur on the metatibia shorter than the external spur, fused with tibia margin. Last sternites slightly concave, lacking projections. Parameres long, thick, with the apical third in an increasingly acute angle, and not fused apically. (
<figureCitation id="209B7987FFFDFFB63DC6FA5FFAB0A781" box="[1255,1313,1454,1475]" captionStart="FIGURES 1 – 2" captionStartId="6.[151,264,1918,1941]" captionTargetBox="[381,1199,1516,1851]" captionTargetId="figure@6.[340,1247,1397,1897]" captionTargetPageId="6" captionText="FIGURES 1 – 2. Phyllophaga luridipennis diagnostic structures (1) Male protarsal claw. (2) Parameres, distal view. Scale: 1 mm." httpUri="https://zenodo.org/record/218999/files/figure.png" pageId="4" pageNumber="105">Fig. 2</figureCitation>
)................................................................................... 
<taxonomicName id="7FA01E81FFFDFFB63D72FA3DFA0DA7A3" authority="Moser" authorityName="Moser" box="[1107,1436,1483,1505]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="luridipennis" subGenus="Phytalus">
<emphasis id="8AD4B910FFFDFFB63D72FA3DFBF2A7A3" box="[1107,1123,1484,1505]" italics="true" pageId="4" pageNumber="105">P.</emphasis>
(
<emphasis id="8AD4B910FFFDFFB63D52FA3AFB56A7A3" box="[1139,1223,1483,1505]" italics="true" pageId="4" pageNumber="105">Phytalus</emphasis>
) 
<emphasis id="8AD4B910FFFDFFB63DF5FA3AFAD9A7A3" box="[1236,1352,1483,1505]" italics="true" pageId="4" pageNumber="105">luridipennis</emphasis>
(Moser)
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</keyLead>
</paragraph>
<paragraph id="B81F6502FFFDFFB639B6FA18FA0AA47B" blockId="4.[151,1437,1365,2033]" pageId="4" pageNumber="105">
<keyLead id="0351C637FFFDFFB639B6FA18FA0AA47B" pageId="4" pageNumber="105">
- Inner spur on the metatibia slightly shorter than the external spur, articulated with tibia margin. Last sternites with projections and abundant setae (
<figureCitation id="209B7987FFFDFFB63885F9F7FD93A459" box="[420,514,1542,1563]" captionStart="FIGURES 3 – 10" captionStartId="7.[151,264,1930,1953]" captionTargetBox="[171,1369,565,1895]" captionTargetId="figure@7.[171,1415,565,1921]" captionTargetPageId="7" captionText="FIGURES 3 – 10. Phyllophaga obsoleta diagnostic structures. Dorsal view of pronotum (3), left antenna (4), pro, meso, and metalegs (5), right protarsal claw (6) dorsal view of abdomen apex (7), frontal view (8), lateral view of male genital capsule (9), transversal view (10). Scale: 1 mm." httpUri="https://zenodo.org/record/219000/files/figure.png" pageId="4" pageNumber="105">Figs. 7–8</figureCitation>
). Parameres long, narrow, with the third apical angle fused apically. (
<figureCitation id="209B7987FFFDFFB63DB8F9F7FA91A459" box="[1177,1280,1542,1563]" captionStart="FIGURES 3 – 10" captionStartId="7.[151,264,1930,1953]" captionTargetBox="[171,1369,565,1895]" captionTargetId="figure@7.[171,1415,565,1921]" captionTargetPageId="7" captionText="FIGURES 3 – 10. Phyllophaga obsoleta diagnostic structures. Dorsal view of pronotum (3), left antenna (4), pro, meso, and metalegs (5), right protarsal claw (6) dorsal view of abdomen apex (7), frontal view (8), lateral view of male genital capsule (9), transversal view (10). Scale: 1 mm." httpUri="https://zenodo.org/record/219000/files/figure.png" pageId="4" pageNumber="105">Figs. 9–10</figureCitation>
)..................................................................................... 
<taxonomicName id="7FA01E81FFFDFFB63D71F9D5FA0AA47B" authority="Blanchard" authorityName="Blanchard" box="[1104,1435,1571,1593]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="obsoleta" subGenus="Phytalus">
<emphasis id="8AD4B910FFFDFFB63D71F9D5FBF2A47B" box="[1104,1123,1572,1593]" italics="true" pageId="4" pageNumber="105">P.</emphasis>
(
<emphasis id="8AD4B910FFFDFFB63D51F9D2FB55A47B" box="[1136,1220,1571,1593]" italics="true" pageId="4" pageNumber="105">Phytalus</emphasis>
) 
<emphasis id="8AD4B910FFFDFFB63DF3F9D2FAB5A47B" box="[1234,1316,1571,1593]" italics="true" pageId="4" pageNumber="105">obsoleta</emphasis>
(Blanchard)
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<paragraph id="B81F6502FFFDFFB639B6F9B0FA0DA414" blockId="4.[151,1437,1365,2033]" box="[151,1436,1600,1622]" pageId="4" pageNumber="105">
<keyLead id="0351C637FFFDFFB639B6F9B0FA0DA414" box="[151,1436,1600,1622]" pageId="4" pageNumber="105">
3. Inner metatibial spur much shorter than external spur, fused with tibia margin................ .. 
<taxonomicName id="7FA01E81FFFDFFB63DB8F9B0FA9BA414" box="[1177,1290,1600,1622]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="rorulenta">
<emphasis id="8AD4B910FFFDFFB63DB8F9B0FA9BA414" box="[1177,1290,1600,1622]" italics="true" pageId="4" pageNumber="105">P. rorulenta</emphasis>
</taxonomicName>
group..... 4
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFDFFB639B6F9AFFA0AA431" blockId="4.[151,1437,1365,2033]" box="[151,1435,1630,1651]" pageId="4" pageNumber="105">
<keyLead id="0351C637FFFDFFB639B6F9AFFA0AA431" box="[151,1435,1630,1651]" pageId="4" pageNumber="105">- Inner metatibial spur slightly shorter or as long as external spur, articulated with tibia margin.......................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFDFFB639B6F98DFA0DA4AB" pageId="4" pageNumber="105">
<paragraph id="B81F6502FFFDFFB639B6F98DFA0DA4EC" blockId="4.[151,1437,1365,2033]" pageId="4" pageNumber="105">
<keyLead id="0351C637FFFDFFB639B6F98DFA0DA4EC" pageId="4" pageNumber="105">
4. Pronotum dull chestnut-brown reddish; elytra dull yellowish chestnut-brown. Pygidial plate convex or with a slight preapical elongate depression. Paramere apex separated by a “v” shaped neckline. (
<figureCitation id="209B7987FFFDFFB63A80F968FC76A4EC" box="[929,999,1689,1710]" captionStart="FIGURES 11 – 12" captionStartId="9.[151,264,1610,1633]" captionTargetBox="[226,1312,1059,1540]" captionTargetId="figure@9.[226,1360,1059,1589]" captionTargetPageId="9" captionText="FIGURES 11 – 12. Phyllophaga chiriquina diagnostic structures, male genital capsule, lateral view (11) parameres, distal view (12). Scale: 1 mm." httpUri="https://zenodo.org/record/219001/files/figure.png" pageId="4" pageNumber="105">Fig. 12</figureCitation>
).................. 
<emphasis id="8AD4B910FFFDFFB63DE7F968FB40A4EC" box="[1222,1233,1689,1710]" italics="true" pageId="4" pageNumber="105">..</emphasis>
<taxonomicName id="7FA01E81FFFDFFB63DF0F968FA0DA4EC" authority="(Bates)" authorityName="Bates" baseAuthorityName="Bates" box="[1233,1436,1688,1710]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="chiriquina">
<emphasis id="8AD4B910FFFDFFB63DF0F968FAC1A4EC" box="[1233,1360,1688,1710]" italics="true" pageId="4" pageNumber="105">P. chiriquina</emphasis>
(Bates)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFDFFB639B6F947FA0DA4AB" blockId="4.[151,1437,1365,2033]" pageId="4" pageNumber="105">
<keyLead id="0351C637FFFDFFB639B6F947FA0DA4AB" pageId="4" pageNumber="105">
- Pronotum dark reddish chestnut-brown; pruinose, elytra grayish red; pygidial plate with wide but slight preapical depression. Paramere apices widely separated by a basal straight bridge (
<figureCitation id="209B7987FFFDFFB63A08F925FCE3A4AB" box="[809,882,1748,1769]" captionStart="FIGURES 13 – 21" captionStartId="10.[151,264,1926,1949]" captionTargetBox="[210,1329,201,1873]" captionTargetId="figure@10.[153,1433,147,1924]" captionTargetPageId="10" captionText="FIGURES 13 – 21. Phyllophaga impressipyga diagnostic structures, pronotum, dorsal view (13), right antenna, lateral view (14), right metatibia, lateral view (15), left anterior claw, lateral view (16), abdomen apex, posterior view (17), frontal view (18) male genital capsule, lateral view (19), frontal view (20), dorsal view (21). Scale: 1 mm." httpUri="https://zenodo.org/record/219002/files/figure.png" pageId="4" pageNumber="105">Fig. 20</figureCitation>
)............................ .. 
<taxonomicName id="7FA01E81FFFDFFB63DF2F925FA0DA4AB" authority="Frey" authorityName="Frey" box="[1235,1436,1748,1769]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="impressipyga">
<emphasis id="8AD4B910FFFDFFB63DF2F925FAFBA4AB" box="[1235,1386,1748,1769]" italics="true" pageId="4" pageNumber="105">P. impressipyga</emphasis>
Frey
</taxonomicName>
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<keyStep id="03547DA7FFFDFFB639B6F900FA0AA561" pageId="4" pageNumber="105">
<paragraph id="B81F6502FFFDFFB639B6F900FA0AA544" blockId="4.[151,1437,1365,2033]" box="[151,1435,1777,1798]" pageId="4" pageNumber="105">
<keyLead id="0351C637FFFDFFB639B6F900FA0AA544" box="[151,1435,1777,1798]" pageId="4" pageNumber="105">
5. Ventral denticles on the tarsal claws located near unguinal base (
<figureCitation id="209B7987FFFDFFB63A6AF900FC00A544" box="[843,913,1777,1798]" captionStart="FIGURES 31 – 33" captionStartId="13.[151,264,769,792]" captionTargetBox="[361,1320,268,716]" captionTargetId="figure@13.[226,1360,193,749]" captionTargetPageId="13" captionText="FIGURES 31 – 33. Phyllophaga brevisetosa diagnostic structures, left metatarsal claw (31) male genital capsule, frontal view (32), lateral view (33). (after Morón 2001). Scale: 1 mm." httpUri="https://zenodo.org/record/219004/files/figure.png" pageId="4" pageNumber="105">Fig. 31</figureCitation>
).......................................... 6
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFDFFB639B6F8FFFA0AA561" blockId="4.[151,1437,1365,2033]" box="[151,1435,1806,1827]" pageId="4" pageNumber="105">
<keyLead id="0351C637FFFDFFB639B6F8FFFA0AA561" box="[151,1435,1806,1827]" pageId="4" pageNumber="105">
- Ventral denticles on the tarsal claws located near the middle of the margin (
<figureCitation id="209B7987FFFDFFB63A92F8FFFC68A561" box="[947,1017,1806,1827]" captionStart="FIGURES 41 – 50" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[232,1365,244,1877]" captionTargetId="figure@16.[173,1414,193,1932]" captionTargetPageId="16" captionText="FIGURES 41 – 50. Phyllophaga schizorhina adult male. Pronotum, dorsal view (41), left antenna (42), right protarsal claw (43); right metatibia (44); distal appearance of abdomen, lateral view (45), dorsal view (46), ventral view (47); male genital capsule, lateral view (48 and 49), transversal view (50). (Figs. 48 – 50 after Morón 2003) Scale: 1 mm." httpUri="https://zenodo.org/record/219008/files/figure.png" pageId="4" pageNumber="105">Fig. 43</figureCitation>
)................................. 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFDFFB639B6F8DDFA0DA5F4" pageId="4" pageNumber="105">
<paragraph id="B81F6502FFFDFFB639B6F8DDFA0DA539" blockId="4.[151,1437,1365,2033]" pageId="4" pageNumber="105">
<keyLead id="0351C637FFFDFFB639B6F8DDFA0DA539" pageId="4" pageNumber="105">
6. Surface of pronotum and elytra bright, almost smooth, reddish chestnut-brown. Tarsal claws with partial ventral and irregular dentate margin, and ventral denticles located close to unguinal base (
<figureCitation id="209B7987FFFDFFB63AA5F8B8FC5FA51C" box="[900,974,1865,1886]" captionStart="FIGURES 80 – 81" captionStartId="24.[151,264,1307,1330]" captionTargetBox="[355,1144,934,1253]" captionTargetId="figure@24.[260,1326,839,1286]" captionTargetPageId="24" captionText="FIGURES 80 – 81. Phyllophaga caviceps. Protarsal claw (80) male genital capsule, lateral view (81). Scale: 1 mm." httpUri="https://zenodo.org/record/219016/files/figure.png" pageId="4" pageNumber="105">Fig. 80</figureCitation>
). Parameres long, fused along their length, and very curved ventrally. (
<figureCitation id="209B7987FFFDFFB6389EF897FD94A539" box="[447,517,1894,1915]" captionStart="FIGURES 80 – 81" captionStartId="24.[151,264,1307,1330]" captionTargetBox="[355,1144,934,1253]" captionTargetId="figure@24.[260,1326,839,1286]" captionTargetPageId="24" captionText="FIGURES 80 – 81. Phyllophaga caviceps. Protarsal claw (80) male genital capsule, lateral view (81). Scale: 1 mm." httpUri="https://zenodo.org/record/219016/files/figure.png" pageId="4" pageNumber="105">Fig. 81</figureCitation>
)............................................................ 
<taxonomicName id="7FA01E81FFFDFFB63DFCF897FA0DA539" authority="Moser" authorityName="Moser" box="[1245,1436,1894,1915]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="caviceps">
<emphasis id="8AD4B910FFFDFFB63DFCF897FAD8A539" box="[1245,1353,1894,1915]" italics="true" pageId="4" pageNumber="105">P. caviceps</emphasis>
(Moser)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFDFFB639B6F875FA0DA5F4" blockId="4.[151,1437,1365,2033]" pageId="4" pageNumber="105">
<keyLead id="0351C637FFFDFFB639B6F875FA0DA5F4" pageId="4" pageNumber="105">
- Surface of pronotum and elytra rather pruinose, dull or with minute scattered setae. Tarsal claws with smooth ventral margin and ventral denticles located near unguinal base and crooked or reclined against it. (
<figureCitation id="209B7987FFFDFFB63D24F850FBDAA5F4" box="[1029,1099,1953,1974]" captionStart="FIGURES 22 – 30" captionStartId="12.[151,264,1947,1970]" captionTargetBox="[218,1368,233,1947]" captionTargetId="figure@12.[218,1369,193,1947]" captionTargetPageId="12" captionText="FIGURES 22 – 30. Phyllophaga densata adult male. Pronotum, dorsal view (22), left antenna (23), right protibia (24); right metatibia (25); right protarsal claw (26) distal appearance of abdomen, frontal view (27), ventral view (28), male genital capsule, lateral view (29), transversal view (30). Scale: 1 mm." httpUri="https://zenodo.org/record/219003/files/figure.png" pageId="4" pageNumber="105">Fig. 26</figureCitation>
).... 
<taxonomicName id="7FA01E81FFFDFFB63DAAF850FA98A5F4" box="[1163,1289,1953,1974]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="8AD4B910FFFDFFB63DAAF850FB06A5F4" box="[1163,1175,1953,1974]" italics="true" pageId="4" pageNumber="105">P</emphasis>
. 
<emphasis id="8AD4B910FFFDFFB63D82F850FA98A5F4" box="[1187,1289,1953,1974]" italics="true" pageId="4" pageNumber="105">rugipennis</emphasis>
</taxonomicName>
group..... 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFDFFB739B6F84FFA0AA288" lastPageId="5" lastPageNumber="106" pageId="4" pageNumber="105">
<paragraph id="B81F6502FFFDFFB639B6F84FFA0DA5B3" blockId="4.[151,1437,1365,2033]" pageId="4" pageNumber="105">
<keyLead id="0351C637FFFDFFB639B6F84FFA0DA5B3" pageId="4" pageNumber="105">
7. Pronotum, elytra and pygidial plate without setae or microtrichia. Surface and legs shiny black. Parameres elongate, stout, slightly curved (
<figureCitation id="209B7987FFFDFFB6385CF82DFE62A5B3" box="[381,499,2012,2033]" captionStart="FIGURES 22 – 30" captionStartId="12.[151,264,1947,1970]" captionTargetBox="[218,1368,233,1947]" captionTargetId="figure@12.[218,1369,193,1947]" captionTargetPageId="12" captionText="FIGURES 22 – 30. Phyllophaga densata adult male. Pronotum, dorsal view (22), left antenna (23), right protibia (24); right metatibia (25); right protarsal claw (26) distal appearance of abdomen, frontal view (27), ventral view (28), male genital capsule, lateral view (29), transversal view (30). Scale: 1 mm." httpUri="https://zenodo.org/record/219003/files/figure.png" pageId="4" pageNumber="105">Figs. 29–30</figureCitation>
). Body length 
<quantity id="7F58C8E7FFFDFFB63B5EF82DFD77A5B3" box="[639,742,2012,2033]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.6" metricValueMin="2.0" pageId="4" pageNumber="105" unit="mm" value="23.0" valueMax="26.0" valueMin="20.0">20–26 mm</quantity>
.......................................... 
<taxonomicName id="7FA01E81FFFDFFB63DC7F82DFA0DA5B3" authority="Moser" authorityName="Moser" box="[1254,1436,2011,2033]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="densata">
<emphasis id="8AD4B910FFFDFFB63DC7F82DFAD9A5B3" box="[1254,1352,2011,2033]" italics="true" pageId="4" pageNumber="105">P. densata</emphasis>
(Moser)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FF66FA0AA288" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FF66FA0AA288" pageId="5" pageNumber="106">- Pronotum, elytra, and pygidial plate with microtrichia and scarce or abundant setae. Body surface dark brown to reddish brown, frequently dull, pruinose; legs shiny black to dark brown................................................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6FF23FA0AA37D" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6FF23FA0DA346" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FF23FA0DA346" pageId="5" pageNumber="106">
8. Pygidial plate with minute, scattered setae. Parameres with lateral-ventral, tooth-like, sharpened projections, clearly angular (
<figureCitation id="209B7987FFFCFFB739C9FF1EFEBFA346" box="[232,302,239,260]" captionStart="FIGURES 31 – 33" captionStartId="13.[151,264,769,792]" captionTargetBox="[361,1320,268,716]" captionTargetId="figure@13.[226,1360,193,749]" captionTargetPageId="13" captionText="FIGURES 31 – 33. Phyllophaga brevisetosa diagnostic structures, left metatarsal claw (31) male genital capsule, frontal view (32), lateral view (33). (after Morón 2001). Scale: 1 mm." httpUri="https://zenodo.org/record/219004/files/figure.png" pageId="5" pageNumber="106">Fig. 33</figureCitation>
)............................................................................ 
<taxonomicName id="7FA01E81FFFCFFB73DE5FF01FA0DA346" authority="Moser" authorityName="Moser" box="[1220,1436,239,261]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="brevisetosa">
<emphasis id="8AD4B910FFFCFFB73DE5FF01FAD8A347" box="[1220,1353,239,261]" italics="true" pageId="5" pageNumber="106">P. brevisetosa</emphasis>
(Moser)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FEFCFA0AA37D" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FEFCFA0AA37D" pageId="5" pageNumber="106">- Pygidial plate with abundant erect, short, or medium-sized setae. Parameres with no angular or sharpened lateral-ventral projections............................................................................................. 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6FEB6FA0AA3D5" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6FEB6FA0DA338" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FEB6FA0DA338" pageId="5" pageNumber="106">
9. Pygidial plate roughly dotted. Parameres with lateral-ventral tooth-like projections, short, rounded. (
<figureCitation id="209B7987FFFCFFB73D83FEB6FB78A31E" box="[1186,1257,327,348]" captionStart="FIGURES 34 – 35" captionStartId="14.[151,264,910,933]" captionTargetBox="[337,1301,592,872]" captionTargetId="figure@14.[226,1360,529,889]" captionTargetPageId="14" captionText="FIGURES 34 – 35. Phyllophaga pruinipennis diagnostic structures, male genital capsule, front view (34), lateral view (35). (after Morón 2001). Scale: 1 mm." httpUri="https://zenodo.org/record/219005/files/figure.png" pageId="5" pageNumber="106">Fig. 35</figureCitation>
)................................................................................................ 
<taxonomicName id="7FA01E81FFFCFFB73D96FE94FA0DA338" authority="Moser" authorityName="Moser" box="[1207,1436,357,378]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="pruinipennis">
<emphasis id="8AD4B910FFFCFFB73D96FE94FAD9A338" box="[1207,1352,357,378]" italics="true" pageId="5" pageNumber="106">P. pruinipennis</emphasis>
(Moser)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FE73FA0AA3D5" blockId="5.[151,1437,151,2021]" box="[151,1435,386,407]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FE73FA0AA3D5" box="[151,1435,386,407]" pageId="5" pageNumber="106">- Pygidial plate dotted. Parameres without tooth-like projections, short and abundant setae............................ 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6FE6EFA0DA3AD" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6FE6EFA0AA3F6" blockId="5.[151,1437,151,2021]" box="[151,1435,415,437]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FE6EFA0AA3F6" box="[151,1435,415,437]" pageId="5" pageNumber="106">
10. Parameres with many, rounded lateral-ventral projections (
<figureCitation id="209B7987FFFCFFB73A32FE6EFCC8A3F6" box="[787,857,415,436]" captionStart="FIGURE 36" captionStartId="14.[151,250,1388,1410]" captionTargetBox="[317,1265,1016,1360]" captionTargetId="figure@14.[309,1278,1013,1366]" captionTargetPageId="14" captionText="FIGURE 36. Phyllophaga pachypyga diagnostic structures, male genital capsule, lateral view (36). Scale: 1 mm." httpUri="https://zenodo.org/record/219006/files/figure.png" pageId="5" pageNumber="106">Fig. 36</figureCitation>
)......................... 
<taxonomicName id="7FA01E81FFFCFFB73DBDFE51FA0AA3F6" authority="Burmeister" authorityName="Burmeister" box="[1180,1435,415,437]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="pachypyga">
<emphasis id="8AD4B910FFFCFFB73DBDFE51FA8AA3F7" box="[1180,1307,415,437]" italics="true" pageId="5" pageNumber="106">P. pachypyga</emphasis>
(Burmeister)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FE4CFA0DA3AD" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FE4CFA0DA3AD" pageId="5" pageNumber="106">
- Parameres with short, rounded, and moderately separated lateral-ventral projections (
<figureCitation id="209B7987FFFCFFB73D2FFE4CFBC7A390" box="[1038,1110,445,466]" captionStart="FIGURES 37 – 40" captionStartId="15.[151,264,1073,1096]" captionTargetBox="[165,1393,261,983]" captionTargetId="figure@15.[151,1436,193,1053]" captionTargetPageId="15" captionText="FIGURES 37 – 40. Phyllophaga lissopyge diagnostic structures, right antenna, (37), metatarsal claw (38), right metatibia (39), male genital capsule, lateral view (after Morón 2001). Scale: 1 mm." httpUri="https://zenodo.org/record/219007/files/figure.png" pageId="5" pageNumber="106">Fig. 40</figureCitation>
). Aedeagus with lateral pre-apical patches of spines....................................................................... 
<taxonomicName id="7FA01E81FFFCFFB73DC1FE2BFA0DA3AD" authority="Bates" authorityName="Bates" box="[1248,1436,473,495]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="lissopyge">
<emphasis id="8AD4B910FFFCFFB73DC1FE2BFAC1A3AD" box="[1248,1360,473,495]" italics="true" pageId="5" pageNumber="106">P. lissopyge</emphasis>
(Bates)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6FE06FA0AA026" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6FE06FA0DA068" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FE06FA0DA068" pageId="5" pageNumber="106">
11. Anterior marginal of clypeus deeply notched or bilobed. Last abdominal sternite narrower than preceding. Aedeagus with strongly sclerotized support and large articulated spines. Parameres not fused ventrally........... 
<taxonomicName id="7FA01E81FFFCFFB73D8DFDE4FAA2A068" box="[1196,1331,532,554]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="schizorhina">
<emphasis id="8AD4B910FFFCFFB73D8DFDE4FB29A068" box="[1196,1208,533,554]" italics="true" pageId="5" pageNumber="106">P</emphasis>
. 
<emphasis id="8AD4B910FFFCFFB73DE5FDE5FAA2A068" box="[1220,1331,532,554]" italics="true" pageId="5" pageNumber="106">schizorhina</emphasis>
</taxonomicName>
group...12
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FDC3FA0AA026" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FDC3FA0AA026" pageId="5" pageNumber="106">- Anterior border of clypeus briefly notched, straight or curved. Last abdominal sternite nearly as wide as preceding one. Aedeagus with diverse structures, hardly sclerotized. Parameres usually fused ventrally................................. 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6FD9CFA0DA098" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6FD9CFA0DA0DD" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FD9CFA0DA0DD" pageId="5" pageNumber="106">
12. Ventral membrane of genital capsule without sclerotized plates (
<figureCitation id="209B7987FFFCFFB73A6EFD9CFC57A0C0" box="[847,966,621,642]" captionStart="FIGURES 41 – 50" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[232,1365,244,1877]" captionTargetId="figure@16.[173,1414,193,1932]" captionTargetPageId="16" captionText="FIGURES 41 – 50. Phyllophaga schizorhina adult male. Pronotum, dorsal view (41), left antenna (42), right protarsal claw (43); right metatibia (44); distal appearance of abdomen, lateral view (45), dorsal view (46), ventral view (47); male genital capsule, lateral view (48 and 49), transversal view (50). (Figs. 48 – 50 after Morón 2003) Scale: 1 mm." httpUri="https://zenodo.org/record/219008/files/figure.png" pageId="5" pageNumber="106">Figs. 48–49</figureCitation>
). Apex of aedeagus with weak sclerotized structures. Body length 
<quantity id="7F58C8E7FFFCFFB738B6FD7BFE6FA0DD" box="[407,510,650,671]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.4" metricValueMin="2.1" pageId="5" pageNumber="106" unit="mm" value="22.5" valueMax="24.0" valueMin="21.0">21–24 mm</quantity>
........................................................... 
<taxonomicName id="7FA01E81FFFCFFB73DE9FD7BFA0DA0DD" authority="Bates" authorityName="Bates" box="[1224,1436,649,671]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="schizorhina">
<emphasis id="8AD4B910FFFCFFB73DE9FD7BFAC1A0DD" box="[1224,1360,649,671]" italics="true" pageId="5" pageNumber="106">P. schizorhina</emphasis>
(Bates)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FD56FA0DA098" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FD56FA0DA098" pageId="5" pageNumber="106">
- Ventral membrane of genital capsule with sclerotized plates (
<figureCitation id="209B7987FFFCFFB73A1CFD56FC3DA0FE" box="[829,940,679,700]" captionStart="FIGURES 51 – 60" captionStartId="18.[151,264,1929,1952]" captionTargetBox="[209,1362,214,1853]" captionTargetId="figure@18.[181,1405,169,1908]" captionTargetPageId="18" captionText="FIGURES 51 – 60. Phyllophaga onoreana adult male. Pronotum, dorsal view (51); left antenna (52); right protarsal claw (53); right protibia (54); right metatibia (55); abdomen, posterior dorsal view (56), ventral view (57); male genital capsule, left lateral view (58), right lateral view (59), frontal view (60). Scale: 1 mm." httpUri="https://zenodo.org/record/219009/files/figure.png" pageId="5" pageNumber="106">Fig. 58–59</figureCitation>
). Apex of aedeagus with asymmetrical sclerotized hooks (
<figureCitation id="209B7987FFFCFFB73806FD34FEFEA098" box="[295,367,709,730]" captionStart="FIGURES 51 – 60" captionStartId="18.[151,264,1929,1952]" captionTargetBox="[209,1362,214,1853]" captionTargetId="figure@18.[181,1405,169,1908]" captionTargetPageId="18" captionText="FIGURES 51 – 60. Phyllophaga onoreana adult male. Pronotum, dorsal view (51); left antenna (52); right protarsal claw (53); right protibia (54); right metatibia (55); abdomen, posterior dorsal view (56), ventral view (57); male genital capsule, left lateral view (58), right lateral view (59), frontal view (60). Scale: 1 mm." httpUri="https://zenodo.org/record/219009/files/figure.png" pageId="5" pageNumber="106">Fig. 60</figureCitation>
). Body length 
<quantity id="7F58C8E7FFFCFFB738DDFD34FDF2A098" box="[508,611,709,730]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="5" pageNumber="106" unit="mm" value="24.5" valueMax="25.0" valueMin="24.0">24–25 mm</quantity>
.................................................... 
<taxonomicName id="7FA01E81FFFCFFB73DC0FD34FA0DA098" authority="Moron" authorityName="Moron" box="[1249,1436,709,730]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="onoreana">
<emphasis id="8AD4B910FFFCFFB73DC0FD34FAC2A098" box="[1249,1363,709,730]" italics="true" pageId="5" pageNumber="106">P. onoreana</emphasis>
Morón
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6FD13FA0AA10D" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6FD13FA0AA156" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FD13FA0AA156" pageId="5" pageNumber="106">
13. Metatibiae with spiniform denticles on superior, inferior, and external margins. Antennae with nine antennomeres. Pronotum and elytra pruinose, dull, reddish chestnut-brown, glabrous. Body length 16.0–18.5 mm............ 
<taxonomicName id="7FA01E81FFFCFFB73D9DFCF1FA0AA156" authority="Burmeister" authorityName="Burmeister" box="[1212,1435,767,789]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="roscida">
<emphasis id="8AD4B910FFFCFFB73D9DFCF1FA8AA157" box="[1212,1307,767,789]" italics="true" pageId="5" pageNumber="106">P. roscida</emphasis>
(Burmeister)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FCECFA0AA10D" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FCECFA0AA10D" pageId="5" pageNumber="106">- Metatibiae with one or two oblique transversal carinae more or less defined in dorsal-lateral region. Antennae with ten antennomeres. Bright or opaque, glabrous or setiferous dorsal surface............................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6FCA6FA0AA1A0" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6FCA6FA0DA1E5" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FCA6FA0DA1E5" pageId="5" pageNumber="106">
14. Posterior region of frons with very evident transversal projection. Dorsal surface glabrous, shiny, dark chestnut-brown. Parameres wide, depressed, independent in dorsal region and fused in ventral region. (Fig. 82) Body length approximately 
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................................................................................... 
<taxonomicName id="7FA01E81FFFCFFB73DC7FC63FA0DA1E5" authority="Saylor" authorityName="Saylor" box="[1254,1436,913,935]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="apolinari">
<emphasis id="8AD4B910FFFCFFB73DC7FC63FAC6A1E5" box="[1254,1367,913,935]" italics="true" pageId="5" pageNumber="106">P. apolinari</emphasis>
Saylor
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FC5EFA0AA1A0" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FC5EFA0AA1A0" pageId="5" pageNumber="106">- Posterior region of frons completely convex or with slight rugosity or discontinuity in texture. Dorsal surface setiferous, bright, or opaque. Parameres with different shapes and proportions than above.......................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6FC1BFA0AA65E" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6FC1BFA0AA1BD" blockId="5.[151,1437,151,2021]" box="[151,1435,1002,1023]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FC1BFA0AA1BD" box="[151,1435,1002,1023]" pageId="5" pageNumber="106">15. Elytra glabrous or with scarce, scattered setae, especially noticeable at base; tegument area with pruinose, dull appearance. 16</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FBF6FA0AA65E" blockId="5.[151,1437,151,2021]" box="[151,1435,1031,1052]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FBF6FA0AA65E" box="[151,1435,1031,1052]" pageId="5" pageNumber="106">- Elytra with abundant, dense, long, short or mixed setae; tegument with bright or opaque appearance................... 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6FBD4FA0AA6D0" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6FBD4FA0DA678" blockId="5.[151,1437,151,2021]" box="[151,1436,1061,1082]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FBD4FA0DA678" box="[151,1436,1061,1082]" pageId="5" pageNumber="106">16. Head and pronotum dark reddish chestnut-brown; elytra yellowish chestnut-brown. Scutellum without long setae........ 18</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FBB3FA0AA6D0" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FBB3FA0AA6D0" pageId="5" pageNumber="106">
- Head, pronotum, and elytra reddish chestnut-brown and chestnut-brown. Scutellum with long setae. Parameres short, with dentate lateral-ventral projections, fused, long apices, directed towards base of genital capsule (
<figureCitation id="209B7987FFFCFFB73D71FBAEFB07A636" box="[1104,1174,1119,1140]" captionStart="FIGURE 62" captionStartId="19.[151,250,1798,1820]" captionTargetBox="[375,1219,1340,1767]" captionTargetId="figure@19.[368,1219,1329,1777]" captionTargetPageId="19" captionText="FIGURE 62. Phyllophaga thoracica, male genital capsule, lateral view (62). Scale: 1 mm." httpUri="https://zenodo.org/record/219011/files/figure.png" pageId="5" pageNumber="106">Fig. 62</figureCitation>
) Body length 
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................................................................................. 
<taxonomicName id="7FA01E81FFFCFFB73D89FB8CFA0AA6D0" authority="Burmeister" authorityName="Burmeister" box="[1192,1435,1148,1170]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="thoracica">
<emphasis id="8AD4B910FFFCFFB73D89FB8CFA8AA6D0" box="[1192,1307,1148,1170]" italics="true" pageId="5" pageNumber="106">P. thoracica</emphasis>
(Burmeister)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6FB6BFA0DA745" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6FB6BFA0DA68E" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FB6BFA0DA68E" pageId="5" pageNumber="106">
17. Scutellum less punctate than elytra. Parameres short, narrow, with curved or rounded lateral-ventral projections, short, fused apices, ventral-apically aligned (
<figureCitation id="209B7987FFFCFFB73B38FB46FD1EA68E" box="[537,655,1207,1228]" captionStart="FIGURES 83 – 90" captionStartId="26.[151,264,1843,1866]" captionTargetBox="[166,1358,340,1791]" captionTargetId="figure@26.[151,1436,337,1822]" captionTargetPageId="26" captionText="FIGURES 83 – 90. Phyllophaga yucana, pronotum, dorsal view (83); right antenna (84); pro, meso, and metatibiae (85); right metatibia (86); abdomen apex, posterior view (87), frontal view (88); male genital capsule, lateral view (89), frontal view (90). Scale: 1 mm." httpUri="https://zenodo.org/record/219017/files/figure.png" pageId="5" pageNumber="106">Figs. 89–90</figureCitation>
). Body length 
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............................... 
<taxonomicName id="7FA01E81FFFCFFB73DDBFB49FA0DA68E" authority="Saylor" authorityName="Saylor" box="[1274,1436,1207,1229]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="yucana">
<emphasis id="8AD4B910FFFCFFB73DDBFB49FAC7A68F" box="[1274,1366,1208,1229]" italics="true" pageId="5" pageNumber="106">P. yucana</emphasis>
Saylor
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FB24FA0DA745" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FB24FA0DA745" pageId="5" pageNumber="106">
- Scutellum dense and finely punctate, greater than elytra. Parameres short, with very short, curved, rounded lateral-ventral projections; apices fused, slightly ventral-apically projected (
<figureCitation id="209B7987FFFCFFB73A2BFB03FCC3A745" box="[778,850,1266,1287]" captionStart="FIGURE 91" captionStartId="27.[151,250,1559,1581]" captionTargetBox="[406,1183,1211,1528]" captionTargetId="figure@27.[393,1194,1202,1537]" captionTargetPageId="27" captionText="FIGURE 91. Phyllophaga bicolor, male genital capsule, lateral view (91). Scale: 1 mm." httpUri="https://zenodo.org/record/219018/files/figure.png" pageId="5" pageNumber="106">Fig. 91</figureCitation>
). Body length 
<quantity id="7F58C8E7FFFCFFB73AFEFB03FBD7A745" box="[991,1094,1266,1287]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="5" pageNumber="106" unit="mm" value="16.5" valueMax="17.0" valueMin="16.0">16–17 mm</quantity>
............. 
<taxonomicName id="7FA01E81FFFCFFB73DCAFB03FA0DA745" authority="Moser" authorityName="Moser" box="[1259,1436,1265,1287]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="8AD4B910FFFCFFB73DCAFB03FAD9A745" box="[1259,1352,1265,1287]" italics="true" pageId="5" pageNumber="106">P. bicolor</emphasis>
(Moser)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6FAFEFA0AA73E" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6FAFEFA0DA700" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FAFEFA0DA700" pageId="5" pageNumber="106">
18. Base of elytra with longer, erect setae. Parameres short, with long, rounded lateral-ventral projections, widely separated from the fused apices, ventral-basally aligned (
<figureCitation id="209B7987FFFCFFB73B4FFADCFD25A700" box="[622,692,1325,1346]" captionStart="FIGURE 61" captionStartId="19.[151,250,809,831]" captionTargetBox="[379,1209,200,777]" captionTargetId="figure@19.[372,1215,193,788]" captionTargetPageId="19" captionText="FIGURE 61. Phyllophaga ruficollis male genital capsule, lateral view (61) (after Morón 2001). Scale: 1 mm." httpUri="https://zenodo.org/record/219010/files/figure.png" pageId="5" pageNumber="106">Fig. 61</figureCitation>
) Body length approximately 
<quantity id="7F58C8E7FFFCFFB73AECFADCFB81A700" box="[973,1040,1325,1346]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="106" unit="mm" value="18.0">18 mm</quantity>
................. 
<taxonomicName id="7FA01E81FFFCFFB73DFBFADCFA0DA700" authority="Moser" authorityName="Moser" box="[1242,1436,1324,1346]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="ruficollis">
<emphasis id="8AD4B910FFFCFFB73DFBFADCFAD9A700" box="[1242,1352,1324,1346]" italics="true" pageId="5" pageNumber="106">P. ruficollis</emphasis>
(Moser)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FABBFA0AA73E" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FABBFA0AA73E" pageId="5" pageNumber="106">
- Base of elytra without longer erect setae. Parameres short with different 
<typeStatus id="671BDBA0FFFCFFB73AB6FABBFC5BA71D" box="[919,970,1354,1375]" pageId="5" pageNumber="106">types</typeStatus>
of lateral-ventral projections.................................................................................................. 
<taxonomicName id="7FA01E81FFFCFFB73D9DFA99FAA5A73F" box="[1212,1332,1383,1405]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="setidorsis">
<emphasis id="8AD4B910FFFCFFB73D9DFA99FB5AA73F" box="[1212,1227,1384,1405]" italics="true" pageId="5" pageNumber="106">P</emphasis>
. 
<emphasis id="8AD4B910FFFCFFB73DF9FA96FAA5A73F" box="[1240,1332,1383,1405]" italics="true" pageId="5" pageNumber="106">setidorsis</emphasis>
</taxonomicName>
group...19
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</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6FA74FA0AA7F5" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6FA74FA0AA7D8" blockId="5.[151,1437,151,2021]" box="[151,1435,1413,1434]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FA74FA0AA7D8" box="[151,1435,1413,1434]" pageId="5" pageNumber="106">
19. Upper margin of clypeus strongly curved (
<figureCitation id="209B7987FFFCFFB73B59FA74FD2FA7D8" box="[632,702,1413,1434]" captionStart="FIGURES 70 – 77" captionStartId="22.[151,264,1953,1976]" captionTargetBox="[336,1219,235,1884]" captionTargetId="figure@22.[270,1316,193,1932]" captionTargetPageId="22" captionText="FIGURES 70 – 77. Phyllophaga menetriesi adult male. Pronotum, dorsal view (70), left antenna (71), pro, meso- and metalegs (72), right metatibia (73), abdomen apex, rear view (74), frontal view (75), male genital capsule, frontal view (76), lateral view (77). Scale: 1 mm." httpUri="https://zenodo.org/record/219013/files/figure.png" pageId="5" pageNumber="106">Fig. 70</figureCitation>
).......................................................... 20
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FA53FA0AA7F5" blockId="5.[151,1437,151,2021]" box="[151,1435,1442,1463]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FA53FA0AA7F5" box="[151,1435,1442,1463]" pageId="5" pageNumber="106">- Upper margin of clypeus slightly sinuous. (Fig. 131)......................................................... 27</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6FA4EFA0AA44D" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6FA4EFA0DA796" blockId="5.[151,1437,151,2021]" box="[151,1436,1471,1493]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FA4EFA0DA796" box="[151,1436,1471,1493]" pageId="5" pageNumber="106">
20. Body length 
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. Pronotum and elytra uniformly covered with abundant, short setae (
<figureCitation id="209B7987FFFCFFB73D78FA4EFB0FA796" box="[1113,1182,1471,1492]" captionStart="FIGURES 63 – 69" captionStartId="20.[151,264,1953,1976]" captionTargetBox="[346,1317,257,1891]" captionTargetId="figure@20.[252,1336,193,1932]" captionTargetPageId="20" captionText="FIGURES 63 – 69. Phyllophaga gigantea adult female. Pronotum, dorsal view (63), right antenna (64), right protarsal claw (65), right metatibia (66), abdomen apex appearance, lateral view (67), frontal view (68), ventral view (69). Scale: 1 mm." httpUri="https://zenodo.org/record/219012/files/figure.png" pageId="5" pageNumber="106">Fig. 63</figureCitation>
)..... 
<taxonomicName id="7FA01E81FFFCFFB73DC0FA31FA0DA796" authority="Bates" authorityName="Bates" box="[1249,1436,1471,1493]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="gigantea">
<emphasis id="8AD4B910FFFCFFB73DC0FA31FAC1A797" box="[1249,1360,1472,1493]" italics="true" pageId="5" pageNumber="106">P. gigantea</emphasis>
(Bates)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6FA2CFA0AA44D" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6FA2CFA0AA44D" pageId="5" pageNumber="106">
- Body length 
<quantity id="7F58C8E7FFFCFFB7387CFA2CFE59A7B0" box="[349,456,1501,1522]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.1" metricValueMin="1.5" pageId="5" pageNumber="106" unit="mm" value="18.0" valueMax="21.0" valueMin="15.0">15–21 mm</quantity>
. Pronotum and elytra with or without long, scattered, erect setae; pronotal vestiture usually less dense than on elytra........................................................................................ 21
</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6F9E6FA0AA4C6" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6F9E6FA0AA425" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6F9E6FA0AA425" pageId="5" pageNumber="106">
21. Pronotum, uniformly, minutely punctate. Parameres short, wide, with sharp, straight lateral-ventral projections; thin sharp apical projection; aedeagus with wide, curved, sclerotized dorsal lamella. (
<figureCitation id="209B7987FFFCFFB73AAEF9C4FB93A408" box="[911,1026,1589,1610]" captionStart="FIGURES 70 – 77" captionStartId="22.[151,264,1953,1976]" captionTargetBox="[336,1219,235,1884]" captionTargetId="figure@22.[270,1316,193,1932]" captionTargetPageId="22" captionText="FIGURES 70 – 77. Phyllophaga menetriesi adult male. Pronotum, dorsal view (70), left antenna (71), pro, meso- and metalegs (72), right metatibia (73), abdomen apex, rear view (74), frontal view (75), male genital capsule, frontal view (76), lateral view (77). Scale: 1 mm." httpUri="https://zenodo.org/record/219013/files/figure.png" pageId="5" pageNumber="106">Figs. 76–77</figureCitation>
) Body length, 
<quantity id="7F58C8E7FFFCFFB73DB0F9C4FB69A408" box="[1169,1272,1589,1610]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="5" pageNumber="106" unit="mm" value="20.5" valueMax="21.0" valueMin="20.0">20–21 mm</quantity>
............................................................................................. 
<taxonomicName id="7FA01E81FFFCFFB73D80F9A3FA0AA425" authority="Blanchard" authorityName="Blanchard" box="[1185,1435,1618,1639]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="menetriesii">
<emphasis id="8AD4B910FFFCFFB73D80F9A3FAB2A425" box="[1185,1315,1618,1639]" italics="true" pageId="5" pageNumber="106">P. menetriesii</emphasis>
(Blanchard)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6F99EFA0AA4C6" blockId="5.[151,1437,151,2021]" box="[151,1435,1647,1668]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6F99EFA0AA4C6" box="[151,1435,1647,1668]" pageId="5" pageNumber="106">- Pronotum irregularly punctate........................................................................... 22</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6F97CFA0AA49E" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6F97CFA0AA4FD" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6F97CFA0AA4FD" pageId="5" pageNumber="106">
22. Anterior border of clypeus slightly curved. Parameres short, with lateral-ventral projections located near articulation at base of phallus with a short apical projection (
<figureCitation id="209B7987FFFCFFB73B70F95BFD09A4FD" box="[593,664,1706,1727]" captionStart="FIGURE 92" captionStartId="28.[151,250,624,646]" captionTargetBox="[472,1116,203,593]" captionTargetId="figure@28.[460,1128,195,602]" captionTargetPageId="28" captionText="FIGURE 92. Phyllophaga longicornis male genital capsule, lateral view (92). Scale: 1 mm." httpUri="https://zenodo.org/record/219019/files/figure.png" pageId="5" pageNumber="106">Fig. 92</figureCitation>
). Body length, 
<quantity id="7F58C8E7FFFCFFB73A0AF95BFCFCA4FD" box="[811,877,1706,1727]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="106" unit="mm" value="20.0">20 mm</quantity>
......................... 
<taxonomicName id="7FA01E81FFFCFFB73DBFF95BFA0AA4FD" authority="Blanchard" authorityName="Blanchard" box="[1182,1435,1705,1727]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="8AD4B910FFFCFFB73DBFF95BFAB2A4FD" box="[1182,1315,1705,1727]" italics="true" pageId="5" pageNumber="106">P. longicornis</emphasis>
(Blanchard)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6F936FA0AA49E" blockId="5.[151,1437,151,2021]" box="[151,1435,1735,1756]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6F936FA0AA49E" box="[151,1435,1735,1756]" pageId="5" pageNumber="106">- Anterior border of clypeus very curved................................................................... 23</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6F914FA0AA510" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6F914FA0AA576" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6F914FA0AA576" pageId="5" pageNumber="106">
23. Lateral border of pronotum smooth, non-crenulate. Pronotum without setae on mid line. Parameres short, with long, curved, sharpened lateral-ventral projections, and long, sharp apical projection (
<figureCitation id="209B7987FFFCFFB73AB0F8F3FC46A555" box="[913,983,1794,1815]" captionStart="FIGURE 78" captionStartId="23.[151,250,1211,1233]" captionTargetBox="[381,1204,810,1183]" captionTargetId="figure@23.[373,1213,806,1190]" captionTargetPageId="23" captionText="FIGURE 78. Phyllophaga fragilipennis, male genital capsule, lateral view (78). Scale: 1 mm." httpUri="https://zenodo.org/record/219014/files/figure.png" pageId="5" pageNumber="106">Fig. 78</figureCitation>
) Body length 
<quantity id="7F58C8E7FFFCFFB73D41F8F3FB56A555" box="[1120,1223,1794,1815]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="5" pageNumber="106" unit="mm" value="16.5" valueMax="17.0" valueMin="16.0">16–17 mm</quantity>
................................................................................................ 
<taxonomicName id="7FA01E81FFFCFFB73DACF8D1FA0AA576" authority="Blanchard" authorityName="Blanchard" box="[1165,1435,1823,1845]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="fragilipennis">
<emphasis id="8AD4B910FFFCFFB73DACF8D1FAB2A577" box="[1165,1315,1823,1845]" italics="true" pageId="5" pageNumber="106">P. fragilipennis</emphasis>
(Blanchard)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6F8CCFA0AA510" blockId="5.[151,1437,151,2021]" box="[151,1435,1853,1874]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6F8CCFA0AA510" box="[151,1435,1853,1874]" pageId="5" pageNumber="106">- Lateral margin of pronotum finely crenulated.............................................................. 24</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFCFFB739B6F8ABFA0DA5A6" pageId="5" pageNumber="106">
<paragraph id="B81F6502FFFCFFB739B6F8ABFA0AA5CE" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6F8ABFA0AA5CE" pageId="5" pageNumber="106">24. Pronotum with abundant setae all over surface. Antennal club as long as length of preceding six antennomeres combined. Tarsal claws with acute basal tooth shorter than apical tooth located near middle of ventral border....................... 25</keyLead>
</paragraph>
<paragraph id="B81F6502FFFCFFB739B6F864FA0DA5A6" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="106">
<keyLead id="0351C637FFFCFFB739B6F864FA0DA5A6" pageId="5" pageNumber="106">
- Pronotum without setae. Antennal club slightly longer than preceding six antennomeres combined (
<figureCitation id="209B7987FFFCFFB73D98F864FA9EA5E8" box="[1209,1295,1941,1962]" captionStart="FIGURES 101 – 109" captionStartId="30.[151,264,1953,1976]" captionTargetBox="[376,1246,237,1876]" captionTargetId="figure@30.[286,1300,193,1932]" captionTargetPageId="30" captionText="FIGURES 101 – 109. Phyllophaga tesorito adult male. Pronotum, dorsal view (101), left antenna (102), left protarsal claw (103) left metatibia (104), abdomen apex appearance, ventral view (105), frontal view (106), male genital capsule, dorsal view (107), lateral view (108), ventral view (109). Scale: 1 mm." httpUri="https://zenodo.org/record/219021/files/figure.png" pageId="5" pageNumber="106">Fig. 102</figureCitation>
). Tarsal claws with truncate basal tooth shorter than apical tooth located toward basal dilatation of ventral border (
<figureCitation id="209B7987FFFCFFB73D9AF843FA80A585" box="[1211,1297,1970,1991]" captionStart="FIGURES 101 – 109" captionStartId="30.[151,264,1953,1976]" captionTargetBox="[376,1246,237,1876]" captionTargetId="figure@30.[286,1300,193,1932]" captionTargetPageId="30" captionText="FIGURES 101 – 109. Phyllophaga tesorito adult male. Pronotum, dorsal view (101), left antenna (102), left protarsal claw (103) left metatibia (104), abdomen apex appearance, ventral view (105), frontal view (106), male genital capsule, dorsal view (107), lateral view (108), ventral view (109). Scale: 1 mm." httpUri="https://zenodo.org/record/219021/files/figure.png" pageId="5" pageNumber="106">Fig. 103</figureCitation>
). Body length approximately 
<quantity id="7F58C8E7FFFCFFB73850F83EFE25A5A6" box="[369,436,1999,2020]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="106" unit="mm" value="24.0">24 mm</quantity>
.............................................................. 
<taxonomicName id="7FA01E81FFFCFFB73DBDF821FA0DA5A6" authority="Vallejo &amp; Wolff" authorityName="Vallejo &amp; Wolff" box="[1180,1436,1999,2021]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="tesorito">
<emphasis id="8AD4B910FFFCFFB73DBDF821FB6CA5A7" box="[1180,1277,2000,2021]" italics="true" pageId="5" pageNumber="106">P. tesorito</emphasis>
Vallejo &amp; Wolff
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFFFFB439B6FF66FA0AA360" pageId="6" pageNumber="107">
<paragraph id="B81F6502FFFFFFB439B6FF66FA0DA2A5" blockId="6.[151,1437,151,759]" pageId="6" pageNumber="107">
<keyLead id="0351C637FFFFFFB439B6FF66FA0DA2A5" pageId="6" pageNumber="107">
25. Pronotum with umbiliform punctures, wider towards the anterior border. Dorsal and ventral color dull dark chestnut-brown. Parameres short, wide, with very short lateral-ventral projections, rounded, directed towards base, and erect apical projection (
<figureCitation id="209B7987FFFFFFB439C9FF23FEFBA2A5" box="[232,362,210,231]" captionStart="FIGURES 93 – 100" captionStartId="29.[151,264,1953,1976]" captionTargetBox="[295,1263,288,1885]" captionTargetId="figure@29.[238,1348,193,1932]" captionTargetPageId="29" captionText="FIGURES 93 – 100. Phyllophaga martinezi adult male. Pronotum, dorsal view (93), left antenna (94), right protarsal claw (95), right metatibia (96) abdomen apex, posterior view (97), frontal view (98), male genital capsule, lateral view (99), frontal view (100). Scale: 1 mm." httpUri="https://zenodo.org/record/219020/files/figure.png" pageId="6" pageNumber="107">Figs. 99–100</figureCitation>
). Body length approximately 
<quantity id="7F58C8E7FFFFFFB43BA7FF23FD58A2A5" box="[646,713,210,231]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="107" unit="mm" value="16.0">16 mm</quantity>
.............................................. 
<taxonomicName id="7FA01E81FFFFFFB43DD6FF23FA0DA2A5" authority="Frey" authorityName="Frey" box="[1271,1436,210,231]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="8AD4B910FFFFFFB43DD6FF23FAF8A2A5" box="[1271,1385,210,231]" italics="true" pageId="6" pageNumber="107">P. martinezi</emphasis>
Frey
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFFFFB439B6FF1EFA0AA360" blockId="6.[151,1437,151,759]" pageId="6" pageNumber="107">
<keyLead id="0351C637FFFFFFB439B6FF1EFA0AA360" pageId="6" pageNumber="107">- Pronotum without umbiliform punctures, covered in small dots. Ventral and dorsal color dark chestnut-brown and light chestnut-brown, respectively................................................................................ 26</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFFFFB439B6FEDBFA0DA3F6" pageId="6" pageNumber="107">
<paragraph id="B81F6502FFFFFFB439B6FEDBFA0DA31E" blockId="6.[151,1437,151,759]" pageId="6" pageNumber="107">
<keyLead id="0351C637FFFFFFB439B6FEDBFA0DA31E" pageId="6" pageNumber="107">
26. Pronotum and elytra dark chestnut-brown. Parameres short, wide, with short, sharp, wide lateral ventral projections, rounded and creased along mid line apical projection (
<figureCitation id="209B7987FFFFFFB43BB4FEB6FD4CA31E" box="[661,733,327,348]" captionStart="FIGURE 79" captionStartId="23.[151,250,1999,2021]" captionTargetBox="[364,1223,1574,1969]" captionTargetId="figure@23.[355,1232,1564,1977]" captionTargetPageId="23" captionText="FIGURE 79. Phyllophaga densepunctata, male genital capsule, lateral view (79) (after Morón 2001). Scale: 1 mm." httpUri="https://zenodo.org/record/219015/files/figure.png" pageId="6" pageNumber="107">Fig. 79</figureCitation>
). Body length 
<quantity id="7F58C8E7FFFFFFB43A4BFEB6FC40A31E" box="[874,977,327,348]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="6" pageNumber="107" unit="mm" value="18.0" valueMax="19.0" valueMin="17.0">17–19 mm</quantity>
.................. 
<taxonomicName id="7FA01E81FFFFFFB43D87FEB9FA0DA31E" authority="Moser" authorityName="Moser" box="[1190,1436,327,349]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="species" species="densepunctata">
<emphasis id="8AD4B910FFFFFFB43D87FEB9FAD9A31F" box="[1190,1352,327,349]" italics="true" pageId="6" pageNumber="107">P. densepunctata</emphasis>
(Moser)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFFFFB439B6FE94FA0DA3F6" blockId="6.[151,1437,151,759]" pageId="6" pageNumber="107">
<keyLead id="0351C637FFFFFFB439B6FE94FA0DA3F6" pageId="6" pageNumber="107">
- Pronotum dark reddish chestnut-brown, elytra chestnut-brown to dark chestnut. Parameres short with long lateral-ventral projections, very narrow, slightly curved, and the apical projection divided in two short, sharpened, separate denticles (
<figureCitation id="209B7987FFFFFFB43C17FE73FE99A3F6" captionStart="FIGURES 110 – 115" captionStartId="32.[151,264,1953,1976]" captionTargetBox="[376,1246,237,1876]" captionTargetId="figure@32.[286,1300,193,1932]" captionTargetPageId="32" captionText="FIGURES 110 – 115. Phyllophaga apolinaria. Pronotum, dorsal view (110), left antenna (111), left metatarsal claw (112), left metatibia (113) male genital capsule, lateral view (114), frontal view (115). Scale: 1 mm." httpUri="https://zenodo.org/record/219022/files/figure.png" pageId="6" pageNumber="107">Figs. 114– 115</figureCitation>
). Body length, 
<quantity id="7F58C8E7FFFFFFB438BBFE6EFD91A3F6" box="[410,512,415,436]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="6" pageNumber="107" unit="mm" value="19.5" valueMax="21.0" valueMin="18.0">18–21 mm</quantity>
............................................................. 
<taxonomicName id="7FA01E81FFFFFFB43DF8FE51FA0DA3F6" authority="Saylor" authorityName="Saylor" box="[1241,1436,415,437]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="species" species="apolinaria">
<emphasis id="8AD4B910FFFFFFB43DF8FE51FAC9A3F7" box="[1241,1368,415,437]" italics="true" pageId="6" pageNumber="107">P. apolinaria</emphasis>
Saylor
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFFFFB439B6FE4CFA0AA068" pageId="6" pageNumber="107">
<paragraph id="B81F6502FFFFFFB439B6FE4CFA0DA04E" blockId="6.[151,1437,151,759]" pageId="6" pageNumber="107">
<keyLead id="0351C637FFFFFFB439B6FE4CFA0DA04E" pageId="6" pageNumber="107">
27. Protarsal claws distinct from mesotarsal and metatarsal claws, lower tooth wider than upper tooth, located near to apex (
<figureCitation id="209B7987FFFFFFB43C54FE4CFE99A3AD" captionStart="FIGURES 116 – 124" captionStartId="33.[151,264,1953,1976]" captionTargetBox="[240,1313,258,1881]" captionTargetId="figure@33.[197,1389,193,1932]" captionTargetPageId="33" captionText="FIGURES 116 – 124. Phyllophaga guanacasteca adult male. Pronotum, dorsal view (116), left antenna (117), right protarsal claw (118), right metatarsal claw (119), right metatibia (120), abdomen apex appearance, frontal view (121), ventral view (122), male genital capsule, transversal view (123), lateral view (124). Scale: 1 mm." httpUri="https://zenodo.org/record/219023/files/figure.png" pageId="6" pageNumber="107">Fig. 118</figureCitation>
). Parameres with an acute tooth on each inner border (
<figureCitation id="209B7987FFFFFFB43A30FE2BFC31A3AD" box="[785,928,474,495]" captionStart="FIGURES 116 – 124" captionStartId="33.[151,264,1953,1976]" captionTargetBox="[240,1313,258,1881]" captionTargetId="figure@33.[197,1389,193,1932]" captionTargetPageId="33" captionText="FIGURES 116 – 124. Phyllophaga guanacasteca adult male. Pronotum, dorsal view (116), left antenna (117), right protarsal claw (118), right metatarsal claw (119), right metatibia (120), abdomen apex appearance, frontal view (121), ventral view (122), male genital capsule, transversal view (123), lateral view (124). Scale: 1 mm." httpUri="https://zenodo.org/record/219023/files/figure.png" pageId="6" pageNumber="107">Figs. 123–124</figureCitation>
). Body length 
<quantity id="7F58C8E7FFFFFFB43D0AFE2BFB07A3AD" box="[1067,1174,474,495]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="6" pageNumber="107" unit="mm" value="21.5" valueMax="22.0" valueMin="21.0">21–22 mm</quantity>
.................................................................................................. 
<taxonomicName id="7FA01E81FFFFFFB43D51FE09FA0DA04E" authority="Moron &amp; Solis" authorityName="Moron &amp; Solis" box="[1136,1436,503,525]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="species" species="guanacasteca">
<emphasis id="8AD4B910FFFFFFB43D51FE09FA94A04F" box="[1136,1285,504,525]" italics="true" pageId="6" pageNumber="107">P.guanacasteca</emphasis>
Morón &amp; Solís
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFFFFB439B6FDE4FA0AA068" blockId="6.[151,1437,151,759]" box="[151,1435,533,554]" pageId="6" pageNumber="107">
<keyLead id="0351C637FFFFFFB439B6FDE4FA0AA068" box="[151,1435,533,554]" pageId="6" pageNumber="107">- Protarsal claws with same shape as mesotarsal and metatarsal claws. Parameres without tooth on inner border........... 28</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFFFFB439B6FDC3FA0AA0DD" pageId="6" pageNumber="107">
<paragraph id="B81F6502FFFFFFB439B6FDC3FA0DA026" blockId="6.[151,1437,151,759]" pageId="6" pageNumber="107">
<keyLead id="0351C637FFFFFFB439B6FDC3FA0DA026" pageId="6" pageNumber="107">
28. First two metatarsomeres similar in length. Parameres short, fused in a ring shape with two denticles on mid-ventral line (
<figureCitation id="209B7987FFFFFFB43C4DFDC3FEA9A026" captionStart="FIGURES 125 – 129" captionStartId="34.[151,264,1937,1960]" captionTargetBox="[257,1353,721,1851]" captionTargetId="figure@34.[151,1436,660,1917]" captionTargetPageId="34" captionText="FIGURES 125 – 129. Phyllophaga aguadita adult male. Pronotum, dorsal view (125), left antenna (126), right metatarsal claw (127), male genital capsule, dorsal view (128), frontal view (129). Scale: 1 mm." httpUri="https://zenodo.org/record/219024/files/figure.png" pageId="6" pageNumber="107">Figs. 128–129</figureCitation>
). Body length approximately 
<quantity id="7F58C8E7FFFFFFB43B74FDBEFD09A026" box="[597,664,591,612]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="107" unit="mm" value="20.0">20 mm</quantity>
................................................. 
<taxonomicName id="7FA01E81FFFFFFB43DC8FDA1FA0DA026" authority="Saylor" authorityName="Saylor" box="[1257,1436,591,613]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="species" species="aguadita">
<emphasis id="8AD4B910FFFFFFB43DC8FDA1FAC7A027" box="[1257,1366,591,613]" italics="true" pageId="6" pageNumber="107">P. aguadita</emphasis>
Saylor
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFFFFB439B6FD9CFA0AA0DD" blockId="6.[151,1437,151,759]" pageId="6" pageNumber="107">
<keyLead id="0351C637FFFFFFB439B6FD9CFA0AA0DD" pageId="6" pageNumber="107">- First two metatarsomeres diferent in length. Parameres elongate, ventrally fused with an almost straight sclerotized bridge, with short lateral-apical projections...................................................................... 29</keyLead>
</paragraph>
</keyStep>
<keyStep id="03547DA7FFFFFFB439B6FD56FA0DA0B5" pageId="6" pageNumber="107">
<paragraph id="B81F6502FFFFFFB439B6FD56FA0DA0FE" blockId="6.[151,1437,151,759]" box="[151,1436,679,701]" pageId="6" pageNumber="107">
<keyLead id="0351C637FFFFFFB439B6FD56FA0DA0FE" box="[151,1436,679,701]" pageId="6" pageNumber="107">
29. Aedeagus with numerous long setae (
<figureCitation id="209B7987FFFFFFB43B69FD56FD0DA0FE" box="[584,668,679,700]" captionStart="FIGURE 130" captionStartId="35.[151,250,1828,1850]" captionTargetBox="[356,1230,1209,1800]" captionTargetId="figure@35.[348,1239,1199,1805]" captionTargetPageId="35" captionText="FIGURE 130. Phyllophaga transversicollis male genital capsule, lateral view (130). Scale: 1 mm." httpUri="https://zenodo.org/record/219025/files/figure.png" pageId="6" pageNumber="107">Fig. 130</figureCitation>
). Body length approximately 
<quantity id="7F58C8E7FFFFFFB43A98FD56FC6DA0FE" box="[953,1020,679,700]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="107" unit="mm" value="18.0">18 mm</quantity>
............. 
<taxonomicName id="7FA01E81FFFFFFB43DB8FD59FA0DA0FE" authority="Moser" authorityName="Moser" box="[1177,1436,679,701]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="species" species="transversicollis">
<emphasis id="8AD4B910FFFFFFB43DB8FD59FAD8A0FF" box="[1177,1353,679,701]" italics="true" pageId="6" pageNumber="107">P. transversicollis</emphasis>
(Moser)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="B81F6502FFFFFFB439B6FD34FA0DA0B5" blockId="6.[151,1437,151,759]" pageId="6" pageNumber="107">
<keyLead id="0351C637FFFFFFB439B6FD34FA0DA0B5" pageId="6" pageNumber="107">
- Aedeagus with lateral sclerotized plates, abundant micro-spines (Figs. 138–139). Body length approximately 
<quantity id="7F58C8E7FFFFFFB43DDAFD34FAAFA098" box="[1275,1342,709,730]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="107" unit="mm" value="20.0">20 mm</quantity>
............................................................................................. .. 
<taxonomicName id="7FA01E81FFFFFFB43DD9FD13FA0DA0B5" authority="Frey" authorityName="Frey" box="[1272,1436,737,759]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="species" species="schneblei">
<emphasis id="8AD4B910FFFFFFB43DD9FD13FAFBA0B5" box="[1272,1386,737,759]" italics="true" pageId="6" pageNumber="107">P. schneblei</emphasis>
Frey
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>