136 lines
17 KiB
XML
136 lines
17 KiB
XML
<document id="B58776D5A15BF4E4B12EE38F321E2567" ID-DOI="10.11646/zootaxa.3722.2.1" ID-GBIF-Dataset="f2c46514-f165-43d3-a574-47c55b89dcd6" ID-ISSN="1175-5326" ID-Zenodo-Dep="218998" ID-ZooBank="29D77A16-096D-4FC1-A5B4-9EEDF2E761A7" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460690754500" checkinUser="plazi" docAuthor="Vallejo, Luís Fernando & Wolff, Martha" docDate="2013" docId="3009D414FFE6FF9039B6F9E2FC9EA3B7" docLanguage="en" docName="zt03722p142.pdf" docOrigin="Zootaxa 3722 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phyllophaga (Phyllophaga) guanacasteca Moron and Solis 2000" docType="treatment" docVersion="8" lastPageNumber="135" masterDocId="CC30AC6CFFF9FFB23921FFF1FF91A242" masterDocTitle="The genus Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) in the Colombian Andean Mountains" masterLastPageNumber="142" masterPageNumber="101" pageNumber="132" updateTime="1698351836216" updateUser="plazi">
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<mods:titleInfo id="3DDCE892DF2370F99A3A1A0A7F989568">
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<mods:title id="10D4826A9ECF9A86C4847E9A826EDFAC">The genus Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) in the Colombian Andean Mountains</mods:title>
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<mods:namePart id="A38CF78CA6A487CEECACE9891E4D1E40">Vallejo, Luís Fernando</mods:namePart>
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<mods:name id="D17C3EB0FF68B82AD891FB95488D56CE" type="personal">
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<mods:namePart id="8E9C8511A7C68C642D1C590C44EB4B39">Wolff, Martha</mods:namePart>
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<mods:title id="33F9391C368F24303AA4CCA9DA44FF23">Zootaxa</mods:title>
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<mods:part id="B6D1F7036F701888F156A6318A366E38">
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<mods:date id="10D15529FBC98B8626919B71F4F178FF">2013</mods:date>
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<mods:number id="8FB2665F75E113731B11C62387DC0C35">3722</mods:number>
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<mods:identifier id="06BF2DEE5657AC455E46C0E1518B417F" type="DOI">10.11646/zootaxa.3722.2.1</mods:identifier>
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<treatment id="3009D414FFE6FF9039B6F9E2FC9EA3B7" ID-DOI="http://doi.org/10.5281/zenodo.6151969" ID-Zenodo-Dep="6151969" LSID="urn:lsid:plazi:treatment:3009D414FFE6FF9039B6F9E2FC9EA3B7" httpUri="http://treatment.plazi.org/id/3009D414FFE6FF9039B6F9E2FC9EA3B7" lastPageId="34" lastPageNumber="135" pageId="31" pageNumber="132">
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<paragraph id="B81F6502FFE6FFAD39B6F9E2FC4AA46F" blockId="31.[151,987,1554,1581]" box="[151,987,1554,1581]" pageId="31" pageNumber="132">
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<heading id="E357D26EFFE6FFAD39B6F9E2FC4AA46F" bold="true" box="[151,987,1554,1581]" fontSize="11" level="1" pageId="31" pageNumber="132" reason="1">
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<key id="CC313233FFE6FFAD39B6F9E2FC4AA46F" box="[151,987,1554,1581]" pageId="31" pageNumber="132">
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<emphasis id="8AD4B910FFE6FFAD39B6F9E2FC4AA46F" bold="true" box="[151,987,1554,1581]" pageId="31" pageNumber="132">
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27.
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<taxonomicName id="7FA01E81FFE6FFAD39E2F9E3FC4AA46F" authority="Moron and Solis, 2000" authorityName="Moron and Solis" authorityYear="2000" box="[195,987,1554,1581]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="132" phylum="Arthropoda" rank="species" species="guanacasteca" subGenus="Phyllophaga">
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<emphasis id="8AD4B910FFE6FFAD39E2F9E3FECDA46E" bold="true" box="[195,348,1554,1580]" italics="true" pageId="31" pageNumber="132">Phyllophaga</emphasis>
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(
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<emphasis id="8AD4B910FFE6FFAD384CF9E3FD97A46E" bold="true" box="[365,518,1554,1580]" italics="true" pageId="31" pageNumber="132">Phyllophaga</emphasis>
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)
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<emphasis id="8AD4B910FFE6FFAD3B36F9E3FD2AA46E" bold="true" box="[535,699,1554,1580]" italics="true" pageId="31" pageNumber="132">guanacasteca</emphasis>
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Morón and Solís, 2000
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</taxonomicName>
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</emphasis>
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</key>
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</keyStep>
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</keyLead>
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</heading>
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</paragraph>
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<paragraph id="B81F6502FFE6FF9039B6F9A8FDFDA35F" blockId="31.[151,1437,1625,2010]" lastBlockId="34.[151,1436,151,501]" lastPageId="34" lastPageNumber="135" pageId="31" pageNumber="132">
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<emphasis id="8AD4B910FFE6FFAD39B6F9A8FED5A430" bold="true" box="[151,324,1625,1650]" pageId="31" pageNumber="132">Redescription.</emphasis>
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Length 21.7 mm in males and 22.2 mm in females. The species has an elongate, oval body; reddish chestnut-brown in color; with a dull, glabrous surface; elytra has longitudinal striations, consisting of fine punctures. Pronotum (
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<figureCitation id="209B7987FFE6FFAD38B2F950FE65A4F8" box="[403,500,1697,1722]" captionStart="FIGURES 116 – 124" captionStartId="33.[151,264,1953,1976]" captionTargetBox="[240,1313,258,1881]" captionTargetId="figure@33.[197,1389,193,1932]" captionTargetPageId="33" captionText="FIGURES 116 – 124. Phyllophaga guanacasteca adult male. Pronotum, dorsal view (116), left antenna (117), right protarsal claw (118), right metatarsal claw (119), right metatibia (120), abdomen apex appearance, frontal view (121), ventral view (122), male genital capsule, transversal view (123), lateral view (124). Scale: 1 mm." httpUri="https://zenodo.org/record/219023/files/figure.png" pageId="31" pageNumber="132">Fig. 116</figureCitation>
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) is wider than it is long, darker than the corrugated elytra, with a surface densely covered in minute punctures; it has straight lateral, non-granulated margins; head has minute punctures, especially on the vertex and the clypeus. Clypeus is wide, with two pronounced lobes and a raised, concave anterior margin. Antennae (
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<figureCitation id="209B7987FFE6FFAD3834F8FCFEE9A564" box="[277,376,1805,1830]" captionStart="FIGURES 116 – 124" captionStartId="33.[151,264,1953,1976]" captionTargetBox="[240,1313,258,1881]" captionTargetId="figure@33.[197,1389,193,1932]" captionTargetPageId="33" captionText="FIGURES 116 – 124. Phyllophaga guanacasteca adult male. Pronotum, dorsal view (116), left antenna (117), right protarsal claw (118), right metatarsal claw (119), right metatibia (120), abdomen apex appearance, frontal view (121), ventral view (122), male genital capsule, transversal view (123), lateral view (124). Scale: 1 mm." httpUri="https://zenodo.org/record/219023/files/figure.png" pageId="31" pageNumber="132">Fig. 117</figureCitation>
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) have 10 antennomeres with short, compact antennal clubs. Protarsal claws (
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<figureCitation id="209B7987FFE6FFAD3C25F8FCFAF7A564" box="[1284,1382,1805,1830]" captionStart="FIGURES 116 – 124" captionStartId="33.[151,264,1953,1976]" captionTargetBox="[240,1313,258,1881]" captionTargetId="figure@33.[197,1389,193,1932]" captionTargetPageId="33" captionText="FIGURES 116 – 124. Phyllophaga guanacasteca adult male. Pronotum, dorsal view (116), left antenna (117), right protarsal claw (118), right metatarsal claw (119), right metatibia (120), abdomen apex appearance, frontal view (121), ventral view (122), male genital capsule, transversal view (123), lateral view (124). Scale: 1 mm." httpUri="https://zenodo.org/record/219023/files/figure.png" pageId="31" pageNumber="132">Fig. 118</figureCitation>
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) are deeply cleft with a wide, sharp tooth very close to the ungual base; flattened protibiae are not very wide, with three toothed projections on the external margin, the third tooth being reduced. Metatibiae (
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<figureCitation id="209B7987FFE6FFAD3D7DF8A4FB2DA52C" box="[1116,1212,1877,1902]" captionStart="FIGURES 116 – 124" captionStartId="33.[151,264,1953,1976]" captionTargetBox="[240,1313,258,1881]" captionTargetId="figure@33.[197,1389,193,1932]" captionTargetPageId="33" captionText="FIGURES 116 – 124. Phyllophaga guanacasteca adult male. Pronotum, dorsal view (116), left antenna (117), right protarsal claw (118), right metatarsal claw (119), right metatibia (120), abdomen apex appearance, frontal view (121), ventral view (122), male genital capsule, transversal view (123), lateral view (124). Scale: 1 mm." httpUri="https://zenodo.org/record/219023/files/figure.png" pageId="31" pageNumber="132">Fig. 120</figureCitation>
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) are long and wide, with two spurs, of which the inner is shorter than the outer and is fused with the tibia. Metatibiae have toothed carinae in the mid region. Pygidium (
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<figureCitation id="209B7987FFE6FFAD3B66F86CFD7DA5F4" box="[583,748,1949,1974]" captionStart="FIGURES 116 – 124" captionStartId="33.[151,264,1953,1976]" captionTargetBox="[240,1313,258,1881]" captionTargetId="figure@33.[197,1389,193,1932]" captionTargetPageId="33" captionText="FIGURES 116 – 124. Phyllophaga guanacasteca adult male. Pronotum, dorsal view (116), left antenna (117), right protarsal claw (118), right metatarsal claw (119), right metatibia (120), abdomen apex appearance, frontal view (121), ventral view (122), male genital capsule, transversal view (123), lateral view (124). Scale: 1 mm." httpUri="https://zenodo.org/record/219023/files/figure.png" pageId="31" pageNumber="132">Figs. 121–122</figureCitation>
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) is wide and evenly convex; fifth abdominal sternites, have long, fine setae on each side. The male genital capsule (
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<figureCitation id="209B7987FFE6FFAD3A26F830FC38A598" box="[775,937,1985,2010]" captionStart="FIGURES 116 – 124" captionStartId="33.[151,264,1953,1976]" captionTargetBox="[240,1313,258,1881]" captionTargetId="figure@33.[197,1389,193,1932]" captionTargetPageId="33" captionText="FIGURES 116 – 124. Phyllophaga guanacasteca adult male. Pronotum, dorsal view (116), left antenna (117), right protarsal claw (118), right metatarsal claw (119), right metatibia (120), abdomen apex appearance, frontal view (121), ventral view (122), male genital capsule, transversal view (123), lateral view (124). Scale: 1 mm." httpUri="https://zenodo.org/record/219023/files/figure.png" pageId="31" pageNumber="132">Figs. 123–124</figureCitation>
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) is short, cylindrical, and curved, with two enormous parameres, slightly longer than two thirds of the genital capsule, each with a symmetrical tooth projected from the mid distal region. The base of the aedeagus has a cylindrical and sclerotized structure from which a membranous sac, widening in the distal zone, is projected. The complete structure is clearly different in appearance to the rest of the abovementioned species.
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</paragraph>
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</subSubSection>
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<caption id="ECDF358AFFD9FF9239B6F850FB8DA594" httpUri="https://zenodo.org/record/219022/files/figure.png" pageId="32" pageNumber="133" targetBox="[376,1246,237,1876]" targetPageId="32">
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<paragraph id="B81F6502FFD9FF9239B6F850FB8DA594" blockId="32.[151,1436,1953,2006]" pageId="32" pageNumber="133">
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<emphasis id="8AD4B910FFD9FF9239B6F850FEFBA5F4" bold="true" box="[151,362,1953,1976]" pageId="32" pageNumber="133">FIGURES 110–115.</emphasis>
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<emphasis id="8AD4B910FFD9FF923853F850FDFCA5F5" box="[370,621,1953,1975]" italics="true" pageId="32" pageNumber="133">
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<taxonomicName id="7FA01E81FFD9FF923853F850FDF9A5F5" box="[370,616,1953,1975]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="133" phylum="Arthropoda" rank="species" species="apolinaria">Phyllophaga apolinaria</taxonomicName>
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.
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</emphasis>
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Pronotum, dorsal view (110), left antenna (111), left metatarsal claw (112), left metatibia (113) male genital capsule, lateral view (114), frontal view (115). Scale: 1 mm.
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</paragraph>
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</caption>
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<caption id="ECDF358AFFD8FF9339B6F850FC7AA5B6" httpUri="https://zenodo.org/record/219023/files/figure.png" pageId="33" pageNumber="134" targetBox="[240,1313,258,1881]" targetPageId="33">
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<paragraph id="B81F6502FFD8FF9339B6F850FC7AA5B6" blockId="33.[151,1436,1953,2036]" pageId="33" pageNumber="134">
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<emphasis id="8AD4B910FFD8FF9339B6F850FEFFA5F4" bold="true" box="[151,366,1953,1976]" pageId="33" pageNumber="134">FIGURES 116–124.</emphasis>
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<taxonomicName id="7FA01E81FFD8FF933854F850FD1BA5F5" box="[373,650,1953,1975]" class="Insecta" family="Melolonthidae" genus="Phyllophaga" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="134" phylum="Arthropoda" rank="species" species="guanacasteca">
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<emphasis id="8AD4B910FFD8FF933854F850FD1BA5F5" box="[373,650,1953,1975]" italics="true" pageId="33" pageNumber="134">Phyllophaga guanacasteca</emphasis>
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</taxonomicName>
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adult male. Pronotum, dorsal view (116), left antenna (117), right protarsal claw (118), right metatarsal claw (119), right metatibia (120), abdomen apex appearance, frontal view (121), ventral view (122), male genital capsule, transversal view (123), lateral view (124). Scale: 1 mm.
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</paragraph>
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</caption>
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<subSubSection id="F0BA3689FFDBFF9039E6FED6FC9EA3B7" pageId="34" pageNumber="135" type="discussion">
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<paragraph id="B81F6502FFDBFF9039E6FED6FC8FA3CA" blockId="34.[151,1436,151,501]" pageId="34" pageNumber="135">
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<emphasis id="8AD4B910FFDBFF9039E6FED6FEDEA302" bold="true" box="[199,335,295,320]" pageId="34" pageNumber="135">Comments.</emphasis>
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The diagnostic features of this species do not correspond to any group of species as described by Morón (1986, 1996, 2003). The Colombian specimens have slight differences with specimens from Guanacaste,
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<collectingCountry id="C0B72592FFDBFF9039B6FE81FE82A3CA" box="[151,275,367,392]" name="Costa Rica" pageId="34" pageNumber="135">Costa Rica</collectingCountry>
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, but these may represent geographic variation.
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</paragraph>
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<paragraph id="B81F6502FFDBFF9039E6FE65FC9EA3B7" blockId="34.[151,1436,151,501]" pageId="34" pageNumber="135">
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<emphasis id="8AD4B910FFDBFF9039E6FE65FC41A3EF" bold="true" box="[199,976,404,429]" pageId="34" pageNumber="135">
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New geographic records. Departamento del Tolima (
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<collectingCountry id="C0B72592FFDBFF903A71FE65FC59A3EF" box="[848,968,404,429]" name="Colombia" pageId="34" pageNumber="135">Colombia</collectingCountry>
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)
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</emphasis>
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: Municipio de Fresno, vereda Alegrías (
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<quantity id="7F58C8E7FFDBFF9039BEFE46FE91A392" box="[159,256,439,464]" metricMagnitude="0" metricUnit="m" metricValue="1.56" pageId="34" pageNumber="135" unit="m" value="1.56">1,560 m</quantity>
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), cafetal,
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<date id="CC1E43C2FFDBFF903851FE46FDBAA392" box="[368,555,439,464]" pageId="34" pageNumber="135" value="2008-09">September 2008</date>
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, Fernando Vallejo,
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<specimenCount id="AEA6AE8BFFDBFF903A26FE49FCF3A392" box="[775,866,439,464]" pageId="34" pageNumber="135" type="male">3 males</specimenCount>
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,
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<specimenCount id="AEA6AE8BFFDBFF903A4FFE49FC4EA392" box="[878,991,439,464]" pageId="34" pageNumber="135" type="female">2 females</specimenCount>
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; verada La Mireya (
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<quantity id="7F58C8E7FFDBFF903DE4FE46FAB6A392" box="[1221,1319,439,464]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="135" unit="m" value="1.5">1,500 m</quantity>
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),
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<date id="CC1E43C2FFDBFF903C1DFE46FEBBA3B6" pageId="34" pageNumber="135" value="2010-07" valueMax="2010-08-31" valueMin="2010-07-00">July and August 2010</date>
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, 2012 Fernando Vallejo,
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<specimenCount id="AEA6AE8BFFDBFF903B63FE2DFD0AA3B7" box="[578,667,476,501]" pageId="34" pageNumber="135" type="male">5 males</specimenCount>
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,
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<specimenCount id="AEA6AE8BFFDBFF903B87FE2DFC9AA3B7" box="[678,779,476,501]" pageId="34" pageNumber="135" type="female">1 female</specimenCount>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |