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<mods:title id="4898FE67C2D3F5A06356FA29CF9CB9BA">An annotated checklist of the South African Acanthodrilidae (Oligochaeta: Acanthodrilidae: Acanthodrilinae, Benhamiinae)</mods:title>
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<mods:namePart id="60D498B09E444046C59A2F0BB5043460">Plisko, Jadwiga Danuta</mods:namePart>
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<mods:date id="6B46978281DD56DC49F72AD025059452">2012</mods:date>
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<treatment id="03F54361FFA5FF91E68BD3F9BC5A6EA5" ID-DOI="http://doi.org/10.5281/zenodo.6171300" ID-GBIF-Taxon="119525015" ID-Zenodo-Dep="6171300" LSID="urn:lsid:plazi:treatment:03F54361FFA5FF91E68BD3F9BC5A6EA5" httpUri="http://treatment.plazi.org/id/03F54361FFA5FF91E68BD3F9BC5A6EA5" lastPageId="30" lastPageNumber="33" pageId="29" pageNumber="33">
<subSubSection id="C346A1FCFFA5FF92E68BD3F9BFCC6B81" box="[151,693,1494,1520]" pageId="29" pageNumber="33" type="nomenclature">
<paragraph id="8BE3F277FFA5FF92E68BD3F9BFCC6B81" blockId="29.[151,693,1494,1520]" box="[151,693,1494,1520]" pageId="29" pageNumber="33">
<heading id="D0AB451BFFA5FF92E68BD3F9BFCC6B81" bold="true" box="[151,693,1494,1520]" fontSize="11" level="1" pageId="29" pageNumber="33" reason="1">
<taxonomicName id="4C5C89F4FFA5FF92E68BD3F9BFCC6B81" ID-CoL="ba9094b6-b66b-48e1-a936-09db9100d29b" authority="Pickford, 1937" authorityName="Pickford" authorityYear="1937" box="[151,693,1494,1520]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="29" pageNumber="33" phylum="Annelida" rank="species" species="michaelsenianus">
<emphasis id="B9282E65FFA5FF92E68BD3F9BFCC6B81" bold="true" box="[151,693,1494,1520]" pageId="29" pageNumber="33">
<emphasis id="B9282E65FFA5FF92E68BD3F9BC8E6B81" bold="true" box="[151,503,1494,1520]" italics="true" pageId="29" pageNumber="33">Parachilota michaelsenianus</emphasis>
<bibRefCitation id="EFCD8F86FFA5FF92E7E2D3F9BFCC6B81" author="Pickford" box="[510,693,1494,1520]" pageId="29" pageNumber="33" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer &amp; Sons Ltd. 612 pp." type="book" year="1937">Pickford, 1937</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C346A1FCFFA5FF92E68BD032BE2768FE" pageId="29" pageNumber="33" type="reference_group">
<paragraph id="8BE3F277FFA5FF92E68BD032BFCA6842" blockId="29.[151,933,1565,1679]" box="[151,691,1565,1587]" pageId="29" pageNumber="33">
<treatmentCitationGroup id="AB4CD559FFA5FF92E68BD032BFCA6842" box="[151,691,1565,1587]" pageId="29" pageNumber="33">
<taxonomicName id="4C5C89F4FFA5FF92E68BD032BFD66842" authority="Michaelsen 1913a: 496" authorityName="Michaelsen" authorityPageNumber="496" authorityYear="1913" box="[151,687,1565,1587]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="29" pageNumber="33" phylum="Annelida" rank="species" species="capensis">
<emphasis id="B9282E65FFA5FF92E68BD032BC396842" box="[151,320,1565,1587]" italics="true" pageId="29" pageNumber="33">Chilota capensis</emphasis>
:
<treatmentCitation id="0AFDD466FFA5FF92E752D032BF456842" author="Michaelsen" box="[334,572,1565,1587]" page="496" pageId="29" pageNumber="33" year="1913">
<bibRefCitation id="EFCD8F86FFA5FF92E752D032BF736842" author="Michaelsen" box="[334,522,1565,1587]" pageId="29" pageNumber="33" refString="Michaelsen, W. (1913 a) Die Oligochaeten des Kaplandes. Zoologische Jahrbucher, Abteilung fur Systematik, 34, 473 - 556." type="journal article" year="1913" yearSuffix="a">Michaelsen 1913a</bibRefCitation>
: 496
</treatmentCitation>
, 1913b: 49
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE3F277FFA5FF92E68BD013BFEF6820" blockId="29.[151,933,1565,1679]" box="[151,662,1595,1618]" pageId="29" pageNumber="33">
<treatmentCitationGroup id="AB4CD559FFA5FF92E68BD013BFEF6820" box="[151,662,1595,1618]" pageId="29" pageNumber="33">
<taxonomicName id="4C5C89F4FFA5FF92E68BD013BFEB6820" ID-CoL="ba9094b6-b66b-48e1-a936-09db9100d29b" authority="Pickford, 1937: 276" authorityName="Pickford" authorityPageNumber="276" authorityYear="1937" box="[151,658,1595,1618]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="29" pageNumber="33" phylum="Annelida" rank="species" species="michaelsenianus">
<emphasis id="B9282E65FFA5FF92E68BD013BCC66823" box="[151,447,1596,1618]" italics="true" pageId="29" pageNumber="33">Parachilota michaelsenianus</emphasis>
<treatmentCitation id="0AFDD466FFA5FF92E7D9D014BFEB6820" author="Pickford" box="[453,658,1595,1617]" page="276" pageId="29" pageNumber="33" year="1937">
<bibRefCitation id="EFCD8F86FFA5FF92E7D9D014BF266820" author="Pickford" box="[453,607,1595,1617]" pageId="29" pageNumber="33" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer &amp; Sons Ltd. 612 pp." type="book" year="1937">Pickford, 1937</bibRefCitation>
: 276
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE3F277FFA5FF92E68BD075BE2768FE" blockId="29.[151,933,1565,1679]" pageId="29" pageNumber="33">
<treatmentCitationGroup id="AB4CD559FFA5FF92E68BD075BE2768FE" pageId="29" pageNumber="33">
<taxonomicName id="4C5C89F4FFA5FF92E68BD075BF8B6801" ID-CoL="ba9094b6-b66b-48e1-a936-09db9100d29b" authority="Reynolds &amp; Cook 1976: 136" authorityName="Reynolds &amp; Cook" authorityPageNumber="136" authorityYear="1976" box="[151,754,1626,1648]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="29" pageNumber="33" phylum="Annelida" rank="species" species="michaelsenianus">
<emphasis id="B9282E65FFA5FF92E68BD075BCC66801" box="[151,447,1626,1648]" italics="true" pageId="29" pageNumber="33">Parachilota michaelsenianus</emphasis>
:
<treatmentCitation id="0AFDD466FFA5FF92E7D0D075BF8B6801" author="Reynolds" box="[460,754,1626,1648]" page="136" pageId="29" pageNumber="33" year="1976">
<bibRefCitation id="EFCD8F86FFA5FF92E7D0D075BFC66801" author="Reynolds" box="[460,703,1626,1648]" pageId="29" pageNumber="33" refString="Reynolds J. W. &amp; Cook, D. G. (1976) Nomenclatura Oligochaetologica. A catalogue of names, descriptions and type specimens of the Oligochaeta. Ottawa: Runge Press. 217 pp." type="book" year="1976">Reynolds &amp; Cook 1976</bibRefCitation>
: 136
</treatmentCitation>
</taxonomicName>
; SAMC; BMNH. nec
<taxonomicName id="4C5C89F4FFA5FF92E6DCD056BF2C68FE" authority="Michaelsen 1900: 147" authorityName="Michaelsen" authorityPageNumber="147" authorityYear="1900" box="[192,597,1657,1679]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="29" pageNumber="33" phylum="Annelida" rank="species" species="capensis">
<emphasis id="B9282E65FFA5FF92E6DCD056BC1368FE" box="[192,362,1657,1679]" italics="true" pageId="29" pageNumber="33">Chilota capensis</emphasis>
<treatmentCitation id="0AFDD466FFA5FF92E76CD056BF2C68FE" author="Michaelsen" box="[368,597,1657,1679]" page="147" pageId="29" pageNumber="33" year="1900">
<bibRefCitation id="EFCD8F86FFA5FF92E76CD056BF5B68FE" author="Michaelsen" box="[368,546,1657,1679]" pageId="29" pageNumber="33" refString="Michaelsen, W. (1900) Oligochaeta. In: Das Tierreich. Lief. 10. Berlin: R. Friedlander &amp; Sohn, 575 pp." type="book" year="1900">Michaelsen 1900</bibRefCitation>
: 147
</treatmentCitation>
</taxonomicName>
: after
<treatmentCitation id="0AFDD466FFA5FF92E48FD056BE2368FE" author="Pickford" box="[659,858,1657,1679]" page="276" pageId="29" pageNumber="33" year="1937">
<bibRefCitation id="EFCD8F86FFA5FF92E48FD056BE5E68FE" author="Pickford" box="[659,807,1657,1679]" pageId="29" pageNumber="33" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer &amp; Sons Ltd. 612 pp." type="book" year="1937">Pickford 1937</bibRefCitation>
: 276
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C346A1FCFFA5FF92E68BD090B86468A7" box="[151,1309,1726,1751]" pageId="29" pageNumber="33" type="materials_examined">
<paragraph id="8BE3F277FFA5FF92E68BD090B86468A7" blockId="29.[151,1436,1726,2003]" box="[151,1309,1726,1751]" pageId="29" pageNumber="33">
<emphasis id="B9282E65FFA5FF92E68BD090BC5468A6" bold="true" box="[151,301,1726,1751]" pageId="29" pageNumber="33">
<typeStatus id="54E74CD5FFA5FF92E68BD090BDAB68A6" box="[151,210,1727,1751]" pageId="29" pageNumber="33">Type</typeStatus>
locality
</emphasis>
.
<emphasis id="B9282E65FFA5FF92E720D091BC1568A6" bold="true" box="[316,364,1726,1751]" pageId="29" pageNumber="33">WC</emphasis>
, Moddergat near Lynedoch, Stellenbosch Dist. (reported by
<bibRefCitation id="EFCD8F86FFA5FF92E238D091B98568A7" author="Michaelsen" box="[1060,1276,1726,1750]" pageId="29" pageNumber="33" refString="Michaelsen, W. (1913 a) Die Oligochaeten des Kaplandes. Zoologische Jahrbucher, Abteilung fur Systematik, 34, 473 - 556." type="journal article" year="1913" yearSuffix="a">Michaelsen, 1913a</bibRefCitation>
,
<bibRefCitation id="EFCD8F86FFA5FF92E31DD091B86968A7" author="Michaelsen" box="[1281,1296,1726,1750]" pageId="29" pageNumber="33" refString="Michaelsen, W. (1913 b) Report upon the Oligochaeta in the South African Museum at Cape Town. Annals of the South African Museum, 13, 43 - 62." type="journal article" year="1913" yearSuffix="b">b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C346A1FCFFA5FF92E6DBD0CDBF99696F" pageId="29" pageNumber="33" type="distribution">
<paragraph id="8BE3F277FFA5FF92E6DBD0CDBF99696F" blockId="29.[151,1436,1726,2003]" pageId="29" pageNumber="33">
<emphasis id="B9282E65FFA5FF92E6DBD0CDBC20688A" bold="true" box="[199,345,1762,1787]" pageId="29" pageNumber="33">Distribution</emphasis>
.
<emphasis id="B9282E65FFA5FF92E777D0CDBCE2688A" bold="true" box="[363,411,1762,1787]" pageId="29" pageNumber="33">WC</emphasis>
, Eastern proximity of Cape Peninsula, Stellenbosch area and at peninsula Kirstenbosch National Botanical Gardens, in the dell flower-beds.
</paragraph>
</subSubSection>
<subSubSection id="C346A1FCFFA5FF91E6DBD104BC5A6EA5" lastPageId="30" lastPageNumber="34" pageId="29" pageNumber="33" type="discussion">
<paragraph id="8BE3F277FFA5FF91E6DBD104BC5A6EA5" blockId="29.[151,1436,1726,2003]" lastBlockId="30.[151,1436,151,212]" lastPageId="30" lastPageNumber="34" pageId="29" pageNumber="33">
<emphasis id="B9282E65FFA5FF92E6DBD104BC4D6932" bold="true" box="[199,308,1835,1859]" pageId="29" pageNumber="33">Remarks</emphasis>
.
<bibRefCitation id="EFCD8F86FFA5FF92E756D105BF7D6933" author="Pickford" box="[330,516,1834,1858]" pageId="29" pageNumber="33" refString="Pickford, G. E. (1937) A monograph of the Acanthodriline earthworms of South Africa. Cambridge: W. Heffer &amp; Sons Ltd. 612 pp." type="book" year="1937">Pickford (1937)</bibRefCitation>
redescribed material collected by
<bibRefCitation id="EFCD8F86FFA5FF92E5B1D105B9ED6933" author="Michaelsen" box="[941,1172,1834,1858]" pageId="29" pageNumber="33" refString="Michaelsen, W. (1913 a) Die Oligochaeten des Kaplandes. Zoologische Jahrbucher, Abteilung fur Systematik, 34, 473 - 556." type="journal article" year="1913" yearSuffix="a">Michaelsen (1913a)</bibRefCitation>
in Stellenbosch area, Moddergat near Lynedoch, misidentified as
<taxonomicName id="4C5C89F4FFA5FF92E49BD161B97C6917" authority="Beddard, 1885" authorityName="Beddard" authorityYear="1885" box="[647,1029,1870,1894]" class="Clitellata" family="Megascolecidae" genus="Chilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="29" pageNumber="33" phylum="Annelida" rank="species" species="capensis">
<emphasis id="B9282E65FFA5FF92E49BD161BE3A6917" box="[647,835,1870,1894]" italics="true" pageId="29" pageNumber="33">Chilota capensis</emphasis>
(
<bibRefCitation id="EFCD8F86FFA5FF92E54FD161BE846917" author="Beddard" box="[851,1021,1870,1894]" pageId="29" pageNumber="33" refString="Beddard, F. E. (1885) Notes on the structure of the new species of earthworm belonging to the genus Acanthodrilus (E. P.). Proceedings of the Royal Physical Society of Edinburgh, 8, 369 - 377." type="journal article" year="1885">Beddard, 1885</bibRefCitation>
)
</taxonomicName>
, and recognized it as a new species,
<taxonomicName id="4C5C89F4FFA5FF92E68BD15CBC9E69FA" box="[151,487,1907,1931]" class="Clitellata" family="Acanthodrilidae" genus="Parachilota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crassiclitellata" pageId="29" pageNumber="33" phylum="Annelida" rank="species" species="michaelsenianus">
<emphasis id="B9282E65FFA5FF92E68BD15CBC9E69FA" box="[151,487,1907,1931]" italics="true" pageId="29" pageNumber="33">Parachilota michaelsenianus</emphasis>
</taxonomicName>
. Some additional systematic observations were made based on Michaelsens specimens and her new material. She made clear that the new species is not identical with Beddards problematic
<taxonomicName id="4C5C89F4FFA5FF92E68BD194BF3669A3" authority="Beddard, 1885" authorityName="Beddard" authorityYear="1885" box="[151,591,1978,2003]" class="Clitellata" family="Acanthodrilidae" genus="Acanthodrilus" kingdom="Animalia" order="Crassiclitellata" pageId="29" pageNumber="33" phylum="Annelida" rank="species" species="capensis">
<emphasis id="B9282E65FFA5FF92E68BD194BCE469A2" box="[151,413,1979,2003]" italics="true" pageId="29" pageNumber="33">Acanthodrilus capensis</emphasis>
<bibRefCitation id="EFCD8F86FFA5FF92E7BAD195BF3669A3" author="Beddard" box="[422,591,1978,2002]" pageId="29" pageNumber="33" refString="Beddard, F. E. (1885) Notes on the structure of the new species of earthworm belonging to the genus Acanthodrilus (E. P.). Proceedings of the Royal Physical Society of Edinburgh, 8, 369 - 377." type="journal article" year="1885">Beddard, 1885</bibRefCitation>
</taxonomicName>
.
<typeStatus id="54E74CD5FFA5FF92E447D195BFBA69A3" box="[603,707,1978,2002]" pageId="29" pageNumber="33" type="holotype">Holotype</typeStatus>
SAMC 147 (from Moddergat near Lynedoch).
<typeStatus id="54E74CD5FFA5FF92E2C2D194B83969A2" box="[1246,1344,1979,2003]" pageId="29" pageNumber="33" type="paratype">Paratype</typeStatus>
BMNH 1938.7.1.1047. Other material: SAMC A21600 (n. sp.?); BMNH 1938.7.1.10481052 with variable hand-written information.
</paragraph>
</subSubSection>
</treatment>
</document>