191 lines
21 KiB
XML
191 lines
21 KiB
XML
<document id="C7990E851572D9120D44353658FB6E15" ID-DOI="10.2478/if-2019-0023" ID-ISSN="2533-4069" ID-Zenodo-Dep="5457521" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630294627634" checkinUser="diego" docAuthor="Hartman, Julian, Van De Weerd, Andrew A., Bruijn, Hans De & Wessels, Wilma" docDate="2019" docId="03E6B8178C38617CFCB6F8837E51FAE5" docLanguage="en" docName="06_Hartman.pdf" docOrigin="Fossil Imprint 75 (3 - 4)" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Sayimys WOOD 1937" docType="treatment" docVersion="3" lastPageNumber="372" masterDocId="FFDFC06F8C346171FFE7FFC67F71FFDB" masterDocTitle="An Exceptional Large Sample Of The Early Miocene Ctenodactyline Rodent Sayimys Giganteus, Specific Variation And Taxonomic Implications" masterLastPageNumber="382" masterPageNumber="359" pageNumber="371" updateTime="1699218057821" updateUser="plazi">
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<mods:title id="AC23028CCD0879000302D63406BED66F">An Exceptional Large Sample Of The Early Miocene Ctenodactyline Rodent Sayimys Giganteus, Specific Variation And Taxonomic Implications</mods:title>
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<mods:name id="72C07D1EE7B2ADF3859F4ED4D10EC987" type="personal">
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<mods:namePart id="426EF8AD1989FD3C7C14E3584EBCBAA5">Hartman, Julian</mods:namePart>
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<mods:affiliation id="A093D3247298E4BF16CCC5C162F79C43">Oranjepoldererf 36, 2807 NE Gouda, the Netherlands; e-mail: juulhartman @ gmail. com. & Department of Earth Sciences, Utrecht University, Princetonlaan 8 A, 3584 CB Utrecht, the Netherlands, P. O. Box 80021 3508 TA Utrecht, the Netherlands; e-mail: a. vandeweerd @ uu. nl, hdbruijn @ uu. nl, w. wessels @ uu. nl.</mods:affiliation>
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<mods:nameIdentifier id="244E9EAD007F7376F20AE26664D97465" type="email">juulhartman@gmail.com.</mods:nameIdentifier>
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<mods:namePart id="25E66BEE35C5773D59862A3A7BE6AC14">Van De Weerd, Andrew A.</mods:namePart>
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<mods:affiliation id="044A7F36D6920912793F0DEDB5983942">Department of Earth Sciences, Utrecht University, Princetonlaan 8 A, 3584 CB Utrecht, the Netherlands, P. O. Box 80021 3508 TA Utrecht, the Netherlands; e-mail: a. vandeweerd @ uu. nl, hdbruijn @ uu. nl, w. wessels @ uu. nl.</mods:affiliation>
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<mods:nameIdentifier id="3C640ABD454E6B2D5088B077D289F205" type="email">a.vandeweerd@uu.nl</mods:nameIdentifier>
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<mods:namePart id="EBE1853108D9BD3B4A18149D2A6D24BC">Bruijn, Hans De</mods:namePart>
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<mods:affiliation id="9BC3E9CEA5B9D39A010B7820E7DA806F">Department of Earth Sciences, Utrecht University, Princetonlaan 8 A, 3584 CB Utrecht, the Netherlands, P. O. Box 80021 3508 TA Utrecht, the Netherlands; e-mail: a. vandeweerd @ uu. nl, hdbruijn @ uu. nl, w. wessels @ uu. nl.</mods:affiliation>
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<mods:nameIdentifier id="F585FB14FBF82CF82B2CB924A5F83CF0" type="email">a.vandeweerd@uu.nl</mods:nameIdentifier>
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<mods:namePart id="5124F38F979A0E21CB9549375611BF17">Wessels, Wilma</mods:namePart>
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<mods:affiliation id="993ABAF88B1C671179DA3CEBB572E34F">Department of Earth Sciences, Utrecht University, Princetonlaan 8 A, 3584 CB Utrecht, the Netherlands, P. O. Box 80021 3508 TA Utrecht, the Netherlands; e-mail: a. vandeweerd @ uu. nl, hdbruijn @ uu. nl, w. wessels @ uu. nl.</mods:affiliation>
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<mods:nameIdentifier id="EF52FB09F4D0C8C6CA92E39E06AC9535" type="email">a.vandeweerd@uu.nl</mods:nameIdentifier>
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<mods:title id="1C2590B467ED075B1CA040F9AE791BF3">Fossil Imprint</mods:title>
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<mods:part id="809E04944EAEB3B3CD68A58DFE2B0243">
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<mods:date id="1A76AC383CC63E17AA4DD576751DBDE7">2019</mods:date>
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<mods:number id="3DC6D19745E9D4AA4156EB87A53F4852">2019-12-30</mods:number>
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<mods:number id="E79AA4C0D24A23AAD33A4D1536FCB2EA">75</mods:number>
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<mods:detail id="F27C223AF95E307B8244A85E132328CA" type="issue">
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<mods:number id="F1B07D2C97C7087369984C33C52989FA">3 - 4</mods:number>
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<mods:start id="4983638DCBFEF76CA063FE7ABD16E4FD">359</mods:start>
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<mods:end id="41B0628E1AEB7A8324AB8B75A9981CDA">382</mods:end>
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<mods:classification id="BD6A23875AD36BF456EF48AA53507113">journal article</mods:classification>
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<mods:identifier id="61D4DB04DF3982ED60BD8167B7659C8F" type="DOI">10.2478/if-2019-0023</mods:identifier>
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<mods:identifier id="88427A6E0FAF66EBF4C9538E62515064" type="ISSN">2533-4069</mods:identifier>
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<mods:identifier id="7BB3A4E2E36E28559EFD696046C857EA" type="Zenodo-Dep">5457521</mods:identifier>
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<treatment id="03E6B8178C38617CFCB6F8837E51FAE5" LSID="urn:lsid:plazi:treatment:03E6B8178C38617CFCB6F8837E51FAE5" httpUri="http://treatment.plazi.org/id/03E6B8178C38617CFCB6F8837E51FAE5" lastPageId="13" lastPageNumber="372" pageId="12" pageNumber="371">
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<subSubSection id="C3555A8A8C38617DFCB6F8837BBBF856" pageId="12" pageNumber="371" type="nomenclature">
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<paragraph id="8BF009018C38617DFCB6F8837AE7F886" blockId="12.[849,1430,1861,1885]" box="[849,1430,1861,1885]" pageId="12" pageNumber="371">
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<heading id="D0B8BE6D8C38617DFCB6F8837AE7F886" box="[849,1430,1861,1885]" centered="true" fontSize="8" level="2" pageId="12" pageNumber="371" reason="2">
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<emphasis id="B93BD5138C38617DFCB6F8837C84F886" bold="true" box="[849,1013,1861,1885]" italics="true" pageId="12" pageNumber="371">
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<taxonomicName id="4C4F72828C38617DFCB6F8837CDAF886" authorityName="WOOD" authorityYear="1937" box="[849,939,1861,1885]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="12" pageNumber="371" phylum="Chordata" rank="genus">Sayimys</taxonomicName>
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minor
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</emphasis>
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<bibRefCitation id="EFDE74F08C38617DFC1CF88C7AE7F886" author="Bruijn, H. de & Hussain, S. T. & Leinders, J. J. M." box="[1019,1430,1861,1885]" pageId="12" pageNumber="371" pagination="71 - 99" refId="ref15995" refString="Bruijn, H. de, Hussain, S. T., Leinders, J. J. M. (1981): Fossil rodents from the Murree formation near Banda Daud Shah, Kohat, Pakistan. - Proceedings of the Koninklijke Nederlandse Akademie van Wetenschappen, B, 84 (1): 71 - 99." type="journal article" year="1981">
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<emphasis id="B93BD5138C38617DFC1CF88C7AE7F886" bold="true" box="[1019,1430,1861,1885]" pageId="12" pageNumber="371">DE BRUIJN, HUSSAIN et LEINDERS, 1981</emphasis>
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</bibRefCitation>
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</heading>
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</paragraph>
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<paragraph id="8BF009018C38617DFCBDF8B37BBBF856" blockId="12.[858,1226,1908,1933]" box="[858,1226,1908,1933]" pageId="12" pageNumber="371">H o l o t y p e. A left m1–2 no 313.</paragraph>
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</subSubSection>
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<subSubSection id="C3555A8A8C38617CFCBDF8627E51FAE5" lastPageId="13" lastPageNumber="372" pageId="12" pageNumber="371" type="description">
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<paragraph id="8BF009018C38617DFCBDF8627CA2F826" blockId="12.[820,1459,1956,2045]" pageId="12" pageNumber="371">
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Ty p e l o c a l i t y. H.-GSP116, Murree Formation near Banda Daud Shah,
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<collectingCountry id="F35849918C38617DFBEFF8027B17F807" box="[1032,1126,1988,2012]" name="Pakistan" pageId="12" pageNumber="371">Pakistan</collectingCountry>
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; age of the site: 18.5–19.5 Ma (
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<bibRefCitation id="EFDE74F08C38617DFCDCF8237CB9F826" author="Baskin, J." box="[827,968,2021,2045]" pageId="12" pageNumber="371" pagination="1 - 49" refId="ref15763" refString="Baskin, J. (1996): Systematic revision of Ctenodactylidae (Mammalia, Rodentia) from the Miocene of Pakistan. - Palaeovertebrata, 25: 1 - 49." type="journal article" year="1996">Baskin 1996</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BF009018C38617DFCBDF7D37AC2F7F7" blockId="12.[858,1459,2068,2093]" box="[858,1459,2068,2093]" pageId="12" pageNumber="371">P a r a t y p e s. One damaged dp4, one m3 and one M1–2.</paragraph>
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<paragraph id="8BF009018C39617CFF41FF5E7C71FF2B" blockId="13.[129,768,151,1343]" pageId="13" pageNumber="372">
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D i s c u s s i o n. A well-developed mesolophid is present in the dp4, but absent in the m1–2 and m3. The sizes of the specimens are like those of
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<emphasis id="B93BD5138C39617CFE42FF1F7D4CFF2B" box="[421,573,216,240]" italics="true" pageId="13" pageNumber="372">
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<taxonomicName id="4C4F72828C39617CFE42FF1F7E8DFF2B" authorityName="WOOD" authorityYear="1937" box="[421,508,217,240]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="372" phylum="Chordata" rank="genus">Sayimys</taxonomicName>
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flynni
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</emphasis>
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, or slightly larger.
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</paragraph>
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<paragraph id="8BF009018C39617CFF41FF3F7E51FAE5" blockId="13.[129,768,151,1343]" pageId="13" pageNumber="372">
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This small collection of only four molars has been the subject of some discussion.
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<bibRefCitation id="EFDE74F08C39617CFE5BFEDF7D28FEEA" author="Baskin, J." box="[444,601,281,305]" pageId="13" pageNumber="372" pagination="1 - 49" refId="ref15763" refString="Baskin, J. (1996): Systematic revision of Ctenodactylidae (Mammalia, Rodentia) from the Miocene of Pakistan. - Palaeovertebrata, 25: 1 - 49." type="journal article" year="1996">Baskin (1996)</bibRefCitation>
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suggested that the rather worn M1–2 is too large to fit in the same species as the other three teeth, without testing the homogeneity of the sample. The m1–2 and M1–2 of all large
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<taxonomicName id="4C4F72828C39617CFD4FFEBD7D8EFE49" authorityName="WOOD" authorityYear="1937" box="[680,767,379,402]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="372" phylum="Chordata" rank="genus">
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<emphasis id="B93BD5138C39617CFD4FFEBD7D8EFE49" box="[680,767,379,402]" italics="true" pageId="13" pageNumber="372">Sayimys</emphasis>
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</taxonomicName>
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populations have a large size variability if first and second molars are not separated (
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<tableCitation id="C6CD3CBA8C39617CFE75FE7D7EA9FE08" box="[402,472,443,467]" captionStart="Table 2" captionStartId="9.[147,203,1514,1535]" captionText="Table 2.Coefficient ofvariation V’ inthree large samples ofSayimys, Keseköy, Daud Kheland H.-GSP82.24. N = numberof specimens." pageId="13" pageNumber="372">Tab. 2</tableCitation>
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). This M1–2 may very well be a large second molar belonging to the same species as the other three teeth.
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<bibRefCitation id="EFDE74F08C39617CFE8AFE3B7E8AFDCF" author="Wang, B." box="[365,507,508,533]" pageId="13" pageNumber="372" pagination="1 - 88" refId="ref17327" refString="Wang, B. (1997): The mid-Tertiary Ctenodactylidae (Rodentia, Mammalia) of Eastern and Central Asia. - Bulletin of the American Museum of Natural History, 234: 1 - 88." type="journal article" year="1997">Wang (1997)</bibRefCitation>
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suggested that
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<emphasis id="B93BD5138C39617CFD4FFE3B7FB0FDEF" italics="true" pageId="13" pageNumber="372">
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<taxonomicName id="4C4F72828C39617CFD4FFE3B7D8EFDCF" authorityName="WOOD" authorityYear="1937" box="[680,767,509,532]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="372" phylum="Chordata" rank="genus">Sayimys</taxonomicName>
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minor
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</emphasis>
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could be a junior synonym of
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<taxonomicName id="4C4F72828C39617CFDC1FDDB7E26FD8E" authority="(Sen and Thomas 1979)" baseAuthorityName="Sen and Thomas" baseAuthorityYear="1979" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="372" phylum="Chordata" rank="species" species="intermedius">
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<emphasis id="B93BD5138C39617CFDC1FDDB7D42FDEF" box="[550,563,541,564]" italics="true" pageId="13" pageNumber="372">S</emphasis>
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.
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<emphasis id="B93BD5138C39617CFDA2FDDB7DB2FDEE" box="[581,707,541,565]" italics="true" pageId="13" pageNumber="372">intermedius</emphasis>
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(
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<bibRefCitation id="EFDE74F08C39617CFD31FDDB7E3EFD8E" author="Sen, S. & Thomas, H." pageId="13" pageNumber="372" pagination="34 - 37" refId="ref17047" refString="Sen, S., Thomas, H. (1979): Decouverte de rongeurs dans le Miocene moyen de la Formation Hofuf (Province du Hasa, Arabie saoudite). - Compte Rendu Sommaire des seances de la Societe Geologique de France, 1979 (1): 34 - 37." type="journal article" year="1979">Sen and Thomas 1979</bibRefCitation>
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)
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</taxonomicName>
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, because these species are similar in size, the only difference being the presence of a mesolophid in the single dp4 of
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<emphasis id="B93BD5138C39617CFE8BFDB87E08FD4E" box="[364,377,638,661]" italics="true" pageId="13" pageNumber="372">S</emphasis>
|
||
.
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<emphasis id="B93BD5138C39617CFE6CFDB97EBAFD4D" box="[395,459,639,662]" italics="true" pageId="13" pageNumber="372">minor</emphasis>
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and absence of this lophid in the single known dp4 of
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<emphasis id="B93BD5138C39617CFE53FD597EB0FD6D" box="[436,449,671,694]" italics="true" pageId="13" pageNumber="372">S</emphasis>
|
||
.
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<emphasis id="B93BD5138C39617CFE36FD587D34FD6D" box="[465,581,670,694]" italics="true" pageId="13" pageNumber="372">intermediu</emphasis>
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||
s. But, as
|
||
<bibRefCitation id="EFDE74F08C39617CFD52FD587F80FD0C" author="Baskin, J." pageId="13" pageNumber="372" pagination="1 - 49" refId="ref15763" refString="Baskin, J. (1996): Systematic revision of Ctenodactylidae (Mammalia, Rodentia) from the Miocene of Pakistan. - Palaeovertebrata, 25: 1 - 49." type="journal article" year="1996">Baskin (1996: 41)</bibRefCitation>
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mentioned: “persistence of a rodent species over 5 million years and
|
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<quantity id="4CB7A4E48C39617CFE8BFD197EA7FD2C" box="[364,470,735,759]" metricMagnitude="6" metricUnit="m" metricValue="3.5" pageId="13" pageNumber="372" unit="km" value="3500.0">3,500 km</quantity>
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distance would be highly unusual”, and we therefore hesitate to follow Wang. López- Antoñanzas and Sen (2003, 2004) subdivided the sample of
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<emphasis id="B93BD5138C39617CFF66FC877FFFFC83" box="[129,142,833,856]" italics="true" pageId="13" pageNumber="372">S</emphasis>
|
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.
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<emphasis id="B93BD5138C39617CFF7BFC877FADFC83" box="[156,220,833,856]" italics="true" pageId="13" pageNumber="372">minor</emphasis>
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from the type locality (only four teeth) into two or three species. The
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<typeStatus id="54F4B7A38C39617CFEA4FCA77ED1FCA2" box="[323,416,865,889]" pageId="13" pageNumber="372" type="holotype">holotype</typeStatus>
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(the m1–2, no 313) was included in
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<taxonomicName id="4C4F72828C39617CFF45FC447D2EFC42" authority="(Sen and Thomas 1979)" baseAuthorityName="Sen and Thomas" baseAuthorityYear="1979" box="[162,607,897,921]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="372" phylum="Chordata" rank="species" species="intermedius">
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<emphasis id="B93BD5138C39617CFF45FC447FDEFC42" box="[162,175,898,921]" italics="true" pageId="13" pageNumber="372">S</emphasis>
|
||
.
|
||
<emphasis id="B93BD5138C39617CFF24FC477E30FC42" box="[195,321,897,921]" italics="true" pageId="13" pageNumber="372">intermedius</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDE74F08C39617CFEB2FC477D29FC42" author="Sen, S. & Thomas, H." box="[341,600,897,921]" pageId="13" pageNumber="372" pagination="34 - 37" refId="ref17047" refString="Sen, S., Thomas, H. (1979): Decouverte de rongeurs dans le Miocene moyen de la Formation Hofuf (Province du Hasa, Arabie saoudite). - Compte Rendu Sommaire des seances de la Societe Geologique de France, 1979 (1): 34 - 37." type="journal article" year="1979">Sen and Thomas 1979</bibRefCitation>
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)
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</taxonomicName>
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because “this molar cannot be differentiated from the equivalent tooth of
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<taxonomicName id="4C4F72828C39617CFF66FC057E6DFC01" baseAuthorityName="Sen and Thomas" baseAuthorityYear="1979" box="[129,284,962,986]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="372" phylum="Chordata" rank="species" species="intermedius">
|
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<emphasis id="B93BD5138C39617CFF66FC057FFFFC01" box="[129,142,963,986]" italics="true" pageId="13" pageNumber="372">S</emphasis>
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.
|
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<emphasis id="B93BD5138C39617CFF79FC047E6DFC01" box="[158,284,962,986]" italics="true" pageId="13" pageNumber="372">intermedius</emphasis>
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</taxonomicName>
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”. The dp4 and m3 were separated from the m1–2 for unknown reasons, and included into
|
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<emphasis id="B93BD5138C39617CFD74FC257DD1FC21" box="[659,672,995,1018]" italics="true" pageId="13" pageNumber="372">S</emphasis>
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.
|
||
<emphasis id="B93BD5138C39617CFD55FC257D8EFC20" box="[690,767,995,1019]" italics="true" pageId="13" pageNumber="372">baskini</emphasis>
|
||
LÓPEZ- ANTOÑANZAS et SEN, 2003, see below. It is not clear in this publication how to classify the fourth tooth in the sample from the type locality (the large M1–2). The clear remark by
|
||
<bibRefCitation id="EFDE74F08C39617CFF15FBA27EE2FBA6" author="Munthe, J." box="[242,403,1124,1149]" pageId="13" pageNumber="372" pagination="1 - 36" refId="ref16962" refString="Munthe, J. (1980): Rodents of the Miocene Daud Khel local fauna, Mianwali district, Pakistan. Part 1. Sciuridae, Gliridae, Ctenodactylidae, and Rhizomyidae. - Contributions in Biology and Geology, Milwauke Public Museum, 34: 1 - 36." type="journal article" year="1980">Munthe (1980)</bibRefCitation>
|
||
on the large variability in
|
||
<taxonomicName id="4C4F72828C39617CFD4FFBA37D8EFBA7" authorityName="WOOD" authorityYear="1937" box="[680,767,1125,1148]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="372" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93BD5138C39617CFD4FFBA37D8EFBA7" box="[680,767,1125,1148]" italics="true" pageId="13" pageNumber="372">Sayimys</emphasis>
|
||
</taxonomicName>
|
||
was here completely forgotten or ignored. In our opinion, the assemblage from H.-GSP116 (Murree Formation) is homogenous, and there is no valid reason to split up the small collection of
|
||
<emphasis id="B93BD5138C39617CFE63FB217EE0FB25" box="[388,401,1255,1278]" italics="true" pageId="13" pageNumber="372">S</emphasis>
|
||
.
|
||
<emphasis id="B93BD5138C39617CFE44FB217E92FB25" box="[419,483,1255,1278]" italics="true" pageId="13" pageNumber="372">minor</emphasis>
|
||
and include the
|
||
<typeStatus id="54F4B7A38C39617CFD45FB207D8EFB25" box="[674,767,1254,1278]" pageId="13" pageNumber="372" type="holotype">holotype</typeStatus>
|
||
in
|
||
<taxonomicName id="4C4F72828C39617CFF45FAC17E31FAC4" baseAuthorityName="Sen and Thomas" baseAuthorityYear="1979" box="[162,320,1287,1311]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="372" phylum="Chordata" rank="species" species="intermedius">
|
||
<emphasis id="B93BD5138C39617CFF45FAC17FDEFAC5" box="[162,175,1287,1310]" italics="true" pageId="13" pageNumber="372">S</emphasis>
|
||
.
|
||
<emphasis id="B93BD5138C39617CFF25FAC17E31FAC4" box="[194,320,1287,1311]" italics="true" pageId="13" pageNumber="372">intermedius</emphasis>
|
||
</taxonomicName>
|
||
, and thus to synonymize
|
||
<emphasis id="B93BD5138C39617CFD8BFAC17D08FAC5" box="[620,633,1287,1310]" italics="true" pageId="13" pageNumber="372">S</emphasis>
|
||
.
|
||
<emphasis id="B93BD5138C39617CFD6AFAC17DBCFAC5" box="[653,717,1287,1310]" italics="true" pageId="13" pageNumber="372">minor</emphasis>
|
||
and
|
||
<taxonomicName id="4C4F72828C39617CFF66FAE17E68FAE4" baseAuthorityName="Sen and Thomas" baseAuthorityYear="1979" box="[129,281,1319,1343]" class="Mammalia" family="Ctenodactylidae" genus="Sayimys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="13" pageNumber="372" phylum="Chordata" rank="species" species="intermedius">
|
||
<emphasis id="B93BD5138C39617CFF66FAE17FFFFAE5" box="[129,142,1319,1342]" italics="true" pageId="13" pageNumber="372">S</emphasis>
|
||
.
|
||
<emphasis id="B93BD5138C39617CFF7CFAE17E68FAE4" box="[155,281,1319,1343]" italics="true" pageId="13" pageNumber="372">intermedius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |