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<document ID-DOI="10.11646/phytotaxa.595.2.4" ID-ISSN="1179-3163" ID-Zenodo-Dep="7905867" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1683527015091" checkinUser="plazi" docAuthor="Büneker, Henrique Mallmann, Pastori, Tamara, Almeida, Pedro Schwambach De &amp; Mariath, Jorge Ernesto De Araujo" docDate="2023" docId="FB6AEA65FF99FFC922F0F9A2E76DFB66" docLanguage="en" docName="phytotaxa.595.2.4.pdf" docOrigin="Phytotaxa 595 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.595.2.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.6:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="6" docTitle="Dyckia selloa Baker 1889" docType="treatment" docVersion="3" lastPageNumber="172" masterDocId="0753921DFF9BFFCA2278FFF6E238FFAA" masterDocTitle="Dyckia magnifica, a new species of Bromeliaceae (Pitcairnioideae) from Southern Brazil, and the re-establishment of D. tomentosa, of the D. selloa complex" masterLastPageNumber="185" masterPageNumber="169" pageNumber="171" updateTime="1683565175284" updateUser="tatiana" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title>Dyckia magnifica, a new species of Bromeliaceae (Pitcairnioideae) from Southern Brazil, and the re-establishment of D. tomentosa, of the D. selloa complex</mods:title>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Büneker, Henrique Mallmann</mods:namePart>
<mods:affiliation>Laboratório de Anatomia Vegetal (LAVeg), Instituto de Biociências, Departamento de Botânica, Universidade Federal do Rio Grande do Sul (UFRGS), Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil. &amp; Programa de Pós-graduação em Botânica, Instituto de Biociências, UFRGS, Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pastori, Tamara</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-2477-0275</mods:nameIdentifier>
<mods:affiliation>Laboratório de Anatomia Vegetal (LAVeg), Instituto de Biociências, Departamento de Botânica, Universidade Federal do Rio Grande do Sul (UFRGS), Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil. &amp; Programa de Pós-graduação em Botânica, Instituto de Biociências, UFRGS, Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil. &amp; tamarapastori @ gmail. com; https: // orcid. org / 0000 - 0002 - 2477 - 0275</mods:affiliation>
<mods:nameIdentifier type="email">tamarapastori@gmail.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Almeida, Pedro Schwambach De</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-2042-3687</mods:nameIdentifier>
<mods:affiliation>Laboratório de Anatomia Vegetal (LAVeg), Instituto de Biociências, Departamento de Botânica, Universidade Federal do Rio Grande do Sul (UFRGS), Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil. &amp; Laboratório de Palinologia, Departamento de Botânica, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, Rio de Janeiro, RJ, Brazil. &amp; Programa de Pós-graduação em Ciências Biológicas (Botânica), Museu Nacional, Universidade Federal do Rio de Janeiro (UFRJ), Quinta da Boa Vista, São Cristóvão, Rio de Janeiro, RJ, Brazil. &amp; pedro. schwambach @ ufrj. br; https: // orcid. org / 0000 - 0003 - 2042 - 3687</mods:affiliation>
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<mods:namePart>Mariath, Jorge Ernesto De Araujo</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-2776-6781</mods:nameIdentifier>
<mods:affiliation>Programa de Pós-graduação em Botânica, Instituto de Biociências, UFRGS, Av. Bento Gonçalves, 9500, Porto Alegre, RS, Brazil. &amp; jorge. mariath @ ufrgs. br; https: // orcid. org / 0000 - 0002 - 2776 - 6781</mods:affiliation>
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<mods:title>Phytotaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-05-05</mods:number>
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<mods:number>595</mods:number>
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<mods:number>2</mods:number>
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<subSubSection box="[136,426,1620,1647]" pageId="2" pageNumber="171" type="nomenclature">
<paragraph blockId="2.[136,1452,1620,2079]" box="[136,426,1620,1647]" pageId="2" pageNumber="171">
<heading box="[136,426,1620,1647]" fontSize="11" level="2" pageId="2" pageNumber="171" reason="4">
1. 
<emphasis bold="true" box="[164,426,1620,1647]" pageId="2" pageNumber="171">
<taxonomicName authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[164,319,1620,1647]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="171" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis bold="true" box="[164,319,1620,1647]" italics="true" pageId="2" pageNumber="171">Dyckia selloa</emphasis>
</taxonomicName>
complex
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="172" pageId="2" pageNumber="171" type="description">
<paragraph blockId="2.[136,1452,1620,2079]" lastBlockId="3.[136,1452,158,1229]" lastPageId="3" lastPageNumber="172" pageId="2" pageNumber="171">
<emphasis bold="true" box="[136,206,1656,1683]" italics="true" pageId="2" pageNumber="171">Herbs</emphasis>
rhizomatous or acaulescent. 
<emphasis bold="true" box="[535,692,1656,1683]" italics="true" pageId="2" pageNumber="171">Inflorescence</emphasis>
axillary, rarely simple (
<taxonomicName authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[963,1088,1656,1682]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="171" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis box="[963,1088,1656,1682]" italics="true" pageId="2" pageNumber="171">D. delicata</emphasis>
</taxonomicName>
), generally composed of first to third order, rarely up to fourth order branches (
<taxonomicName authorityName="Strehl" authorityYear="2008" box="[677,878,1692,1718]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="171" phylum="Tracheophyta" rank="species" species="nigrospinulata">
<emphasis box="[677,878,1692,1718]" italics="true" pageId="2" pageNumber="171">D. nigrospinulata</emphasis>
</taxonomicName>
); 
<emphasis box="[903,1084,1692,1718]" italics="true" pageId="2" pageNumber="171">peduncle bracts</emphasis>
foliaceous; 
<emphasis box="[1224,1452,1692,1718]" italics="true" pageId="2" pageNumber="171">first order branches</emphasis>
numerous, usually more than 10, spirally arranged along the main axis of the fertile part, occupying 
<emphasis box="[1277,1310,1729,1754]" italics="true" pageId="2" pageNumber="171">ca.</emphasis>
2/3 or more of the fertile portion of inflorescence; 
<emphasis box="[584,847,1764,1790]" italics="true" pageId="2" pageNumber="171">second order branches</emphasis>
when present usually less than 5, distichous (rarely polystichous as in 
<taxonomicName authorityName="Strehl" authorityYear="2008" box="[342,540,1800,1826]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="171" phylum="Tracheophyta" rank="species" species="nigrospinulata">
<emphasis box="[342,540,1800,1826]" italics="true" pageId="2" pageNumber="171">D. nigrospinulata</emphasis>
</taxonomicName>
) at the base of the first order branches; 
<emphasis box="[980,1210,1800,1826]" italics="true" pageId="2" pageNumber="171">third order branches</emphasis>
when present usually underdeveloped at the base of the second order branches, with few or no flowers; 
<emphasis box="[1094,1236,1836,1862]" italics="true" pageId="2" pageNumber="171">floral bracts</emphasis>
usually triangular, reduced, shorter than the sepals. 
<emphasis bold="true" box="[518,611,1872,1899]" italics="true" pageId="2" pageNumber="171">Flowers</emphasis>
numerous (more than 100), androgynosporangiate, protandric, sessile or pedicellate; 
<emphasis box="[272,404,1908,1934]" italics="true" pageId="2" pageNumber="171">hypanthium</emphasis>
absent or reduced; 
<emphasis box="[621,701,1908,1934]" italics="true" pageId="2" pageNumber="171">corolla</emphasis>
tubular to rarely campanulate (
<taxonomicName authorityName="Strehl" authorityYear="2004" box="[1051,1153,1908,1934]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="171" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis box="[1051,1153,1908,1934]" italics="true" pageId="2" pageNumber="171">D. rigida</emphasis>
</taxonomicName>
) at anthesis; 
<emphasis box="[1300,1367,1908,1934]" italics="true" pageId="2" pageNumber="171">petals</emphasis>
oblong or oblanceolate, not unguiculate, generally abaxially indumented or with trichomes only along the margins (rarely glabrescent as in 
<taxonomicName authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[328,429,1980,2006]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="171" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis box="[328,429,1980,2006]" italics="true" pageId="2" pageNumber="171">D. selloa</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Baker" authorityYear="1889" box="[482,667,1980,2006]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="171" phylum="Tracheophyta" rank="species" species="myriostachya">
<emphasis box="[482,667,1980,2006]" italics="true" pageId="2" pageNumber="171">D. myriostachya</emphasis>
</taxonomicName>
); 
<emphasis box="[690,780,1981,2006]" italics="true" pageId="2" pageNumber="171">stamens</emphasis>
reaching the apex of the petals or exserted; 
<emphasis box="[1269,1372,1980,2006]" italics="true" pageId="2" pageNumber="171">filaments</emphasis>
free or inconspicuously connate at the base, the antepetalous ones briefly adnate to the petals or free; 
<emphasis box="[1168,1252,2016,2042]" italics="true" pageId="2" pageNumber="171">anthers</emphasis>
generally rostrate at the apex; 
<emphasis box="[271,350,2052,2078]" italics="true" pageId="2" pageNumber="171">locules</emphasis>
with less than 18 ovules; 
<emphasis box="[640,712,2052,2078]" italics="true" pageId="2" pageNumber="171">ovules</emphasis>
generally suberectly positioned in relation to the longitudinal axis of the ovary, disposed in two rows per locule (rarely four as in 
<taxonomicName authorityName="Larocca &amp; Sobral" authorityYear="2002" box="[825,948,158,184]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis box="[825,948,158,184]" italics="true" pageId="3" pageNumber="172">D. delicata</emphasis>
</taxonomicName>
and 
<taxonomicName authorityName="Irgang &amp; Sobral" authorityYear="1987" box="[1000,1145,158,184]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="agudensis">
<emphasis box="[1000,1145,158,184]" italics="true" pageId="3" pageNumber="172">D. agudensis</emphasis>
</taxonomicName>
); 
<emphasis box="[1168,1243,159,184]" italics="true" pageId="3" pageNumber="172">stigma</emphasis>
at anthesis usually with erect to suberect stigmatic lobes, reaching the apex of the petals or exserted. 
<emphasis bold="true" box="[1064,1129,194,221]" italics="true" pageId="3" pageNumber="172">Seeds</emphasis>
oblanceoloid with tegument equally distributed in raphe and antiraphe, chalaza slightly expanded and asymmetrical or as developed as the raphe and the antiraphe.
</paragraph>
<paragraph blockId="3.[136,1452,158,1229]" pageId="3" pageNumber="172">
<emphasis bold="true" box="[189,286,302,328]" pageId="3" pageNumber="172">Species:</emphasis>
—The species that possess these morphological characteristics and 
<taxonomicName box="[1043,1145,302,328]" form="this" pageId="3" pageNumber="172" rank="form">form this</taxonomicName>
complex are: 
<taxonomicName authority="Irgang &amp; Sobral (1987: 5)" authorityName="Irgang &amp; Sobral" authorityPageNumber="5" authorityYear="1987" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="agudensis">
<emphasis box="[1305,1451,302,328]" italics="true" pageId="3" pageNumber="172">D. agudensis</emphasis>
<bibRefCitation author="Irgang, B. E. &amp; Sobral, M." box="[136,429,338,365]" pageId="3" pageNumber="172" pagination="5 - 7" refId="ref13008" refString="Irgang, B. E. &amp; Sobral, M. (1987) Dyckia agudensis (Bromeliaceae), nova especie do Rio Grande do Sul, Brasil. Napaea 3: 5 - 7." type="journal article" year="1987">Irgang &amp; Sobral (1987: 5)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Winkler (1982: 33)" authorityName="Winkler" authorityPageNumber="33" authorityYear="1982" box="[440,747,338,364]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="alba">
<emphasis box="[440,524,338,364]" italics="true" pageId="3" pageNumber="172">D. alba</emphasis>
<bibRefCitation author="Winkler, S." box="[530,747,338,364]" pageId="3" pageNumber="172" pagination="1 - 90" refId="ref15835" refString="Winkler, S. (1982) Die Bromeliaceae von Rio Grande do Sul. Documenta Naturae 3: 1 - 90." type="journal article" year="1982">Winkler (1982: 33)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Larocca &amp; Sobral (2002: 234)" authorityName="Larocca &amp; Sobral" authorityPageNumber="234" authorityYear="2002" box="[758,1227,338,365]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="delicata">
<emphasis box="[758,882,338,364]" italics="true" pageId="3" pageNumber="172">D. delicata</emphasis>
<bibRefCitation author="Larocca, J. &amp; Sobral, M." box="[889,1227,338,365]" pageId="3" pageNumber="172" pagination="234 - 236" refId="ref13337" refString="Larocca, J. &amp; Sobral, M. (2002) Dyckia delicata (Bromeliaceae), a new species from Rio Grande do Sul, Brazil. Novon 12: 234 - 236. https: // doi. org / 10.2307 / 3392959" type="journal article" year="2002">Larocca &amp; Sobral (2002: 234)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Smith (1971: 90)" authorityName="Smith" authorityPageNumber="90" authorityYear="1971" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="hebdingii">
<emphasis box="[1238,1378,338,364]" italics="true" pageId="3" pageNumber="172">D. hebdingii</emphasis>
<bibRefCitation author="Smith, L. B." pageId="3" pageNumber="172" pagination="73 - 96" refId="ref15062" refString="Smith, L. B. (1971) Notes on Bromeliaceae, XXXII. Phytologia 21: 73 - 96." type="journal article" year="1971">Smith (1971: 90)</bibRefCitation>
</taxonomicName>
, 
<emphasis box="[265,499,374,400]" italics="true" pageId="3" pageNumber="172">
<taxonomicName authority="Büneker &amp; Pastori &amp; Almeida &amp; Mariath, 2023" authorityName="Büneker &amp; Pastori &amp; Almeida &amp; Mariath" authorityYear="2023" box="[265,410,374,400]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="magnifica" status="sp. nov.">D. magnifica</taxonomicName>
<taxonomicNameLabel box="[415,499,375,400]" pageId="3" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(described here), 
<taxonomicName authority="Baker (1889: 136)" authorityName="Baker" authorityPageNumber="136" authorityYear="1889" box="[696,1043,374,400]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="maritima">
<emphasis box="[696,832,375,400]" italics="true" pageId="3" pageNumber="172">D. maritima</emphasis>
<bibRefCitation author="Baker, J. G." box="[837,1043,374,400]" pageId="3" pageNumber="172" refId="ref11313" refString="Baker, J. G. (1889) Handbook of the Bromeliaceae. G. Bell &amp; Sons, London, 243 pp. https: // doi. org / 10.5962 / bhl. title. 51681" type="book" year="1889">Baker (1889: 136)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Baker (1889: 137)" authorityName="Baker" authorityPageNumber="137" authorityYear="1889" box="[1052,1447,374,400]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="myriostachya">
<emphasis box="[1052,1236,374,400]" italics="true" pageId="3" pageNumber="172">D. myriostachya</emphasis>
<bibRefCitation author="Baker, J. G." box="[1242,1447,374,400]" pageId="3" pageNumber="172" refId="ref11313" refString="Baker, J. G. (1889) Handbook of the Bromeliaceae. G. Bell &amp; Sons, London, 243 pp. https: // doi. org / 10.5962 / bhl. title. 51681" type="book" year="1889">Baker (1889: 137)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Strehl (2008: 9)" authorityName="Strehl" authorityPageNumber="9" authorityYear="2008" box="[136,529,410,436]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="nigrospinulata">
<emphasis box="[136,337,410,436]" italics="true" pageId="3" pageNumber="172">D. nigrospinulata</emphasis>
<bibRefCitation author="Strehl, T." box="[346,529,410,436]" pageId="3" pageNumber="172" pagination="8 - 22" refId="ref15391" refString="Strehl, T. (2008) New bromeliads, genus Dyckia, from Rio Grande do Sul, Brazil. Bromeliaceae 42: 8 - 22." type="journal article" year="2008">Strehl (2008: 9)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName box="[544,751,410,436]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="pseudodelicata">
<emphasis box="[544,751,410,436]" italics="true" pageId="3" pageNumber="172">D. pseudodelicata</emphasis>
</taxonomicName>
B̧neker &amp; 
<bibRefCitation author="Mariath, J. E. A." box="[895,1115,410,436]" pageId="3" pageNumber="172" pagination="59 - 70" refId="ref11651" refString="B ¸ neker, H. M. &amp; Mariath, J. E. A. (2022) Dyckia pseudodelicata, a new species of the D. selloa complex (Bromeliaceae, Pitcairnioideae) from Rio Grande do Sul, Brazil. Phytotaxa 550: 59 - 70. https: // doi. org / 10.11646 / phytotaxa. 550.1.4" type="journal article" year="2022">Mariath (2022: 60)</bibRefCitation>
, 
<taxonomicName authority="Winkler (1982: 42)" authorityName="Winkler" authorityPageNumber="42" authorityYear="1982" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="retardata">
<emphasis box="[1129,1270,410,436]" italics="true" pageId="3" pageNumber="172">D. retardata</emphasis>
<bibRefCitation author="Winkler, S." pageId="3" pageNumber="172" pagination="1 - 90" refId="ref15835" refString="Winkler, S. (1982) Die Bromeliaceae von Rio Grande do Sul. Documenta Naturae 3: 1 - 90." type="journal article" year="1982">Winkler (1982: 42)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Winkler (1982: 44)" authorityName="Winkler" authorityPageNumber="44" authorityYear="1982" box="[187,551,446,472]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="retroflexa">
<emphasis box="[187,329,446,472]" italics="true" pageId="3" pageNumber="172">D. retroflexa</emphasis>
<bibRefCitation author="Winkler, S." box="[335,551,446,472]" pageId="3" pageNumber="172" pagination="1 - 90" refId="ref15835" refString="Winkler, S. (1982) Die Bromeliaceae von Rio Grande do Sul. Documenta Naturae 3: 1 - 90." type="journal article" year="1982">Winkler (1982: 44)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="Strehl (2004: 32)" authorityName="Strehl" authorityPageNumber="32" authorityYear="2004" box="[563,865,446,472]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="rigida">
<emphasis box="[563,664,446,472]" italics="true" pageId="3" pageNumber="172">D. rigida</emphasis>
<bibRefCitation author="Strehl, T." box="[671,865,446,472]" pageId="3" pageNumber="172" pagination="19 - 25" refId="ref15365" refString="Strehl, T. (2004) Novas especies de Bromeliaceae do Rio Grande do Sul, Brasil. Vidalia 2: 19 - 25." type="journal article" year="2004">Strehl (2004: 32)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName authority="(Koch 1873: 7) Baker (1889: 136)" authorityName="Baker" authorityPageNumber="136" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityPageNumber="7" baseAuthorityYear="1873" box="[875,1366,446,472]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis box="[875,976,446,472]" italics="true" pageId="3" pageNumber="172">D. selloa</emphasis>
(
<bibRefCitation author="Koch, K." box="[990,1144,446,472]" pageId="3" pageNumber="172" pagination="1 - 7" refId="ref13117" refString="Koch, K. (1873) Index seminum in horto botanico berolinensi anno 1873 collectorum. Appendix quarta ad indicem seminum horti botanici Berolinenensis anni 1873. Berlin, pp. 1 - 7." type="book chapter" year="1873">Koch 1873: 7</bibRefCitation>
) 
<bibRefCitation author="Baker, J. G." box="[1159,1366,446,472]" pageId="3" pageNumber="172" refId="ref11313" refString="Baker, J. G. (1889) Handbook of the Bromeliaceae. G. Bell &amp; Sons, London, 243 pp. https: // doi. org / 10.5962 / bhl. title. 51681" type="book" year="1889">Baker (1889: 136)</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName authority="Mez (1896: 515)" authorityName="Mez" authorityPageNumber="515" authorityYear="1896" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="tomentosa">
<emphasis italics="true" pageId="3" pageNumber="172">D. tomentosa</emphasis>
<bibRefCitation author="Mez, C." box="[258,449,482,509]" pageId="3" pageNumber="172" refId="ref13742" refString="Mez, C. (1896) Bromeliaceae. In: De Candolle, C. (Ed.) Monographiae Phanerogamarum 9. Sumptibus G. Masson &amp; Cie., Paris, 990 pp." type="book" year="1896">Mez (1896: 515)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph blockId="3.[136,1452,158,1229]" pageId="3" pageNumber="172">
<emphasis bold="true" box="[189,357,518,544]" pageId="3" pageNumber="172">Observations:</emphasis>
—The 
<taxonomicName authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[437,539,518,544]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis box="[437,539,518,544]" italics="true" pageId="3" pageNumber="172">D. selloa</emphasis>
</taxonomicName>
complex is characterized and basically distinct 
<taxonomicName box="[1081,1201,518,544]" form="other" pageId="3" pageNumber="172" rank="form">form other</taxonomicName>
species of 
<emphasis box="[1328,1451,518,544]" italics="true" pageId="3" pageNumber="172">
<taxonomicName baseAuthorityName="Smith" baseAuthorityYear="1943" box="[1328,1407,518,544]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="genus" sensu="stricto">Dyckia</taxonomicName>
<taxonomicNameLabel box="[1416,1451,519,544]" pageId="3" pageNumber="172" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
by the following set of characters: inflorescences usually composed of first to third order (
<emphasis box="[1177,1200,555,580]" italics="true" pageId="3" pageNumber="172">vs</emphasis>
. simple or compound to first order), with numerous first order branches (usually more than 10 
<emphasis box="[970,1001,591,616]" italics="true" pageId="3" pageNumber="172">vs.</emphasis>
less than 10 branches), generally small and inconspicuous floral bracts, numerous flowers (more than 100 
<emphasis box="[889,912,627,652]" italics="true" pageId="3" pageNumber="172">vs</emphasis>
. less than 80) with non-unguiculate (
<emphasis box="[1333,1356,627,652]" italics="true" pageId="3" pageNumber="172">vs</emphasis>
. usually unguiculate) petals, stigma at anthesis usually with erect to suberect (
<emphasis box="[918,941,663,688]" italics="true" pageId="3" pageNumber="172">vs</emphasis>
. twisted) stigmatic lobes, pauciovulate ovary locules (less than 18 
<emphasis box="[371,394,699,724]" italics="true" pageId="3" pageNumber="172">vs</emphasis>
. more than 30 ovules) and oblanceoloid (
<emphasis box="[861,884,699,724]" italics="true" pageId="3" pageNumber="172">vs</emphasis>
. flattened) seeds with a poorly developed wing.
</paragraph>
<paragraph blockId="3.[136,1452,158,1229]" pageId="3" pageNumber="172">
<bibRefCitation author="Mez, C." box="[189,314,734,761]" pageId="3" pageNumber="172" refId="ref13742" refString="Mez, C. (1896) Bromeliaceae. In: De Candolle, C. (Ed.) Monographiae Phanerogamarum 9. Sumptibus G. Masson &amp; Cie., Paris, 990 pp." type="book" year="1896">Mez (1896</bibRefCitation>
, 
<bibRefCitation author="Mez, C." box="[325,382,734,761]" pageId="3" pageNumber="172" refId="ref13779" refString="Mez, C. (1935) Bromeliaceae. In: Engler, A. (Ed.) Das Pflanzenreich, regni vegetabilis conspectus 4 (32). W. Engelmann, Leipzig, 667 pp." type="book" year="1935">1935</bibRefCitation>
) presents as main diagnostic character of 
<taxonomicName authorityName="Koch" authorityYear="1873" box="[851,1019,734,760]" class="Liliopsida" family="Bromeliaceae" genus="Prionophyllum" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="genus">
<emphasis box="[851,1019,734,760]" italics="true" pageId="3" pageNumber="172">Prionophyllum</emphasis>
</taxonomicName>
the dimorphic flowers, the larger ones would be hermaphrodites and the smaller ones with reduced androecium. This is reproduced in several taxonomic treatments (
<emphasis box="[272,313,807,832]" italics="true" pageId="3" pageNumber="172">e.g.</emphasis>
<bibRefCitation author="Smith, L. B. &amp; Downs, R. J." box="[323,582,806,833]" pageId="3" pageNumber="172" pagination="1 - 662" refId="ref15134" refString="Smith, L. B. &amp; Downs, R. J. (1974) Bromeliaceae (Pitcairnioideae). Flora Neotropica Monograph 14: 1 - 662." type="journal article" year="1974">Smith &amp; Downs 1974</bibRefCitation>
; 
<bibRefCitation author="Krapp, F. &amp; Eggli, U." box="[597,842,806,833]" pageId="3" pageNumber="172" pagination="1 - 71" refId="ref13183" refString="Krapp, F. &amp; Eggli, U. (2019) Dyckia, Bromeliaceae. In: Eggli, U. &amp; Nyffeler, R. (Eds.) Illustrated Handbook of Succulent Plants: Monocotyledons, ed. 2. Springer, Berlin, pp. 1 - 71. https: // doi. org / 10.1007 / 978 - 3 - 662 - 56324 - 3 _ 80 - 1" type="book chapter" year="2019">Krapp &amp; Eggli 2019</bibRefCitation>
). However, we never checked specimens with really dimorphic flowers. We found some flowers galled by insects in 
<taxonomicName authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[859,961,842,868]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis box="[859,961,842,868]" italics="true" pageId="3" pageNumber="172">D. selloa</emphasis>
</taxonomicName>
specimens, that generated anomalies in the size of flowers and floral parts, and we suspect that this may be related to what Mez reported. Koch (1874: 7), when describing 
<taxonomicName authority="Koch (1873: 7)" authorityName="Koch" authorityPageNumber="7" authorityYear="1873" box="[261,705,914,940]" class="Liliopsida" family="Bromeliaceae" genus="Prionophyllum" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="selloum">
<emphasis box="[261,524,914,940]" italics="true" pageId="3" pageNumber="172">Prionophyllum selloum</emphasis>
<bibRefCitation author="Koch, K." box="[532,705,914,940]" pageId="3" pageNumber="172" pagination="1 - 7" refId="ref13117" refString="Koch, K. (1873) Index seminum in horto botanico berolinensi anno 1873 collectorum. Appendix quarta ad indicem seminum horti botanici Berolinenensis anni 1873. Berlin, pp. 1 - 7." type="book chapter" year="1873">Koch (1873: 7)</bibRefCitation>
</taxonomicName>
as a new monospecific genus, describes “hermaphrodite” flowers and points out that the habit resembles 
<taxonomicName authority="Zuccarini (1838: 258)" authorityName="Zuccarini" authorityPageNumber="258" authorityYear="1838" box="[571,943,950,977]" class="Liliopsida" family="Asparagaceae" genus="Dasylirion" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="genus">
<emphasis box="[571,691,950,976]" italics="true" pageId="3" pageNumber="172">Dasylirion</emphasis>
<bibRefCitation author="Zuccarini, J. G." box="[698,943,950,977]" pageId="3" pageNumber="172" pagination="257 - 259" refId="ref15888" refString="Zuccarini, J. G. (1838) Ueber eine neue Gattung aus der Familie der Bromeliaceae nebst Bemerkungen ¸ ber einige mexikanische Agaven. Allgemeine Gartenzeitung 6: 257 - 259." type="journal article" year="1838">Zuccarini (1838: 258)</bibRefCitation>
</taxonomicName>
(
<taxonomicName authorityName="Salisbury" authorityYear="1866" box="[957,1116,950,976]" class="Liliopsida" family="Dracaenaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="family">Dracaenaceae</taxonomicName>
) and 
<taxonomicName authority="Klotzsch (1835: 401)" authorityName="Klotzsch" authorityPageNumber="401" authorityYear="1835" class="Liliopsida" family="Bromeliaceae" genus="Hechtia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="genus">
<emphasis box="[1177,1266,950,976]" italics="true" pageId="3" pageNumber="172">Hechtia</emphasis>
<bibRefCitation author="Klotzsch, J. F." pageId="3" pageNumber="172" pagination="401 - 403" refId="ref13092" refString="Klotzsch, J. F. (1835) Hechtia. Eine neue Gattung der Bromeliaceen. Allgemeine Gartenzeitung 3: 401 - 403." type="journal article" year="1835">Klotzsch (1835: 401)</bibRefCitation>
</taxonomicName>
(
<taxonomicName authorityName="Jussieu" authorityYear="1789" box="[204,363,986,1012]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="family">Bromeliaceae</taxonomicName>
, 
<taxonomicName authorityName="Givnish" authorityYear="2007" box="[377,530,986,1012]" class="Liliopsida" family="Bromeliaceae" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="subFamily" subFamily="Hechtioideae">Hechtioideae</taxonomicName>
), but differs greatly by its hermaphrodite flowers (“ 
<emphasis italics="true" pageId="3" pageNumber="172">Planta Dasylirii et Hechtiae specierum habitum prae se fert, sed floribus hermaphroditis longe difert.</emphasis>
”).
</paragraph>
<paragraph blockId="3.[136,1452,158,1229]" pageId="3" pageNumber="172">
<bibRefCitation author="Krapp, F. &amp; Eggli, U." box="[189,434,1058,1085]" pageId="3" pageNumber="172" pagination="1 - 71" refId="ref13183" refString="Krapp, F. &amp; Eggli, U. (2019) Dyckia, Bromeliaceae. In: Eggli, U. &amp; Nyffeler, R. (Eds.) Illustrated Handbook of Succulent Plants: Monocotyledons, ed. 2. Springer, Berlin, pp. 1 - 71. https: // doi. org / 10.1007 / 978 - 3 - 662 - 56324 - 3 _ 80 - 1" type="book chapter" year="2019">Krapp &amp; Eggli (2019)</bibRefCitation>
include 
<taxonomicName authority="Smith (1988: 249)" authorityName="Smith" authorityPageNumber="249" authorityYear="1988" box="[529,856,1058,1084]" class="Liliopsida" family="Asparagaceae" genus="Dasylirion" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="racinae">
<emphasis box="[529,648,1059,1084]" italics="true" pageId="3" pageNumber="172">D. racinae</emphasis>
<bibRefCitation author="Smith, L. B." box="[653,856,1058,1084]" pageId="3" pageNumber="172" pagination="248 - 249" refId="ref15084" refString="Smith, L. B. (1988) A giant Dyckia mystery. Journal of the Bromeliad Society 38: 248 - 249." type="journal article" year="1988">Smith (1988: 249)</bibRefCitation>
</taxonomicName>
in this complex, however, this taxon does not possess several of the diagnostic characteristics mentioned above (
<emphasis box="[780,820,1095,1120]" italics="true" pageId="3" pageNumber="172">e.g.</emphasis>
petals oblong to oblanceolate, not unguiculate, and seeds oblanceoloid). B̧neker 
<emphasis box="[408,465,1130,1156]" italics="true" pageId="3" pageNumber="172">et al.</emphasis>
(2020, 2021) regard the taxon as 
<taxonomicName authorityName="Smith" authorityYear="1988" box="[850,989,1131,1157]" class="Liliopsida" family="Asparagaceae" genus="Dasylirion" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="racinae">
<emphasis box="[850,877,1131,1156]" italics="true" pageId="3" pageNumber="172">D.</emphasis>
× 
<emphasis box="[901,989,1131,1156]" italics="true" pageId="3" pageNumber="172">racinae</emphasis>
</taxonomicName>
<emphasis box="[989,993,1131,1156]" italics="true" pageId="3" pageNumber="172">,</emphasis>
arguing that there are strong indications that this taxon may be a natural hybrid between 
<taxonomicName authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[672,773,1166,1192]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis box="[672,773,1166,1192]" italics="true" pageId="3" pageNumber="172">D. selloa</emphasis>
</taxonomicName>
and 
<taxonomicName authority="Smith (1932: 98)" authorityName="Smith" authorityPageNumber="98" authorityYear="1932" box="[826,1138,1166,1192]" class="Liliopsida" family="Asparagaceae" genus="Dasylirion" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="dusenii">
<emphasis box="[826,941,1166,1192]" italics="true" pageId="3" pageNumber="172">D. dusenii</emphasis>
<bibRefCitation author="Smith, L. B." box="[948,1138,1166,1192]" pageId="3" pageNumber="172" pagination="3 - 42" refId="ref14828" refString="Smith, L. B. (1932) Studies in the Bromeliaceae - III. Contributions from the Gray Herbarium of Harvard University 98: 3 - 42. https: // doi. org / 10.5962 / p. 336146" type="journal article" year="1932">Smith (1932: 98)</bibRefCitation>
</taxonomicName>
and therefore has similarity with the species of the 
<taxonomicName authorityName="Baker" authorityYear="1889" baseAuthorityName="Koch" baseAuthorityYear="1873" box="[393,495,1202,1228]" class="Liliopsida" family="Bromeliaceae" genus="Dyckia" kingdom="Plantae" order="Poales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="selloa">
<emphasis box="[393,495,1202,1228]" italics="true" pageId="3" pageNumber="172">D. selloa</emphasis>
</taxonomicName>
complex. Here we do not consider 
<taxonomicName authorityName="Smith" authorityYear="1988" box="[893,1029,1203,1229]" class="Liliopsida" family="Asparagaceae" genus="Dasylirion" isHybrid="true" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="172" phylum="Tracheophyta" rank="species" species="racinae">
<emphasis box="[893,920,1203,1228]" italics="true" pageId="3" pageNumber="172">D.</emphasis>
× 
<emphasis box="[943,1029,1203,1228]" italics="true" pageId="3" pageNumber="172">racinae</emphasis>
</taxonomicName>
as belonging to this complex.
</paragraph>
</subSubSection>
</treatment>
</document>