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<document ID-DOI="10.5252/a2015n2a2" ID-ISSN="1639-4798" ID-Zenodo-Dep="5208903" approvalRequired="50" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="44" approvalRequired_for_treatments="5" checkinTime="1629150308770" checkinUser="carolina" docAuthor="Daly, Douglas C., Raharimampionona, Jeannie &amp; Federman, Sarah" docDate="2015" docId="03F5FA39BC57FFEFB1D4F9EB336BFAAC" docLanguage="en" docName="Adansonia.37.2.277-345.pdf" docOrigin="Adansonia 37 (2)" docSource="https://www.mendeley.com/catalogue/2522e05a-0ab0-3e53-96aa-d45147a2ca6c/" docStyle="DocumentStyle:E7A860AFEA63D44BCC37955D07FCAFC7.2:Adansonia.2015-2017.journal_article" docStyleId="E7A860AFEA63D44BCC37955D07FCAFC7" docStyleName="Adansonia.2015-2017.journal_article" docStyleVersion="2" docTitle="Canarium arcuatum Daly, Raharim. &amp; Federman 2015, sp. nov." docType="treatment" docVersion="2" lastPageId="17" lastPageNumber="294" masterDocId="FFCC8241BC58FFFEB157FF99310AFFA1" masterDocTitle="A revision of Canarium L. (Burseraceae) in Madagascar" masterLastPageNumber="345" masterPageNumber="277" pageId="15" pageNumber="292" updateTime="1629150974147" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>A revision of Canarium L. (Burseraceae) in Madagascar</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Daly, Douglas C.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Raharimampionona, Jeannie</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Federman, Sarah</mods:namePart>
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<mods:title>Adansonia</mods:title>
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<mods:date>2015</mods:date>
<mods:detail type="pubDate">
<mods:number>2015-12-31</mods:number>
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<mods:detail type="volume">
<mods:number>37</mods:number>
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<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:identifier type="DOI">10.5252/a2015n2a2</mods:identifier>
<mods:identifier type="ISSN">1639-4798</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:03F5FA39BC57FFEFB1D4F9EB336BFAAC" httpUri="http://treatment.plazi.org/id/03F5FA39BC57FFEFB1D4F9EB336BFAAC" lastPageId="17" lastPageNumber="294" pageId="15" pageNumber="292">
<subSubSection box="[131,775,1650,1677]" pageId="15" pageNumber="292" type="nomenclature">
<paragraph blockId="15.[131,775,1650,1709]" box="[131,775,1650,1677]" pageId="15" pageNumber="292">
<heading box="[131,775,1650,1677]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="292" reason="3">
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[131,680,1650,1677]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="292" phylum="Tracheophyta" rank="species" species="arcuatum" status="sp. nov.">
<emphasis bold="true" box="[131,356,1650,1677]" italics="true" pageId="15" pageNumber="292">Canarium arcuatum</emphasis>
Daly, Raharim. &amp; Federman
</taxonomicName>
,
<taxonomicNameLabel box="[690,775,1651,1677]" pageId="15" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="292" type="description">
<paragraph blockId="15.[131,775,1650,1709]" box="[398,509,1682,1709]" pageId="15" pageNumber="292">
(
<figureCitation box="[405,474,1682,1708]" captionStart="FIG" captionStartId="11.[132,143,1985,2003]" captionTargetBox="[203,1375,215,1863]" captionTargetId="figure-18@11.[200,1387,211,1874]" captionTargetPageId="11" captionText="FIG. 2. — Canarium ampasindavae Daly, Raharim. &amp; Federman, sp. nov.: A, leaf, with detail of leaflet (A) and of petiole with stipular scar (A); B, infructescence, with detail of fruit surface (B). From Tahinarivony et al. 179 (NY). Scale bars: A, B, 2 cm; A, 1 cm; A, B, not to scale." figureDoi="http://doi.org/10.5281/zenodo.5208905" httpUri="https://zenodo.org/record/5208905/files/figure.png" pageId="15" pageNumber="292">Figs 2</figureCitation>
;
<figureCitation box="[486,500,1682,1709]" captionStart="FIG" captionStartId="14.[132,143,1800,1818]" captionTargetBox="[132,1450,215,1754]" captionTargetId="figure-81@14.[132,1455,207,1765]" captionTargetPageId="14" captionText="FIG. 5. — Canarium arcuatum Daly, Raharim. &amp; Federman, sp. nov.: A, flowering branchlet; B, pistillate flower; C, fruiting branchlet, with detail of leaflet (A) and of fruit surface (A); D, petiole and stipules; E, cross-section of fruit; A, B, Carlson 46 (NY); C, D, Schatz &amp; Miller 2520 (NY); E, after field photo by D. Daly. Scale bars: A, C, E, 2 cm; B, 3 mm; D, 1 cm; A, A, not to scale." figureDoi="http://doi.org/10.5281/zenodo.5208911" httpUri="https://zenodo.org/record/5208911/files/figure.png" pageId="15" pageNumber="292">5</figureCitation>
)
</paragraph>
<paragraph blockId="15.[131,777,1737,2025]" pageId="15" pageNumber="292">
<emphasis italics="true" pageId="15" pageNumber="292">Small trees, leaves 2-3-jugate, rachis with only sparse to scattered hairs, stipules</emphasis>
c.
<emphasis box="[296,488,1763,1787]" italics="true" pageId="15" pageNumber="292">
<quantity box="[296,374,1763,1787]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="15" pageNumber="292" unit="mm" value="4.2">4.2 mm</quantity>
long, fruits
</emphasis>
c.
<emphasis italics="true" pageId="15" pageNumber="292">
4.2 ×
<quantity box="[585,656,1763,1787]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="292" unit="cm" value="2.5">2.5 cm</quantity>
, oblong, the apex rounded, the base truncate, surface with lenticels dense, relatively small, not raised, ferrugineous; distinguished from
</emphasis>
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="292" phylum="Tracheophyta" rank="species" species="lamianum" status="sp. nov.">
C. lamianum
<emphasis italics="true" pageId="15" pageNumber="292">Daly, Raharim. &amp; Federman</emphasis>
</taxonomicName>
<emphasis italics="true" pageId="15" pageNumber="292">
,
<taxonomicNameLabel box="[373,449,1843,1866]" pageId="15" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
by the lateral leaflet base usually rounded (vs obtuse to truncate, rarely some leaflet bases acute or cordate in
</emphasis>
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[158,287,1896,1920]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="292" phylum="Tracheophyta" rank="species" species="lamianum" status="sp. nov.">C. lamianum</taxonomicName>
,
<emphasis italics="true" pageId="15" pageNumber="292">
<taxonomicNameLabel box="[299,373,1896,1919]" pageId="15" pageNumber="292" rank="species">sp. nov.</taxonomicNameLabel>
), the margin often slightly revolute (vs flat), leaflet secondary veins strongly arcuate (vs spreading to nearly straight, rarely slightly arcuate), and the fruit oblong and red-beige (vs broadly (oblong-)ovoid to slightly obovoid, sometimes trigonous, and (grayish-) green to (light) brown).
</emphasis>
</paragraph>
<paragraph blockId="15.[811,1457,215,779]" pageId="15" pageNumber="292">
<materialsCitation collectingDate="1990-02-22" collectionCode="NY, MO, TAN, US" collectorName="B. Carlson" country="Madagascar" latitude="-15.483334" location="Bay of Antongil" longLatPrecision="1284" longitude="49.75" municipality="Island of Nosy Mangabe" pageId="15" pageNumber="292" specimenCount="1" stateProvince="Toamasina">
TYPUS. —
<emphasis bold="true" box="[916,1039,215,239]" pageId="15" pageNumber="292">
<collectingCountry box="[916,1035,215,239]" name="Madagascar" pageId="15" pageNumber="292">Madagascar</collectingCountry>
.
</emphasis>
<collectingRegion box="[1042,1149,215,239]" country="Madagascar" name="Toamasina" pageId="15" pageNumber="292">Toamasina</collectingRegion>
,
<collectingMunicipality box="[1158,1395,215,240]" pageId="15" pageNumber="292">Island of Nosy Mangabe</collectingMunicipality>
,
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<quantity box="[1404,1454,215,239]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="292" unit="km" value="5.0">5 km</quantity>
S of Maroantsetra in
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of Antongil,
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,
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,
<date box="[1349,1454,242,266]" pageId="15" pageNumber="292" value="1990-02-22">
<collectingDate box="[1349,1454,242,266]" pageId="15" pageNumber="292" value="1990-02-22">22.II.1990</collectingDate>
</date>
(passed f fl),
<emphasis box="[935,1064,268,292]" italics="true" pageId="15" pageNumber="292">
<collectorName box="[935,1034,268,292]" pageId="15" pageNumber="292">B. Carlson</collectorName>
46
</emphasis>
(holo-,
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!; iso-,
<collectionCode box="[1242,1288,268,292]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="15" pageNumber="292" type="Herbarium">MO</collectionCode>
, P[P00501663]!,
<collectionCode box="[813,866,295,319]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="15" pageNumber="292" type="Herbarium">TAN</collectionCode>
,
<collectionCode box="[876,913,295,319]" country="South Africa" name="University of Stellenbosch" pageId="15" pageNumber="292">US</collectionCode>
!)
</materialsCitation>
.
</paragraph>
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<materialsCitation collectingDate="1990-02-22" collectionCode="MO, NY" collectorName="B. Carlson" country="Madagascar" latitude="-15.483334" location="Nosy Mangabe" longLatPrecision="1284" longitude="49.75" pageId="15" pageNumber="292" specimenCount="1" stateProvince="Toamasina" typeStatus="paratype">
<typeStatus box="[813,903,343,366]" pageId="15" pageNumber="292">PARATYPI</typeStatus>
. —
<emphasis bold="true" box="[953,1078,342,366]" pageId="15" pageNumber="292">
<collectingCountry box="[953,1074,342,366]" name="Madagascar" pageId="15" pageNumber="292">Madagascar</collectingCountry>
.
</emphasis>
<collectingRegion box="[1084,1193,342,366]" country="Madagascar" name="Toamasina" pageId="15" pageNumber="292">Toamasina</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC57FFEFB1D4F9EB336BFAAC:8E831DF4BC57FFF1B5E2FECE3445FECF" box="[1205,1359,342,367]" country="Madagascar" latitude="-15.483334" longLatPrecision="1284" longitude="49.75" name="Nosy Mangabe" pageId="15" pageNumber="292" stateProvince="Toamasina">Nosy Mangabe</location>
,
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,
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,
<date box="[907,1015,369,393]" pageId="15" pageNumber="292" value="1990-02-22">
<collectingDate box="[907,1015,369,393]" pageId="15" pageNumber="292" value="1990-02-22">22.II.1990</collectingDate>
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,
<emphasis box="[1026,1198,368,392]" italics="true" pageId="15" pageNumber="292">
<collectorName box="[1026,1127,368,392]" pageId="15" pageNumber="292">B. Carlson</collectorName>
CB 46
</emphasis>
(
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,
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)
</materialsCitation>
;
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,
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<elevation box="[814,885,395,419]" metricMagnitude="2" metricUnit="m" metricValue="4.75" pageId="15" pageNumber="292" unit="m" value="475.0">475 m</elevation>
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,
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,
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<collectingDate box="[1163,1284,395,419]" pageId="15" pageNumber="292" value="2004-06-10">10.VI.2004</collectingDate>
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,
<emphasis italics="true" pageId="15" pageNumber="292">
<collectorName box="[1298,1457,395,419]" pageId="15" pageNumber="292">P. Antilahimena</collectorName>
AP 2483
</emphasis>
(
<collectionCode box="[913,952,422,446]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="15" pageNumber="292" type="Herbarium">NY</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="2006-01-18" collectionCode="NY" collectorName="D. C. Daly &amp; Raharimampionona &amp; Razanatsoa &amp; J. Aridy &amp; Senario" country="Madagascar" elevation="300" latitude="-15.506111" location="Nosy Mangabe" longLatPrecision="21" longitude="49.75028" pageId="15" pageNumber="292" specimenCount="1" stateProvince="Toamasina" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC57FFEFB1D4F9EB336BFAAC:8E831DF4BC57FFF1B29DFE3F356BFE1C" box="[970,1121,421,446]" country="Madagascar" latitude="-15.506111" longLatPrecision="21" longitude="49.75028" name="Nosy Mangabe" pageId="15" pageNumber="292" stateProvince="Toamasina">Nosy Mangabe</location>
,
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<elevation box="[1131,1198,422,446]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="292" unit="m" value="300.0">300 m</elevation>
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,
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,
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,
<date box="[813,913,448,472]" pageId="15" pageNumber="292" value="2006-01-18">
<collectingDate box="[813,913,448,472]" pageId="15" pageNumber="292" value="2006-01-18">18.I.2006</collectingDate>
</date>
,
<emphasis italics="true" pageId="15" pageNumber="292">
<collectorName box="[922,1027,448,472]" pageId="15" pageNumber="292">D. C. Daly</collectorName>
, J.
<collectorName box="[1056,1248,448,472]" pageId="15" pageNumber="292">Raharimampionona</collectorName>
, J.
<collectorName box="[1277,1387,449,472]" pageId="15" pageNumber="292">Razanatsoa</collectorName>
,
<collectorName pageId="15" pageNumber="292">J. Aridy</collectorName>
&amp;
<collectorName box="[864,934,475,498]" pageId="15" pageNumber="292">Senario</collectorName>
12973
</emphasis>
(
<collectionCode box="[1016,1055,474,498]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="15" pageNumber="292" type="Herbarium">NY</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1988-02-10" collectionCode="MO" collectorName="G. E. Schatz &amp; J. Dransfield &amp; Dransfield" country="Madagascar" elevation="400" latitude="-15.283334" location="Analanjirofo" longLatPrecision="1285" longitude="49.833332" pageId="15" pageNumber="292" specimenCount="1" stateProvince="Toamasina" typeStatus="paratype">
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,
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<elevation box="[1206,1271,474,498]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="292" unit="m" value="400.0">400 m</elevation>
</quantity>
,
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,
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,
<date box="[812,920,501,525]" pageId="15" pageNumber="292" value="1988-02-10">
<collectingDate box="[812,920,501,525]" pageId="15" pageNumber="292" value="1988-02-10">10.II.1988</collectingDate>
</date>
,
<emphasis box="[930,1384,501,525]" italics="true" pageId="15" pageNumber="292">
<collectorName box="[930,1050,501,525]" pageId="15" pageNumber="292">G. E. Schatz</collectorName>
,
<collectorName box="[1059,1177,501,525]" pageId="15" pageNumber="292">J. Dransfield</collectorName>
&amp; S.
<collectorName box="[1231,1328,501,525]" pageId="15" pageNumber="292">Dransfield</collectorName>
1873
</emphasis>
(
<collectionCode box="[1397,1443,501,525]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="15" pageNumber="292" type="Herbarium">MO</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1989-01-09" collectionCode="MO, NY" collectorName="G. E. Schatz &amp; J. S. Miller" country="Madagascar" elevation="165" latitude="-15.5" location="Nosy Mangabe" longLatPrecision="1284" longitude="49.766666" pageId="15" pageNumber="292" specimenCount="1" stateProvince="Toamasina" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC57FFEFB1D4F9EB336BFAAC:8E831DF4BC57FFF1B27BFD8932B6FD86" box="[812,956,527,552]" country="Madagascar" latitude="-15.5" longLatPrecision="1284" longitude="49.766666" name="Nosy Mangabe" pageId="15" pageNumber="292" stateProvince="Toamasina">Nosy Mangabe</location>
,
<quantity box="[964,1047,527,551]" metricMagnitude="2" metricUnit="m" metricValue="1.65" metricValueMax="3.3" metricValueMin="0.0" pageId="15" pageNumber="292" unit="m" value="165.0" valueMax="330.0" valueMin="0.0">
<elevation box="[964,1047,527,551]" metricMagnitude="2" metricUnit="m" metricValue="1.65" metricValueMax="3.3" metricValueMin="0.0" pageId="15" pageNumber="292" unit="m" value="165.0" valueMax="330.0" valueMin="0.0">0-330 m</elevation>
</quantity>
,
<geoCoordinate box="[1055,1131,527,551]" degrees="15" direction="south" minutes="30" orientation="latitude" pageId="15" pageNumber="292" precision="925" value="-15.5">15°30S</geoCoordinate>
,
<geoCoordinate box="[1140,1219,527,551]" degrees="49" direction="east" minutes="46" orientation="longitude" pageId="15" pageNumber="292" precision="925" value="49.766666">49°46E</geoCoordinate>
,
<date box="[1227,1310,527,551]" pageId="15" pageNumber="292" value="1989-01-09">
<collectingDate box="[1227,1310,527,551]" pageId="15" pageNumber="292" value="1989-01-09">9.I.1989</collectingDate>
</date>
,
<emphasis italics="true" pageId="15" pageNumber="292">
<collectorName box="[1318,1429,527,551]" pageId="15" pageNumber="292">G. E. Schatz</collectorName>
&amp;
<collectorName box="[811,910,554,578]" pageId="15" pageNumber="292">J. S. Miller</collectorName>
2520
</emphasis>
(
<collectionCode box="[976,1021,554,578]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="15" pageNumber="292" type="Herbarium">MO</collectionCode>
,
<collectionCode box="[1030,1068,554,578]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="15" pageNumber="292" type="Herbarium">NY</collectionCode>
)
</materialsCitation>
;
<materialsCitation collectingDate="1964-10-24" collectionCode="NY, TEF" collectorName="Service Forestier" country="Madagascar" latitude="-17.625" location="Analanjirofo" longLatPrecision="21" longitude="49.19167" pageId="15" pageNumber="292" specimenCount="1" stateProvince="Toamasina" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:03F5FA39BC57FFEFB1D4F9EB336BFAAC:8E831DF4BC57FFF1B56CFDB335BCFDE3" box="[1083,1206,554,578]" country="Madagascar" latitude="-17.625" longLatPrecision="21" longitude="49.19167" name="Analanjirofo" pageId="15" pageNumber="292" stateProvince="Toamasina">Analanjirofo</location>
,
<geoCoordinate box="[1214,1326,554,578]" degrees="17" direction="south" minutes="37" orientation="latitude" pageId="15" pageNumber="292" precision="15" seconds="30" value="-17.625">17°3730”S</geoCoordinate>
,
<geoCoordinate box="[1334,1449,554,578]" degrees="49" direction="east" minutes="11" orientation="longitude" pageId="15" pageNumber="292" precision="15" seconds="30" value="49.19167">49°1130”E</geoCoordinate>
,
<date box="[811,918,580,604]" pageId="15" pageNumber="292" value="1964-10-24">
<collectingDate box="[811,918,580,604]" pageId="15" pageNumber="292" value="1964-10-24">24.X.1964</collectingDate>
</date>
,
<emphasis box="[930,1149,580,604]" italics="true" pageId="15" pageNumber="292">
<collectorName box="[930,1079,581,604]" pageId="15" pageNumber="292">Service Forestier</collectorName>
21912
</emphasis>
(
<collectionCode box="[1162,1198,580,604]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="15" pageNumber="292" type="Herbarium">NY</collectionCode>
,
<collectionCode box="[1207,1252,581,604]" country="Madagascar" lsid="urn:lsid:biocol.org:col:12774" name="Centre National de la Recherche Appliquée au Developement Rural" pageId="15" pageNumber="292" type="Herbarium">TEF</collectionCode>
[not seen])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="292" type="distribution">
<paragraph blockId="15.[811,1457,215,779]" pageId="15" pageNumber="292">DISTRIBUTION AND ECOLOGY. — A relatively small tree, to date confirmed only from the small island of Nosy Mangabe in the Bay of Antongil. Known to flower in Feb. and fruit in Jan.</paragraph>
</subSubSection>
<subSubSection pageId="15" pageNumber="292" type="etymology">
<paragraph blockId="15.[811,1457,215,779]" pageId="15" pageNumber="292">ETYMOLOGY. — The specific epithet refers to the strongly arcuate leaflet secondary veins.</paragraph>
</subSubSection>
<subSubSection lastPageId="17" lastPageNumber="294" pageId="15" pageNumber="292" type="description">
<paragraph blockId="15.[810,1457,822,2028]" box="[813,957,822,848]" pageId="15" pageNumber="292">DESCRIPTION</paragraph>
<paragraph blockId="15.[810,1457,822,2028]" lastBlockId="17.[130,777,215,624]" lastPageId="17" lastPageNumber="294" pageId="15" pageNumber="292">
Trees, reproductive size
<emphasis box="[1081,1098,854,879]" italics="true" pageId="15" pageNumber="292">c.</emphasis>
<quantity box="[1105,1205,853,880]" metricMagnitude="1" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="15" pageNumber="292" unit="m" value="15.5" valueMax="16.0" valueMin="15.0">15-16 m</quantity>
, with small buttresses. Leaves
<quantity box="[890,996,885,912]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="15" pageNumber="292" unit="cm" value="22.0" valueMax="25.0" valueMin="19.0">19-25 cm</quantity>
long, 2-3-jugate; petiole
<quantity box="[1273,1392,885,911]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="6.2" metricValueMin="2.9" pageId="15" pageNumber="292" unit="cm" value="4.55" valueMax="6.2" valueMin="2.9">2.9-6.2 cm</quantity>
long, petiole and rachis with erect golden hairs to
<quantity box="[1273,1360,917,944]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="292" unit="mm" value="0.5">0.5 mm</quantity>
, dense to scattered on rachis but often absent on petiole, also with dense to scattered snail-shaped glands; stipules
<quantity box="[1234,1322,981,1007]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="15" pageNumber="292" unit="mm" value="5.5" valueMax="8.0" valueMin="3.0">3-8 mm</quantity>
from petiole insertion,
<emphasis box="[922,939,1013,1038]" italics="true" pageId="15" pageNumber="292">c.</emphasis>
<quantity box="[945,1031,1013,1039]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="15" pageNumber="292" unit="mm" value="4.2">4.2 mm</quantity>
long, broadly ovate (but revolute when dry), slightly constricted at base, the scar
<quantity box="[1220,1323,1044,1071]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="15" pageNumber="292" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5-2 mm</quantity>
long, surface with dense erect to appressed white hairs to
<quantity box="[1266,1366,1076,1103]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="292" unit="mm" value="0.05">0.05 mm</quantity>
long; all lateral petiolules
<quantity box="[989,1075,1108,1135]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="9.0" metricValueMin="3.0" pageId="15" pageNumber="292" unit="mm" value="6.0" valueMax="9.0" valueMin="3.0">3-9 mm</quantity>
long, terminal one
<quantity box="[1280,1394,1108,1135]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="15" pageNumber="292" unit="mm" value="20.5" valueMax="22.0" valueMin="19.0">19-22 mm</quantity>
long, petiolules very slightly canaliculate, lateral pulvinuli usually inconspicuous; basal leaflets 4.4-6.7
<emphasis box="[1206,1222,1173,1198]" italics="true" pageId="15" pageNumber="292">×</emphasis>
<quantity box="[1229,1330,1172,1198]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="4.0" metricValueMin="2.9" pageId="15" pageNumber="292" unit="cm" value="3.45" valueMax="4.0" valueMin="2.9">2.9-4 cm</quantity>
, obovate or less often ovate, other laterals 5.1-10.6
<emphasis box="[1223,1239,1205,1230]" italics="true" pageId="15" pageNumber="292">×</emphasis>
<quantity box="[1242,1363,1204,1230]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.8" metricValueMin="2.1" pageId="15" pageNumber="292" unit="cm" value="2.95" valueMax="3.8" valueMin="2.1">2.1-3.8 cm</quantity>
, broadly ovate or less often elliptic, or slightly obovate; the terminal 5.6-10
<emphasis box="[895,911,1268,1293]" italics="true" pageId="15" pageNumber="292">×</emphasis>
<quantity box="[921,1046,1268,1294]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" metricValueMax="4.9" metricValueMin="2.3" pageId="15" pageNumber="292" unit="cm" value="3.6" valueMax="4.9" valueMin="2.3">2.3-4.9 cm</quantity>
, slightly obovate; leaflet apex usually abruptly and usually broadly short-acuminate, the acumen
<quantity box="[814,901,1331,1358]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="15" pageNumber="292" unit="mm" value="4.0" valueMax="6.0" valueMin="2.0">2-6 mm</quantity>
long, leaflet base symmetrical, rounded or less often obtuse, margin usually slightly revolute; leaflets coriaceous, drying light brown, sometimes glossy abaxially when dry; secondary vein framework brochidodromous but looping at margin, secondaries in 4-7 pairs, strongly arcuate, the angle uniform, spacing sometimes decreasing toward apex and decreasing abruptly toward base; tertiaries alternate-percurrent and irregular-reticulate, perpendicular epimedial tertiaries present, quaternaries irregular-reticulate; on abaxial side the midvein broadly prominent, secondaries prominulous, rest of veins broadly prominulous, the surface with scattered snail-shaped glands; on adaxial side the midvein narrowly prominulous but sunk in a groove, rest of veins broadly prominulous to almost flat, the surface glabrous. Staminate flowers unknown. Pistillate inflorescences
<quantity box="[1294,1390,1778,1804]" metricMagnitude="-1" metricUnit="m" metricValue="1.15" metricValueMax="1.6" metricValueMin="0.7" pageId="15" pageNumber="292" unit="cm" value="11.5" valueMax="16.0" valueMin="7.0">7-16 cm</quantity>
long, with few flowers or fruits, secondary axes to
<quantity box="[1315,1393,1809,1836]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="292" unit="cm" value="2.5">2.5 cm</quantity>
long, axes with dense flexuous to straight golden hairs to
<quantity box="[1364,1453,1842,1868]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="292" unit="mm" value="0.2">0.2 mm</quantity>
; bracts caducous, semi-clasping (scars very broad); pedicel 4-5
<emphasis box="[855,871,1906,1931]" italics="true" pageId="15" pageNumber="292">×</emphasis>
<quantity box="[874,962,1905,1932]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="15" pageNumber="292" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2-3 mm</quantity>
, cylindrical. Pistillate flowers (only incomplete post-anthesis flowers known)
<emphasis box="[1127,1144,1938,1963]" italics="true" pageId="15" pageNumber="292">c.</emphasis>
<quantity box="[1150,1251,1937,1964]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="15" pageNumber="292" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9-10 mm</quantity>
long; calyx 5.5-6
<emphasis box="[1439,1455,1938,1963]" italics="true" pageId="15" pageNumber="292">×</emphasis>
<quantity box="[811,922,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="6.0" metricValueMin="5.3" pageId="15" pageNumber="292" unit="mm" value="5.65" valueMax="6.0" valueMin="5.3">5.3-6 mm</quantity>
overall, campanulate, the lobes
<quantity box="[1279,1390,1969,1996]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="15" pageNumber="292" unit="mm" value="2.25" valueMax="2.5" valueMin="2.0">2-2.5 mm</quantity>
long, (depressed) rounded-deltate, abaxial surface with dense de- scending thick ferrugineous hairs to
<quantity box="[521,609,215,242]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="294" unit="mm" value="0.1">0.1 mm</quantity>
, adaxial surface with dense appressed whitish hairs to
<quantity box="[528,626,247,274]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="294" unit="mm" value="0.25">0.25 mm</quantity>
long; petals
<emphasis box="[757,773,248,273]" italics="true" pageId="17" pageNumber="294">c.</emphasis>
7-8
<emphasis box="[174,190,280,305]" italics="true" pageId="17" pageNumber="294">×</emphasis>
<quantity box="[192,300,279,306]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" metricValueMax="4.0" metricValueMin="3.8" pageId="17" pageNumber="294" unit="mm" value="3.9" valueMax="4.0" valueMin="3.8">3.8-4 mm</quantity>
, ovate and slightly acuminate, with a strongly inflexed apiculum
<quantity box="[333,437,311,338]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="294" unit="mm" value="0.25">0.25 mm</quantity>
long; abaxial surface with pale golden flexuous hairs to
<quantity box="[401,488,343,370]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="294" unit="mm" value="0.5">0.5 mm</quantity>
long, ascending near apex and retrorse near base, adaxial surface subglabrous; androecium unknown; pistil with dense appressed whitish hairs to
<quantity box="[133,232,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="294" unit="mm" value="0.25">0.25 mm</quantity>
long. Fruiting pedicel
<emphasis box="[478,495,439,464]" italics="true" pageId="17" pageNumber="294">c.</emphasis>
6
<emphasis box="[522,538,439,464]" italics="true" pageId="17" pageNumber="294">×</emphasis>
<quantity box="[545,634,438,465]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="17" pageNumber="294" unit="mm" value="3.5">3.5 mm</quantity>
, terete, fruiting calyx
<emphasis box="[236,253,471,496]" italics="true" pageId="17" pageNumber="294">c.</emphasis>
<quantity box="[260,369,470,497]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="17" pageNumber="294" unit="mm" value="3.75" valueMax="4.0" valueMin="3.5">3.5-4 mm</quantity>
long, patent, the lobes distinct. Fruit
<emphasis box="[132,149,503,528]" italics="true" pageId="17" pageNumber="294">c.</emphasis>
4.2
<emphasis box="[195,211,503,528]" italics="true" pageId="17" pageNumber="294">×</emphasis>
<quantity box="[215,291,502,529]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="294" unit="cm" value="2.5">2.5 cm</quantity>
, oblong, the apex rounded, the base truncate, surface with lenticels dense, small, not raised, ferrugineous, also with appressed golden hairs to
<quantity box="[501,599,566,593]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="294" unit="mm" value="0.25">0.25 mm</quantity>
long, dense near apex but scattered elsewhere; mesocarp moderately thick.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5208915" ID-Zenodo-Dep="5208915" httpUri="https://zenodo.org/record/5208915/files/figure.png" pageId="16" pageNumber="293" startId="16.[132,143,1939,1957]" targetBox="[206,1378,214,1866]" targetPageId="16">
<paragraph blockId="16.[132,1456,1939,2025]" pageId="16" pageNumber="293">
FIG. 7. — A-D,
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[258,687,1939,1957]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="16" pageNumber="293" phylum="Tracheophyta" rank="species" species="betamponae" status="sp. nov.">
<emphasis box="[258,448,1939,1957]" italics="true" pageId="16" pageNumber="293">Canarium betamponae</emphasis>
Daly, Raharim. &amp; Federman
</taxonomicName>
,
<taxonomicNameLabel box="[695,766,1939,1957]" pageId="16" pageNumber="293" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis bold="true" box="[772,785,1939,1957]" pageId="16" pageNumber="293">A</emphasis>
, fruiting branchlet, with detail of leaflet (
<emphasis bold="true" box="[1124,1141,1939,1957]" pageId="16" pageNumber="293">A</emphasis>
);
<emphasis bold="true" box="[1157,1170,1939,1957]" pageId="16" pageNumber="293">B</emphasis>
, stipules;
<emphasis bold="true" box="[1257,1271,1939,1957]" pageId="16" pageNumber="293">C</emphasis>
, cross-section of fruit (above) and pyrene;
<emphasis bold="true" box="[302,316,1962,1980]" pageId="16" pageNumber="293">D</emphasis>
, detail of infructescence;
<emphasis bold="true" box="[530,542,1962,1980]" pageId="16" pageNumber="293">E</emphasis>
,
<emphasis bold="true" box="[552,563,1962,1980]" pageId="16" pageNumber="293">F</emphasis>
,
<taxonomicName authority="Daly, Raharim. &amp; Federman" authorityName="Daly, Raharim. &amp; Federman" authorityYear="2015" box="[573,1001,1962,1980]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="16" pageNumber="293" phylum="Tracheophyta" rank="species" species="moramangae" status="sp. nov.">
<emphasis box="[573,767,1962,1980]" italics="true" pageId="16" pageNumber="293">Canarium moramangae</emphasis>
Daly, Raharim. &amp; Federman
</taxonomicName>
,
<taxonomicNameLabel box="[1009,1078,1962,1980]" pageId="16" pageNumber="293" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis bold="true" box="[1084,1096,1962,1980]" pageId="16" pageNumber="293">E</emphasis>
, fruiting branchlet, with detail of leaflet (
<emphasis bold="true" box="[1428,1445,1962,1980]" pageId="16" pageNumber="293">E</emphasis>
);
<emphasis bold="true" box="[132,143,1985,2003]" pageId="16" pageNumber="293">F</emphasis>
, stipule and scar;
<emphasis bold="true" box="[299,312,1985,2003]" pageId="16" pageNumber="293">A</emphasis>
,
<emphasis bold="true" box="[322,336,1984,2002]" pageId="16" pageNumber="293">C</emphasis>
,
<emphasis box="[347,487,1985,2003]" italics="true" pageId="16" pageNumber="293">Daly et al. 13079</emphasis>
(NY sheet and field photos);
<emphasis bold="true" box="[732,751,1985,2003]" pageId="16" pageNumber="293">B,</emphasis>
<emphasis box="[756,923,1985,2003]" italics="true" pageId="16" pageNumber="293">Federman et al. 116</emphasis>
(NY);
<emphasis bold="true" box="[975,989,1985,2003]" pageId="16" pageNumber="293">D</emphasis>
,
<emphasis box="[1000,1140,1985,2003]" italics="true" pageId="16" pageNumber="293">Daly et al. 13073</emphasis>
(field photo by D. Daly);
<emphasis bold="true" box="[1349,1361,1985,2003]" pageId="16" pageNumber="293">E</emphasis>
,
<emphasis bold="true" box="[1372,1383,1985,2003]" pageId="16" pageNumber="293">F</emphasis>
,
<emphasis italics="true" pageId="16" pageNumber="293">Service Forestier-21894 no. 1748-R-100</emphasis>
. Scale bars: A, C-E, 2 cm; B, F, 0.5 cm; A, E, not to scale.
</paragraph>
</caption>
<paragraph blockId="17.[130,777,662,1294]" box="[132,206,662,688]" pageId="17" pageNumber="294">NOTES</paragraph>
<paragraph blockId="17.[130,777,662,1294]" pageId="17" pageNumber="294">
Like
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[187,327,693,720]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="17" pageNumber="294" phylum="Tracheophyta" rank="species" species="lamianum" status="sp. nov.">
<emphasis box="[187,327,693,720]" italics="true" pageId="17" pageNumber="294">C. lamianum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[341,424,694,720]" pageId="17" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[437,569,694,720]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="17" pageNumber="294" phylum="Tracheophyta" rank="species" species="arcuatum" status="sp. nov.">
<emphasis box="[437,569,694,720]" italics="true" pageId="17" pageNumber="294">C. arcuatum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[583,665,694,720]" pageId="17" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
has leaves with few juga, the petiole and rachis with only sparse to scattered hairs, stipules
<quantity box="[346,435,757,784]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="17" pageNumber="294" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3-7 mm</quantity>
long, and their fruits similar in size and shape, but
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[332,463,790,816]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="17" pageNumber="294" phylum="Tracheophyta" rank="species" species="arcuatum" status="sp. nov.">
<emphasis box="[332,463,790,816]" italics="true" pageId="17" pageNumber="294">C. arcuatum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[475,555,790,816]" pageId="17" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished by the stipule pubescence of fine (not thick) hairs, the petiole and rachis with long erect golden hairs to
<quantity box="[599,687,853,880]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="294" unit="mm" value="0.5">0.5 mm</quantity>
long at least on the rachis (vs scattered thick, appressed golden hairs to
<quantity box="[163,265,917,944]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="294" unit="mm" value="0.05">0.05 mm</quantity>
long in
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[365,507,917,943]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="17" pageNumber="294" phylum="Tracheophyta" rank="species" species="lamianum" status="sp. nov.">
<emphasis box="[365,507,917,943]" italics="true" pageId="17" pageNumber="294">C. lamianum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[523,611,917,943]" pageId="17" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
), the pulvinuli usually inconspicuous (vs usually conspicuous), lateral leaflet base usually rounded (vs obtuse to truncate, rarely some leaflet bases acute or cordate), the margin often slightly revolute (vs flat), the secondary veins usually fewer (4-7 vs 6-10), abruptly crowded toward the base (vs often irregularly spaced), and strongly arcuate (vs spreading to nearly straight, rarely slightly arcuate), the infructescence more slender, and the fruit oblong and red-beige (vs broadly (oblong-)ovoid to slightly obovoid, sometimes trigonous, and (grayish-) green to (light) brown).
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[131,341,1236,1262]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="17" pageNumber="294" phylum="Tracheophyta" rank="species" species="arcuatum" status="sp. nov.">
<emphasis box="[131,341,1236,1262]" italics="true" pageId="17" pageNumber="294">Canarium arcuatum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[354,434,1236,1262]" pageId="17" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
is contrasted with
<taxonomicName authority="Daly &amp; Raharimampionona &amp; Federman, 2015" authorityName="Daly &amp; Raharimampionona &amp; Federman" authorityYear="2015" box="[625,766,1236,1262]" class="Magnoliopsida" family="Burseraceae" genus="Canarium" kingdom="Plantae" order="Sapindales" pageId="17" pageNumber="294" phylum="Tracheophyta" rank="species" species="angulatum" status="sp. nov.">
<emphasis box="[625,766,1236,1262]" italics="true" pageId="17" pageNumber="294">C. angulatum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[132,215,1268,1294]" pageId="17" pageNumber="294" rank="species">sp. nov.</taxonomicNameLabel>
in the discussion under that species.
</paragraph>
</subSubSection>
</treatment>
</document>