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<document id="15008E00CEC1F9E787129D1AA09C2BBE" ID-CLB-Dataset="29406" ID-DOI="10.11646/zootaxa.4462.2.1" ID-GBIF-Dataset="b52189b5-2b57-474a-86b6-40580b016ecc" ID-ISSN="1175-5326" ID-Zenodo-Dep="1441457" ID-ZooBank="FA92FB55-EA2E-4193-946E-F780A16EB3CC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538402499097" checkinUser="plazi" docAuthor="Janák, Jiří" docDate="2018" docId="03DD5963FFF8FFD1FF7AFE86FB90EAFC" docLanguage="en" docName="zootaxa.4462.2.1.pdf" docOrigin="Zootaxa 4462 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acylophorus delphinus Fauvel 1905" docType="treatment" docVersion="7" lastPageNumber="164" masterDocId="FFE4211BFFF5FFDCFFEDFFE7FFFCED62" masterDocTitle="Revision of the genus Acylophorus Nordmann, 1837 from Madagascar and Mascarene Islands (Coleoptera: Staphylinidae: Staphylininae: Acylophorina)" masterLastPageNumber="191" masterPageNumber="151" pageNumber="164" updateTime="1698561012784" updateUser="ExternalLinkService">
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<mods:title id="1AA64AFBBA4BDA14EAA97AC0787DABC6">Revision of the genus Acylophorus Nordmann, 1837 from Madagascar and Mascarene Islands (Coleoptera: Staphylinidae: Staphylininae: Acylophorina)</mods:title>
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<mods:namePart id="0C80D013C0FB339553D6AF4C2F5B568F">Janák, Jiří</mods:namePart>
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<mods:date id="1D3E91F41441EAFC7E814BBF360D4129">2018</mods:date>
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<mods:number id="1060F832542D3C2919E10585A8BF8FDB">2018-08-27</mods:number>
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<treatment id="03DD5963FFF8FFD1FF7AFE86FB90EAFC" ID-DOI="http://doi.org/10.5281/zenodo.5960295" ID-GBIF-Taxon="148424833" ID-Zenodo-Dep="5960295" LSID="urn:lsid:plazi:treatment:03DD5963FFF8FFD1FF7AFE86FB90EAFC" httpUri="http://treatment.plazi.org/id/03DD5963FFF8FFD1FF7AFE86FB90EAFC" lastPageNumber="164" pageId="13" pageNumber="164">
<subSubSection id="C36EBBFEFFF8FFD1FF7AFE86FE99ECFF" pageId="13" pageNumber="164" type="nomenclature">
<paragraph id="8BCBE875FFF8FFD1FF7AFE86FDA8EC19" blockId="13.[151,596,353,413]" box="[151,596,353,379]" pageId="13" pageNumber="164">
<heading id="D0835F19FFF8FFD1FF7AFE86FDA8EC19" bold="true" box="[151,596,353,379]" fontSize="11" level="1" pageId="13" pageNumber="164" reason="1">
<taxonomicName id="4C7493F6FFF8FFD1FF7AFE86FDA8EC19" ID-CoL="9W6Y" authority="Fauvel, 1905" authorityName="Fauvel" authorityYear="1905" box="[151,596,353,379]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="delphinus">
<emphasis id="B9003467FFF8FFD1FF7AFE86FDA8EC19" bold="true" box="[151,596,353,379]" pageId="13" pageNumber="164">
<emphasis id="B9003467FFF8FFD1FF7AFE86FE52EC19" bold="true" box="[151,430,353,379]" italics="true" pageId="13" pageNumber="164">Acylophorus delphinus</emphasis>
<bibRefCitation id="EFE59584FFF8FFD1FE58FE86FDA8EC19" box="[437,596,353,379]" pageId="13" pageNumber="164" refId="ref17474" refString="Fauvel, A. (1905) Staphylinides exotiques nouveaux. 3 e Partie. Revue d'Entomologie, 24, 113 - 147." type="journal article">Fauvel, 1905</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCBE875FFF8FFD1FF7AFE63FE99ECFF" blockId="13.[151,596,353,413]" box="[151,357,388,413]" pageId="13" pageNumber="164">
(
<figureCitation id="134FF4F0FFF8FFD1FF72FE63FEDBECFF" box="[159,295,388,413]" captionStart-0="FIGURES 5260" captionStart-1="FIGURES 6166" captionStartId-0="14.[151,264,1907,1929]" captionStartId-1="15.[151,264,1933,1955]" captionTargetBox-0="[160,1423,212,1874]" captionTargetBox-1="[179,1374,273,1811]" captionTargetId-0="figure@14.[151,1435,193,1885]" captionTargetId-1="figure@15.[152,1435,240,1854]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 5260. Acylophorus delphinus Fauvel. 52, 53, 5660, Kirindi; 54, 55, Ampatika. 60, holotype. 52, habitus; 53, head and pronotum; 54, mandibles; 55, maxillary palpus; 56, aedeagus lateral; 57, aedeagus ventro-lateral; 5860, aedeagus ventral. 5660, same scale bars." captionText-1="FIGURES 6166. Acylophorus delphinus Fauvel. 6164, Kirindy; 65, 66, Ampatika. 61, aedeagus lateral; 62, aedeagus ventral; 63, male tergites 9, 10; 64, male sternite 9; 65, female tergites 9, 10; 66, valves. 61, 62, transmitted light. 6162, 6566, same scale bars." httpUri-0="https://zenodo.org/record/1441473/files/figure.png" httpUri-1="https://zenodo.org/record/1441475/files/figure.png" pageId="13" pageNumber="164">Figs. 5266</figureCitation>
,
<figureCitation id="134FF4F0FFF8FFD1FEDFFE63FEA1ECFF" box="[306,349,388,413]" captionStart="FIGURES 172177" captionStartId="38.[151,264,1927,1949]" captionTargetBox="[265,1321,625,1895]" captionTargetId="figure@38.[265,1321,625,1909]" captionTargetPageId="38" captionText="FIGURES 172177. Distribution maps. 172, Acylophorus tristis Jarrige; 173, A. derougemonti sp. nov.; 174, A. muscorum Jarrige; 175, A. delphinus Fauvel (D), A. madecassus sp. nov. (Ŧ); 176, A. minutus Jarrige (D), A. hova sp. nov. (Ɨ), A. montanus sp. nov. (Δ); 177, A. sogai Jarrige." httpUri="https://zenodo.org/record/1441507/files/figure.png" pageId="13" pageNumber="164">175</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36EBBFEFFF8FFD1FF7AFE2BFE2BEF62" pageId="13" pageNumber="164" type="reference_group">
<paragraph id="8BCBE875FFF8FFD1FF7AFE2BFE2BEF62" blockId="13.[151,1436,460,512]" pageId="13" pageNumber="164">
<treatmentCitationGroup id="AB64CF5BFFF8FFD1FF7AFE2BFE2BEF62" pageId="13" pageNumber="164">
<taxonomicName id="4C7493F6FFF8FFD1FF7AFE2BFDBFEC80" ID-CoL="9W6Y" authority="Fauvel, 1905: 181" authorityName="Fauvel" authorityPageNumber="181" authorityYear="1905" box="[151,579,460,482]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="delphinus">
<emphasis id="B9003467FFF8FFD1FF7AFE2BFE7CEC80" box="[151,384,460,482]" italics="true" pageId="13" pageNumber="164">Acylophorus delphinus</emphasis>
<treatmentCitation id="0AD5CE64FFF8FFD1FE6AFE2BFDBFEC80" author="" box="[391,579,460,482]" page="181" pageId="13" pageNumber="164">
<bibRefCitation id="EFE59584FFF8FFD1FE6AFE2BFDECEC80" box="[391,528,460,482]" pageId="13" pageNumber="164" refId="ref17474" refString="Fauvel, A. (1905) Staphylinides exotiques nouveaux. 3 e Partie. Revue d'Entomologie, 24, 113 - 147." type="journal article">Fauvel, 1905</bibRefCitation>
: 181
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0AD5CE64FFF8FFD1FDA0FE2BFC46EC80" author="" box="[589,954,460,482]" page="438" pageId="13" pageNumber="164">
<bibRefCitation id="EFE59584FFF8FFD1FDA0FE2BFC7BEC80" box="[589,903,460,482]" pageId="13" pageNumber="164" refId="ref17193" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. 5 (57). Junk, Berlin, pp. 289 - 408." type="book chapter">Bernhauer and Schubert, 1914</bibRefCitation>
: 438
</treatmentCitation>
;
<treatmentCitation id="0AD5CE64FFF8FFD1FC28FE2BFB7CEC80" author="" box="[965,1152,460,482]" page="141" pageId="13" pageNumber="164">
<bibRefCitation id="EFE59584FFF8FFD1FC28FE2BFBB1EC80" box="[965,1101,460,482]" pageId="13" pageNumber="164" refId="ref17650" refString="Jarrige, J. (1965) Les Acylophorus malgaches (Col. Staphylinidae). Bullettin de la Societ entomologique de France, 70, 139 - 143." type="journal article">Jarrige, 1965</bibRefCitation>
: 141
</treatmentCitation>
;
<treatmentCitation id="0AD5CE64FFF8FFD1FB66FE2AFA9DEC80" author="" box="[1163,1377,460,482]" page="3028" pageId="13" pageNumber="164">
<bibRefCitation id="EFE59584FFF8FFD1FB66FE2AFADCEC80" box="[1163,1312,460,482]" pageId="13" pageNumber="164" refId="ref17555" refString="Herman (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article">Herman, 2001</bibRefCitation>
: 3028
</treatmentCitation>
;
<treatmentCitation id="0AD5CE64FFF8FFD1FA86FE2AFEE8EF62" author="" page="9" pageId="13" pageNumber="164">
<bibRefCitation id="EFE59584FFF8FFD1FA86FE2AFF02EF62" pageId="13" pageNumber="164" refId="ref17680" refString="Lott, D. A. (2010) The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub- Saharan Africa. Zootaxa, 2401, 1 - 51." type="journal article">Lott, 2010</bibRefCitation>
: 9
</treatmentCitation>
(characters, Figs.).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36EBBFEFFF8FFD1FF7AFDCAFA83EE7C" pageId="13" pageNumber="164" type="materials_examined">
<paragraph id="8BCBE875FFF8FFD1FF7AFDCAFC9FEF27" blockId="13.[151,1437,557,1950]" box="[151,867,557,582]" pageId="13" pageNumber="164">
<materialsCitation id="3B1CE228FFF8FFD1FF7AFDCAFC9FEF27" ID-GBIF-Occurrence="1913892206" box="[151,867,557,582]" country="Madagascar" latitude="-25.166666" location="Ambovombe" longLatPrecision="1247" longitude="46.083332" pageId="13" pageNumber="164" specimenCount="1" typeStatus="holotype">
<emphasis id="B9003467FFF8FFD1FF7AFDCAFEC9EF24" bold="true" box="[151,309,557,582]" pageId="13" pageNumber="164">
<typeStatus id="54CF56D7FFF8FFD1FF7AFDCAFF2EEF24" box="[151,210,557,582]" pageId="13" pageNumber="164">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F363A8E5FFF8FFD1FED1FDCAFE39EF24" box="[316,453,557,582]" name="Madagascar" pageId="13" pageNumber="164">Madagascar</collectingCountry>
,
<location id="8EABBEAEFFF8FFD1FE3DFDCAFD99EF24" LSID="urn:lsid:plazi:treatment:03DD5963FFF8FFD1FF7AFE86FB90EAFC:8EABBEAEFFF8FFD1FE3DFDCAFD99EF24" box="[464,613,557,582]" country="Madagascar" latitude="-25.166666" longLatPrecision="1247" longitude="46.083332" name="Ambovombe" pageId="13" pageNumber="164">Ambovombé</location>
, ca.
<geoCoordinate id="EE408EB2FFF8FFD1FD7AFDCAFD0FEF24" box="[663,755,557,582]" direction="south" orientation="latitude" pageId="13" pageNumber="164" precision="925" value="-25.166666">25 10' S</geoCoordinate>
,
<geoCoordinate id="EE408EB2FFF8FFD1FCEDFDCAFCA3EF27" box="[768,863,557,582]" direction="east" orientation="longitude" pageId="13" pageNumber="164" precision="925" value="46.083332">46 05' E</geoCoordinate>
.
</materialsCitation>
</paragraph>
<paragraph id="8BCBE875FFF8FFD1FF2AFDB6FD21EFD0" blockId="13.[151,1437,557,1950]" pageId="13" pageNumber="164">
<emphasis id="B9003467FFF8FFD1FF2AFDB6FD6FEF08" bold="true" box="[199,659,593,618]" pageId="13" pageNumber="164">
<typeStatus id="54CF56D7FFF8FFD1FF2AFDB6FEFEEF08" box="[199,258,593,618]" pageId="13" pageNumber="164">Type</typeStatus>
material examined.
<typeStatus id="54CF56D7FFF8FFD1FDCBFDB6FD6FEF08" box="[550,659,593,618]" pageId="13" pageNumber="164" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D7223FCFFF8FFD1FD47FDB5FD3EEF0B" box="[682,706,594,617]" pageId="13" pageNumber="164" type="male"></specimenCount>
:
<collectingCountry id="F363A8E5FFF8FFD1FD33FDB6FC57EF08" box="[734,939,593,618]" name="Madagascar" pageId="13" pageNumber="164">
<emphasis id="B9003467FFF8FFD1FD33FDB6FC57EF08" bold="true" box="[734,939,593,618]" pageId="13" pageNumber="164">MADAGASCAR</emphasis>
</collectingCountry>
: “Ambovombé Madag.”, “
<taxonomicName id="4C7493F6FFF8FFD1FAE4FDB6FEE8EFEC" authority="Nordmann" authorityName="Nordmann" authorityYear="1837" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">Acylophorus Nordmann</taxonomicName>
”, “
<taxonomicName id="4C7493F6FFF8FFD1FEDEFD92FE2BEFEC" authority="Fvl." authorityName="Fvl." box="[307,471,629,654]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="delphinus">delphinus Fvl.</taxonomicName>
”, “R.I.Sc.N.B. 17.479 Coll. et det. A. Fauvel”, “
<collectionCode id="ED6570B0FFF8FFD1FBECFD91FBB2EFEC" box="[1025,1102,630,654]" pageId="13" pageNumber="164">
<typeStatus id="54CF56D7FFF8FFD1FBECFD91FBB2EFEC" box="[1025,1102,630,654]" pageId="13" pageNumber="164">TYPE</typeStatus>
</collectionCode>
”, “
<typeStatus id="54CF56D7FFF8FFD1FB83FD91FAF8EFEC" box="[1134,1284,630,654]" pageId="13" pageNumber="164" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C7493F6FFF8FFD1FAE1FD92FEABEFD0" authority="Fauvel" authorityName="Fauvel" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="delphinus">Acylophorus delphinus Fauvel</taxonomicName>
m# det. DA Lott, 2009” (
<collectionCode id="ED6570B0FFF8FFD1FD94FD7DFD2EEFD0" box="[633,722,666,690]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="164">IRSNB</collectionCode>
).
</paragraph>
<paragraph id="8BCBE875FFF8FFD1FF2AFD5AFA83EE7C" blockId="13.[151,1437,557,1950]" pageId="13" pageNumber="164">
<materialsCitation id="3B1CE228FFF8FFD1FF2AFD5AFA83EE7C" ID-GBIF-Occurrence="1913892203" collectingDate="1996-12-12" collectingDateMax="2004-01-07" collectingDateMin="1996-12-12" collectionCode="JJRC" collectorName="NW Madagascar &amp; J. Rolcik &amp; A. Shamaev" country="Madagascar" county="Morondava distr." elevation="100" location="Mahajamba" municipality="Kirindi Forest" pageId="13" pageNumber="164" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Mahajanga">
<emphasis id="B9003467FFF8FFD1FF2AFD5AFCF0EFB4" bold="true" box="[199,780,701,726]" pageId="13" pageNumber="164">
Additional material examined.
<collectingCountry id="F363A8E5FFF8FFD1FDD2FD5AFCF0EFB4" box="[575,780,701,726]" name="Madagascar" pageId="13" pageNumber="164">MADAGASCAR</collectingCountry>
</emphasis>
:
<specimenCount id="9D7223FCFFF8FFD1FCF1FD59FCB7EFB7" box="[796,843,702,726]" pageId="13" pageNumber="164" type="male">2 ♂</specimenCount>
:
<collectorName id="26818DA3FFF8FFD1FCB5FD59FBE5EFB4" box="[856,1049,701,726]" pageId="13" pageNumber="164">NW Madagascar</collectorName>
,
<collectingCounty id="62AA90F9FFF8FFD1FBC8FD5AFB15EFB4" box="[1061,1257,701,726]" pageId="13" pageNumber="164">Morondava distr.</collectingCounty>
,
<collectingMunicipality id="6BAF720FFFF8FFD1FB1BFD5AFA64EFB7" box="[1270,1432,701,726]" pageId="13" pageNumber="164">Kirindi Forest</collectingMunicipality>
, 4.
<collectingDate id="EF8E375DFFF8FFD1FF5AFD05FEE3EF98" box="[183,287,738,762]" pageId="13" pageNumber="164" value="2002-01-08">
<date id="FFCACEB5FFF8FFD1FF5AFD05FEE3EF98" box="[183,287,738,762]" pageId="13" pageNumber="164" value="2002-01-08">8.1.2002</date>
</collectingDate>
,
<collectorName id="26818DA3FFF8FFD1FEC0FD05FE68EF98" box="[301,404,737,762]" pageId="13" pageNumber="164">J. Rolčík</collectorName>
lgt. (
<collectionCode id="ED6570B0FFF8FFD1FE38FD05FDEAEF98" box="[469,534,738,762]" pageId="13" pageNumber="164">JJRC</collectionCode>
);
<specimenCount id="9D7223FCFFF8FFD1FDC1FD05FDA7EF9B" box="[556,603,738,762]" pageId="13" pageNumber="164" type="female">1 ♀</specimenCount>
:
<collectingRegion id="49B02697FFF8FFD1FD86FD06FD14EF98" box="[619,744,737,762]" country="Madagascar" name="Mahajanga" pageId="13" pageNumber="164">Mahajanga</collectingRegion>
prov.,
<location id="8EABBEAEFFF8FFD1FCD1FD06FC3CEF98" LSID="urn:lsid:plazi:treatment:03DD5963FFF8FFD1FF7AFE86FB90EAFC:8EABBEAEFFF8FFD1FCD1FD06FC3CEF98" box="[828,960,737,762]" country="Madagascar" county="Morondava distr." municipality="Kirindi Forest" name="Mahajamba" pageId="13" pageNumber="164" stateProvince="Mahajanga">Mahajamba</location>
riv.,
<location id="8EABBEAEFFF8FFD1FC12FD06FB91EF98" LSID="urn:lsid:plazi:treatment:03DD5963FFF8FFD1FF7AFE86FB90EAFC:8EABBEAEFFF8FFD1FC12FD06FB91EF98" box="[1023,1133,737,762]" country="Madagascar" county="Morondava distr." municipality="Kirindi Forest" name="Ampatika" pageId="13" pageNumber="164" stateProvince="Mahajanga">Ampatika</location>
env., 10.
<collectingDate id="EF8E375DFFF8FFD1FB09FD06FA95EF98" box="[1252,1385,737,762]" pageId="13" pageNumber="164" value="1996-12-12">
<date id="FFCACEB5FFF8FFD1FB09FD06FA95EF98" box="[1252,1385,737,762]" pageId="13" pageNumber="164" value="1996-12-12">12.12.1996</date>
</collectingDate>
,
<location id="8EABBEAEFFF8FFD1FA9AFD05FF33EE7C" LSID="urn:lsid:plazi:treatment:03DD5963FFF8FFD1FF7AFE86FB90EAFC:8EABBEAEFFF8FFD1FA9AFD05FF33EE7C" country="Madagascar" county="Morondava distr." municipality="Kirindi Forest" name="Ivo Jenis" pageId="13" pageNumber="164" stateProvince="Mahajanga">Ivo Jeniš</location>
lgt. (
<collectionCode id="ED6570B0FFF8FFD1FEE4FCE1FEB6EE7C" box="[265,330,774,798]" pageId="13" pageNumber="164">JJRC</collectionCode>
);
<specimenCount id="9D7223FCFFF8FFD1FEB3FCE1FE77EE7F" box="[350,395,774,798]" pageId="13" pageNumber="164" type="male">1 ♂</specimenCount>
:
<locationDeviation id="9A79F157FFF8FFD1FE7AFCE2FD78EE7C" box="[407,644,773,798]" pageId="13" pageNumber="164">
<quantity id="4C8C4590FFF8FFD1FE7AFCE2FE22EE7C" box="[407,478,773,798]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="164" unit="km" value="60.0">60 km</quantity>
N Morondava
</locationDeviation>
,
<location id="8EABBEAEFFF8FFD1FD62FCE2FD1FEE7C" LSID="urn:lsid:plazi:treatment:03DD5963FFF8FFD1FF7AFE86FB90EAFC:8EABBEAEFFF8FFD1FD62FCE2FD1FEE7C" box="[655,739,773,798]" country="Madagascar" county="Morondava distr." municipality="Kirindi Forest" name="Kirindi" pageId="13" pageNumber="164" stateProvince="Mahajanga">Kirindi</location>
,
<elevation id="00590F46FFF8FFD1FD03FCE1FCCBEE7F" box="[750,823,774,798]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="164" unit="m" value="100.0">
<quantity id="4C8C4590FFF8FFD1FD03FCE1FCCBEE7F" box="[750,823,774,798]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="164" unit="m" value="100.0">100 m</quantity>
</elevation>
, 5.
<collectingDate id="EF8E375DFFF8FFD1FC8CFCE2FC36EE7C" box="[865,970,773,798]" pageId="13" pageNumber="164" value="2004-01-07">
<date id="FFCACEB5FFF8FFD1FC8CFCE2FC36EE7C" box="[865,970,773,798]" pageId="13" pageNumber="164" value="2004-01-07">7.1.2004</date>
</collectingDate>
, S. Murzin &amp;
<collectorName id="26818DA3FFF8FFD1FB84FCE1FB0EEE7C" box="[1129,1266,773,798]" pageId="13" pageNumber="164">A. Shamaev</collectorName>
leg. (
<collectionCode id="ED6570B0FFF8FFD1FADFFCE1FA8FEE7C" box="[1330,1395,774,798]" pageId="13" pageNumber="164">JJRC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36EBBFEFFF8FFD1FF2AFCCEFC57EBC0" pageId="13" pageNumber="164" type="description">
<paragraph id="8BCBE875FFF8FFD1FF2AFCCEFC71EECC" blockId="13.[151,1437,557,1950]" pageId="13" pageNumber="164">
<emphasis id="B9003467FFF8FFD1FF2AFCCEFE3CEE20" bold="true" box="[199,448,809,834]" pageId="13" pageNumber="164">Redescription (n=5).</emphasis>
Body length
<quantity id="4C8C4590FFF8FFD1FD85FCCEFD08EE23" box="[616,756,809,834]" metricMagnitude="-3" metricUnit="m" metricValue="7.199999999999999" metricValueMax="7.6" metricValueMin="6.8" pageId="13" pageNumber="164" unit="mm" value="7.199999999999999" valueMax="7.6" valueMin="6.8">6.87.6 mm</quantity>
(M=
<quantity id="4C8C4590FFF8FFD1FCDCFCCDFC73EE23" box="[817,911,810,834]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="13" pageNumber="164" unit="mm" value="7.2">7.2 mm</quantity>
, HT=
<quantity id="4C8C4590FFF8FFD1FC38FCCDFBC9EE20" box="[981,1077,810,834]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="13" pageNumber="164" unit="mm" value="7.2">7.2 mm</quantity>
), forebody length 3.0
<quantity id="4C8C4590FFF8FFD1FAAFFCCDFA60EE23" box="[1346,1436,810,834]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="164" unit="mm" value="3.2">3.2 mm</quantity>
(M=
<quantity id="4C8C4590FFF8FFD1FF24FCA9FEDAEE07" box="[201,294,846,870]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="13" pageNumber="164" unit="mm" value="3.1">3.1 mm</quantity>
, HT=
<quantity id="4C8C4590FFF8FFD1FE81FCA9FE30EE04" box="[364,460,846,870]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="164" unit="mm" value="3.2">3.2 mm</quantity>
). Body (
<figureCitation id="134FF4F0FFF8FFD1FDD8FCAAFD77EE04" box="[565,651,845,870]" captionStart="FIGURES 5260" captionStartId="14.[151,264,1907,1929]" captionTargetBox="[160,1423,212,1874]" captionTargetId="figure@14.[151,1435,193,1885]" captionTargetPageId="14" captionText="FIGURES 5260. Acylophorus delphinus Fauvel. 52, 53, 5660, Kirindi; 54, 55, Ampatika. 60, holotype. 52, habitus; 53, head and pronotum; 54, mandibles; 55, maxillary palpus; 56, aedeagus lateral; 57, aedeagus ventro-lateral; 5860, aedeagus ventral. 5660, same scale bars." httpUri="https://zenodo.org/record/1441473/files/figure.png" pageId="13" pageNumber="164">Fig. 52</figureCitation>
) brown, apical margins of abdominal segment reddish, abdominal tergites markedly iridescent; antennae brown with base of first two segments and segment 11 light reddish; legs reddish yellow, maxillary palpi reddish yellow, last segment brown.
</paragraph>
<paragraph id="8BCBE875FFF8FFD1FF2AFC5EFC6CE990" blockId="13.[151,1437,557,1950]" pageId="13" pageNumber="164">
Head subrounded (
<figureCitation id="134FF4F0FFF8FFD1FE49FC5EFE07EEB0" box="[420,507,953,978]" captionStart="FIGURES 5260" captionStartId="14.[151,264,1907,1929]" captionTargetBox="[160,1423,212,1874]" captionTargetId="figure@14.[151,1435,193,1885]" captionTargetPageId="14" captionText="FIGURES 5260. Acylophorus delphinus Fauvel. 52, 53, 5660, Kirindi; 54, 55, Ampatika. 60, holotype. 52, habitus; 53, head and pronotum; 54, mandibles; 55, maxillary palpus; 56, aedeagus lateral; 57, aedeagus ventro-lateral; 5860, aedeagus ventral. 5660, same scale bars." httpUri="https://zenodo.org/record/1441473/files/figure.png" pageId="13" pageNumber="164">Fig. 53</figureCitation>
), about as long as wide (L/W=0.971.04, M=1.00, HT=1.01), slightly widened behind eyes, with two pairs of interocular setae. Temples markedly shorter than eyes (LT/LE=0.630.69, M=0.65, HT=0.67). Dorsal side of head shining, without micropunctures. Dense pale grey pubescence behind eyes. Four postocular setae visible on each side, additional seta on hind margin of eye absent. Left mandible with two sharp teeth, right mandible with three sharp teeth (
<figureCitation id="134FF4F0FFF8FFD1FD46FBAEFCF9E900" box="[683,773,1097,1122]" captionStart="FIGURES 5260" captionStartId="14.[151,264,1907,1929]" captionTargetBox="[160,1423,212,1874]" captionTargetId="figure@14.[151,1435,193,1885]" captionTargetPageId="14" captionText="FIGURES 5260. Acylophorus delphinus Fauvel. 52, 53, 5660, Kirindi; 54, 55, Ampatika. 60, holotype. 52, habitus; 53, head and pronotum; 54, mandibles; 55, maxillary palpus; 56, aedeagus lateral; 57, aedeagus ventro-lateral; 5860, aedeagus ventral. 5660, same scale bars." httpUri="https://zenodo.org/record/1441473/files/figure.png" pageId="13" pageNumber="164">Fig. 54</figureCitation>
). Maxillary palpi with terminal segment wide and long, pubescent, asymmetric, markedly wider and much longer than glabrous penultimate segment which is triangular (
<figureCitation id="134FF4F0FFF8FFD1FF72FB76FF0AE9C8" box="[159,246,1169,1194]" captionStart="FIGURES 5260" captionStartId="14.[151,264,1907,1929]" captionTargetBox="[160,1423,212,1874]" captionTargetId="figure@14.[151,1435,193,1885]" captionTargetPageId="14" captionText="FIGURES 5260. Acylophorus delphinus Fauvel. 52, 53, 5660, Kirindi; 54, 55, Ampatika. 60, holotype. 52, habitus; 53, head and pronotum; 54, mandibles; 55, maxillary palpus; 56, aedeagus lateral; 57, aedeagus ventro-lateral; 5860, aedeagus ventral. 5660, same scale bars." httpUri="https://zenodo.org/record/1441473/files/figure.png" pageId="13" pageNumber="164">Fig. 55</figureCitation>
). First segment of antenna nearly as long as next four; segments 1 to 7 elongate, segment 5 markedly elongate (R=1.581.69, M=1.62, HT=1.69), segment 8 about as long as wide, segment 9 slightly transverse, segment 10 markedly transverse (R=0.680.82, M=0.75, HT=0.82).
</paragraph>
<paragraph id="8BCBE875FFF8FFD1FF2AFB19FC4CE8A8" blockId="13.[151,1437,557,1950]" pageId="13" pageNumber="164">
Pronotum markedly transverse (R=1.141.21, M=1.16, HT=1.12), markedly widened behind with sides slightly rounded, widest in basal half (
<figureCitation id="134FF4F0FFF8FFD1FDA4FAC6FD61E858" box="[585,669,1313,1338]" captionStart="FIGURES 5260" captionStartId="14.[151,264,1907,1929]" captionTargetBox="[160,1423,212,1874]" captionTargetId="figure@14.[151,1435,193,1885]" captionTargetPageId="14" captionText="FIGURES 5260. Acylophorus delphinus Fauvel. 52, 53, 5660, Kirindi; 54, 55, Ampatika. 60, holotype. 52, habitus; 53, head and pronotum; 54, mandibles; 55, maxillary palpus; 56, aedeagus lateral; 57, aedeagus ventro-lateral; 5860, aedeagus ventral. 5660, same scale bars." httpUri="https://zenodo.org/record/1441473/files/figure.png" pageId="13" pageNumber="164">Fig. 53</figureCitation>
), shining, without micropunctures. One pair of dorsal setae and one pair of lateral setae. Marginal setae short. Elytra transverse (R=1.431.52, M=1.47, HT=1.52) with short, pale, but not shining pubescence arising from dense aspirate punctures. Apical fringe of bristles slightly longer than pubescence on other parts of the elytra. Abdominal tergites with relativelly short, moderately dense pubescence, arising from aspirate punctures more coarse and sparser than on elytra.
</paragraph>
<paragraph id="8BCBE875FFF8FFD1FF2AFA31FB7AE88C" blockId="13.[151,1437,557,1950]" box="[199,1158,1493,1518]" pageId="13" pageNumber="164">
Meso- and metatarsi with empodial setae longer than claws (similar as in
<figureCitation id="134FF4F0FFF8FFD1FC12FA32FB84E88C" box="[1023,1144,1493,1518]" captionStart="FIGURES 917" captionStartId="4.[151,264,1929,1951]" captionTargetBox="[153,1433,215,1902]" captionTargetId="figure@4.[151,1435,210,1902]" captionTargetPageId="4" captionText="FIGURES 917. Acylophorus tristis Jarrige. 9, 10, 1517, Col de Marotaolana; 1113, Sendrisoa; 14, Manindray. 9, metatarsus lateral; 10, metatarsus dorsal; 11, 16, aedeagus lateral; 12, aedeagus ventro-lateral; 1315, aedeagus ventral. ES, empodial setae. 16, 17, transmitted light. 910, 1117, same scale bars." httpUri="https://zenodo.org/record/1441461/files/figure.png" pageId="13" pageNumber="164">Figs. 9, 10</figureCitation>
).
</paragraph>
<paragraph id="8BCBE875FFF8FFD1FF2AFA1EFB61EB1C" blockId="13.[151,1437,557,1950]" pageId="13" pageNumber="164">
Male (n=5). Sternite 9 long and narrow (
<figureCitation id="134FF4F0FFF8FFD1FD7AFA1EFD10EB70" box="[663,748,1529,1554]" captionStart="FIGURES 6166" captionStartId="15.[151,264,1933,1955]" captionTargetBox="[179,1374,273,1811]" captionTargetId="figure@15.[152,1435,240,1854]" captionTargetPageId="15" captionText="FIGURES 6166. Acylophorus delphinus Fauvel. 6164, Kirindy; 65, 66, Ampatika. 61, aedeagus lateral; 62, aedeagus ventral; 63, male tergites 9, 10; 64, male sternite 9; 65, female tergites 9, 10; 66, valves. 61, 62, transmitted light. 6162, 6566, same scale bars." httpUri="https://zenodo.org/record/1441475/files/figure.png" pageId="13" pageNumber="164">Fig. 64</figureCitation>
), apex of tergite 10 narrowly rounded, with very long apical setae (
<figureCitation id="134FF4F0FFF8FFD1FF0DF9FAFEC8EB54" box="[224,308,1565,1590]" captionStart="FIGURES 6166" captionStartId="15.[151,264,1933,1955]" captionTargetBox="[179,1374,273,1811]" captionTargetId="figure@15.[152,1435,240,1854]" captionTargetPageId="15" captionText="FIGURES 6166. Acylophorus delphinus Fauvel. 6164, Kirindy; 65, 66, Ampatika. 61, aedeagus lateral; 62, aedeagus ventral; 63, male tergites 9, 10; 64, male sternite 9; 65, female tergites 9, 10; 66, valves. 61, 62, transmitted light. 6162, 6566, same scale bars." httpUri="https://zenodo.org/record/1441475/files/figure.png" pageId="13" pageNumber="164">Fig. 63</figureCitation>
). Aedeagus (
<figureCitation id="134FF4F0FFF8FFD1FE24F9FAFDADEB54" box="[457,593,1565,1590]" captionStart-0="FIGURES 5260" captionStart-1="FIGURES 6166" captionStartId-0="14.[151,264,1907,1929]" captionStartId-1="15.[151,264,1933,1955]" captionTargetBox-0="[160,1423,212,1874]" captionTargetBox-1="[179,1374,273,1811]" captionTargetId-0="figure@14.[151,1435,193,1885]" captionTargetId-1="figure@15.[152,1435,240,1854]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 5260. Acylophorus delphinus Fauvel. 52, 53, 5660, Kirindi; 54, 55, Ampatika. 60, holotype. 52, habitus; 53, head and pronotum; 54, mandibles; 55, maxillary palpus; 56, aedeagus lateral; 57, aedeagus ventro-lateral; 5860, aedeagus ventral. 5660, same scale bars." captionText-1="FIGURES 6166. Acylophorus delphinus Fauvel. 6164, Kirindy; 65, 66, Ampatika. 61, aedeagus lateral; 62, aedeagus ventral; 63, male tergites 9, 10; 64, male sternite 9; 65, female tergites 9, 10; 66, valves. 61, 62, transmitted light. 6162, 6566, same scale bars." httpUri-0="https://zenodo.org/record/1441473/files/figure.png" httpUri-1="https://zenodo.org/record/1441475/files/figure.png" pageId="13" pageNumber="164">Figs. 5662</figureCitation>
)
<quantity id="4C8C4590FFF8FFD1FD8FF9F9FCFAEB57" box="[610,774,1566,1590]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" metricValueMax="8.7" metricValueMin="8.1" pageId="13" pageNumber="164" unit="mm" value="0.84" valueMax="0.87" valueMin="0.81">0.810.87 mm</quantity>
long (M =
<quantity id="4C8C4590FFF8FFD1FC61F9F9FC0FEB57" box="[908,1011,1566,1590]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="13" pageNumber="164" unit="mm" value="0.84">0.84 mm</quantity>
, HT =
<quantity id="4C8C4590FFF8FFD1FBABF9F9FB4CEB54" box="[1094,1200,1566,1590]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="13" pageNumber="164" unit="mm" value="0.87">0.87 mm</quantity>
). Parameres bilobed, lobes widely separated, apically slightly widened, sensory pegs confusedly arranged at the very apex (
<figureCitation id="134FF4F0FFF8FFD1FAD4F9A6FA6CEB38" box="[1337,1424,1601,1626]" captionStart="FIGURES 6166" captionStartId="15.[151,264,1933,1955]" captionTargetBox="[179,1374,273,1811]" captionTargetId="figure@15.[152,1435,240,1854]" captionTargetPageId="15" captionText="FIGURES 6166. Acylophorus delphinus Fauvel. 6164, Kirindy; 65, 66, Ampatika. 61, aedeagus lateral; 62, aedeagus ventral; 63, male tergites 9, 10; 64, male sternite 9; 65, female tergites 9, 10; 66, valves. 61, 62, transmitted light. 6162, 6566, same scale bars." httpUri="https://zenodo.org/record/1441475/files/figure.png" pageId="13" pageNumber="164">Fig. 62</figureCitation>
). Median lobe of aedeagus markedly overreaching paramere, apex rounded (
<figureCitation id="134FF4F0FFF8FFD1FC33F982FB99EB1C" box="[990,1125,1637,1662]" captionStart="FIGURES 5260" captionStartId="14.[151,264,1907,1929]" captionTargetBox="[160,1423,212,1874]" captionTargetId="figure@14.[151,1435,193,1885]" captionTargetPageId="14" captionText="FIGURES 5260. Acylophorus delphinus Fauvel. 52, 53, 5660, Kirindi; 54, 55, Ampatika. 60, holotype. 52, habitus; 53, head and pronotum; 54, mandibles; 55, maxillary palpus; 56, aedeagus lateral; 57, aedeagus ventro-lateral; 5860, aedeagus ventral. 5660, same scale bars." httpUri="https://zenodo.org/record/1441473/files/figure.png" pageId="13" pageNumber="164">Figs. 5860</figureCitation>
,
<figureCitation id="134FF4F0FFF8FFD1FB9CF982FB6DEB1C" box="[1137,1169,1637,1662]" captionStart="FIGURES 6166" captionStartId="15.[151,264,1933,1955]" captionTargetBox="[179,1374,273,1811]" captionTargetId="figure@15.[152,1435,240,1854]" captionTargetPageId="15" captionText="FIGURES 6166. Acylophorus delphinus Fauvel. 6164, Kirindy; 65, 66, Ampatika. 61, aedeagus lateral; 62, aedeagus ventral; 63, male tergites 9, 10; 64, male sternite 9; 65, female tergites 9, 10; 66, valves. 61, 62, transmitted light. 6162, 6566, same scale bars." httpUri="https://zenodo.org/record/1441475/files/figure.png" pageId="13" pageNumber="164">62</figureCitation>
).
</paragraph>
<paragraph id="8BCBE875FFF8FFD1FF2AF96EFC57EBC0" blockId="13.[151,1437,557,1950]" box="[199,939,1673,1698]" pageId="13" pageNumber="164">
Female. Tergite 10 pointed apically (
<figureCitation id="134FF4F0FFF8FFD1FD8EF96EFD4AEBC0" box="[611,694,1673,1698]" captionStart="FIGURES 6166" captionStartId="15.[151,264,1933,1955]" captionTargetBox="[179,1374,273,1811]" captionTargetId="figure@15.[152,1435,240,1854]" captionTargetPageId="15" captionText="FIGURES 6166. Acylophorus delphinus Fauvel. 6164, Kirindy; 65, 66, Ampatika. 61, aedeagus lateral; 62, aedeagus ventral; 63, male tergites 9, 10; 64, male sternite 9; 65, female tergites 9, 10; 66, valves. 61, 62, transmitted light. 6162, 6566, same scale bars." httpUri="https://zenodo.org/record/1441475/files/figure.png" pageId="13" pageNumber="164">Fig. 65</figureCitation>
), valves as in
<figureCitation id="134FF4F0FFF8FFD1FCB9F96EFC5AEBC0" box="[852,934,1673,1698]" captionStart="FIGURES 6166" captionStartId="15.[151,264,1933,1955]" captionTargetBox="[179,1374,273,1811]" captionTargetId="figure@15.[152,1435,240,1854]" captionTargetPageId="15" captionText="FIGURES 6166. Acylophorus delphinus Fauvel. 6164, Kirindy; 65, 66, Ampatika. 61, aedeagus lateral; 62, aedeagus ventral; 63, male tergites 9, 10; 64, male sternite 9; 65, female tergites 9, 10; 66, valves. 61, 62, transmitted light. 6162, 6566, same scale bars." httpUri="https://zenodo.org/record/1441475/files/figure.png" pageId="13" pageNumber="164">Fig. 66</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36EBBFEFFF8FFD1FF2AF94AFCC0EA6C" pageId="13" pageNumber="164" type="diagnosis">
<paragraph id="8BCBE875FFF8FFD1FF2AF94AFCC0EA6C" blockId="13.[151,1437,557,1950]" pageId="13" pageNumber="164">
<emphasis id="B9003467FFF8FFD1FF2AF94AFE31EBA4" bold="true" box="[199,461,1709,1734]" pageId="13" pageNumber="164">Differential diagnosis.</emphasis>
<taxonomicName id="4C7493F6FFF8FFD1FE3EF94AFD28EBA7" box="[467,724,1709,1733]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="delphinus">
<emphasis id="B9003467FFF8FFD1FE3EF94AFD28EBA7" box="[467,724,1709,1733]" italics="true" pageId="13" pageNumber="164">Acylophorus delphinus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="4C7493F6FFF8FFD1FC89F949FB7BEBA4" authority="Fauvel, 1907" authorityName="Fauvel" authorityYear="1907" box="[868,1159,1709,1734]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B9003467FFF8FFD1FC89F949FC13EBA7" box="[868,1007,1709,1733]" italics="true" pageId="13" pageNumber="164">A. orientalis</emphasis>
Fauvel, 1907
</taxonomicName>
from continental Africa, but can be distinguished by the pale terminal segment of the maxillary palpi, the longer additional postocular seta on the hind margin of the eye and the shape of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C36EBBFEFFF8FFD1FF2AF8FEFC1DEA34" pageId="13" pageNumber="164" type="distribution">
<paragraph id="8BCBE875FFF8FFD1FF2AF8FEFB2BEA50" blockId="13.[151,1437,557,1950]" box="[199,1239,1817,1842]" pageId="13" pageNumber="164">
<emphasis id="B9003467FFF8FFD1FF2AF8FEFE9CEA50" bold="true" box="[199,352,1817,1842]" pageId="13" pageNumber="164">Distribution.</emphasis>
The species is known only from South and South-west
<collectingCountry id="F363A8E5FFF8FFD1FC3CF8FEFBA4EA50" box="[977,1112,1817,1842]" name="Madagascar" pageId="13" pageNumber="164">Madagascar</collectingCountry>
(
<figureCitation id="134FF4F0FFF8FFD1FB85F8FEFB36EA50" box="[1128,1226,1817,1842]" captionStart="FIGURES 172177" captionStartId="38.[151,264,1927,1949]" captionTargetBox="[265,1321,625,1895]" captionTargetId="figure@38.[265,1321,625,1909]" captionTargetPageId="38" captionText="FIGURES 172177. Distribution maps. 172, Acylophorus tristis Jarrige; 173, A. derougemonti sp. nov.; 174, A. muscorum Jarrige; 175, A. delphinus Fauvel (D), A. madecassus sp. nov. (Ŧ); 176, A. minutus Jarrige (D), A. hova sp. nov. (Ɨ), A. montanus sp. nov. (Δ); 177, A. sogai Jarrige." httpUri="https://zenodo.org/record/1441507/files/figure.png" pageId="13" pageNumber="164">Fig. 175</figureCitation>
).
</paragraph>
<paragraph id="8BCBE875FFF8FFD1FF2AF8DAFC1DEA34" blockId="13.[151,1437,557,1950]" box="[199,993,1853,1878]" pageId="13" pageNumber="164">
<emphasis id="B9003467FFF8FFD1FF2AF8DAFEB6EA34" bold="true" box="[199,330,1853,1878]" pageId="13" pageNumber="164">Bionomics.</emphasis>
Nothing is known about the bionomics of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C36EBBFEFFF8FFD1FF2AF886FB90EAFC" pageId="13" pageNumber="164" type="discussion">
<paragraph id="8BCBE875FFF8FFD1FF2AF886FB90EAFC" blockId="13.[151,1437,557,1950]" pageId="13" pageNumber="164">
<emphasis id="B9003467FFF8FFD1FF2AF886FEB0EA18" bold="true" box="[199,332,1889,1914]" pageId="13" pageNumber="164">Discussion.</emphasis>
The unique male
<typeStatus id="54CF56D7FFF8FFD1FDC9F885FDA8EA18" box="[548,596,1890,1914]" pageId="13" pageNumber="164">type</typeStatus>
from Ambovombe was studied by
<bibRefCitation id="EFE59584FFF8FFD1FC18F886FF10EAFC" pageId="13" pageNumber="164" refId="ref17650" refString="Jarrige, J. (1965) Les Acylophorus malgaches (Col. Staphylinidae). Bullettin de la Societ entomologique de France, 70, 139 - 143." type="journal article">Jarrige (1965, wrongly reported as a female)</bibRefCitation>
and
<bibRefCitation id="EFE59584FFF8FFD1FECEF861FE5AEAFC" box="[291,422,1926,1950]" pageId="13" pageNumber="164" refId="ref17680" refString="Lott, D. A. (2010) The species of Acylophorus Nordmann (Coleoptera: Staphylinidae: Staphylininae) in continental sub- Saharan Africa. Zootaxa, 2401, 1 - 51." type="journal article">Lott (2010)</bibRefCitation>
. Additional material is from south-western part of
<collectingCountry id="F363A8E5FFF8FFD1FC33F862FB94EAFC" box="[990,1128,1925,1950]" name="Madagascar" pageId="13" pageNumber="164">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>