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<document id="D3D1C97526497242B5736BBF59B77914" ID-DOI="10.5281/zenodo.179899" ID-GBIF-Dataset="4ae64d2e-fea3-4fbc-9cd8-538a7396e89b" ID-ISSN="1175-5326" ID-Zenodo-Dep="179899" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460010654862" checkinUser="plazi" docAuthor="Means, Bruce &amp; Savage, Jay M." docDate="2007" docId="03AA879CFF854435C1861DCB3B6DE877" docLanguage="en" docName="zt01658p055.pdf" docOrigin="Zootaxa 1658" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Pristimantis saltissimus Means &amp; Savage, 2007, n. sp." docType="treatment" docVersion="10" lastPageNumber="53" masterDocId="FF93FFE4FF8C443BC1111B583E14E90D" masterDocTitle="Three New Malodorous Rainfrogs of the Genus Pristimantis (Anura: Brachycephalidae) from the Wokomung Massif in west-central Guyana, South America" masterLastPageNumber="55" masterPageNumber="39" pageNumber="48" updateTime="1698233360999" updateUser="plazi">
<mods:mods id="1D126848ED78E010EF410B02AEB2C7CF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="76BF39A20CD715CACD684ABDF07F2076">
<mods:title id="6A3619E233C6F3F91D199AA35D08EDE1">Three New Malodorous Rainfrogs of the Genus Pristimantis (Anura: Brachycephalidae) from the Wokomung Massif in west-central Guyana, South America</mods:title>
</mods:titleInfo>
<mods:name id="11BA9221AE151266E6DE73D739632128" type="personal">
<mods:role id="F27B5BC2700D24A0AF8CC17365C1F455">
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</mods:role>
<mods:namePart id="AEAFED0448566090DE2D76B9B0423364">Means, Bruce</mods:namePart>
</mods:name>
<mods:name id="961808511F38D098DBF83720B228B09A" type="personal">
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</mods:role>
<mods:namePart id="95847792D8F5BD1705E767A59A8D5B5C">Savage, Jay M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2EF25CCF5FB164775A557408162924F1">text</mods:typeOfResource>
<mods:relatedItem id="DA608B0412905776423EF6A2B46409D8" type="host">
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<mods:title id="28B7A61CD4D9A377DA0BF4ED6D85DD60">Zootaxa</mods:title>
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<mods:part id="6B9B0AE801AAAE51EADFFAA9BF9FBCC7">
<mods:date id="7F3821C2436506B31D66FF0BBED24931">2007</mods:date>
<mods:detail id="9A06070369C554CAE0221251BA99C1B8" type="volume">
<mods:number id="829145CB0F883C16F6E4A9AFE5FFF37D">1658</mods:number>
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<mods:classification id="63FAE4B16FC67382B144680A1F9347FC">journal article</mods:classification>
<mods:identifier id="D8ADF41A0B9562E9693596DDB418A19D" type="DOI">10.5281/zenodo.179899</mods:identifier>
<mods:identifier id="B401C7D975F87F0162C9B11257BD3DC1" type="GBIF-Dataset">4ae64d2e-fea3-4fbc-9cd8-538a7396e89b</mods:identifier>
<mods:identifier id="E7FD259F78E40BD5D3F71970113132CB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FFD4431EDB7CA90CFD7A5D981525AC20" type="Zenodo-Dep">179899</mods:identifier>
</mods:mods>
<treatment id="03AA879CFF854435C1861DCB3B6DE877" ID-DOI="http://doi.org/10.5281/zenodo.5681046" ID-GBIF-Taxon="119381326" ID-Zenodo-Dep="5681046" LSID="urn:lsid:plazi:treatment:03AA879CFF854435C1861DCB3B6DE877" httpUri="http://treatment.plazi.org/id/03AA879CFF854435C1861DCB3B6DE877" lastPageId="14" lastPageNumber="53" pageId="9" pageNumber="48">
<subSubSection id="C3196501FF854432C1861DCB3F20EFDD" pageId="9" pageNumber="48" type="nomenclature">
<paragraph id="8BBC368AFF854432C1861DCB3FEAEFA0" blockId="9.[151,510,1683,1784]" box="[151,510,1683,1709]" pageId="9" pageNumber="48">
<heading id="D0F481E6FF854432C1861DCB3FEAEFA0" bold="true" box="[151,510,1683,1709]" fontSize="11" level="1" pageId="9" pageNumber="48" reason="1">
<emphasis id="B977EA98FF854432C1861DCB3FEAEFA0" bold="true" box="[151,510,1683,1709]" pageId="9" pageNumber="48">
<taxonomicName id="4C034D09FF854432C1861DCB3FA1EFA0" box="[151,437,1683,1709]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="48" phylum="Chordata" rank="species" species="saltissimus" status="sp. nov.">
<emphasis id="B977EA98FF854432C1861DCB3FA1EFA0" bold="true" box="[151,437,1683,1709]" italics="true" pageId="9" pageNumber="48">Pristimantis saltissimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457E3FF854432C0AD1DCB3FEAEFA0" box="[444,510,1683,1709]" pageId="9" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBC368AFF854432C1861DEE3F20EFDD" blockId="9.[151,510,1683,1784]" box="[151,308,1718,1744]" pageId="9" pageNumber="48">
(
<figureCitation id="13382A0FFF854432C18E1DEE3EE6EFDD" box="[159,242,1718,1744]" captionStart="FIGURE 3" captionStartId="5.[151,255,1305,1329]" captionTargetBox="[388,1200,208,1304]" captionTargetId="figure@5.[388,1200,194,1351]" captionTargetPageId="5" captionText="FIGURE 3. Map of Guyana with the location of the Wokomung Massif in the Pakaraima Mountains (gray shading) and in relation to some other teuis in Guyana. Map modified from one supplied by Philippe J. R. Kok." httpUri="https://zenodo.org/record/179902/files/figure.png" pageId="9" pageNumber="48">Figs. 3</figureCitation>
,
<figureCitation id="13382A0FFF854432C1EE1DEE3F1BEFDD" box="[255,271,1718,1744]" captionStart="FIGURE 6" captionStartId="11.[151,255,1578,1602]" captionTargetBox="[153,1433,274,1554]" captionTargetId="figure@11.[153,1433,274,1554]" captionTargetPageId="11" captionText="FIGURE 6. Pristimantis saltissimus new species. a = plain or slightly spotted morph (43 % in a sample of 30); b = middorsally orange-striped morph (10 %); c = dorsally tan morph (17 %); d = thin, white-striped morph (10 %); e = dorsal views of the mottled morph (20 %) and thin, white-striped morph; f = ventral views of same morphs as in e. Note that the striping also occurs on the belly of the thin, white-striped morph." httpUri="https://zenodo.org/record/179905/files/figure.png" pageId="9" pageNumber="48">6</figureCitation>
,
<figureCitation id="13382A0FFF854432C00D1DEE3F3FEFDD" box="[284,299,1718,1744]" captionStart="FIGURE 7" captionStartId="13.[151,255,1402,1426]" captionTargetBox="[172,1393,204,1378]" captionTargetId="figure@13.[155,1432,194,1378]" captionTargetPageId="13" captionText="FIGURE 7. Pristimantis saltissimus new species. Hand (left) and foot (right). USNM 563639 holotype. Black bars = 5.0 mm." httpUri="https://zenodo.org/record/274028/files/figure.png" pageId="9" pageNumber="48">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3196501FF854432C1861D863F4DEFF5" box="[151,345,1758,1784]" pageId="9" pageNumber="48" type="description">
<paragraph id="8BBC368AFF854432C1861D863F4DEFF5" blockId="9.[151,510,1683,1784]" box="[151,345,1758,1784]" pageId="9" pageNumber="48">Rocket Rainfrog</paragraph>
</subSubSection>
<subSubSection id="C3196501FF854431C1861C753D7BE827" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48" type="materials_examined">
<paragraph id="8BBC368AFF854432C1861C753CC0EE95" blockId="9.[151,1437,1837,2024]" pageId="9" pageNumber="48">
<emphasis id="B977EA98FF854432C1861C753F1DEE4A" bold="true" box="[151,265,1837,1863]" pageId="9" pageNumber="48">
<typeStatus id="54B88828FF854432C1861C753F1DEE4A" box="[151,265,1837,1863]" pageId="9" pageNumber="48" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectionCode id="ED12AE4FFF854432C00B1C763F7BEE45" LSID="urn:lsid:biocol.org:col:34871" box="[282,367,1838,1864]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="48">USNM</collectionCode>
563639, an adult female from the Wokomung Massif, near Falls Camp, Potaro-Siparuni District, west-central
<collectingCountry id="F314761AFF854432C0851C0E3FE7EE7D" box="[404,499,1878,1904]" name="Guyana" pageId="9" pageNumber="48">Guyana</collectingCountry>
;
<geoCoordinate id="EE37504DFF854432C0EE1C0E3CBCEE7D" box="[511,680,1878,1904]" direction="north" orientation="latitude" pageId="9" pageNumber="48" precision="15" value="5.090278">05° 05 25” N</geoCoordinate>
,
<geoCoordinate id="EE37504DFF854432C3A51C0E3D76EE7D" box="[692,866,1878,1904]" direction="west" orientation="longitude" pageId="9" pageNumber="48" precision="15" value="-59.838333">59° 50 18” W</geoCoordinate>
,
<quantity id="4CFB9B6FFF854432C27C1C0E3DDDEE7D" box="[877,969,1878,1904]" metricMagnitude="3" metricUnit="m" metricValue="1.385" pageId="9" pageNumber="48" unit="m" value="1385.0">1385 m</quantity>
; collected on
<date id="FFBD104AFF854432C57D1C0E3B24EE7D" box="[1132,1328,1878,1904]" pageId="9" pageNumber="48" value="2003-07-19" valueMax="2003-07-23" valueMin="2003-07-19">1923 July 2003</date>
by D. B. Means (field collection DBM-3152; CPI 10335).
</paragraph>
<paragraph id="8BBC368AFF854431C1D71CFD3D7BE827" blockId="9.[151,1437,1837,2024]" lastBlockId="10.[151,1437,152,2018]" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48">
<emphasis id="B977EA98FF854432C1D71CFD3F6FEEB2" bold="true" box="[198,379,1957,1983]" pageId="9" pageNumber="48">Paratopotypes</emphasis>
.
<collectionCode id="ED12AE4FFF854432C0811CFE3FF1EECD" LSID="urn:lsid:biocol.org:col:34871" box="[400,485,1958,1984]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="48">USNM</collectionCode>
563634, an adult from the Wokomung Massif, summit of Mt. Wokomung, Potaro-Siparuni District, west-central
<collectingCountry id="F314761AFF854432C34B1C963CADEEE5" box="[602,697,1998,2024]" name="Guyana" pageId="9" pageNumber="48">Guyana</collectingCountry>
;
<geoCoordinate id="EE37504DFF854432C3D71C963D67EEE5" box="[710,883,1998,2024]" direction="north" orientation="latitude" pageId="9" pageNumber="48" precision="15" value="5.0675">05° 04 03” N</geoCoordinate>
,
<geoCoordinate id="EE37504DFF854432C2901C963A27EEE5" box="[897,1075,1998,2024]" direction="west" orientation="longitude" pageId="9" pageNumber="48" precision="15" value="-59.861664">59° 51 42” W</geoCoordinate>
,
<quantity id="4CFB9B6FFF854432C5511C963A8AEEE5" box="[1088,1182,1998,2024]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="9" pageNumber="48" unit="m" value="1560.0">1560 m</quantity>
; collected on
<date id="FFBD104AFF854431C4541C963EC6E9BF" lastPageId="10" lastPageNumber="49" pageId="9" pageNumber="48" value="2003-07-25">25 July 2003</date>
by D. B. Means (field collection DBM-3154; CPI 10329).
<collectionCode id="ED12AE4FFF864431C2B91BC03DE9E9BF" LSID="urn:lsid:biocol.org:col:34871" box="[936,1021,152,178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="49">USNM</collectionCode>
563635-563637, 563640, 563641, 563644, 563645, all from the Wokomung Massif, near Falls Camp, Potaro-Siparuni District, west-central
<collectingCountry id="F314761AFF864431C1861BB03EE2E80F" box="[151,246,232,258]" name="Guyana" pageId="10" pageNumber="49">Guyana</collectingCountry>
;
<geoCoordinate id="EE37504DFF864431C0121BB03FBAE80F" box="[259,430,232,258]" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="15" value="5.090278">05° 05 25” N</geoCoordinate>
,
<geoCoordinate id="EE37504DFF864431C0AA1BB03C7FE80F" box="[443,619,232,258]" direction="west" orientation="longitude" pageId="10" pageNumber="49" precision="15" value="-59.838333">59° 50 18” W</geoCoordinate>
,
<quantity id="4CFB9B6FFF864431C3691BB03CC1E80F" box="[632,725,232,258]" metricMagnitude="3" metricUnit="m" metricValue="1.385" pageId="10" pageNumber="49" unit="m" value="1385.0">1385 m</quantity>
; collected on
<date id="FFBD104AFF864431C26B1BB03A00E80F" box="[890,1044,232,258]" pageId="10" pageNumber="49" value="2003-07-19">19 July 2003</date>
by D. B. Means (field collection DBM-3152; CPI 10331-10333, 10336, 10337, 10340, 10341).
</paragraph>
</subSubSection>
<subSubSection id="C3196501FF864431C1D71A6F3C48EB4F" pageId="10" pageNumber="49" type="description">
<paragraph id="8BBC368AFF864431C1D71A6F3C48EB4F" blockId="10.[151,1437,152,2018]" pageId="10" pageNumber="49">
<emphasis id="B977EA98FF864431C1D71A6F3F81E85C" bold="true" box="[198,405,311,337]" pageId="10" pageNumber="49">
Other
<typeStatus id="54B88828FF864431C0081A6F3F81E85C" box="[281,405,311,337]" pageId="10" pageNumber="49" type="paratype">paratypes</typeStatus>
</emphasis>
. ROM 43302, from north slope of Mt. Wokomung, Potaro-Siparuni District, west-central
<collectingCountry id="F314761AFF864431C1D71A383F31E877" box="[198,293,352,378]" name="Guyana" pageId="10" pageNumber="49">Guyana</collectingCountry>
;
<geoCoordinate id="EE37504DFF864431C0231A383FC9E877" box="[306,477,352,378]" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="15" value="5.1294446">05° 07 46” N</geoCoordinate>
,
<geoCoordinate id="EE37504DFF864431C0FB1A383C8EE877" box="[490,666,352,378]" direction="west" orientation="longitude" pageId="10" pageNumber="49" precision="15" value="-59.82111">59° 49 16” W</geoCoordinate>
,
<quantity id="4CFB9B6FFF864431C3B71A383D17E877" box="[678,771,352,378]" metricMagnitude="3" metricUnit="m" metricValue="1.234" pageId="10" pageNumber="49" unit="m" value="1234.0">1234 m</quantity>
; collected
<date id="FFBD104AFF864431C2921A383A62E877" box="[899,1142,352,378]" pageId="10" pageNumber="49" value="2004-10-27" valueMax="2004-10-31" valueMin="2004-10-27">2731 October 2004</date>
by A. Lathrop, S. Khan, and R. MacCulloch. ROM 43309 &amp; 43310, from the base of the Wokomung Massif, Potaro-Siparuni District, west-central
<collectingCountry id="F314761AFF864431C03F1AE83F99E8C7" box="[302,397,432,458]" name="Guyana" pageId="10" pageNumber="49">Guyana</collectingCountry>
;
<geoCoordinate id="EE37504DFF864431C0881AE83C57E8C7" box="[409,579,432,458]" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="15" value="5.109722">05° 06 35” N</geoCoordinate>
,
<geoCoordinate id="EE37504DFF864431C35E1AE83CE9E8C7" box="[591,765,432,458]" direction="west" orientation="longitude" pageId="10" pageNumber="49" precision="15" value="-59.810276">59° 48 37” W</geoCoordinate>
,
<quantity id="4CFB9B6FFF864431C2181AE83D43E8C7" box="[777,855,432,458]" metricMagnitude="2" metricUnit="m" metricValue="6.9799999999999995" pageId="10" pageNumber="49" unit="m" value="698.0">698 m</quantity>
; collected
<date id="FFBD104AFF864431C2C41AE83AD3E8C7" box="[981,1223,432,458]" pageId="10" pageNumber="49" value="2004-10-23" valueMax="2004-10-26" valueMin="2004-10-23">2326 October 2004</date>
by A. Lathrop, S. Khan, and R. MacCulloch. ROM 43307, 43313, and 43314, from N slope of Mt. Wokomung, Potaro-Siparuni District, west-central
<collectingCountry id="F314761AFF864431C08919583FE3EB17" box="[408,503,512,538]" name="Guyana" pageId="10" pageNumber="49">Guyana</collectingCountry>
;
<geoCoordinate id="EE37504DFF864431C31419583CA7EB17" box="[517,691,512,538]" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="15" value="5.0925">05° 05 33” N</geoCoordinate>
,
<geoCoordinate id="EE37504DFF864431C3D019583D60EB17" box="[705,884,512,538]" direction="west" orientation="longitude" pageId="10" pageNumber="49" precision="15" value="-59.843056">59° 50 35” W</geoCoordinate>
,
<quantity id="4CFB9B6FFF864431C29319583DCAEB17" box="[898,990,512,538]" metricMagnitude="3" metricUnit="m" metricValue="1.411" pageId="10" pageNumber="49" unit="m" value="1411.0">1411 m</quantity>
, collected
<date id="FFBD104AFF864431C54E19583B44EB17" box="[1119,1360,512,538]" pageId="10" pageNumber="49" value="2004-11-03" valueMax="2004-11-05" valueMin="2004-11-03">3-5 November 2004</date>
by A. Lathrop, S. Khan, and R. MacCulloch.
</paragraph>
</subSubSection>
<subSubSection id="C3196501FF864431C1D719173A5CEF5F" pageId="10" pageNumber="49" type="materials_examined">
<paragraph id="8BBC368AFF864431C1D719173A5CEF5F" blockId="10.[151,1437,152,2018]" pageId="10" pageNumber="49">
<emphasis id="B977EA98FF864431C1D719173FBAEB64" bold="true" box="[198,430,591,617]" pageId="10" pageNumber="49">Referred material.</emphasis>
<collectionCode id="ED12AE4FFF864431C0A419083C1EEB67" LSID="urn:lsid:biocol.org:col:34871" box="[437,522,592,618]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="49">USNM</collectionCode>
563638, 563642-563643, 563646-563651, from near Falls Camp on the Wokomung Massif, Potaro-Siparuni District, west-central
<collectingCountry id="F314761AFF864431C20619203D62EB9F" box="[791,886,632,658]" name="Guyana" pageId="10" pageNumber="49">Guyana</collectingCountry>
;
<geoCoordinate id="EE37504DFF864431C29519203A27EB9F" box="[900,1075,632,658]" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="15" value="5.090278">05° 05 25” N</geoCoordinate>
,
<geoCoordinate id="EE37504DFF864431C55019203AE0EB9F" box="[1089,1268,632,658]" direction="west" orientation="longitude" pageId="10" pageNumber="49" precision="15" value="-59.838333">59° 50 18” W</geoCoordinate>
,
<quantity id="4CFB9B6FFF864431C41319203B74EB9F" box="[1282,1376,632,658]" metricMagnitude="3" metricUnit="m" metricValue="1.385" pageId="10" pageNumber="49" unit="m" value="1385.0">1385 m</quantity>
; collected
<date id="FFBD104AFF864431C1F919F83FA1EBB7" box="[232,437,672,698]" pageId="10" pageNumber="49" value="2003-07-19" valueMax="2003-07-23" valueMin="2003-07-19">1923 July 2003</date>
by D. B. Means (field collection DBM-3152; CPI 10334, 10338, 10339, 10342- 10347).
<collectionCode id="ED12AE4FFF864431C1EB19903F5BEBEF" LSID="urn:lsid:biocol.org:col:34871" box="[250,335,712,738]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="49">USNM</collectionCode>
564165 &amp; 564166, vicinity of a small, first-order creek on a terrace of the southern slope of Mt. Kopinang, SW part of the Wokomung Massif, Potaro-Siparuni District, west-central
<collectingCountry id="F314761AFF864431C5B819A83B13EA07" box="[1193,1287,752,778]" name="Guyana" pageId="10" pageNumber="49">Guyana</collectingCountry>
,
<geoCoordinate id="EE37504DFF864431C40219A83EBBEA3F" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="15" value="4.0805554">04° 04 50” N</geoCoordinate>
,
<geoCoordinate id="EE37504DFF864431C1AB18403F71EA3F" box="[186,357,792,818]" direction="west" orientation="longitude" pageId="10" pageNumber="49" precision="15" value="-59.87861">59° 52 43” W</geoCoordinate>
, ca.
<quantity id="4CFB9B6FFF864431C08918403FE0EA3F" box="[408,500,792,818]" metricMagnitude="3" metricUnit="m" metricValue="1.385" pageId="10" pageNumber="49" unit="m" value="1385.0">1385 m</quantity>
; collected
<date id="FFBD104AFF864431C36118403D54EA3F" box="[624,832,792,818]" pageId="10" pageNumber="49" value="2006-12-06">6 December 2006</date>
by D. B. Means and M. Kalamandeen (field collection DBM-3371; CPI 10357 &amp; 10358).
<collectionCode id="ED12AE4FFF864431C36418183CDEEA57" LSID="urn:lsid:biocol.org:col:34871" box="[629,714,832,858]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="49">USNM</collectionCode>
564167-564169, cloud forest summit of Mt. Kopinang, SW part of the Wokomung Massif, along a trail paralleling Kamana Creek for about
<quantity id="4CFB9B6FFF864431C54318303A8AEA8F" box="[1106,1182,872,898]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="49" unit="m" value="100.0">100 m</quantity>
west of the top of its cascade off of the summit at the E end of the trail, Potaro-Siparuni District, west-central
<collectingCountry id="F314761AFF864431C5B918C83B12EAA7" box="[1192,1286,912,938]" name="Guyana" pageId="10" pageNumber="49">Guyana</collectingCountry>
,
<geoCoordinate id="EE37504DFF864431C40318C83EBBEADF" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="15" value="5.002222">05° 00 08” N</geoCoordinate>
,
<geoCoordinate id="EE37504DFF864431C1AA18E03F7DEADF" box="[187,361,952,978]" direction="west" orientation="longitude" pageId="10" pageNumber="49" precision="15" value="-59.87972">59° 52 47” W</geoCoordinate>
, ca.
<quantity id="4CFB9B6FFF864431C08E18E03FEFEADF" box="[415,507,952,978]" metricMagnitude="3" metricUnit="m" metricValue="1.538" pageId="10" pageNumber="49" unit="m" value="1538.0">1538 m</quantity>
; collected
<date id="FFBD104AFF864431C36818E03D5FEADF" box="[633,843,952,978]" pageId="10" pageNumber="49" value="2006-12-07">7 December 2006</date>
by D. B. Means (field collection DBM-3372; CPI 10362, 10363, 10366).
<collectionCode id="ED12AE4FFF864431C0A118B83C11EAF7" LSID="urn:lsid:biocol.org:col:34871" box="[432,517,992,1018]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="49">USNM</collectionCode>
564170 &amp; 564171, cloud forested summit of Mt. Kopinang, SW part of the Wokomung Massif, along a trail paralleling Kamana Creek for about
<quantity id="4CFB9B6FFF864431C2DB1F503A01ED2F" box="[970,1045,1032,1058]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="49" unit="m" value="300.0">300 m</quantity>
west of the top of its cascade off of the summit at the E end of the trail, Potaro-Siparuni District, west-central
<collectingCountry id="F314761AFF864431C5331F683A94ED47" box="[1058,1152,1072,1098]" name="Guyana" pageId="10" pageNumber="49">Guyana</collectingCountry>
,
<geoCoordinate id="EE37504DFF864431C59C1F683B23ED47" box="[1165,1335,1072,1098]" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="15" value="5.002222">05° 00 08” N</geoCoordinate>
,
<geoCoordinate id="EE37504DFF864431C4551F683EF2ED7F" direction="west" orientation="longitude" pageId="10" pageNumber="49" precision="15" value="-59.87972">59° 52 47” W</geoCoordinate>
, ca.
<quantity id="4CFB9B6FFF864431C00F1F003F6FED7F" box="[286,379,1112,1138]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="10" pageNumber="49" unit="m" value="1570.0">1570 m</quantity>
; collected
<date id="FFBD104AFF864431C0ED1F003CC4ED7F" box="[508,720,1112,1138]" pageId="10" pageNumber="49" value="2006-12-08">8 December 2006</date>
by D. B. Means (field collection DBM-3373; CPI 10377 &amp; 10378).
<collectionCode id="ED12AE4FFF864431C1E81FD83F5AED97" LSID="urn:lsid:biocol.org:col:34871" box="[249,334,1152,1178]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="49">USNM</collectionCode>
564172 &amp; 564173, on leaves
<quantity id="4CFB9B6FFF864431C3A11FD83D36ED97" box="[688,802,1152,1178]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" metricValueMax="13.0" metricValueMin="3.0" pageId="10" pageNumber="49" unit="m" value="0.8" valueMax="1.3" valueMin="0.3">0.3-1.3 m</quantity>
off the ground in cloud forested summit of Mt. Kopinang, SW part of the Wokomung Massif, along a trail paralleling Kamana Creek for about
<quantity id="4CFB9B6FFF864431C5DF1FF03B0DEDCF" box="[1230,1305,1192,1218]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="49" unit="m" value="300.0">300 m</quantity>
west of its cascade off the top of the summit at the E end of the trail, Potaro-Siparuni District, west-central
<collectingCountry id="F314761AFF864431C4191F883B72EDE7" box="[1288,1382,1232,1258]" name="Guyana" pageId="10" pageNumber="49">Guyana</collectingCountry>
,
<geoCoordinate id="EE37504DFF864431C4621F883F04EC1F" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="15" value="5.002222">05° 00 08” N</geoCoordinate>
,
<geoCoordinate id="EE37504DFF864431C00D1FA03FDEEC1F" box="[284,458,1272,1298]" direction="west" orientation="longitude" pageId="10" pageNumber="49" precision="15" value="-59.87972">59° 52 47” W</geoCoordinate>
, ca.
<quantity id="4CFB9B6FFF864431C0EE1FA03C4FEC1F" box="[511,603,1272,1298]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="10" pageNumber="49" unit="m" value="1570.0">1570 m</quantity>
; collected
<date id="FFBD104AFF864431C3C81FA03DAEEC1F" box="[729,954,1272,1298]" pageId="10" pageNumber="49" value="2006-12-10">10 December 2006</date>
by D. B. Means, M. Kalamandeen (field collection DBM-3376; CPI 10384 &amp; 10390).
<collectionCode id="ED12AE4FFF864431C3A51E783D1DEC37" LSID="urn:lsid:biocol.org:col:34871" box="[692,777,1312,1338]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="49">USNM</collectionCode>
564174-564176, cloud forest summit of Mt. Kopinang, SW part of the Wokomung Massif, in the vicinity of the top of Kamana Creek Falls, Potaro-Siparuni District, west-central
<collectingCountry id="F314761AFF864431C03C1E283F9FEC87" box="[301,395,1392,1418]" name="Guyana" pageId="10" pageNumber="49">Guyana</collectingCountry>
,
<geoCoordinate id="EE37504DFF864431C0871E283C29EC87" box="[406,573,1392,1418]" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="15" value="5.002222">05° 00 08” N</geoCoordinate>
,
<geoCoordinate id="EE37504DFF864431C3581E283CE0EC87" box="[585,756,1392,1418]" direction="west" orientation="longitude" pageId="10" pageNumber="49" precision="15" value="-59.87972">59° 52 47” W</geoCoordinate>
, ca.
<quantity id="4CFB9B6FFF864431C2361E283D97EC87" box="[807,899,1392,1418]" metricMagnitude="3" metricUnit="m" metricValue="1.538" pageId="10" pageNumber="49" unit="m" value="1538.0">1538 m</quantity>
; collected
<date id="FFBD104AFF864431C2EE1E283ACAEC87" box="[1023,1246,1392,1418]" pageId="10" pageNumber="49" value="2006-12-10">10 December 2006</date>
by D. B. Means (field collection DBM-3377; CPI 10395, 10397, 103403).
<collectionCode id="ED12AE4FFF864431C27B1EC03DABECBF" LSID="urn:lsid:biocol.org:col:34871" box="[874,959,1432,1458]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="49">USNM</collectionCode>
564177-564179, all found at night on leaves 0.6-2.0 m off the ground in transitional forest (rainforest to cloud forest) next to a 300-m long trail on a terrace of the southern slope of Mt. Kopinang, SW part of the Wokomung Massif, Potaro-Siparuni District, west-central
<collectingCountry id="F314761AFF864431C03C1D483F9FEF27" box="[301,395,1552,1578]" name="Guyana" pageId="10" pageNumber="49">Guyana</collectingCountry>
,
<geoCoordinate id="EE37504DFF864431C0871D483C2AEF27" box="[406,574,1552,1578]" direction="north" orientation="latitude" pageId="10" pageNumber="49" precision="15" value="4.0805554">04° 04 50” N</geoCoordinate>
,
<geoCoordinate id="EE37504DFF864431C3581D483CE0EF27" box="[585,756,1552,1578]" direction="west" orientation="longitude" pageId="10" pageNumber="49" precision="15" value="-59.87861">59° 52 43” W</geoCoordinate>
, ca.
<quantity id="4CFB9B6FFF864431C2391D483D90EF27" box="[808,900,1552,1578]" metricMagnitude="3" metricUnit="m" metricValue="1.385" pageId="10" pageNumber="49" unit="m" value="1385.0">1385 m</quantity>
; collected
<date id="FFBD104AFF864431C5111D483AC9EF27" box="[1024,1245,1552,1578]" pageId="10" pageNumber="49" value="2006-12-11">11 December 2006</date>
by D. B. Means and M. Kalamandeen (field collection DBM-3378; CPI 10414, 10416 &amp; 10417).
</paragraph>
</subSubSection>
<subSubSection id="C3196501FF864430C1D71D073C28E9D7" lastPageId="11" lastPageNumber="50" pageId="10" pageNumber="49" type="diagnosis">
<paragraph id="8BBC368AFF864430C1D71D073C28E9D7" blockId="10.[151,1437,152,2018]" lastBlockId="11.[151,1436,152,218]" lastPageId="11" lastPageNumber="50" pageId="10" pageNumber="49">
<emphasis id="B977EA98FF864431C1D71D073F2BEF74" bold="true" box="[198,319,1631,1657]" pageId="10" pageNumber="49">Diagnosis</emphasis>
. A small species (SL 16.0
<quantity id="4CFB9B6FFF864431C3971D383CE6EF77" box="[646,754,1632,1658]" metricMagnitude="-2" metricUnit="m" metricValue="2.71" pageId="10" pageNumber="49" unit="mm" value="27.1">27.1 mm</quantity>
), one of eight
<collectingCountry id="F314761AFF864431C28D1D383DE5EF77" box="[924,1009,1632,1658]" name="Guyana" pageId="10" pageNumber="49">Guiana</collectingCountry>
Shield species in the
<taxonomicName id="4C034D09FF864431C5E31D383B88EF74" box="[1266,1436,1632,1658]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="unistrigatus">
<emphasis id="B977EA98FF864431C5E31D383B15EF74" box="[1266,1281,1632,1657]" italics="true" pageId="10" pageNumber="49">P</emphasis>
.
<emphasis id="B977EA98FF864431C4011D383B88EF74" box="[1296,1436,1632,1657]" italics="true" pageId="10" pageNumber="49">unistrigatus</emphasis>
</taxonomicName>
group having the auditory apparatus present but lacking toe webbing (
<figureCitation id="13382A0FFF864431C2C31DD03A0FEFAF" box="[978,1051,1672,1698]" captionStart="FIGURE 6" captionStartId="11.[151,255,1578,1602]" captionTargetBox="[153,1433,274,1554]" captionTargetId="figure@11.[153,1433,274,1554]" captionTargetPageId="11" captionText="FIGURE 6. Pristimantis saltissimus new species. a = plain or slightly spotted morph (43 % in a sample of 30); b = middorsally orange-striped morph (10 %); c = dorsally tan morph (17 %); d = thin, white-striped morph (10 %); e = dorsal views of the mottled morph (20 %) and thin, white-striped morph; f = ventral views of same morphs as in e. Note that the striping also occurs on the belly of the thin, white-striped morph." httpUri="https://zenodo.org/record/179905/files/figure.png" pageId="10" pageNumber="49">Fig. 6</figureCitation>
). The principal features that distinguish
<taxonomicName id="4C034D09FF864431C0101DE83FB0EFC4" box="[257,420,1712,1738]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="saltissimus">
<emphasis id="B977EA98FF864431C0101DE83F04EFC4" box="[257,272,1712,1737]" italics="true" pageId="10" pageNumber="49">P</emphasis>
.
<emphasis id="B977EA98FF864431C0351DE83FB0EFC4" box="[292,420,1712,1737]" italics="true" pageId="10" pageNumber="49">saltissimus</emphasis>
</taxonomicName>
from these taxa are the character state of the tympanum and coloration in life as detailed below (features of
<taxonomicName id="4C034D09FF864431C0CC1D803C69EFFC" box="[477,637,1752,1778]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="saltissimus">
<emphasis id="B977EA98FF864431C0CC1D803FFFEFFC" box="[477,491,1752,1777]" italics="true" pageId="10" pageNumber="49">P</emphasis>
.
<emphasis id="B977EA98FF864431C0ED1D803C69EFFC" box="[508,637,1752,1777]" italics="true" pageId="10" pageNumber="49">saltissimus</emphasis>
</taxonomicName>
in parentheses).
<taxonomicName id="4C034D09FF864431C2561D803B6DEFFF" authority="Myers and Donnelly, 1997" authorityName="Myers and Donnelly" authorityYear="1997" box="[839,1401,1752,1778]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="avius">
<emphasis id="B977EA98FF864431C2561D803A09EFFC" box="[839,1053,1752,1777]" italics="true" pageId="10" pageNumber="49">Pristimantis avius</emphasis>
(
<bibRefCitation id="EF924B7BFF864431C53F1D803B64EFFF" author="Myers" box="[1070,1392,1752,1778]" pageId="10" pageNumber="55" refString="Myers, C. W. &amp; Donnelly, M. A. (1997) A tepui herpetofauna on a granitic mountain (Tamacuari) in the borderland between Venezuela and Brazil: Report from the Phipps Tapirapeco Expedition. American Museum Novitates, 3213, 1 - 71." type="journal article" year="1997">Myers and Donnelly, 1997</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C034D09FF864431C4971D803C79EE17" authority="Myers and Donnelly, 1997" authorityName="Myers and Donnelly" authorityYear="1997" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="memorans">
<emphasis id="B977EA98FF864431C4971D803B81EFFC" box="[1414,1429,1752,1777]" italics="true" pageId="10" pageNumber="49">P</emphasis>
.
<emphasis id="B977EA98FF864431C1861C583F05EE14" box="[151,273,1792,1817]" italics="true" pageId="10" pageNumber="49">memorans</emphasis>
(
<bibRefCitation id="EF924B7BFF864431C0331C583C70EE17" author="Myers" box="[290,612,1792,1818]" pageId="10" pageNumber="55" refString="Myers, C. W. &amp; Donnelly, M. A. (1997) A tepui herpetofauna on a granitic mountain (Tamacuari) in the borderland between Venezuela and Brazil: Report from the Phipps Tapirapeco Expedition. American Museum Novitates, 3213, 1 - 71." type="journal article" year="1997">Myers and Donnelly, 1997</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C034D09FF864431C36A1C583DD9EE17" authority="Rivero, 1968" authorityName="Rivero" authorityYear="1968" box="[635,973,1792,1818]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="pulvinatus">
<emphasis id="B977EA98FF864431C36A1C583C9EEE14" box="[635,650,1792,1817]" italics="true" pageId="10" pageNumber="49">P</emphasis>
.
<emphasis id="B977EA98FF864431C3881C583D07EE14" box="[665,787,1792,1817]" italics="true" pageId="10" pageNumber="49">pulvinatus</emphasis>
(
<bibRefCitation id="EF924B7BFF864431C2341C583DD0EE17" author="Rivero" box="[805,964,1792,1818]" pageId="10" pageNumber="55" refString="Rivero, J. A. (1968) A new species of Eleutherodactylus (Amphibia, Salientia) from the Guayana region, Edo. Bolivar, Venezuela. Breviora, Museum of Comparative Zoology, (306), 1 - 11." type="journal article" year="1968">Rivero, 1968</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C034D09FF864431C2CA1C583A6FEE14" box="[987,1147,1792,1818]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="saltissimus">
<emphasis id="B977EA98FF864431C2CA1C583DFDEE14" box="[987,1001,1792,1817]" italics="true" pageId="10" pageNumber="49">P</emphasis>
.
<emphasis id="B977EA98FF864431C2EB1C583A6FEE14" box="[1018,1147,1792,1817]" italics="true" pageId="10" pageNumber="49">saltissimus</emphasis>
</taxonomicName>
(see below),
<taxonomicName id="4C034D09FF864431C40C1C583C07EE4F" authority="Heyer and Hardy, 1991" authorityName="Heyer and Hardy" authorityYear="1991" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="zimmermanae">
<emphasis id="B977EA98FF864431C40C1C583B38EE14" box="[1309,1324,1792,1817]" italics="true" pageId="10" pageNumber="49">P</emphasis>
.
<emphasis id="B977EA98FF864431C42D1C583EF1EE4C" italics="true" pageId="10" pageNumber="49">zimmermanae</emphasis>
(
<bibRefCitation id="EF924B7BFF864431C1E51C703C1EEE4F" author="Heyer" box="[244,522,1832,1858]" pageId="10" pageNumber="55" refString="Heyer, W. R. &amp; Hardy, L. M. (1991) A new species of frog of the Eleutherodactylus lacrimosus assembly from Amazonia, South America (Amphibia: Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 104 (3), 436 - 447." type="journal article" year="1991">Heyer and Hardy, 1991</bibRefCitation>
)
</taxonomicName>
, and sp. 2 of
<bibRefCitation id="EF924B7BFF864431C3BE1C703DF6EE4F" author="Lescure" box="[687,994,1832,1858]" pageId="10" pageNumber="55" refString="Lescure, J. &amp; Marty, C. (2000) Atlas des amphibiens de Guyane. Museum National D'Histoire Naturelle, Paris. 388 pp." type="book" year="2000">Lescure and Marty (2000)</bibRefCitation>
have prominent, large tympana and a distinct oticus tympanicus (indistinct tympanum and oticus tympanicus visible through the skin).
<taxonomicName id="4C034D09FF864431C4971C083CAAEE9F" authority="Fuentes and Barrio-Amoros 2000" authorityName="Fuentes and Barrio-Amoros" authorityYear="2000" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="marahuaka">
<emphasis id="B977EA98FF864431C4971C083B81EE64" box="[1414,1429,1872,1897]" italics="true" pageId="10" pageNumber="49">P</emphasis>
.
<emphasis id="B977EA98FF864431C1861C203F08EE9C" box="[151,284,1912,1937]" italics="true" pageId="10" pageNumber="49">marahuaka</emphasis>
(Fuentes and Barrio-Amorós 2000)
</taxonomicName>
of Cerro
<taxonomicName id="4C034D09FF864431C23C1C203DA0EE9F" box="[813,948,1912,1938]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="49" phylum="Chordata" rank="species" species="marahuaka">Marahuaka</taxonomicName>
,
<collectingCountry id="F314761AFF864431C2AE1C203A2EEE9F" box="[959,1082,1912,1938]" name="Venezuela" pageId="10" pageNumber="49">Venezuela</collectingCountry>
, has a small, indistinct tympanum with the oticus tympanicus visible through the skin but is uniform brown above with abundant miniscule white spots, especially on the flanks and upper surfaces of the forearms and thighs, and the venter is pale yellow or dirty white in preservative, without dark markings (dorsum with dark markings and venter usually heavily marked with dark pigment).
</paragraph>
</subSubSection>
<caption id="DF7C6602FF874430C1861D723D59EFA4" httpUri="https://zenodo.org/record/179905/files/figure.png" pageId="11" pageNumber="50" targetBox="[153,1433,274,1554]" targetPageId="11">
<paragraph id="8BBC368AFF874430C1861D723D59EFA4" blockId="11.[151,1436,1578,1705]" pageId="11" pageNumber="50">
<emphasis id="B977EA98FF874430C1861D723F06EF4F" bold="true" box="[151,274,1578,1602]" pageId="11" pageNumber="50">FIGURE 6</emphasis>
.
<taxonomicName id="4C034D09FF874430C0311D723C0BEF4C" box="[288,543,1578,1601]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="50" phylum="Chordata" rank="species" species="saltissimus" status="sp. nov.">
<emphasis id="B977EA98FF874430C0311D723C0BEF4C" box="[288,543,1578,1601]" italics="true" pageId="11" pageNumber="50">Pristimantis saltissimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457E3FF874430C3371D723CBDEF4C" box="[550,681,1578,1601]" pageId="11" pageNumber="50" rank="species">new species</taxonomicNameLabel>
. a = plain or slightly spotted morph (43% in a sample of 30); b = middorsally orange-striped morph (10%); c = dorsally tan morph (17%); d = thin, white-striped morph (10%); e = dorsal views of the mottled morph (20%) and thin, white-striped morph; f = ventral views of same morphs as in e. Note that the striping also occurs on the belly of the thin, white-striped morph.
</paragraph>
</caption>
<subSubSection id="C3196501FF874437C1D71D853EF7EA57" lastPageId="12" lastPageNumber="51" pageId="11" pageNumber="50" type="etymology">
<paragraph id="8BBC368AFF874430C1D71D853AB1EE12" blockId="11.[151,1437,1757,2023]" pageId="11" pageNumber="50">
<emphasis id="B977EA98FF874430C1D71D853F5FEFFA" bold="true" box="[198,331,1757,1783]" pageId="11" pageNumber="50">Etymology</emphasis>
. The name is from the Latin
<emphasis id="B977EA98FF874430C3AE1D853CECEFFB" box="[703,760,1757,1782]" italics="true" pageId="11" pageNumber="50">salto</emphasis>
(to jump) + the superlative
<emphasis id="B977EA98FF874430C5481D853ABBEFFB" box="[1113,1199,1757,1782]" italics="true" pageId="11" pageNumber="50">issimus</emphasis>
in reference to the extreme jumping ability of this diminutive frog. In the field it was called the rocket frog.
</paragraph>
<paragraph id="8BBC368AFF874437C1D71C753EF7EA57" blockId="11.[151,1437,1757,2023]" lastBlockId="12.[151,1437,152,1978]" lastPageId="12" lastPageNumber="51" pageId="11" pageNumber="50">
<emphasis id="B977EA98FF874430C1D71C753FFAEE4A" bold="true" box="[198,494,1837,1863]" pageId="11" pageNumber="50">General characteristics</emphasis>
. Head longer than broad; snout subelliptical in dorsal outline; snout profile acuminate. Canthus rostralis concave. Loreal region concave, upper lip not flared in cross-section; weak cloacal tubercle present. Choanae round, not concealed by maxillary arch, small, about the same size as tiny oblique patches of vomerine teeth lying posterior to and between choanae. No vocal slits or sac. Surface of head shagreened, dorsum and upper limb surfaces shagreened with widely scattered dorsal pustules and/or a few short ridges. Upper eyelid with several large pustules. EW/IOD = 73150%. Tympanum small, indistinct; annulus tympanicus visible under skin, width less than horizontal diameter of eye; TY/E = 2137%; tiny ostia pharyngea present; no distinct supra- or post-tympanic fold. Finger II much longer than Finger I when adpressed together; relative finger lengths III&gt;IV&gt;II&gt;I. No nuptial pads. Disc pad on Finger I only slightly wider than finger. Disc covers on Fingers II to IV expanded, even. Disc pads on Fingers IIIIV about equal in size to that on Toe IV; width of disc on Finger III equals width of tympanum; disc pads on all fingers broadened. No fringes, ridges or webbings on fingers. Subarticular tubercles under fingers and toes low, round, and globular in profile; no supernumerary tubercles; thenar tubercle low, elongate, smaller than bifid palmar tubercle; several accessory palmar tubercles. No distinct ulnar tubercles or fold. Heel smooth or with very weak tubercles. Toe disc pads moderate, disc on Toe IV slightly smaller to about same size as on Finger III; disc cover expanded, even; disc pads broadened. Relative toe lengths IV&gt;V&gt;III&gt;II=I; Toe V much longer than Toe III when adpressed against Toe IV, reaching distal subarticular tubercle; Toe III extends to the penultimate subarticular tubercle when adpressed against Toe IV. No fringes, ridges or webs on toes. No supernumerary tubercles under toes; plantar surface with numerous, small low tubercles; inner metatarsal tubercle elongate, outer metatarsal tubercle tiny, round; no tarsal fold or tubercle (
<figureCitation id="13382A0FFF804437C26919903DABEBEF" box="[888,959,712,738]" captionStart="FIGURE 7" captionStartId="13.[151,255,1402,1426]" captionTargetBox="[172,1393,204,1378]" captionTargetId="figure@13.[155,1432,194,1378]" captionTargetPageId="13" captionText="FIGURE 7. Pristimantis saltissimus new species. Hand (left) and foot (right). USNM 563639 holotype. Black bars = 5.0 mm." httpUri="https://zenodo.org/record/274028/files/figure.png" pageId="12" pageNumber="51">Fig. 7</figureCitation>
). No inguinal gland; venter coarsely areolate; throat and under surfaces of limbs smooth; pericloacal area granular. Legs relatively short, heels barely overlapping when legs folded at right angles to sagittal plane; C/SL = 5364%. See
<tableCitation id="C6810331FF804437C56518403AD8EA3F" box="[1140,1228,792,818]" captionStart="TABLE 1" captionStartId="4.[151,239,831,855]" captionTargetBox="[159,1260,922,1969]" captionTargetPageId="4" captionText="TABLE 1. Measurements (in mm) and proportions of adult Pristimantis dendrobatoides, P. j e s t e r, and P. saltissimus; ranges followed by means and one standard deviation in parentheses. For abbreviations, see Methods." httpUri="http://table.plazi.org/id/DF7C6602FF88443FC18618673AC1EA74" pageId="12" pageNumber="51" tableUuid="DF7C6602FF88443FC18618673AC1EA74">Table 1</tableCitation>
for summary statistics.
</paragraph>
</subSubSection>
<subSubSection id="C3196501FF804435C1D7183F3D98E85F" lastPageId="14" lastPageNumber="53" pageId="12" pageNumber="51" type="description">
<paragraph id="8BBC368AFF804437C1D7183F3F52EF0F" blockId="12.[151,1437,152,1978]" pageId="12" pageNumber="51">
<emphasis id="B977EA98FF804437C1D7183F3F77EA8C" bold="true" box="[198,355,871,897]" pageId="12" pageNumber="51">Color in life.</emphasis>
Dorsal color pattern extremely variable due to pattern polymorphism as well as metachrosis (
<figureCitation id="13382A0FFF804437C18E18C83EF3EAA7" box="[159,231,912,938]" captionStart="FIGURE 6" captionStartId="11.[151,255,1578,1602]" captionTargetBox="[153,1433,274,1554]" captionTargetId="figure@11.[153,1433,274,1554]" captionTargetPageId="11" captionText="FIGURE 6. Pristimantis saltissimus new species. a = plain or slightly spotted morph (43 % in a sample of 30); b = middorsally orange-striped morph (10 %); c = dorsally tan morph (17 %); d = thin, white-striped morph (10 %); e = dorsal views of the mottled morph (20 %) and thin, white-striped morph; f = ventral views of same morphs as in e. Note that the striping also occurs on the belly of the thin, white-striped morph." httpUri="https://zenodo.org/record/179905/files/figure.png" pageId="12" pageNumber="51">Fig. 6</figureCitation>
). Frogs may be darker or lighter overall, as a result of handling, background coloration, temperature, or time of day. Upper surfaces of head and body light brown to tan with extremely variable dark brown markings that range from small spots or irregular blotches (
<figureCitation id="13382A0FFF804437C3FD18B83D20EAF7" box="[748,820,992,1018]" captionStart="FIGURE 6" captionStartId="11.[151,255,1578,1602]" captionTargetBox="[153,1433,274,1554]" captionTargetId="figure@11.[153,1433,274,1554]" captionTargetPageId="11" captionText="FIGURE 6. Pristimantis saltissimus new species. a = plain or slightly spotted morph (43 % in a sample of 30); b = middorsally orange-striped morph (10 %); c = dorsally tan morph (17 %); d = thin, white-striped morph (10 %); e = dorsal views of the mottled morph (20 %) and thin, white-striped morph; f = ventral views of same morphs as in e. Note that the striping also occurs on the belly of the thin, white-striped morph." httpUri="https://zenodo.org/record/179905/files/figure.png" pageId="12" pageNumber="51">Fig. 6</figureCitation>
a), through single transverse suprascapular bar, to a series of several oblique bars that extend laterally onto the flanks (
<figureCitation id="13382A0FFF804437C2BC1F503DE1ED2F" box="[941,1013,1032,1058]" captionStart="FIGURE 6" captionStartId="11.[151,255,1578,1602]" captionTargetBox="[153,1433,274,1554]" captionTargetId="figure@11.[153,1433,274,1554]" captionTargetPageId="11" captionText="FIGURE 6. Pristimantis saltissimus new species. a = plain or slightly spotted morph (43 % in a sample of 30); b = middorsally orange-striped morph (10 %); c = dorsally tan morph (17 %); d = thin, white-striped morph (10 %); e = dorsal views of the mottled morph (20 %) and thin, white-striped morph; f = ventral views of same morphs as in e. Note that the striping also occurs on the belly of the thin, white-striped morph." httpUri="https://zenodo.org/record/179905/files/figure.png" pageId="12" pageNumber="51">Fig. 6</figureCitation>
e). Many specimens have an unbroken, cream to light tannish orange colored middorsal stripe highlighted by a suffusion of black pigment laterally (
<figureCitation id="13382A0FFF804437C1C41F003F02ED7F" box="[213,278,1112,1138]" captionStart="FIGURE 6" captionStartId="11.[151,255,1578,1602]" captionTargetBox="[153,1433,274,1554]" captionTargetId="figure@11.[153,1433,274,1554]" captionTargetPageId="11" captionText="FIGURE 6. Pristimantis saltissimus new species. a = plain or slightly spotted morph (43 % in a sample of 30); b = middorsally orange-striped morph (10 %); c = dorsally tan morph (17 %); d = thin, white-striped morph (10 %); e = dorsal views of the mottled morph (20 %) and thin, white-striped morph; f = ventral views of same morphs as in e. Note that the striping also occurs on the belly of the thin, white-striped morph." httpUri="https://zenodo.org/record/179905/files/figure.png" pageId="12" pageNumber="51">Fig 6</figureCitation>
b, d, e); the stripe may be narrower than the width of a finger or half as wide as the interorbital distance; others have a broad middorsal light tan field bordered laterally by a black margin (
<figureCitation id="13382A0FFF804437C5D31FD83B1CED97" box="[1218,1288,1152,1178]" captionStart="FIGURE 6" captionStartId="11.[151,255,1578,1602]" captionTargetBox="[153,1433,274,1554]" captionTargetId="figure@11.[153,1433,274,1554]" captionTargetPageId="11" captionText="FIGURE 6. Pristimantis saltissimus new species. a = plain or slightly spotted morph (43 % in a sample of 30); b = middorsally orange-striped morph (10 %); c = dorsally tan morph (17 %); d = thin, white-striped morph (10 %); e = dorsal views of the mottled morph (20 %) and thin, white-striped morph; f = ventral views of same morphs as in e. Note that the striping also occurs on the belly of the thin, white-striped morph." httpUri="https://zenodo.org/record/179905/files/figure.png" pageId="12" pageNumber="51">Fig. 6</figureCitation>
c), and a few have the dorsum marked with slightly irregular dark and light longitudinal stripes. Head in many individuals with a dark brown interorbital bar set off by tan color on snout (
<figureCitation id="13382A0FFF804437C2821F883DCEEDE7" box="[915,986,1232,1258]" captionStart="FIGURE 6" captionStartId="11.[151,255,1578,1602]" captionTargetBox="[153,1433,274,1554]" captionTargetId="figure@11.[153,1433,274,1554]" captionTargetPageId="11" captionText="FIGURE 6. Pristimantis saltissimus new species. a = plain or slightly spotted morph (43 % in a sample of 30); b = middorsally orange-striped morph (10 %); c = dorsally tan morph (17 %); d = thin, white-striped morph (10 %); e = dorsal views of the mottled morph (20 %) and thin, white-striped morph; f = ventral views of same morphs as in e. Note that the striping also occurs on the belly of the thin, white-striped morph." httpUri="https://zenodo.org/record/179905/files/figure.png" pageId="12" pageNumber="51">Fig. 6</figureCitation>
a, e); canthus rostralis usually marked with a strong dark stripe; upper surfaces of arms and legs the same color as dorsum; finger and toe discs pinkish tan; upper and lower legs and crus on some specimens have a hint of three to four light gray and tan alternating crossbands and on others there are three dark brown bands; throat of males white with black smudging or mottling, often white in females; belly and undersurfaces of legs heavily marked with dark pigment sometimes mottled as on throat, but lighter overall (
<figureCitation id="13382A0FFF804437C3D41EC03D18ECBF" box="[709,780,1432,1458]" captionStart="FIGURE 6" captionStartId="11.[151,255,1578,1602]" captionTargetBox="[153,1433,274,1554]" captionTargetId="figure@11.[153,1433,274,1554]" captionTargetPageId="11" captionText="FIGURE 6. Pristimantis saltissimus new species. a = plain or slightly spotted morph (43 % in a sample of 30); b = middorsally orange-striped morph (10 %); c = dorsally tan morph (17 %); d = thin, white-striped morph (10 %); e = dorsal views of the mottled morph (20 %) and thin, white-striped morph; f = ventral views of same morphs as in e. Note that the striping also occurs on the belly of the thin, white-striped morph." httpUri="https://zenodo.org/record/179905/files/figure.png" pageId="12" pageNumber="51">Fig. 6</figureCitation>
f); individuals with a middorsal stripe also have a midventral, off-white streak (
<figureCitation id="13382A0FFF804437C0D91E983C04ECD7" box="[456,528,1472,1498]" captionStart="FIGURE 6" captionStartId="11.[151,255,1578,1602]" captionTargetBox="[153,1433,274,1554]" captionTargetId="figure@11.[153,1433,274,1554]" captionTargetPageId="11" captionText="FIGURE 6. Pristimantis saltissimus new species. a = plain or slightly spotted morph (43 % in a sample of 30); b = middorsally orange-striped morph (10 %); c = dorsally tan morph (17 %); d = thin, white-striped morph (10 %); e = dorsal views of the mottled morph (20 %) and thin, white-striped morph; f = ventral views of same morphs as in e. Note that the striping also occurs on the belly of the thin, white-striped morph." httpUri="https://zenodo.org/record/179905/files/figure.png" pageId="12" pageNumber="51">Fig. 6</figureCitation>
f); upper two-fifths of iris dirty gold color, lower three-fifths dark brown to black (
<figureCitation id="13382A0FFF804437C1F71EB03F38EF0F" box="[230,300,1512,1538]" captionStart="FIGURE 6" captionStartId="11.[151,255,1578,1602]" captionTargetBox="[153,1433,274,1554]" captionTargetId="figure@11.[153,1433,274,1554]" captionTargetPageId="11" captionText="FIGURE 6. Pristimantis saltissimus new species. a = plain or slightly spotted morph (43 % in a sample of 30); b = middorsally orange-striped morph (10 %); c = dorsally tan morph (17 %); d = thin, white-striped morph (10 %); e = dorsal views of the mottled morph (20 %) and thin, white-striped morph; f = ventral views of same morphs as in e. Note that the striping also occurs on the belly of the thin, white-striped morph." httpUri="https://zenodo.org/record/179905/files/figure.png" pageId="12" pageNumber="51">Fig. 6</figureCitation>
f).
</paragraph>
<paragraph id="8BBC368AFF804437C1D71D573D0DEEB7" blockId="12.[151,1437,152,1978]" pageId="12" pageNumber="51">
<emphasis id="B977EA98FF804437C1D71D573FCCEF24" bold="true" box="[198,472,1551,1577]" pageId="12" pageNumber="51">Color in preservative.</emphasis>
Upper surfaces of head and body light gray to light tan with dark brown to black marks in various shapes from dashes to Us, the dark marks paired on either side of the midline of the back in individuals with a dirty white, narrow, black bordered, middorsal stripe; usually a black interorbital triangle on the head set off by a tan rostrum; a prominent canthus rostralis with black stripe from eye to snout; upper arm uniformly dirty white, sometimes with a few black specks; forearm with one or two wide, alternating black and dirty white crossbands; hand and fingers with smaller black and white crossbands; finger and toe discs with a dark gray center, lighter distally; upper leg surface with three or four alternating dark and light bands, often faint or absent; crus, foot, and toes slightly banded as in the hands and fingers. Lower lip and throat black with dirty white speckling; belly dirty white in some females, dirty white and mottled or speckled with dark brown in males; posterior surface of thigh and inner surfaces of crus and legs uniformly rusty brown; palmar and plantar surfaces light gray to black.
</paragraph>
<caption id="DF7C6602FF814436C1861E223EDCECB9" httpUri="https://zenodo.org/record/274028/files/figure.png" pageId="13" pageNumber="52" targetBox="[172,1393,204,1378]" targetPageId="13">
<paragraph id="8BBC368AFF814436C1861E223EDCECB9" blockId="13.[151,1435,1402,1460]" pageId="13" pageNumber="52">
<emphasis id="B977EA98FF814436C1861E223F06EC9F" bold="true" box="[151,274,1402,1426]" pageId="13" pageNumber="52">FIGURE 7</emphasis>
.
<taxonomicName id="4C034D09FF814436C00E1E223C09EC9C" box="[287,541,1402,1425]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="52" phylum="Chordata" rank="species" species="saltissimus" status="sp. nov.">
<emphasis id="B977EA98FF814436C00E1E223C09EC9C" box="[287,541,1402,1425]" italics="true" pageId="13" pageNumber="52">Pristimantis saltissimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24457E3FF814436C3321E223CB2EC9C" box="[547,678,1402,1425]" pageId="13" pageNumber="52" rank="species">new species</taxonomicNameLabel>
. Hand (left) and foot (right). USNM 563639 holotype. Black bars = 5.0 mm.
</paragraph>
</caption>
<paragraph id="8BBC368AFF814436C1D71EBF3F2AEF27" blockId="13.[151,1436,1511,2018]" pageId="13" pageNumber="52">
<emphasis id="B977EA98FF814436C1D71EBF3C96EF0C" bold="true" box="[198,642,1511,1537]" pageId="13" pageNumber="52">
Measurements of
<typeStatus id="54B88828FF814436C0B91EBF3C07EF0C" box="[424,531,1511,1537]" pageId="13" pageNumber="52" type="holotype">holotype</typeStatus>
(in mm)
</emphasis>
. SL 26.6; HL 11.8; HW 10.8; EW 2.8; IOD 2.4; E 4.9; TY 1.3; C 17.5; FL 14.5.
</paragraph>
<paragraph id="8BBC368AFF814436C1D71D6F3B88EEEF" blockId="13.[151,1436,1511,2018]" pageId="13" pageNumber="52">
<emphasis id="B977EA98FF814436C1D71D6F3FA2EF5C" bold="true" box="[198,438,1591,1617]" pageId="13" pageNumber="52">Habitat and habits</emphasis>
. Leafy shrubbery, branches, tree buttresses, exposed roots, and herbaceous ground vegetation usually&gt;
<quantity id="4CFB9B6FFF814436C09E1D383FC6EF77" box="[399,466,1632,1658]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="52" unit="m" value="0.5">0.5 m</quantity>
off the ground to
<quantity id="4CFB9B6FFF814436C3BA1D383CCDEF77" box="[683,729,1632,1658]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="52" unit="m" value="2.0">2 m</quantity>
high, in dense cloud forest. The most productive sites were the leafy banks of small cascades plunging down the steep sides of escarpments, often in the identical microhabitats of a species of arboreal toad,
<taxonomicName id="4C034D09FF814436C34E1DE83D1EEFC4" box="[607,778,1712,1737]" class="Amphibia" family="Bufonidae" genus="Oreophrynella" kingdom="Animalia" order="Anura" pageId="13" pageNumber="52" phylum="Chordata" rank="genus">
<emphasis id="B977EA98FF814436C34E1DE83D1EEFC4" box="[607,778,1712,1737]" italics="true" pageId="13" pageNumber="52">Oreophrynella</emphasis>
</taxonomicName>
sp. Vegetation near Falls Camp (
<geoCoordinate id="EE37504DFF814436C58D1DE83B5CEFC7" box="[1180,1352,1712,1738]" direction="north" orientation="latitude" pageId="13" pageNumber="52" precision="1" value="5.09035">05° 05.421 N</geoCoordinate>
<date id="FFBD104AFF814436C4401DE83B99EFC7" box="[1361,1421,1712,1738]" pageId="13" pageNumber="52" value="1959-10">X 59</date>
û 50.296 W) along the stream, which steepened its incline and became a rocky, mossy, tumbling, cascading stream whose valley ran through huge sandstone boulders, covered in wet mosses and lichens. Also found on the leaves of understory vegetation in the cloud forest away from streams and cascades. This little frog seems to be a “leaf-sitter,” foraging for ants and insects on leaves off the ground. Individuals were seen within a few meters of
<taxonomicName id="4C034D09FF814436C01E1C203FF6EE9C" box="[271,482,1912,1937]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="52" phylum="Chordata" rank="species" species="dendrobatoides">
<emphasis id="B977EA98FF814436C01E1C203FF6EE9C" box="[271,482,1912,1937]" italics="true" pageId="13" pageNumber="52">P. dendrobatoides</emphasis>
</taxonomicName>
, also sitting on leaves. During the 2003 expedition on the eastern portion of the Wokomung Massif, every specimen of
<taxonomicName id="4C034D09FF814436C3711CF83CE9EEB4" box="[608,765,1952,1977]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="13" pageNumber="52" phylum="Chordata" rank="species" species="saltissimus">
<emphasis id="B977EA98FF814436C3711CF83CE9EEB4" box="[608,765,1952,1977]" italics="true" pageId="13" pageNumber="52">P. saltissimus</emphasis>
</taxonomicName>
had a distinctive stinky smell when handled, and a bittersweet taste. The skin secretions also made the back of the collectors tongue feel affected for some minutes.
</paragraph>
<paragraph id="8BBC368AFF824435C1861BC03D98E85F" blockId="14.[151,1436,152,378]" pageId="14" pageNumber="53">The sensation is difficult to describe, although numbness comes closest. During the 2006 expedition to the SW side of the massif (on the slopes and summit of Mt. Kopinang), DBM made a special effort to collect specimens of this species, smelling and tasting every one. Interestingly, individuals were quite variable in degree of noxiousness. Some were neither malodorous nor distasteful, but others distinctly smelled and tasted to varying intensities like those on the eastern side of the massif.</paragraph>
</subSubSection>
<subSubSection id="C3196501FF824435C1D71A073B6DE877" box="[198,1401,351,378]" pageId="14" pageNumber="53" type="distribution">
<paragraph id="8BBC368AFF824435C1D71A073B6DE877" blockId="14.[151,1436,152,378]" box="[198,1401,351,378]" pageId="14" pageNumber="53">
<emphasis id="B977EA98FF824435C1D71A073F4BE874" bold="true" box="[198,351,351,377]" pageId="14" pageNumber="53">Distribution</emphasis>
. Known only from the Wokomung Massif at elevations between 698 and
<quantity id="4CFB9B6FFF824435C5D01A383B0EE877" box="[1217,1306,352,378]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="14" pageNumber="53" unit="m" value="1560.0">1560 m</quantity>
(
<figureCitation id="13382A0FFF824435C4381A383B7DE877" box="[1321,1385,352,378]" captionStart="FIGURE 3" captionStartId="5.[151,255,1305,1329]" captionTargetBox="[388,1200,208,1304]" captionTargetId="figure@5.[388,1200,194,1351]" captionTargetPageId="5" captionText="FIGURE 3. Map of Guyana with the location of the Wokomung Massif in the Pakaraima Mountains (gray shading) and in relation to some other teuis in Guyana. Map modified from one supplied by Philippe J. R. Kok." httpUri="https://zenodo.org/record/179902/files/figure.png" pageId="14" pageNumber="53">Fig 3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>