599 lines
82 KiB
XML
599 lines
82 KiB
XML
<document id="01B0814329F39654C88497D3DFDFED99" ID-CLB-Dataset="28397" ID-DOI="10.11646/zootaxa.4565.2.2" ID-GBIF-Dataset="b289d291-5da8-48a8-b283-a456bce4ec76" ID-ISSN="1175-5326" ID-Zenodo-Dep="2589826" ID-ZooBank="1E01FCF7-0D6D-4E8C-BDF2-E2D8068934B5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552279673760" checkinUser="plazi" docAuthor="Kovačić, Marcelo, Bogorodsky, Sergey V., Troyer, Emily M. & Tornabene, Luke" docDate="2019" docId="DE205C17FF8CFFB9FF3AFAD94B5FDA7F" docLanguage="en" docName="zootaxa.4565.2.2.pdf" docOrigin="Zootaxa 4565 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cerogobius Kovačić & Bogorodsky & Troyer & Tornabene 2019, gen. nov." docType="treatment" docVersion="8" lastPageNumber="177" masterDocId="2219246FFF8EFFBFFFADFF4E4821DD30" masterDocTitle="Cerogobius petrophilus (Perciformes: Gobiidae), a new gobiid genus and species from the Red Sea" masterLastPageNumber="189" masterPageNumber="171" pageNumber="173" updateTime="1698701428881" updateUser="ExternalLinkService">
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<mods:title id="0891EAC1E51CA9E1526A3461E0FD235B">Cerogobius petrophilus (Perciformes: Gobiidae), a new gobiid genus and species from the Red Sea</mods:title>
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<mods:name id="BB8E670E6F5B6D737F70D92114B23340" type="personal">
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<mods:namePart id="669462063906C3474E1D3B36FDA0B940">Kovačić, Marcelo</mods:namePart>
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<mods:namePart id="A054DE0700DFB5875886F03C11AD6EC5">Bogorodsky, Sergey V.</mods:namePart>
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<mods:namePart id="14E3E9DCDF1A2205906E5F487235701D">Troyer, Emily M.</mods:namePart>
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<mods:namePart id="DD732F068F66F2B8540DA8DD538CA015">Tornabene, Luke</mods:namePart>
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<mods:title id="85809B5813EBB7DE11D4642B501DDEEC">Zootaxa</mods:title>
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<mods:date id="346BF6F8179DEAB1CCB2073A978BF68D">2019</mods:date>
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<mods:number id="1FFC1FDA0780C6B046382A1564432220">2019-03-08</mods:number>
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<mods:number id="179935ABEC7A419E804AEE473FC49D76">4565</mods:number>
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<treatment id="DE205C17FF8CFFB9FF3AFAD94B5FDA7F" ID-DOI="http://doi.org/10.5281/zenodo.5928725" ID-GBIF-Taxon="154366881" ID-Zenodo-Dep="5928725" LSID="urn:lsid:plazi:treatment:DE205C17FF8CFFB9FF3AFAD94B5FDA7F" httpUri="http://treatment.plazi.org/id/DE205C17FF8CFFB9FF3AFAD94B5FDA7F" lastPageId="6" lastPageNumber="177" pageId="2" pageNumber="173">
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<subSubSection id="1E93BE8AFF8CFFBDFF3AFAD949B7D882" box="[151,406,1431,1458]" pageId="2" pageNumber="173" type="nomenclature">
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<paragraph id="5636ED01FF8CFFBDFF3AFAD949B7D882" blockId="2.[151,406,1431,1458]" box="[151,406,1431,1458]" pageId="2" pageNumber="173">
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<heading id="0D7E5A6DFF8CFFBDFF3AFAD949B7D882" bold="true" box="[151,406,1431,1458]" fontSize="11" level="1" pageId="2" pageNumber="173" reason="1">
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<emphasis id="64FD3113FF8CFFBDFF3AFAD949B7D882" bold="true" box="[151,406,1431,1458]" pageId="2" pageNumber="173">
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<taxonomicName id="91899682FF8CFFBDFF3AFAD94903D881" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[151,290,1431,1457]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="64FD3113FF8CFFBDFF3AFAD94903D881" bold="true" box="[151,290,1431,1457]" italics="true" pageId="2" pageNumber="173">Cerogobius</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7FCE8C68FF8CFFBDFE85FAD649B7D882" box="[296,406,1432,1458]" pageId="2" pageNumber="173" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="1E93BE8AFF8CFFBDFF3AFA904B1EDAE4" pageId="2" pageNumber="173" type="diagnosis">
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<paragraph id="5636ED01FF8CFFBDFF3AFA904B1EDAE4" blockId="2.[151,1436,1502,2004]" pageId="2" pageNumber="173">
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<emphasis id="64FD3113FF8CFFBDFF3AFA90492BD8C7" bold="true" box="[151,266,1502,1527]" pageId="2" pageNumber="173">Diagnosis</emphasis>
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. The new genus has the following combination of characters unique among known gobiid genera: 1) unpaired horn-like tentacle at the level of nostrils on the snout midline (
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<figureCitation id="CEB2F184FF8CFFBDFC69F94C4C3FDB2B" box="[964,1054,1538,1563]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="2" pageNumber="173">Fig. 1A</figureCitation>
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); 2) snout with steep profile from lateral view and very short, about 0.4–0.5 of eye diameter; 3) orbit projecting forward and elevated, obviously extending above dorsal profile of the head; 4) posterior angle of jaws extends behind vertical through posterior edge of eye; 5) lower limb of first gill arch joined to gill cover by membrane; 6) caudal peduncle deep and short, 0.8-0.9 of body depth at anal-fin origin, and caudal-peduncle depth slightly to distinctly greater than its length; 7) segmented caudal-fin rays 17; 8) pelvic fins completely separated and without frenum (
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<figureCitation id="CEB2F184FF8CFFBDFBC1F9F84CE5DBFF" box="[1132,1220,1718,1743]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="2" pageNumber="173">Fig. 1C</figureCitation>
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); 9) head and body completely scaleless; 10) no head canals; 11) preorbital rows
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<emphasis id="64FD3113FF8CFFBDFCE4F9904BAADBD8" box="[841,907,1754,1781]" italics="true" pageId="2" pageNumber="173">
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r, s, s
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<superScript id="A1FC4049FF8CFFBDFC2EF9944BAADBD8" attach="left" box="[899,907,1754,1768]" fontSize="6" pageId="2" pageNumber="173">3</superScript>
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</emphasis>
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each reduced to a single papilla; 12) suborbital sensory papillae rows
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<emphasis id="64FD3113FF8CFFBDFE3EF84C4980DA29" box="[403,417,1794,1817]" italics="true" pageId="2" pageNumber="173">a</emphasis>
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and
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<emphasis id="64FD3113FF8CFFBDFE77F84F49C8DA2A" box="[474,489,1793,1818]" italics="true" pageId="2" pageNumber="173">b</emphasis>
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absent; four very short transverse suborbital rows of sensory papillae (
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<emphasis id="64FD3113FF8CFFBDFAA0F84C4D12DA2A" box="[1293,1331,1794,1818]" italics="true" pageId="2" pageNumber="173">1-4</emphasis>
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) present, row
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<emphasis id="64FD3113FF8CFFBDFF64F86848F6DA0E" box="[201,215,1830,1854]" italics="true" pageId="2" pageNumber="173">1</emphasis>
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oblique, reaching upper lip, row
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<emphasis id="64FD3113FF8CFFBDFDE6F8684A78DA0E" box="[587,601,1830,1854]" italics="true" pageId="2" pageNumber="173">2</emphasis>
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distant from eye, close to row
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<emphasis id="64FD3113FF8CFFBDFC1FF86B4BE1DA0E" box="[946,960,1829,1854]" italics="true" pageId="2" pageNumber="173">d</emphasis>
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, rows
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<emphasis id="64FD3113FF8CFFBDFBA6F8684C38DA0E" box="[1035,1049,1830,1854]" italics="true" pageId="2" pageNumber="173">3</emphasis>
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and
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<emphasis id="64FD3113FF8CFFBDFBE3F8684C7DDA0E" box="[1102,1116,1830,1854]" italics="true" pageId="2" pageNumber="173">4</emphasis>
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close to posteroventral edge of eye and distant from row
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<emphasis id="64FD3113FF8CFFBDFE75F80249C7DA55" box="[472,486,1868,1893]" italics="true" pageId="2" pageNumber="173">d</emphasis>
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, row
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<emphasis id="64FD3113FF8CFFBDFD8AF8024A14DA55" box="[551,565,1868,1893]" italics="true" pageId="2" pageNumber="173">d</emphasis>
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short; 13) oculoscapular rows reduced to rows
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<emphasis id="64FD3113FF8CFFBDFBE0F8034CA4DA67" box="[1101,1157,1865,1892]" italics="true" pageId="2" pageNumber="173">
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<date id="2237CBC1FF8CFFBDFBE0F8034C43DA67" box="[1101,1122,1865,1892]" pageId="2" pageNumber="173">
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x
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<superScript id="A1FC4049FF8CFFBDFBF7F8074C43DA67" attach="left" box="[1114,1122,1865,1879]" fontSize="6" pageId="2" pageNumber="173">1</superScript>
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</date>
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,
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<date id="2237CBC1FF8CFFBDFBDCF8034C5CDA54" box="[1137,1149,1869,1892]" pageId="2" pageNumber="173">x</date>
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||
<superScript id="A1FC4049FF8CFFBDFBD0F8074CA4DA67" attach="left" box="[1149,1157,1865,1879]" fontSize="6" pageId="2" pageNumber="173">2</superScript>
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</emphasis>
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,
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<emphasis id="64FD3113FF8CFFBDFB3EF8034CEFDA67" box="[1171,1230,1865,1892]" italics="true" pageId="2" pageNumber="173">
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z, as
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<superScript id="A1FC4049FF8CFFBDFB6BF8074CEFDA67" attach="left" box="[1222,1230,1865,1879]" fontSize="6" pageId="2" pageNumber="173">1</superScript>
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</emphasis>
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and
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<emphasis id="64FD3113FF8CFFBDFAAFF8034D03DA67" box="[1282,1314,1865,1892]" italics="true" pageId="2" pageNumber="173">
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as
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<superScript id="A1FC4049FF8CFFBDFAB7F8074D03DA67" attach="left" box="[1306,1314,1865,1879]" fontSize="6" pageId="2" pageNumber="173">2</superScript>
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</emphasis>
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, with only
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<date id="2237CBC1FF8CFFBDFF3AF83A488ADA4D" box="[151,171,1903,1931]" pageId="2" pageNumber="173">
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<emphasis id="64FD3113FF8CFFBDFF3AF83A488ADA4D" box="[151,171,1903,1931]" italics="true" pageId="2" pageNumber="173">
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x
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<superScript id="A1FC4049FF8CFFBDFF0EF821488ADA4D" attach="left" box="[163,171,1903,1917]" fontSize="6" pageId="2" pageNumber="173">1</superScript>
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</emphasis>
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</date>
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having more than 2 papillae (
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<figureCitation id="CEB2F184FF8CFFBDFDAEF83D4A68DABB" box="[515,585,1907,1932]" captionStart="FIGURE 2" captionStartId="4.[151,250,1022,1044]" captionTargetBox="[201,1426,297,947]" captionTargetId="figure@4.[161,1426,193,1001]" captionTargetPageId="4" captionText="FIGURE 2. Cerogobius petrophilus sp. nov. Cephalic sensory papillae, lateral view of paratype PMR VP4501, female, 15.36 + 3.21 mm, and dorsal view of holotype, SMF 35961, female, 16.32 + 3.54 mm. Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/2589830/files/figure.png" pageId="2" pageNumber="173">Fig. 2</figureCitation>
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); 14) dorsal-fin pterygiophore insertion pattern 3-22110; 15) 10 precaudal vertebrae + 17 caudal vertebrae; 16) hypurals 1–2 and 3–4 not fused, distinctly separate along entire length; 17) two anal-fin pterygiophores inserted anterior to haemal arch.
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</paragraph>
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</subSubSection>
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<caption id="02F6BD89FF8DFFBCFF3AF84049A0DAEC" httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="3" pageNumber="174" startId="3.[151,250,1806,1828]" targetBox="[188,1398,190,1789]" targetPageId="3">
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<paragraph id="5636ED01FF8DFFBCFF3AF84049A0DAEC" blockId="3.[151,1436,1806,2012]" pageId="3" pageNumber="174">
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<emphasis id="64FD3113FF8DFFBCFF3AF8404934DA14" bold="true" box="[151,277,1806,1828]" pageId="3" pageNumber="174">FIGURE 1.</emphasis>
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<taxonomicName id="91899682FF8DFFBCFEB3F8404A2EDA14" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[286,527,1806,1828]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="174" phylum="Chordata" rank="species" species="petrophilus" status="sp. nov.">
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<emphasis id="64FD3113FF8DFFBCFEB3F8404A2EDA14" box="[286,527,1806,1828]" italics="true" pageId="3" pageNumber="174">Cerogobius petrophilus</emphasis>
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</taxonomicName>
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<emphasis id="64FD3113FF8DFFBCFDB5F8414A4ADA14" bold="true" box="[536,619,1807,1828]" pageId="3" pageNumber="174">
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<taxonomicNameLabel id="7FCE8C68FF8DFFBCFDB5F8414A4ADA14" box="[536,619,1807,1828]" pageId="3" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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:
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<emphasis id="64FD3113FF8DFFBCFDD6F8404AACDA13" bold="true" box="[635,653,1806,1827]" pageId="3" pageNumber="174">A</emphasis>
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: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm);
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<emphasis id="64FD3113FF8DFFBCFE02F80249E1DA51" bold="true" box="[431,448,1868,1889]" pageId="3" pageNumber="174">B</emphasis>
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: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm);
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<emphasis id="64FD3113FF8DFFBCFD02F8244AE0DAB0" bold="true" box="[687,705,1898,1920]" pageId="3" pageNumber="174">C</emphasis>
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: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained;
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<emphasis id="64FD3113FF8DFFBCFC83F8C74B61DAAE" bold="true" box="[814,832,1929,1950]" pageId="3" pageNumber="174">D</emphasis>
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: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić.
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</paragraph>
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</caption>
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<paragraph id="5636ED01FF8AFFBBFF3AFCB04DA9D903" blockId="4.[151,1436,1022,1075]" pageId="4" pageNumber="175">
|
||
<emphasis id="64FD3113FF8AFFBBFF3AFCB04933D923" bold="true" box="[151,274,1022,1044]" pageId="4" pageNumber="175">FIGURE 2.</emphasis>
|
||
<taxonomicName id="91899682FF8AFFBBFEB5FCB04A27D924" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[280,518,1022,1044]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="175" phylum="Chordata" rank="species" species="petrophilus" status="sp. nov.">
|
||
<emphasis id="64FD3113FF8AFFBBFEB5FCB04A27D924" box="[280,518,1022,1044]" italics="true" pageId="4" pageNumber="175">Cerogobius petrophilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FD3113FF8AFFBBFDA6FCB14A7AD924" bold="true" box="[523,603,1023,1044]" pageId="4" pageNumber="175">
|
||
<taxonomicNameLabel id="7FCE8C68FF8AFFBBFDA6FCB14A7AD924" box="[523,603,1023,1044]" pageId="4" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Cephalic sensory papillae, lateral view of paratype PMR VP4501, female, 15.36 + 3.21 mm, and dorsal view of holotype, SMF 35961, female, 16.32 + 3.54 mm. Terminology in text. Drawing by M. Kovačić.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="1E93BE8AFF8AFFBAFF6AFB144D4FDC70" lastPageId="5" lastPageNumber="176" pageId="4" pageNumber="175" type="description">
|
||
<paragraph id="5636ED01FF8AFFBAFF6AFB144D4FDC70" blockId="4.[151,1437,1114,2009]" lastBlockId="5.[151,1436,151,320]" lastPageId="5" lastPageNumber="176" pageId="4" pageNumber="175">
|
||
<emphasis id="64FD3113FF8AFFBBFF6AFB144970D943" bold="true" box="[199,337,1114,1139]" pageId="4" pageNumber="175">Description</emphasis>
|
||
. Body elongate, laterally compressed posteriorly, caudal peduncle short and deep, its depth 0.8– 0.9 of body depth at anal-fin origin, caudal peduncle 1.04–1.17 times deeper than its length (
|
||
<figureCitation id="CEB2F184FF8AFFBBFB06FB304CCED9A7" box="[1195,1263,1150,1175]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[159,1427,373,1730]" captionTargetId="figure@5.[159,1427,373,1730]" captionTargetPageId="5" captionText="FIGURE 3. Cerogobius petrophilus sp. nov. A: paratype, SMF 35963, female, 14.81 + 2.97 mm, freshly collected specimen, Thuwal, Saudi Arabia. Holotype, SMF 35961, female, 16.32 + 3.54 mm, Thuwal, Saudi Arabia, B: freshly collected specimen; C: preserved specimen. Photos by E.M. Troyer (A & B), M. Kovačić (C)." httpUri="https://zenodo.org/record/2589832/files/figure.png" pageId="4" pageNumber="175">Fig. 3</figureCitation>
|
||
). Head slightly depressed, head depth 0.9 of width to almost equal to width. Snout with steep profile from lateral view, very short, about 0.4-0.5 of eye diameter. Unpaired horn-like tentacle at the level of nostrils on the snout midline, length of tentacle slightly shorter than diameter of eye when measured from the anterior side. Anterior nostril a tube without process on rim, posterior nostril a short tube (
|
||
<figureCitation id="CEB2F184FF8AFFBBFD36FA404AD4D817" box="[667,757,1294,1319]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="4" pageNumber="175">Fig. 1A</figureCitation>
|
||
). Orbit projecting forward and upper part of orbit obviously elevated above dorsal profile of the head. Interorbital narrow, about 0.2 of eye diameter. No tentacle above eye. Mouth terminal, oblique. Posterior angle of jaws behind vertical through posterior margin of eye. Upper lip wider than cheek (
|
||
<figureCitation id="CEB2F184FF8AFFBBFEB2FA344990D8A3" box="[287,433,1402,1427]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="4" pageNumber="175">Figs. 1B, 1D</figureCitation>
|
||
), covered with a thin dermal flap, extending upwards over cheek and nearly reaching eye (
|
||
<figureCitation id="CEB2F184FF8AFFBBFF32FAD048D4D887" box="[159,245,1438,1463]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="4" pageNumber="175">Fig. 1B</figureCitation>
|
||
) (only in
|
||
<typeStatus id="893253A3FF8AFFBBFEC5FAD149E9D887" box="[360,456,1439,1463]" pageId="4" pageNumber="175" type="paratype">paratype</typeStatus>
|
||
SMF 35963 dermal flap shorter,
|
||
<figureCitation id="CEB2F184FF8AFFBBFCE9FAD04BBCD887" box="[836,925,1438,1463]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="4" pageNumber="175">Fig. 1D</figureCitation>
|
||
). Cheek very narrow. Teeth in both jaws in a single row posteriorly, becoming 3–4 rows anteriorly, with outer row as of enlarged recurved canines, and inner rows of smaller, pointed, and irregularly spaced teeth. Branchiostegal membranes fused to isthmus, gill openings restricted to pectoral-fin base. Lower limb of first gill arch joined to gill cover by membrane. Small mental frenum, at midventral behind lower lip. No spines on preopercle. Two dorsal fins, first fin with VI spines, not elongated or filamentous, second dorsal fin I,12–13. Pectoral-fin rays count low, 13–14 (usually 13). Pectoral girdle without flaps on anterior edge. Uppermost rays of pectoral fin within membrane. Pelvic fins I,5 + 5,I, left and right fin completely separated, lacking both an anterior frenum and membrane connecting innermost rays (
|
||
<figureCitation id="CEB2F184FF8AFFBBFB44F9D44D60DB83" box="[1257,1345,1690,1715]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="4" pageNumber="175">Fig. 1C</figureCitation>
|
||
); pelvic fins short, the length 0.6–0.8 of distance between origin of fins and anus. Caudal fin rounded, shorter than head length, with 17 segmented rays. Head and body completely scaleless. No head canals. Rows of head sensory papillae greatly reduced, with rows missing or reduced to one or a few papillae, and additional individual papillae present on the position of absent head canal pores. Preorbital rows
|
||
<emphasis id="64FD3113FF8AFFBBFC2BF8604BE9DA08" box="[902,968,1834,1861]" italics="true" pageId="4" pageNumber="175">
|
||
r, s, s
|
||
<superScript id="A1FC4049FF8AFFBBFC6DF8644BE9DA08" attach="left" box="[960,968,1834,1848]" fontSize="6" pageId="4" pageNumber="175">3</superScript>
|
||
</emphasis>
|
||
each reduced to single papilla. Suborbital rows of sensory papillae with rows
|
||
<emphasis id="64FD3113FF8AFFBBFD80F81C4A1ADA59" box="[557,571,1874,1897]" italics="true" pageId="4" pageNumber="175">a</emphasis>
|
||
and
|
||
<emphasis id="64FD3113FF8AFFBBFDD8F81F4AA4DA5A" box="[629,645,1873,1898]" italics="true" pageId="4" pageNumber="175">b</emphasis>
|
||
absent; four very short transverse suborbital rows of sensory papillae (
|
||
<emphasis id="64FD3113FF8AFFBBFF0DF83848EBDABE" box="[160,202,1910,1934]" italics="true" pageId="4" pageNumber="175">1–4</emphasis>
|
||
) present, row
|
||
<emphasis id="64FD3113FF8AFFBBFED9F83849A3DABE" box="[372,386,1910,1934]" italics="true" pageId="4" pageNumber="175">1</emphasis>
|
||
oblique, reaching upper lip, row
|
||
<emphasis id="64FD3113FF8AFFBBFCAAF8384B34DABE" box="[775,789,1910,1934]" italics="true" pageId="4" pageNumber="175">2</emphasis>
|
||
distant from eye, close to row
|
||
<emphasis id="64FD3113FF8AFFBBFB2EF83B4CB0DABE" box="[1155,1169,1909,1934]" italics="true" pageId="4" pageNumber="175">d</emphasis>
|
||
, rows
|
||
<emphasis id="64FD3113FF8AFFBBFB4CF8384CCEDABE" box="[1249,1263,1910,1934]" italics="true" pageId="4" pageNumber="175">3</emphasis>
|
||
and
|
||
<emphasis id="64FD3113FF8AFFBBFA86F8384D18DABE" box="[1323,1337,1910,1934]" italics="true" pageId="4" pageNumber="175">4</emphasis>
|
||
close to posteroventral edge of eye and distant from row
|
||
<emphasis id="64FD3113FF8AFFBBFD6BF8D74AF5DA82" box="[710,724,1945,1970]" italics="true" pageId="4" pageNumber="175">d</emphasis>
|
||
, row
|
||
<emphasis id="64FD3113FF8AFFBBFCB5F8D74B07DA82" box="[792,806,1945,1970]" italics="true" pageId="4" pageNumber="175">d</emphasis>
|
||
short, above posterior part of upper lip and continuing shortly on cheek. Oculoscapular rows reduced to rows
|
||
<emphasis id="64FD3113FF8AFFBBFCACF88F4B19DAFB" box="[769,824,1981,2008]" italics="true" pageId="4" pageNumber="175">
|
||
<date id="2237CBC1FF8AFFBBFCACF88F4B34DAFB" box="[769,789,1981,2008]" pageId="4" pageNumber="175">
|
||
x
|
||
<superScript id="A1FC4049FF8AFFBBFCA0F8F34B34DAFB" attach="left" box="[781,789,1981,1995]" fontSize="6" pageId="4" pageNumber="175">1</superScript>
|
||
</date>
|
||
,
|
||
<date id="2237CBC1FF8AFFBBFC8EF88F4B0EDAE8" box="[803,815,1985,2008]" pageId="4" pageNumber="175">x</date>
|
||
<superScript id="A1FC4049FF8AFFBBFC9DF8F34B19DAFB" attach="right" box="[816,824,1981,1995]" fontSize="6" pageId="4" pageNumber="175">2</superScript>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="64FD3113FF8AFFBBFCEBF88F4BA1DAFB" box="[838,896,1981,2008]" italics="true" pageId="4" pageNumber="175">
|
||
z, as
|
||
<superScript id="A1FC4049FF8AFFBBFCD5F8F34BA1DAFB" attach="left" box="[888,896,1981,1995]" fontSize="6" pageId="4" pageNumber="175">1</superScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="64FD3113FF8AFFBBFC19F88F4BF5DAFB" box="[948,980,1981,2008]" italics="true" pageId="4" pageNumber="175">
|
||
as
|
||
<superScript id="A1FC4049FF8AFFBBFC61F8F34BF5DAFB" attach="left" box="[972,980,1981,1995]" fontSize="6" pageId="4" pageNumber="175">2</superScript>
|
||
</emphasis>
|
||
, only
|
||
<date id="2237CBC1FF8AFFBBFBB1F88F4C11DAFB" box="[1052,1072,1981,2008]" pageId="4" pageNumber="175">
|
||
<emphasis id="64FD3113FF8AFFBBFBB1F88F4C11DAFB" box="[1052,1072,1981,2008]" italics="true" pageId="4" pageNumber="175">
|
||
x
|
||
<superScript id="A1FC4049FF8AFFBBFB85F8F34C11DAFB" attach="left" box="[1064,1072,1981,1995]" fontSize="6" pageId="4" pageNumber="175">1</superScript>
|
||
</emphasis>
|
||
</date>
|
||
of more than 2 papillae (
|
||
<figureCitation id="CEB2F184FF8AFFBBFAE4F88E4DAEDAE8" box="[1353,1423,1984,2009]" captionStart="FIGURE 2" captionStartId="4.[151,250,1022,1044]" captionTargetBox="[201,1426,297,947]" captionTargetId="figure@4.[161,1426,193,1001]" captionTargetPageId="4" captionText="FIGURE 2. Cerogobius petrophilus sp. nov. Cephalic sensory papillae, lateral view of paratype PMR VP4501, female, 15.36 + 3.21 mm, and dorsal view of holotype, SMF 35961, female, 16.32 + 3.54 mm. Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/2589830/files/figure.png" pageId="4" pageNumber="175">Fig. 2</figureCitation>
|
||
). Head and body to below origin of the second dorsal fin densely covered with very small, irregular, black spots some within blackish background, body behind the vertical of origin of the second dorsal fin pale with sparsely scattered melanophores; head with oblique, irregular, white band behind orbit. Dorsal-fin pterygiophore insertion pattern 3-22110. Vertebrae (including urostyle) 27 (precaudal + caudal vertebrae: 10 + 17). Hypurals 1–2 and 3–4 not fused, distinctly separate along entire length. Two anal-fin pterygiophores inserted anterior to haemal arch.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="02F6BD89FF8BFFBAFF3AF9994BA0DA1B" httpUri="https://zenodo.org/record/2589832/files/figure.png" pageId="5" pageNumber="176" startId="5.[151,250,1751,1773]" targetBox="[159,1427,373,1730]" targetPageId="5">
|
||
<paragraph id="5636ED01FF8BFFBAFF3AF9994BA0DA1B" blockId="5.[151,1436,1751,1835]" pageId="5" pageNumber="176">
|
||
<emphasis id="64FD3113FF8BFFBAFF3AF9994932DBDC" bold="true" box="[151,275,1751,1773]" pageId="5" pageNumber="176">FIGURE 3.</emphasis>
|
||
<taxonomicName id="91899682FF8BFFBAFEB6F9994A28DBDD" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[283,521,1751,1773]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="176" phylum="Chordata" rank="species" species="petrophilus" status="sp. nov.">
|
||
<emphasis id="64FD3113FF8BFFBAFEB6F9994A28DBDD" box="[283,521,1751,1773]" italics="true" pageId="5" pageNumber="176">Cerogobius petrophilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FD3113FF8BFFBAFDBCF9964A5DDBDC" bold="true" box="[529,636,1751,1773]" pageId="5" pageNumber="176">
|
||
<taxonomicNameLabel id="7FCE8C68FF8BFFBAFDBCF9964A43DBDD" box="[529,610,1752,1773]" pageId="5" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A
|
||
</emphasis>
|
||
: paratype, SMF 35963, female, 14.81 + 2.97 mm, freshly collected specimen, Thuwal, Saudi Arabia. Holotype, SMF 35961, female, 16.32 + 3.54 mm, Thuwal, Saudi Arabia,
|
||
<emphasis id="64FD3113FF8BFFBAFBC4F9B94C5BDA3C" bold="true" box="[1129,1146,1783,1804]" pageId="5" pageNumber="176">B</emphasis>
|
||
: freshly collected specimen;
|
||
<emphasis id="64FD3113FF8BFFBAFF3AF85B4888DA1B" bold="true" box="[151,169,1813,1835]" pageId="5" pageNumber="176">C</emphasis>
|
||
: preserved specimen. Photos by E.M. Troyer (A & B), M. Kovačić (C).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="1E93BE8AFF8BFFB9FF6AF8194BC1DCC4" lastPageId="6" lastPageNumber="177" pageId="5" pageNumber="176" type="discussion">
|
||
<paragraph id="5636ED01FF8BFFB9FF6AF8194BC1DCC4" blockId="5.[151,1436,1879,2012]" lastBlockId="6.[151,1436,151,1871]" lastPageId="6" lastPageNumber="177" pageId="5" pageNumber="176">
|
||
<emphasis id="64FD3113FF8BFFBAFF6AF8194915DA40" bold="true" box="[199,308,1879,1904]" pageId="5" pageNumber="176">Remarks</emphasis>
|
||
. Our phylogenetic analysis (
|
||
<figureCitation id="CEB2F184FF8BFFBAFDD0F8194AE3DA40" box="[637,706,1879,1904]" captionStart="FIGURE 4" captionStartId="7.[151,250,1946,1968]" captionTargetBox="[205,1292,205,1916]" captionTargetId="figure@7.[201,1389,193,1926]" captionTargetPageId="7" captionText="FIGURE 4. Bayesian inference molecular phylogeny of Gobiidae inferred from 5704 bp of combined nuclear genes and mtDNA. “Priolepis lineage” and “Gobiosomatini” refer to clades from Agorreta et al. (2013). Support values are posterior probabilities." httpUri="https://zenodo.org/record/2589834/files/figure.png" pageId="5" pageNumber="176">Fig. 4</figureCitation>
|
||
) shows that
|
||
<taxonomicName id="91899682FF8BFFBAFCF8F8194BF6DA40" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[853,983,1879,1904]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF8BFFBAFCF8F8194BF6DA40" box="[853,983,1879,1904]" italics="true" pageId="5" pageNumber="176">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
is closely related to
|
||
<taxonomicName id="91899682FF8BFFBAFB6AF8194D7BDA40" authorityName="Bleeker" authorityYear="1874" box="[1223,1370,1879,1904]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="176" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF8BFFBAFB6AF8194D7BDA40" box="[1223,1370,1879,1904]" italics="true" pageId="5" pageNumber="176">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
, with strong support. The clade is sister to the
|
||
<taxonomicName id="91899682FF8BFFBAFD1AF8354B29DAA4" box="[695,776,1915,1940]" class="Actinopterygii" family="Gobiidae" genus="Gobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="176" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF8BFFBAFD1AF8354B29DAA4" box="[695,776,1915,1940]" italics="true" pageId="5" pageNumber="176">Gobius</emphasis>
|
||
</taxonomicName>
|
||
-lineage
|
||
<emphasis id="64FD3113FF8BFFBAFCD8F8334B92DAA4" box="[885,947,1917,1940]" italics="true" pageId="5" pageNumber="176">sensu</emphasis>
|
||
<bibRefCitation id="321890F0FF8BFFBAFC68F8324CFEDAA4" author="Agorreta, A. & San Mauro, D. & Schliewen, U. & Van Tassell, J. L. & Kovacic, M. & Zardoya, R. & Ruber, L." box="[965,1247,1915,1940]" pageId="5" pageNumber="176" pagination="619 - 633" refId="ref8017" refString="Agorreta, A., San Mauro, D., Schliewen, U., Van Tassell, J. L., Kovacic, M., Zardoya, R. & Ruber, L. (2013) Molecular phylogenetics of Gobioidei and phylogenetic placement of European gobies. Molecular Phylogenetics and Evolution, 69, 619 - 633. https: // doi. org / 10.1016 / j. ympev. 2013.07.017" type="journal article" year="2013">
|
||
Agorreta
|
||
<emphasis id="64FD3113FF8BFFBAFB96F8334C59DAA4" box="[1083,1144,1915,1940]" italics="true" pageId="5" pageNumber="176">et al</emphasis>
|
||
. (2013)
|
||
</bibRefCitation>
|
||
.
|
||
<taxonomicName id="91899682FF8BFFBAFB58F8354D56DAA4" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[1269,1399,1915,1940]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF8BFFBAFB58F8354D56DAA4" box="[1269,1399,1915,1940]" italics="true" pageId="5" pageNumber="176">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
is morphologically clearly distinguished from
|
||
<taxonomicName id="91899682FF8BFFBAFD3CF8D14B05DA88" authorityName="Bleeker" authorityYear="1874" box="[657,804,1951,1976]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="176" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF8BFFBAFD3CF8D14B05DA88" box="[657,804,1951,1976]" italics="true" pageId="5" pageNumber="176">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
species and from genera in the
|
||
<taxonomicName id="91899682FF8BFFBAFB33F8D14CCEDA88" box="[1182,1263,1951,1976]" class="Actinopterygii" family="Gobiidae" genus="Gobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="176" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF8BFFBAFB33F8D14CCEDA88" box="[1182,1263,1951,1976]" italics="true" pageId="5" pageNumber="176">Gobius</emphasis>
|
||
</taxonomicName>
|
||
-lineage by the combination of characters listed in the genus diagnosis. Despite high diversity and morphological variability within the
|
||
<taxonomicName id="91899682FF88FFB9FF6FFFD94932DD80" box="[194,275,151,176]" class="Actinopterygii" family="Gobiidae" genus="Gobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FF6FFFD94932DD80" box="[194,275,151,176]" italics="true" pageId="6" pageNumber="177">Gobius</emphasis>
|
||
</taxonomicName>
|
||
-lineage, none of
|
||
<taxonomicName id="91899682FF88FFB9FE73FFD94A0EDD80" box="[478,559,151,176]" class="Actinopterygii" family="Gobiidae" genus="Gobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FE73FFD94A0EDD80" box="[478,559,151,176]" italics="true" pageId="6" pageNumber="177">Gobius</emphasis>
|
||
</taxonomicName>
|
||
-lineage species have an unpaired, horn-like tentacle at snout midline, at the level of nostrils (character
|
||
<quantity id="917140E4FF88FFB9FE67FFF249D6DDE5" box="[458,503,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="6" pageNumber="177" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="91899682FF88FFB9FE52FFF24AA0DDE5" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[511,641,188,213]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FE52FFF24AA0DDE5" box="[511,641,188,213]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
diagnosis), and none have suborbital transverse rows reduced to four rows (character 12) or lack row
|
||
<emphasis id="64FD3113FF88FFB9FE53FF914A2DDDC8" box="[510,524,223,248]" italics="true" pageId="6" pageNumber="177">b</emphasis>
|
||
(character 12) (
|
||
<bibRefCitation id="321890F0FF88FFB9FD12FF914B68DDC8" author="Miller, P. J." box="[703,841,223,248]" pageId="6" pageNumber="177" pagination="1019 - 1085" refId="ref9016" refString="Miller, P. J. (1986) Gobiidae. In: Whitehead, P. J. P., Bauchot, M. - L., Hureau, J. - C., Nielsen, J. & Tortonese, E. (Eds.), Fishes of the North-eastern Atlantic and the Mediterranean 3. UNESCO, Paris, pp. 1019 - 1085." type="book chapter" year="1986">Miller 1986</bibRefCitation>
|
||
). The lower limb of first gill arch joined to gill cover by membrane (character 5) has to date not been reported for any
|
||
<taxonomicName id="91899682FF88FFB9FCCAFE4A4B99DC2D" box="[871,952,260,285]" class="Actinopterygii" family="Gobiidae" genus="Gobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FCCAFE4A4B99DC2D" box="[871,952,260,285]" italics="true" pageId="6" pageNumber="177">Gobius</emphasis>
|
||
</taxonomicName>
|
||
-lineage genera except
|
||
<taxonomicName id="91899682FF88FFB9FB14FE4A4D6BDC2D" authorityName="Smith" authorityYear="1958" box="[1209,1354,260,285]" class="Actinopterygii" family="Gobiidae" genus="Coryogalops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FB14FE4A4D6BDC2D" box="[1209,1354,260,285]" italics="true" pageId="6" pageNumber="177">Coryogalops</emphasis>
|
||
</taxonomicName>
|
||
, which was mentioned without original data or references in
|
||
<bibRefCitation id="321890F0FF88FFB9FC89FE664BE4DC70" author="Hoese, D. F." box="[804,965,295,320]" pageId="6" pageNumber="177" pagination="1 - 25" refId="ref8592" refString="Hoese, D. F. (1986) Descriptions of two new species of Hetereleotris (Pisces: Gobiidae) from the western Indian Ocean, with discussion of related species. J. L. B. Smith Institute of Ichthyology, Special Publication, 41, 1 - 25." type="journal article" year="1986">Hoese (1986)</bibRefCitation>
|
||
. Some of the diagnostic characters for
|
||
<taxonomicName id="91899682FF88FFB9FF3AFE024938DC55" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[151,281,332,357]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FF3AFE024938DC55" box="[151,281,332,357]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
(e.g. characters 8, 9, 10 but also others) rarely appears among species of the
|
||
<taxonomicName id="91899682FF88FFB9FB3FFE024CC2DC55" box="[1170,1251,332,357]" class="Actinopterygii" family="Gobiidae" genus="Gobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FB3FFE024CC2DC55" box="[1170,1251,332,357]" italics="true" pageId="6" pageNumber="177">Gobius</emphasis>
|
||
</taxonomicName>
|
||
-lineage, but are never combined in a single species. The combination of 3-22110 dorsal-fin pterygiophore insertion pattern and 10 + 17 vertebral count in
|
||
<taxonomicName id="91899682FF88FFB9FE0EFEDA4A04DC9D" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[419,549,404,429]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FE0EFEDA4A04DC9D" box="[419,549,404,429]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
differentiate it from many genera in the
|
||
<taxonomicName id="91899682FF88FFB9FC58FEDA4C7FDC9D" box="[1013,1118,404,429]" class="Actinopterygii" family="Gobiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
|
||
, although several species of
|
||
<taxonomicName id="91899682FF88FFB9FF3AFEF9490BDCE0" authorityName="Bleeker" authorityYear="1874" box="[151,298,439,464]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FF3AFEF9490BDCE0" box="[151,298,439,464]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
, several genera in the
|
||
<taxonomicName id="91899682FF88FFB9FD8FFEF94A53DCE0" box="[546,626,439,464]" class="Actinopterygii" family="Gobiidae" genus="Gobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FD8FFEF94A53DCE0" box="[546,626,439,464]" italics="true" pageId="6" pageNumber="177">Gobius</emphasis>
|
||
</taxonomicName>
|
||
-lineage (e.g.
|
||
<taxonomicName id="91899682FF88FFB9FCA1FEF94BAEDCE0" authorityName="Miller" authorityYear="1966" box="[780,911,439,464]" class="Actinopterygii" family="Gobiidae" genus="Didogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FCA1FEF94BAEDCE0" box="[780,911,439,464]" italics="true" pageId="6" pageNumber="177">Didogobius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="91899682FF88FFB9FC31FEF94C60DCE0" authorityName="de Buen" authorityYear="1930" box="[924,1089,439,464]" class="Actinopterygii" family="Gobiidae" genus="Chromogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FC31FEF94C60DCE0" box="[924,1089,439,464]" italics="true" pageId="6" pageNumber="177">Chromogobius</emphasis>
|
||
</taxonomicName>
|
||
) and a few other gobiid genera also possess this combination (
|
||
<bibRefCitation id="321890F0FF88FFB9FE5DFE924A5BDCC5" author="Hoese, D. F." box="[496,634,476,501]" pageId="6" pageNumber="177" pagination="1 - 25" refId="ref8592" refString="Hoese, D. F. (1986) Descriptions of two new species of Hetereleotris (Pisces: Gobiidae) from the western Indian Ocean, with discussion of related species. J. L. B. Smith Institute of Ichthyology, Special Publication, 41, 1 - 25." type="journal article" year="1986">Hoese 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="321890F0FF88FFB9FD2BFE924B4CDCC4" author="Birdsong, R. S. & Murdy, E. O. & Pezold, F. L." box="[646,877,476,501]" pageId="6" pageNumber="177" pagination="174 - 214" refId="ref8094" refString="Birdsong, R. S., Murdy, E. O. & Pezold, F. L. (1988) A study of the vertebral column and median fish osteology in gobioid fishes with comments on gobioid relationships. Bulletin of Marine Science, 42, 174 - 214." type="journal article" year="1988">
|
||
Birdsong
|
||
<emphasis id="64FD3113FF88FFB9FD5EFE934B05DCC5" box="[755,804,476,501]" italics="true" pageId="6" pageNumber="177">et al</emphasis>
|
||
. 1988
|
||
</bibRefCitation>
|
||
, table 11).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E93BE8AFF88FFB9FF6AFEB14B5FDA7F" pageId="6" pageNumber="177" type="reference_group">
|
||
<paragraph id="5636ED01FF88FFB9FF6AFEB14996D920" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="177">
|
||
<taxonomicName id="91899682FF88FFB9FF6AFEB14968DF28" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[199,329,511,536]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FF6AFEB14968DF28" box="[199,329,511,536]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
shares several external characters with
|
||
<taxonomicName id="91899682FF88FFB9FCBEFEB14B87DF28" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[787,934,511,536]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FCBEFEB14B87DF28" box="[787,934,511,536]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
that occur broadly across the
|
||
<taxonomicName id="91899682FF88FFB9FAA9FEB14D4ADF28" ID-CoL="623LH" box="[1284,1387,511,536]" class="Actinopterygii" family="Gobiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
|
||
and are less useful in diagnosing
|
||
<taxonomicName id="91899682FF88FFB9FE4BFD6A4A49DF0D" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[486,616,548,573]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FE4BFD6A4A49DF0D" box="[486,616,548,573]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
, and thus are not included in the present
|
||
<taxonomicName id="91899682FF88FFB9FBE5FD6A4CEBDF0D" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[1096,1226,548,573]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FBE5FD6A4CEBDF0D" box="[1096,1226,548,573]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
diagnosis, despite some being used to diagnose
|
||
<taxonomicName id="91899682FF88FFB9FE46FD094A5FDF50" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[491,638,583,608]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FE46FD094A5FDF50" box="[491,638,583,608]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
by Kovačić & Bogorodsky (2014) and
|
||
<bibRefCitation id="321890F0FF88FFB9FBE7FD094D6EDF50" author="Kovacic, M. & Bogorodsky, S. V. & Mal, A. O." box="[1098,1359,583,608]" pageId="6" pageNumber="177" pagination="119 - 126" refId="ref8826" refString="Kovacic, M., Bogorodsky, S. V. & Mal, A. O. (2014 a) A new species of Hetereleotris (Perciformes: Gobiidae) from Farasan Island (Red Sea). Zootaxa, 3846 (1), 119 - 126. https: // doi. org / 10.11646 / zootaxa. 3846.1.6" type="journal article" year="2014">
|
||
Kovačić
|
||
<emphasis id="64FD3113FF88FFB9FB1DFD074CC2DF50" box="[1200,1251,583,608]" italics="true" pageId="6" pageNumber="177">et al</emphasis>
|
||
. (2014a)
|
||
</bibRefCitation>
|
||
: broad isthmus and gill membrane attached to the base of the pectoral fin, first dorsal fin with VI spines, segmented caudal-fin rays 17; posterior nostril a tube of variable length (among
|
||
<taxonomicName id="91899682FF88FFB9FC31FDC14C0EDF98" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[924,1071,655,680]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FC31FDC14C0EDF98" box="[924,1071,655,680]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
species only in
|
||
<taxonomicName id="91899682FF88FFB9FB4FFDDF490FDFFD" ID-CoL="6LR9K" authority="(Smith 1958)" authorityName="Smith" authorityYear="1958" baseAuthorityName="Smith" baseAuthorityYear="1958" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="species" species="zanzibarensis">
|
||
<emphasis id="64FD3113FF88FFB9FB4FFDDF4DBADF98" box="[1250,1435,655,680]" italics="true" pageId="6" pageNumber="177">H. zanzibarensis</emphasis>
|
||
(
|
||
<bibRefCitation id="321890F0FF88FFB9FF33FDFA4907DFFD" author="Smith, J. L. B." box="[158,294,692,717]" pageId="6" pageNumber="177" pagination="137 - 163" refId="ref9273" refString="Smith, J. L. B. (1958) The fishes of the family Eleotridae in the western Indian Ocean. Ichthyological Bulletin, Department of Ichthyology, Rhodes University, 11, 137 - 163." type="journal article" year="1958">Smith 1958</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
posterior nostril subtubular).
|
||
<taxonomicName id="91899682FF88FFB9FD24FDFA4B2ADFFD" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[649,779,692,717]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FD24FDFA4B2ADFFD" box="[649,779,692,717]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
also shares with
|
||
<taxonomicName id="91899682FF88FFB9FC75FDFA4C4ADFFD" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[984,1131,692,717]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FC75FDFA4C4ADFFD" box="[984,1131,692,717]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
characters that are rare in
|
||
<taxonomicName id="91899682FF88FFB9FF3AFD9948DFDFC0" ID-CoL="623LH" box="[151,254,727,752]" class="Actinopterygii" family="Gobiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
|
||
but not exclusive only to
|
||
<taxonomicName id="91899682FF88FFB9FD9EFD994AE7DFC0" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[563,710,727,752]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FD9EFD994AE7DFC0" box="[563,710,727,752]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="91899682FF88FFB9FCAEFD994BA4DFC0" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[771,901,727,752]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FCAEFD994BA4DFC0" box="[771,901,727,752]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
: pelvic fins completely separated and without frenum (only
|
||
<taxonomicName id="91899682FF88FFB9FE9CFDB24A40DE25" ID-CoL="6LR9K" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[305,609,764,789]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="species" species="zanzibarensis">
|
||
<emphasis id="64FD3113FF88FFB9FE9CFDB24A40DE25" box="[305,609,764,789]" italics="true" pageId="6" pageNumber="177">Hetereleotris zanzibarensis</emphasis>
|
||
</taxonomicName>
|
||
exhibits variability in this character); head and body completely scaleless (some species of
|
||
<taxonomicName id="91899682FF88FFB9FEF5FC5149CADE08" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[344,491,799,824]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FEF5FC5149CADE08" box="[344,491,799,824]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
are scaled or partly scaled), no head canals (there are head canals in some species of
|
||
<taxonomicName id="91899682FF88FFB9FF3AFC0A490BDE6D" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[151,298,836,861]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FF3AFC0A490BDE6D" box="[151,298,836,861]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
) (characters 7, 8, 9, 10 in
|
||
<taxonomicName id="91899682FF88FFB9FDDBFC0A4AD9DE6D" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[630,760,836,861]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FDDBFC0A4AD9DE6D" box="[630,760,836,861]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
diagnosis respectively). At present, without new data of sufficient quality for these characters from many other
|
||
<taxonomicName id="91899682FF88FFB9FC87FC294BB0DEB0" ID-CoL="623LH" box="[810,913,871,896]" class="Actinopterygii" family="Gobiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
|
||
genera, synapomorphies that are unique for
|
||
<taxonomicName id="91899682FF88FFB9FF3AFCC2490BDE95" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[151,298,908,933]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FF3AFCC2490BDE95" box="[151,298,908,933]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="91899682FF88FFB9FECCFCC249C2DE95" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[353,483,908,933]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FECCFCC249C2DE95" box="[353,483,908,933]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
are: lower limb of first gill arch joined to gill cover by membrane and the suborbital papillae pattern with four transverse suborbital rows (present in all
|
||
<taxonomicName id="91899682FF88FFB9FC61FCE14C7EDEF8" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[972,1119,943,968]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FC61FCE14C7EDEF8" box="[972,1119,943,968]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
species with described or illustrated cephalic sensory systems, except
|
||
<taxonomicName id="91899682FF88FFB9FD24FC9B4B07DEDD" ID-CoL="3L7HZ" authorityName="Gill" authorityYear="1998" box="[649,806,980,1005]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="species" species="georgegilli">
|
||
<emphasis id="64FD3113FF88FFB9FD24FC9B4B07DEDD" box="[649,806,980,1005]" italics="true" pageId="6" pageNumber="177">H. georgegilli</emphasis>
|
||
</taxonomicName>
|
||
with six rows) (character 5 and the part of character
|
||
<quantity id="917140E4FF88FFB9FA2DFC9A488CD920" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="6" pageNumber="177" unit="in" value="12.0">12 in</quantity>
|
||
<taxonomicName id="91899682FF88FFB9FF19FCB94917D920" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[180,310,1015,1040]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FF19FCB94917D920" box="[180,310,1015,1040]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
diagnosis).
|
||
</paragraph>
|
||
<paragraph id="5636ED01FF88FFB9FF6AFB524952DB1F" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="177">
|
||
<taxonomicName id="91899682FF88FFB9FF6AFB524968D905" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[199,329,1052,1077]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FF6AFB524968D905" box="[199,329,1052,1077]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
differs from all species presently recognized in
|
||
<taxonomicName id="91899682FF88FFB9FCC9FB524BD6D905" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[868,1015,1052,1077]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FCC9FB524BD6D905" box="[868,1015,1052,1077]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
(except
|
||
<taxonomicName id="91899682FF88FFB9FBFAFB534DBDD905" ID-CoL="3L7J9" authority="(Smith 1958))" authorityName="" authorityYear="1958" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[1111,1436,1052,1077]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="64FD3113FF88FFB9FBFAFB534CD6D905" box="[1111,1271,1052,1077]" italics="true" pageId="6" pageNumber="177">H. tentaculata</emphasis>
|
||
<bibRefCitation id="321890F0FF88FFB9FB53FB524DBDD905" author="Smith, J. L. B." box="[1278,1436,1052,1077]" pageId="6" pageNumber="177" pagination="137 - 163" refId="ref9273" refString="Smith, J. L. B. (1958) The fishes of the family Eleotridae in the western Indian Ocean. Ichthyological Bulletin, Department of Ichthyology, Rhodes University, 11, 137 - 163." type="journal article" year="1958">(Smith 1958))</bibRefCitation>
|
||
</taxonomicName>
|
||
by the following: unpaired horn-like tentacle at the level of nostrils in the middle of snout (
|
||
<figureCitation id="CEB2F184FF88FFB9FB24FB714CFFD968" box="[1161,1246,1087,1112]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="6" pageNumber="177">Fig. 1A</figureCitation>
|
||
)
|
||
<emphasis id="64FD3113FF88FFB9FB43FB0F4D2CD968" box="[1262,1293,1089,1112]" italics="true" pageId="6" pageNumber="177">vs.</emphasis>
|
||
no horn-like tentacle on snout in
|
||
<taxonomicName id="91899682FF88FFB9FE28FB2A4A39D94D" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[389,536,1124,1149]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FE28FB2A4A39D94D" box="[389,536,1124,1149]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
; snout with steep profile in lateral view and very short, about 0.4–0.5 of eye diameter
|
||
<emphasis id="64FD3113FF88FFB9FEACFBC7493ED990" box="[257,287,1161,1184]" italics="true" pageId="6" pageNumber="177">vs.</emphasis>
|
||
snout moderately steep and always longer than half of eye diameter in
|
||
<taxonomicName id="91899682FF88FFB9FBE6FBC94CFFD990" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[1099,1246,1159,1184]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FBE6FBC94CFFD990" box="[1099,1246,1159,1184]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
; orbit projecting forward and elevated above dorsal profile
|
||
<emphasis id="64FD3113FF88FFB9FDDAFBE34AB7D9F4" box="[631,662,1197,1220]" italics="true" pageId="6" pageNumber="177">vs.</emphasis>
|
||
orbit not projecting forward and only slightly elevated above dorsal profile in some species of
|
||
<taxonomicName id="91899682FF88FFB9FE71FB814A4ED9D8" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[476,623,1231,1256]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FE71FB814A4ED9D8" box="[476,623,1231,1256]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
; mouth moderately large, posterior angle of jaws behind vertical at posterior edge of eye
|
||
<emphasis id="64FD3113FF88FFB9FE27FBBB4988D83C" box="[394,425,1269,1292]" italics="true" pageId="6" pageNumber="177">vs.</emphasis>
|
||
posterior angle of jaws below eye; caudal peduncle deep and short, the depth greater than its length
|
||
<emphasis id="64FD3113FF88FFB9FEA4FA574909D800" box="[265,296,1305,1328]" italics="true" pageId="6" pageNumber="177">vs.</emphasis>
|
||
caudal peduncle depth clearly less than length in
|
||
<taxonomicName id="91899682FF88FFB9FCC5FA594BDAD800" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[872,1019,1303,1328]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FCC5FA594BDAD800" box="[872,1019,1303,1328]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
; suborbital rows of sensory papillae with row
|
||
<emphasis id="64FD3113FF88FFB9FEAFFA724931D865" box="[258,272,1340,1365]" italics="true" pageId="6" pageNumber="177">b</emphasis>
|
||
absent
|
||
<emphasis id="64FD3113FF88FFB9FEC8FA7349A2D864" box="[357,387,1341,1364]" italics="true" pageId="6" pageNumber="177">vs.</emphasis>
|
||
row
|
||
<emphasis id="64FD3113FF88FFB9FE10FA7249EAD865" box="[445,459,1340,1365]" italics="true" pageId="6" pageNumber="177">b</emphasis>
|
||
present in
|
||
<taxonomicName id="91899682FF88FFB9FDEBFA724AF8D865" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[582,729,1340,1365]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FDEBFA724AF8D865" box="[582,729,1340,1365]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
; four very short transverse suborbital rows of sensory papillae, row
|
||
<emphasis id="64FD3113FF88FFB9FF67FA2E48F9D848" box="[202,216,1376,1400]" italics="true" pageId="6" pageNumber="177">2</emphasis>
|
||
distant from eye, close to row
|
||
<emphasis id="64FD3113FF88FFB9FD99FA114A63D848" box="[564,578,1375,1400]" italics="true" pageId="6" pageNumber="177">d</emphasis>
|
||
, rows
|
||
<emphasis id="64FD3113FF88FFB9FD23FA2E4ABDD848" box="[654,668,1376,1400]" italics="true" pageId="6" pageNumber="177">3</emphasis>
|
||
and
|
||
<emphasis id="64FD3113FF88FFB9FD7EFA2E4AC0D848" box="[723,737,1376,1400]" italics="true" pageId="6" pageNumber="177">4</emphasis>
|
||
close to posteroventral edge of eye and distant from row
|
||
<emphasis id="64FD3113FF88FFB9FAC5FA114DBDD848" box="[1384,1436,1375,1400]" italics="true" pageId="6" pageNumber="177">d vs.</emphasis>
|
||
four transverse suborbital rows well developed over cheek, and row
|
||
<emphasis id="64FD3113FF88FFB9FC32FACA4B8CD8AC" box="[927,941,1412,1436]" italics="true" pageId="6" pageNumber="177">3</emphasis>
|
||
always close and perpendicular to row
|
||
<emphasis id="64FD3113FF88FFB9FADDFACA4D5FD8AD" box="[1392,1406,1412,1437]" italics="true" pageId="6" pageNumber="177">d</emphasis>
|
||
in
|
||
<taxonomicName id="91899682FF88FFB9FF3AFAE4490BD8F3" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[151,298,1450,1475]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FF3AFAE4490BD8F3" box="[151,298,1450,1475]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
; oculoscapular rows reduced, with only
|
||
<date id="2237CBC1FF88FFB9FCBBFAE24B0AD885" box="[790,811,1447,1475]" pageId="6" pageNumber="177">
|
||
<emphasis id="64FD3113FF88FFB9FCBBFAE24B0AD885" box="[790,811,1447,1475]" italics="true" pageId="6" pageNumber="177">
|
||
x
|
||
<superScript id="A1FC4049FF88FFB9FC8EFAE94B0AD885" attach="left" box="[803,811,1447,1461]" fontSize="6" pageId="6" pageNumber="177">1</superScript>
|
||
</emphasis>
|
||
</date>
|
||
of more than 2 papillae (
|
||
<figureCitation id="CEB2F184FF88FFB9FBC9FAE44C8FD8F3" box="[1124,1198,1450,1475]" captionStart="FIGURE 2" captionStartId="4.[151,250,1022,1044]" captionTargetBox="[201,1426,297,947]" captionTargetId="figure@4.[161,1426,193,1001]" captionTargetPageId="4" captionText="FIGURE 2. Cerogobius petrophilus sp. nov. Cephalic sensory papillae, lateral view of paratype PMR VP4501, female, 15.36 + 3.21 mm, and dorsal view of holotype, SMF 35961, female, 16.32 + 3.54 mm. Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/2589830/files/figure.png" pageId="6" pageNumber="177">Fig. 2</figureCitation>
|
||
)
|
||
<emphasis id="64FD3113FF88FFB9FB69FAE24CC3D8F3" box="[1220,1250,1452,1475]" italics="true" pageId="6" pageNumber="177">vs.</emphasis>
|
||
more than one oculoscapular rows with several papillae in
|
||
<taxonomicName id="91899682FF88FFB9FD24FA804B3DD8D7" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[649,796,1486,1511]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FD24FA804B3DD8D7" box="[649,796,1486,1511]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
(characters 1, 2, 3, 4, 6, part of 12, 13 from
|
||
<taxonomicName id="91899682FF88FFB9FAB7FA804DBDD8D7" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[1306,1436,1486,1511]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FAB7FA804DBDD8D7" box="[1306,1436,1486,1511]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
diagnosis respectively; note that only seven
|
||
<taxonomicName id="91899682FF88FFB9FD28FABC4B39DB3B" ID-CoL="62V3V" authorityName="Bleeker" authorityYear="1874" box="[645,792,1522,1547]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="genus">
|
||
<emphasis id="64FD3113FF88FFB9FD28FABC4B39DB3B" box="[645,792,1522,1547]" italics="true" pageId="6" pageNumber="177">Hetereleotris</emphasis>
|
||
</taxonomicName>
|
||
species had available data or at least rough illustration of lateral line system).
|
||
</paragraph>
|
||
<paragraph id="5636ED01FF88FFB9FF6AF9744B5FDA7F" blockId="6.[151,1436,151,1871]" pageId="6" pageNumber="177">
|
||
<taxonomicName id="91899682FF88FFB9FF6AF9744968DB63" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[199,329,1594,1619]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF88FFB9FF6AF9744968DB63" box="[199,329,1594,1619]" italics="true" pageId="6" pageNumber="177">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="91899682FF88FFB9FE71F9724A5CDB63" ID-CoL="3L7J9" authorityName="" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[476,637,1594,1619]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="64FD3113FF88FFB9FE71F9724A5CDB63" box="[476,637,1594,1619]" italics="true" pageId="6" pageNumber="177">H. tentaculata</emphasis>
|
||
</taxonomicName>
|
||
in the shape of head (i.e. head with very short snout, orbit far forward, and posterior angle of mouth behind a vertical at posterior edge of eye), having a scaleless head and body, and in having a short and deep caudal peduncle. It differs from
|
||
<taxonomicName id="91899682FF88FFB9FCBBF9CA4B96DBAB" ID-CoL="3L7J9" authorityName="" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[790,951,1666,1691]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="64FD3113FF88FFB9FCBBF9CA4B96DBAB" box="[790,951,1666,1691]" italics="true" pageId="6" pageNumber="177">H. tentaculata</emphasis>
|
||
</taxonomicName>
|
||
in having an unpaired horn-like tentacle at the level of nostrils on snout midline (
|
||
<figureCitation id="CEB2F184FF88FFB9FD93F9E84AB5DB8F" box="[574,660,1702,1727]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="6" pageNumber="177">Fig. 1A</figureCitation>
|
||
)
|
||
<emphasis id="64FD3113FF88FFB9FD09F9E64AE2DB8F" box="[676,707,1704,1727]" italics="true" pageId="6" pageNumber="177">vs.</emphasis>
|
||
no tentacle on snout in
|
||
<taxonomicName id="91899682FF88FFB9FC61F9E64C4DDB8F" ID-CoL="3L7J9" authorityName="" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[972,1132,1702,1727]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="64FD3113FF88FFB9FC61F9E64C4DDB8F" box="[972,1132,1702,1727]" italics="true" pageId="6" pageNumber="177">H. tentaculata</emphasis>
|
||
</taxonomicName>
|
||
; no cirri or tentacles above eye
|
||
<emphasis id="64FD3113FF88FFB9FF68F98248C2DBD3" box="[197,227,1740,1763]" italics="true" pageId="6" pageNumber="177">vs.</emphasis>
|
||
large fleshy tentacle above each eye, 0.8 of eye diameter, with trilobate apex in
|
||
<taxonomicName id="91899682FF88FFB9FBC7F9824D2ADBD3" ID-CoL="3L7J9" authorityName="" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[1130,1291,1738,1763]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="64FD3113FF88FFB9FBC7F9824D2ADBD3" box="[1130,1291,1738,1763]" italics="true" pageId="6" pageNumber="177">H. tentaculata</emphasis>
|
||
</taxonomicName>
|
||
; head canals absent
|
||
<emphasis id="64FD3113FF88FFB9FF44F9BE4929DA37" box="[233,264,1776,1799]" italics="true" pageId="6" pageNumber="177">vs.</emphasis>
|
||
head canals present in
|
||
<taxonomicName id="91899682FF88FFB9FD8DF9BE4AE5DA37" ID-CoL="3L7J9" authorityName="" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[544,708,1774,1799]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="64FD3113FF88FFB9FD8DF9BE4AE5DA37" box="[544,708,1774,1799]" italics="true" pageId="6" pageNumber="177">H. tentaculata</emphasis>
|
||
</taxonomicName>
|
||
; posterior nostril a short tube, shorter than anterior nostril
|
||
<emphasis id="64FD3113FF88FFB9FAD0F9BE4DBDDA37" box="[1405,1436,1776,1799]" italics="true" pageId="6" pageNumber="177">vs.</emphasis>
|
||
posterior tube elongate, much longer than anterior nostril, about half eye diameter in
|
||
<taxonomicName id="91899682FF88FFB9FB28F85A4D08DA1B" ID-CoL="3L7J9" authorityName="" baseAuthorityName="Smith" baseAuthorityYear="1958" box="[1157,1321,1810,1835]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="177" phylum="Chordata" rank="species" species="tentaculata">
|
||
<emphasis id="64FD3113FF88FFB9FB28F85A4D08DA1B" box="[1157,1321,1810,1835]" italics="true" pageId="6" pageNumber="177">H. tentaculata</emphasis>
|
||
</taxonomicName>
|
||
. The two species also differ in fin meristics, morphometrics, and coloration.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |