570 lines
80 KiB
XML
570 lines
80 KiB
XML
<document id="A40654A88A40790EBFA5856F7ACB18FD" ID-CLB-Dataset="28397" ID-DOI="10.11646/zootaxa.4565.2.2" ID-GBIF-Dataset="b289d291-5da8-48a8-b283-a456bce4ec76" ID-ISSN="1175-5326" ID-Zenodo-Dep="2589826" ID-ZooBank="1E01FCF7-0D6D-4E8C-BDF2-E2D8068934B5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552279673760" checkinUser="plazi" docAuthor="Kovačić, Marcelo, Bogorodsky, Sergey V., Troyer, Emily M. & Tornabene, Luke" docDate="2019" docId="DE205C17FF86FFB1FF3AFFD94A77DDC3" docLanguage="en" docName="zootaxa.4565.2.2.pdf" docOrigin="Zootaxa 4565 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cerogobius petrophilus Kovačić & Bogorodsky & Troyer & Tornabene 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="185" masterDocId="2219246FFF8EFFBFFFADFF4E4821DD30" masterDocTitle="Cerogobius petrophilus (Perciformes: Gobiidae), a new gobiid genus and species from the Red Sea" masterLastPageNumber="189" masterPageNumber="171" pageNumber="179" updateTime="1698701428881" updateUser="ExternalLinkService">
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<mods:mods id="160B644F391004CE10EACD41D613EEA9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="FDC81304AE1CEC09191B5878EA920326">Cerogobius petrophilus (Perciformes: Gobiidae), a new gobiid genus and species from the Red Sea</mods:title>
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</mods:titleInfo>
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<mods:name id="0EC4DF4E687803F739B8638E2494CF2B" type="personal">
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<mods:namePart id="76AE13AD755C37B1912CA723BD2CF2F8">Kovačić, Marcelo</mods:namePart>
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<mods:namePart id="6120407E7CA7271DF61A98407F8D578E">Bogorodsky, Sergey V.</mods:namePart>
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<mods:namePart id="767DD10D8889355781C1928C5445A9F3">Troyer, Emily M.</mods:namePart>
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<mods:namePart id="2B7453BFCE71AB682816FA3C3FFC3917">Tornabene, Luke</mods:namePart>
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<mods:typeOfResource id="B041A59EFE9FED7B1C84C1C6877633D9">text</mods:typeOfResource>
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<mods:title id="4BE6DBB8CD0B17B6C744D5DA27129A6C">Zootaxa</mods:title>
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<mods:part id="DE19B8071D3FCAD01C5589B512B0012D">
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<mods:date id="14BE81F7764A1CEBC47A385E94890580">2019</mods:date>
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<mods:number id="42D2DB76EF3D4E79C43F71026635FF72">2019-03-08</mods:number>
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<mods:number id="284DA7C5F6D90DECE234DAD37492BB89">4565</mods:number>
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<treatment id="DE205C17FF86FFB1FF3AFFD94A77DDC3" ID-DOI="http://doi.org/10.5281/zenodo.5928731" ID-GBIF-Taxon="154366882" ID-Zenodo-Dep="5928731" LSID="urn:lsid:plazi:treatment:DE205C17FF86FFB1FF3AFFD94A77DDC3" httpUri="http://treatment.plazi.org/id/DE205C17FF86FFB1FF3AFFD94A77DDC3" lastPageId="14" lastPageNumber="185" pageId="8" pageNumber="179">
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<subSubSection id="1E93BE8AFF86FFB7FF3AFFD9490BDDE3" pageId="8" pageNumber="179" type="nomenclature">
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<paragraph id="5636ED01FF86FFB7FF3AFFD94A37DD82" blockId="8.[151,534,151,247]" box="[151,534,151,178]" pageId="8" pageNumber="179">
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<heading id="0D7E5A6DFF86FFB7FF3AFFD94A37DD82" bold="true" box="[151,534,151,178]" fontSize="11" level="1" pageId="8" pageNumber="179" reason="1">
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<emphasis id="64FD3113FF86FFB7FF3AFFD94A37DD82" bold="true" box="[151,534,151,178]" pageId="8" pageNumber="179">
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<taxonomicName id="91899682FF86FFB7FF3AFFD94990DD81" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[151,433,151,177]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="179" phylum="Chordata" rank="species" species="petrophilus" status="sp. nov.">
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<emphasis id="64FD3113FF86FFB7FF3AFFD94990DD81" bold="true" box="[151,433,151,177]" italics="true" pageId="8" pageNumber="179">Cerogobius petrophilus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7FCE8C68FF86FFB7FE14FFD64A37DD82" box="[441,534,152,178]" pageId="8" pageNumber="179" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="5636ED01FF86FFB7FF3AFFF4490BDDE3" blockId="8.[151,534,151,247]" box="[151,298,186,211]" pageId="8" pageNumber="179">Horned goby</paragraph>
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</subSubSection>
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<subSubSection id="1E93BE8AFF86FFB7FF3AFF9049BCDDC7" box="[151,413,222,247]" pageId="8" pageNumber="179" type="description">
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<paragraph id="5636ED01FF86FFB7FF3AFF9049BCDDC7" blockId="8.[151,534,151,247]" box="[151,413,222,247]" pageId="8" pageNumber="179">
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(
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<figureCitation id="CEB2F184FF86FFB7FF33FF904924DDC7" box="[158,261,222,247]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="8" pageNumber="179">Figures 1</figureCitation>
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–12,
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||
<tableCitation id="1B0BD8BAFF86FFB7FEECFF9049B3DDC7" box="[321,402,222,247]" captionStart="TABLE 1" captionStartId="1.[151,239,546,569]" captionText="TABLE 1. Morphometric characters (% SL) of Cerogobius petrophilus sp. nov. Characters in alphabetic order." pageId="8" pageNumber="179">Table 1</tableCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="1E93BE8AFF86FFB7FF3AFE684B21DCC3" pageId="8" pageNumber="179" type="materials_examined">
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<paragraph id="5636ED01FF86FFB7FF3AFE684C45DC53" blockId="8.[151,1437,294,2017]" pageId="8" pageNumber="179">
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<materialsCitation id="E6E1E75CFF86FFB7FF3AFE684C45DC53" ID-GBIF-Occurrence="2012823388" collectingDate="2017-10-22" collectionCode="SMF" collectorName="E. M. Troyer" country="Saudi Arabia" county="Al Fahal" elevation="8" latitude="22.2276" location="Red Sea" longLatPrecision="1" longitude="38.96975" municipality="Thuwal" pageId="8" pageNumber="179" specimenCount="1" typeStatus="holotype">
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<emphasis id="64FD3113FF86FFB7FF3AFE684925DC0F" bold="true" box="[151,260,294,319]" pageId="8" pageNumber="179">
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<typeStatus id="893253A3FF86FFB7FF3AFE684925DC0F" box="[151,260,294,319]" pageId="8" pageNumber="179" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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||
<collectionCode id="309875C4FF86FFB7FEBEFE69496ADC0F" box="[275,331,295,319]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="179">SMF</collectionCode>
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35961, female, 16.32 +
|
||
<quantity id="917140E4FF86FFB7FDCFFE684AE8DC0E" box="[610,713,294,319]" metricMagnitude="-3" metricUnit="m" metricValue="3.54" pageId="8" pageNumber="179" unit="mm" value="3.54">3.54 mm</quantity>
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,
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<location id="5356BBDAFF86FFB7FD78FE684B14DC0F" LSID="urn:lsid:plazi:treatment:DE205C17FF86FFB1FF3AFFD94A77DDC3:5356BBDAFF86FFB7FD78FE684B14DC0F" box="[725,821,294,319]" country="Saudi Arabia" county="Al Fahal" latitude="22.2276" longLatPrecision="1" longitude="38.96975" municipality="Thuwal" name="Red Sea" pageId="8" pageNumber="179">Red Sea</location>
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,
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<collectingCountry id="2E9EAD91FF86FFB7FCECFE684BF9DC0F" box="[833,984,294,319]" name="Saudi Arabia" pageId="8" pageNumber="179">Saudi Arabia</collectingCountry>
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,
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<collectingMunicipality id="B652777BFF86FFB7FC49FE684C1CDC0F" box="[996,1085,294,319]" pageId="8" pageNumber="179">Thuwal</collectingMunicipality>
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,
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||
<collectingCounty id="BF57958DFF86FFB7FBE4FE684C8ADC0F" box="[1097,1195,294,319]" pageId="8" pageNumber="179">Al Fahal</collectingCounty>
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reef,
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||
<geoCoordinate id="33BD8BC6FF86FFB7FB43FE694DA4DC0F" box="[1262,1413,294,319]" direction="north" orientation="latitude" pageId="8" pageNumber="179" precision="1" value="22.2276">N 22°13.656'</geoCoordinate>
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<geoCoordinate id="33BD8BC6FF86FFB7FA26FE694935DC53" direction="east" orientation="longitude" pageId="8" pageNumber="179" precision="1" value="38.96975">E 38°58.185'</geoCoordinate>
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||
, rubble-sand patch at base of reef,
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||
<quantity id="917140E4FF86FFB7FD34FE054AE7DC52" box="[665,710,331,355]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="179" unit="m" value="8.0">
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<elevation id="DDA40A32FF86FFB7FD34FE054AE7DC52" box="[665,710,331,355]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="179" unit="m" value="8.0">8 m</elevation>
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</quantity>
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, coll.
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||
<collectorName id="FB7C88D7FF86FFB7FCA4FE054BB7DC53" box="[777,918,331,355]" pageId="8" pageNumber="179">E.M. Troyer</collectorName>
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,
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<date id="2237CBC1FF86FFB7FC0DFE054C40DC53" box="[928,1121,330,355]" pageId="8" pageNumber="179" value="2017-10-22">
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<collectingDate id="32733229FF86FFB7FC0DFE054C40DC53" box="[928,1121,330,355]" pageId="8" pageNumber="179" value="2017-10-22">22 October 2017</collectingDate>
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</date>
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.
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||
</materialsCitation>
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</paragraph>
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||
<paragraph id="5636ED01FF86FFB7FF6AFE204B21DCC3" blockId="8.[151,1437,294,2017]" pageId="8" pageNumber="179">
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<materialsCitation id="E6E1E75CFF86FFB7FF6AFE204A49DCB7" ID-GBIF-Occurrence="2012823387" box="[199,616,366,391]" collectingDate="2017-10-22" collectionCode="SMF" collectorName="E. M. Troyer" country="Saudi Arabia" county="Al Fahal" elevation="8" latitude="22.2276" location="Red Sea" longLatPrecision="1" longitude="38.96975" municipality="Thuwal" pageId="8" pageNumber="179" specimenCount="1" typeStatus="paratype">
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<emphasis id="64FD3113FF86FFB7FF6AFE20491EDCB7" bold="true" box="[199,319,366,391]" pageId="8" pageNumber="179">
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<typeStatus id="893253A3FF86FFB7FF6AFE20491EDCB7" box="[199,319,366,391]" pageId="8" pageNumber="179" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
(collected with holotype)
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||
</materialsCitation>
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||
.
|
||
<materialsCitation id="E6E1E75CFF86FFB7FDDEFE214C83DC9A" ID-GBIF-Occurrence="2012823389" collectionCode="PMR, SMF" country="Saudi Arabia" pageId="8" pageNumber="179" specimenCode="VP4501" specimenCount="1" typeStatus="paratype">
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||
<collectionCode id="309875C4FF86FFB7FDDEFE214A8FDCB6" box="[627,686,367,390]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="8" pageNumber="179">PMR</collectionCode>
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||
<specimenCode id="062F457AFF86FFB7FD1AFE204B34DCB7" box="[695,789,366,391]" collectionCode="PMR" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="8" pageNumber="179">VP4501</specimenCode>
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||
, female, 15.36 +
|
||
<quantity id="917140E4FF86FFB7FC73FE214C67DCB6" box="[990,1094,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="3.21" pageId="8" pageNumber="179" unit="mm" value="3.21">3.21 mm</quantity>
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;
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<collectionCode id="309875C4FF86FFB7FBFEFE214CAADCB7" box="[1107,1163,367,391]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="179">SMF</collectionCode>
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||
35962, female, 16.72 +
|
||
<quantity id="917140E4FF86FFB7FF3AFEDD4920DC9A" box="[151,257,403,427]" metricMagnitude="-3" metricUnit="m" metricValue="3.47" pageId="8" pageNumber="179" unit="mm" value="3.47">3.47 mm</quantity>
|
||
;
|
||
<collectionCode id="309875C4FF86FFB7FEA2FEDD4966DC9B" box="[271,327,403,427]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="179">SMF</collectionCode>
|
||
35963, female, 14.81 +
|
||
<quantity id="917140E4FF86FFB7FDCAFEDD4AF0DC9A" box="[615,721,403,427]" metricMagnitude="-3" metricUnit="m" metricValue="2.97" pageId="8" pageNumber="179" unit="mm" value="2.97">2.97 mm</quantity>
|
||
;
|
||
<collectionCode id="309875C4FF86FFB7FD4DFEDD4B39DC9B" box="[736,792,403,427]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="179">SMF</collectionCode>
|
||
35964, female, 14.09 +
|
||
<quantity id="917140E4FF86FFB7FB95FEDD4C83DC9A" box="[1080,1186,403,427]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="179" unit="mm" value="2.8">2.80 mm</quantity>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="E6E1E75CFF86FFB7FB1DFEDD4ADDDCC3" ID-GBIF-Occurrence="2012823390" collectionCode="UW" country="Saudi Arabia" pageId="8" pageNumber="179" specimenCode="SMF 3, 3, 14.1" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="062F457AFF86FFB7FB1DFEDD4CDEDC9B" box="[1200,1279,403,427]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="8" pageNumber="179">SMF 3</specimenCode>
|
||
5965, female, 12.97 +
|
||
<quantity id="917140E4FF86FFB7FF5EFEF94978DCFE" box="[243,345,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="2.84" pageId="8" pageNumber="179" unit="mm" value="2.84">2.84 mm</quantity>
|
||
;
|
||
<collectionCode id="309875C4FF86FFB7FEC9FEF949B2DCFE" box="[356,403,439,462]" country="USA" name="University of Washington Fish Collection" pageId="8" pageNumber="179" type="Museum">UW</collectionCode>
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||
158279,
|
||
<specimenCode id="062F457AFF86FFB7FE56FEF94A2ADCFF" box="[507,523,439,463]" collectionCode="UW" country="USA" name="University of Washington Fish Collection" pageId="8" pageNumber="179" type="Museum">3</specimenCode>
|
||
,
|
||
<specimenCode id="062F457AFF86FFB7FDBBFEF94A66DCFF" box="[534,583,439,463]" collectionCode="UW" country="USA" name="University of Washington Fish Collection" pageId="8" pageNumber="179" type="Museum">14.1</specimenCode>
|
||
+ 3.0 mm (cleared and stained), 12.6 + 2.0 mm (cleared and stained), 12.4 +
|
||
<quantity id="917140E4FF86FFB7FF3AFE9548CCDCC2" box="[151,237,475,499]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="179" unit="mm" value="2.8">2.8 mm</quantity>
|
||
(preserved in 95% ethanol), all tissue sampled
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E93BE8AFF86FFB7FF6AFEB049FEDF27" box="[199,479,510,535]" pageId="8" pageNumber="179" type="diagnosis">
|
||
<paragraph id="5636ED01FF86FFB7FF6AFEB049FEDF27" blockId="8.[151,1437,294,2017]" box="[199,479,510,535]" pageId="8" pageNumber="179">
|
||
<emphasis id="64FD3113FF86FFB7FF6AFEB0491BDF27" bold="true" box="[199,314,510,535]" pageId="8" pageNumber="179">Diagnosis</emphasis>
|
||
. As for genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E93BE8AFF86FFB6FF6AFD6C4CC4DB4D" lastPageId="9" lastPageNumber="180" pageId="8" pageNumber="179" type="description">
|
||
<paragraph id="5636ED01FF86FFB7FF6AFD6C498FD9F3" blockId="8.[151,1437,294,2017]" pageId="8" pageNumber="179">
|
||
<emphasis id="64FD3113FF86FFB7FF6AFD6C4973DF0B" bold="true" box="[199,338,546,571]" pageId="8" pageNumber="179">Description</emphasis>
|
||
(morphometric values in the text are presented first for the
|
||
<typeStatus id="893253A3FF86FFB7FB8CFD6C4CA2DF0B" box="[1057,1155,546,571]" pageId="8" pageNumber="179" type="holotype">holotype</typeStatus>
|
||
followed by ranges for
|
||
<typeStatus id="893253A3FF86FFB7FF3AFD094927DF6F" box="[151,262,583,607]" pageId="8" pageNumber="179" type="paratype">paratypes</typeStatus>
|
||
; meristic values, if variable, the same). Body elongate, its depth at anal-fin origin 8.2 (7.2–7.9) in SL, laterally compressed posteriorly, with caudal peduncle deep and short, caudal-peduncle depth 0.9 (0.8–0.9) of body depth at anal-fin origin, caudal peduncle depth 1.12 (1.04–1.17) of caudal peduncle length (
|
||
<figureCitation id="CEB2F184FF86FFB7FB77FDC04D02DF97" box="[1242,1315,654,679]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[159,1427,373,1730]" captionTargetId="figure@5.[159,1427,373,1730]" captionTargetPageId="5" captionText="FIGURE 3. Cerogobius petrophilus sp. nov. A: paratype, SMF 35963, female, 14.81 + 2.97 mm, freshly collected specimen, Thuwal, Saudi Arabia. Holotype, SMF 35961, female, 16.32 + 3.54 mm, Thuwal, Saudi Arabia, B: freshly collected specimen; C: preserved specimen. Photos by E.M. Troyer (A & B), M. Kovačić (C)." httpUri="https://zenodo.org/record/2589832/files/figure.png" pageId="8" pageNumber="179">Fig. 3</figureCitation>
|
||
). Head of moderate size, the length 3.0 (3.1–3.3) in SL, width 5.4 (5.3–5.8) in SL, depth 6.1 (5.8–6.3) in SL, and slightly depressed, its depth 0.9 (0.9–1.0) of width. Snout very steep in profile from lateral view, and short, its length 0.5 (0.4–0.5) of eye diameter, 9.4 (8.1–10.4) in head length, with unpaired horn-like tentacle at the level of nostrils in snout midline, slightly shorter in height measured from the anterior side than the eye diameter (
|
||
<figureCitation id="CEB2F184FF86FFB7FB63FC504D09DE07" box="[1230,1320,798,823]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="8" pageNumber="179">Fig. 1A</figureCitation>
|
||
). Anterior nostril a tube without process from rim; posterior nostril a short tube (
|
||
<figureCitation id="CEB2F184FF86FFB7FC33FC0C4BD6DE6B" box="[926,1015,834,859]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="8" pageNumber="179">Fig. 1A</figureCitation>
|
||
). Eyes dorsolateral, moderately large, eye diameter is 4.5 (4.0–4.5) in head length, orbit projecting far forward and distinctly elevated above dorsal profile. Interorbital narrow, 4.4 (4.5–5.6) of eye diameter. No tentacle above eye. Mouth terminal, oblique, jaws ending anteriorly about equally, lower jaw not projecting anteriorly. Mouth moderately large, posterior angle of jaws behind vertical through posterior edge of eye. Upper lip wider than cheek, covered with a thin dermal flap, extending upwards over cheek and nearly reaching eye (
|
||
<figureCitation id="CEB2F184FF86FFB7FCA6FCB84B41D93F" box="[779,864,1014,1039]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="8" pageNumber="179">Fig. 1B</figureCitation>
|
||
) (only in
|
||
<typeStatus id="893253A3FF86FFB7FC62FCB94C0ED93F" box="[975,1071,1015,1039]" pageId="8" pageNumber="179" type="paratype">paratype</typeStatus>
|
||
SMF 35963 dermal flap shorter,
|
||
<figureCitation id="CEB2F184FF86FFB7FF3AFB5448D5D903" box="[151,244,1050,1075]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="8" pageNumber="179">Fig. 1D</figureCitation>
|
||
). Cheek very narrow, depthless than upper lip width. Teeth in both jaws in a single row posteriorly, becoming 3-4 rows anteriorly, with outer row as enlarged recurved canines, inner rows smaller, pointed, and irregularly spaced. Small mental frenum at mid-ventral behind lower lip. Branchiostegal membranes fused to isthmus, gill openings restricted to pectoral-fin base. Lower limb of first gill arch joined to gill cover by membrane. No spines on preopercle.
|
||
</paragraph>
|
||
<paragraph id="5636ED01FF86FFB7FF6AFB8148C5DBA7" blockId="8.[151,1437,294,2017]" pageId="8" pageNumber="179">
|
||
<emphasis id="64FD3113FF86FFB7FF6AFB8148D8D9D6" box="[199,249,1231,1254]" italics="true" pageId="8" pageNumber="179">Fins</emphasis>
|
||
. First dorsal fin VI, second dorsal fin I,12–13 (I,12:2, I,13:5); anal fin I,12–13 (I,12:2, I,13:5); branched caudal-fin rays 13, segmented 17. Pectoral-fin rays 13–14 (13:8, 14:3, both sides, right side cut in
|
||
<typeStatus id="893253A3FF86FFB7FB56FBBD4D7DD83B" box="[1275,1372,1267,1291]" pageId="8" pageNumber="179" type="paratype">paratype</typeStatus>
|
||
SMF 35964), all rays branched except lowermost and uppermost upper rays not free at tips. Pectoral girdle without flaps on anterior edge. Pelvic fins I,5 + 5,I, left and right fin completely separated, distant and without frenum, rays branched (except fifth ray), fourth ray the longest, rays progressively shorter towards the first ray and pelvic spine (
|
||
<figureCitation id="CEB2F184FF86FFB7FF32FACC48D8D8AB" box="[159,249,1410,1435]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="8" pageNumber="179">Fig. 1C</figureCitation>
|
||
). Spines of first dorsal fin not elongate or filamentous, first to fifth spine of first dorsal fin subequal in length, third and fourth only slightly longer, sixth spine short; fourth or fifth to sixth spine of first dorsal fin nearly reaching to origin of second dorsal fin when folded down. Origin of first dorsal fin behind vertical at pectoral-fin base. Origin of anal fin posterior to vertical at origin of second dorsal fin i.e. opposite about first segmented ray of second dorsal fin or slightly behind. Pectoral fins not reaching posteriorly to below origin of second dorsal fin. Pelvic fins short, ending distantly in front of anus, 0.7 (0.6–0.8) of distance between origin of fins and anus, shorter than pectoral fins, 0.8 (0.8–0.9) of pectoral-fin length. Caudal fin rounded, shorter than head, 1.5 (1.5) in head length.
|
||
</paragraph>
|
||
<paragraph id="5636ED01FF86FFB7FF6AF9ED49A5DBEF" blockId="8.[151,1437,294,2017]" pageId="8" pageNumber="179">
|
||
<emphasis id="64FD3113FF86FFB7FF6AF9ED496CDB8A" box="[199,333,1699,1722]" italics="true" pageId="8" pageNumber="179">Squamation</emphasis>
|
||
. Head and body completely scaleless. Cyanine Blue stain confirmed lack of any scales or scale pouches on the body.
|
||
</paragraph>
|
||
<paragraph id="5636ED01FF86FFB6FF6AF9A449BFDFC9" blockId="8.[151,1437,294,2017]" lastBlockId="9.[151,1437,151,1877]" lastPageId="9" lastPageNumber="180" pageId="8" pageNumber="179">
|
||
<emphasis id="64FD3113FF86FFB7FF6AF9A449C5DA33" box="[199,484,1770,1795]" italics="true" pageId="8" pageNumber="179">Cephalic sensory systems</emphasis>
|
||
(
|
||
<figureCitation id="CEB2F184FF86FFB7FE5FF9A44A16DA33" box="[498,567,1770,1795]" captionStart="FIGURE 2" captionStartId="4.[151,250,1022,1044]" captionTargetBox="[201,1426,297,947]" captionTargetId="figure@4.[161,1426,193,1001]" captionTargetPageId="4" captionText="FIGURE 2. Cerogobius petrophilus sp. nov. Cephalic sensory papillae, lateral view of paratype PMR VP4501, female, 15.36 + 3.21 mm, and dorsal view of holotype, SMF 35961, female, 16.32 + 3.54 mm. Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/2589830/files/figure.png" pageId="8" pageNumber="179">Fig. 2</figureCitation>
|
||
). No head canals. Rows of head sensory papillae were counted on the left side of all type material, except
|
||
<typeStatus id="893253A3FF86FFB7FE6BF8414A07DA17" box="[454,550,1807,1831]" pageId="8" pageNumber="179" type="paratype">paratype</typeStatus>
|
||
SMF
|
||
<quantity id="917140E4FF86FFB7FDC6F8404A90DA17" box="[619,689,1806,1831]" metricMagnitude="2" metricUnit="m" metricValue="9.134348" pageId="8" pageNumber="179" unit="in" value="35962.0">35962</quantity>
|
||
in which papillae were hard to notice on the damaged skin surface. In addition, some rows were not visible in every specimen. Rows of head sensory papillae were heavily reduced, with rows missing or reduced to one to few papillae and additional individual papillae present on the position of absent head canals. Preorbital rows: snout with three median preorbital series, each as single papilla, upper row
|
||
<emphasis id="64FD3113FF86FFB7FA3CF8324DBDDAA3" box="[1425,1436,1916,1939]" italics="true" pageId="8" pageNumber="179">r</emphasis>
|
||
(1) at middorsal between eyes and posterior nostrils, and row
|
||
<emphasis id="64FD3113FF86FFB7FCFAF8EC4B43DA89" box="[855,866,1954,1977]" italics="true" pageId="8" pageNumber="179">s</emphasis>
|
||
(1) at anterior nostril, and with
|
||
<emphasis id="64FD3113FF86FFB7FB7CF8EC4CC5DA9C" box="[1233,1252,1950,1977]" italics="true" pageId="8" pageNumber="179">
|
||
s
|
||
<superScript id="A1FC4049FF86FFB7FB71F8D04CC5DA9C" attach="left" box="[1244,1252,1950,1964]" fontSize="6" pageId="8" pageNumber="179">3</superScript>
|
||
</emphasis>
|
||
(1) above upper lip. Lateral series
|
||
<emphasis id="64FD3113FF86FFB7FEC8F8874950DAD0" box="[357,369,1993,2016]" italics="true" pageId="8" pageNumber="179">c</emphasis>
|
||
on snout in three parts: superior horizontal row
|
||
<emphasis id="64FD3113FF86FFB7FC0CF8874B94DAE3" box="[929,949,1989,2016]" italics="true" pageId="8" pageNumber="179">
|
||
c
|
||
<superScript id="A1FC4049FF86FFB7FC00F88B4B94DAE3" attach="left" box="[941,949,1989,2003]" fontSize="6" pageId="8" pageNumber="179">2</superScript>
|
||
</emphasis>
|
||
between anterior and posterior nostril (2); inferior rows: upper horizontal
|
||
<emphasis id="64FD3113FF87FFB6FE52FFD74A32DD84" box="[511,531,153,180]" italics="true" pageId="9" pageNumber="180">
|
||
c
|
||
<subScript id="CA0DEF44FF87FFB6FDA6FFE84A32DD84" attach="left" box="[523,531,166,180]" fontSize="6" pageId="9" pageNumber="180">2</subScript>
|
||
</emphasis>
|
||
(2) and lower horizontal
|
||
<emphasis id="64FD3113FF87FFB6FC9AFFD74B6ADD84" box="[823,843,153,180]" italics="true" pageId="9" pageNumber="180">
|
||
c
|
||
<subScript id="CA0DEF44FF87FFB6FCEEFFE84B6ADD84" attach="left" box="[835,843,166,180]" fontSize="6" pageId="9" pageNumber="180">1</subScript>
|
||
</emphasis>
|
||
(2) above upper lip. Suborbital rows: rows
|
||
<emphasis id="64FD3113FF87FFB6FAE8FFD74D72DD80" box="[1349,1363,153,176]" italics="true" pageId="9" pageNumber="180">a</emphasis>
|
||
and
|
||
<emphasis id="64FD3113FF87FFB6FA23FFD94DBDDD80" box="[1422,1436,151,176]" italics="true" pageId="9" pageNumber="180">b</emphasis>
|
||
absent; four very short transverse suborbital rows of sensory papillae (
|
||
<emphasis id="64FD3113FF87FFB6FC02FFF14BFBDDE7" box="[943,986,191,215]" italics="true" pageId="9" pageNumber="180">1–4</emphasis>
|
||
) present, row
|
||
<emphasis id="64FD3113FF87FFB6FBD1FFF14CABDDE7" box="[1148,1162,191,215]" italics="true" pageId="9" pageNumber="180">1</emphasis>
|
||
oblique, reaching upper lip, row
|
||
<emphasis id="64FD3113FF87FFB6FF54FFAD4926DDCB" box="[249,263,227,251]" italics="true" pageId="9" pageNumber="180">2</emphasis>
|
||
short and distant from eye, close to row
|
||
<emphasis id="64FD3113FF87FFB6FD4FFFAC4AD1DDCB" box="[738,752,226,251]" italics="true" pageId="9" pageNumber="180">d</emphasis>
|
||
, row
|
||
<emphasis id="64FD3113FF87FFB6FC99FFAD4B63DDCB" box="[820,834,227,251]" italics="true" pageId="9" pageNumber="180">3</emphasis>
|
||
and
|
||
<emphasis id="64FD3113FF87FFB6FCD1FFAD4BABDDCB" box="[892,906,227,251]" italics="true" pageId="9" pageNumber="180">4</emphasis>
|
||
short, close to posteroventral edge of eye and distant from row
|
||
<emphasis id="64FD3113FF87FFB6FECCFE48494EDC2F" box="[353,367,262,287]" italics="true" pageId="9" pageNumber="180">d</emphasis>
|
||
(
|
||
<emphasis id="64FD3113FF87FFB6FE2EFE4949B0DC2F" box="[387,401,263,287]" italics="true" pageId="9" pageNumber="180">1</emphasis>
|
||
: 4–5,
|
||
<emphasis id="64FD3113FF87FFB6FE70FE4949CADC2F" box="[477,491,263,287]" italics="true" pageId="9" pageNumber="180">2</emphasis>
|
||
: 2,
|
||
<emphasis id="64FD3113FF87FFB6FDB1FE494A0BDC2F" box="[540,554,263,287]" italics="true" pageId="9" pageNumber="180">3</emphasis>
|
||
: 2–3,
|
||
<emphasis id="64FD3113FF87FFB6FDDAFE494AA4DC2F" box="[631,645,263,287]" italics="true" pageId="9" pageNumber="180">4</emphasis>
|
||
: 3); row
|
||
<emphasis id="64FD3113FF87FFB6FD58FE484B22DC2F" box="[757,771,262,287]" italics="true" pageId="9" pageNumber="180">d</emphasis>
|
||
(3–4 + 2–3) short, above posterior part of upper lip, and continuing shortly on cheek. Preoperculo
|
||
<emphasis id="64FD3113FF87FFB6FDCAFE624A51DC73" box="[615,624,300,323]" italics="true" pageId="9" pageNumber="180">-</emphasis>
|
||
mandibular rows: external row
|
||
<emphasis id="64FD3113FF87FFB6FC7EFE624BFEDC73" box="[979,991,300,323]" italics="true" pageId="9" pageNumber="180">e</emphasis>
|
||
(8–9 + 2–6) longitudinal and uniserial, divided into anterior and posterior sections; internal row
|
||
<emphasis id="64FD3113FF87FFB6FC9FFE1E4B1BDC57" box="[818,826,336,359]" italics="true" pageId="9" pageNumber="180">i</emphasis>
|
||
(4–6 + 2–4) longitudinal and uniserial, divided into anterior and posterior sections, mental row
|
||
<emphasis id="64FD3113FF87FFB6FDD0FE3D4AA4DCBB" box="[637,645,371,395]" italics="true" pageId="9" pageNumber="180">f</emphasis>
|
||
(2–3) behind frenum. Oculoscapular rows: anterior upper longitudinal row
|
||
<date id="2237CBC1FF87FFB6FF66FED448FEDC94" box="[203,223,406,433]" pageId="9" pageNumber="180">
|
||
<emphasis id="64FD3113FF87FFB6FF66FED448FEDC94" box="[203,223,406,433]" italics="true" pageId="9" pageNumber="180">
|
||
x
|
||
<superScript id="A1FC4049FF87FFB6FF7AFED848FEDC94" attach="left" box="[215,223,406,420]" fontSize="6" pageId="9" pageNumber="180">1</superScript>
|
||
</emphasis>
|
||
</date>
|
||
divided (3 + 1–2); posterior longitudinal row
|
||
<date id="2237CBC1FF87FFB6FD41FED44B20DC94" box="[748,769,406,433]" pageId="9" pageNumber="180">
|
||
<emphasis id="64FD3113FF87FFB6FD41FED44B20DC94" box="[748,769,406,433]" italics="true" pageId="9" pageNumber="180">
|
||
x
|
||
<superScript id="A1FC4049FF87FFB6FD54FED84B20DC94" attach="left" box="[761,769,406,420]" fontSize="6" pageId="9" pageNumber="180">2</superScript>
|
||
</emphasis>
|
||
</date>
|
||
(2); posterior part of row
|
||
<date id="2237CBC1FF87FFB6FB81FED44C61DC94" box="[1068,1088,406,433]" pageId="9" pageNumber="180">
|
||
<emphasis id="64FD3113FF87FFB6FB81FED44C61DC94" box="[1068,1088,406,433]" italics="true" pageId="9" pageNumber="180">
|
||
x
|
||
<superScript id="A1FC4049FF87FFB6FB95FED84C61DC94" attach="left" box="[1080,1088,406,420]" fontSize="6" pageId="9" pageNumber="180">1</superScript>
|
||
</emphasis>
|
||
</date>
|
||
and row
|
||
<date id="2237CBC1FF87FFB6FB05FED44C9CDC94" box="[1192,1213,406,433]" pageId="9" pageNumber="180">
|
||
<emphasis id="64FD3113FF87FFB6FB05FED44C9CDC94" box="[1192,1213,406,433]" italics="true" pageId="9" pageNumber="180">
|
||
x
|
||
<superScript id="A1FC4049FF87FFB6FB18FED84C9CDC94" attach="left" box="[1205,1213,406,420]" fontSize="6" pageId="9" pageNumber="180">2</superScript>
|
||
</emphasis>
|
||
</date>
|
||
not visible in every specimen; anterior lower transverse row
|
||
<emphasis id="64FD3113FF87FFB6FDF0FEF04A49DCE5" box="[605,616,446,469]" italics="true" pageId="9" pageNumber="180">z</emphasis>
|
||
as single papilla, not visible in every specimen; rows
|
||
<emphasis id="64FD3113FF87FFB6FB6FFEF04CF1DCE5" box="[1218,1232,446,469]" italics="true" pageId="9" pageNumber="180">q</emphasis>
|
||
and
|
||
<emphasis id="64FD3113FF87FFB6FAABFEF04D33DCE5" box="[1286,1298,446,469]" italics="true" pageId="9" pageNumber="180">y</emphasis>
|
||
not present; transverse axillary rows
|
||
<emphasis id="64FD3113FF87FFB6FE1CFEAB49F3DCDF" box="[433,466,481,508]" italics="true" pageId="9" pageNumber="180">
|
||
as
|
||
<superScript id="A1FC4049FF87FFB6FE67FEAF49F3DCDF" attach="left" box="[458,466,481,495]" fontSize="6" pageId="9" pageNumber="180">1</superScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="64FD3113FF87FFB6FDA4FEAB4A0BDCDF" box="[521,554,481,508]" italics="true" pageId="9" pageNumber="180">
|
||
as
|
||
<superScript id="A1FC4049FF87FFB6FD8FFEAF4A0BDCDF" attach="left" box="[546,554,481,495]" fontSize="6" pageId="9" pageNumber="180">2</superScript>
|
||
</emphasis>
|
||
as single papilla, not visible in every specimen;
|
||
<emphasis id="64FD3113FF87FFB6FBF3FEAB4C5EDCDF" box="[1118,1151,481,508]" italics="true" pageId="9" pageNumber="180">
|
||
as
|
||
<superScript id="A1FC4049FF87FFB6FBDAFEAF4C5EDCDF" attach="left" box="[1143,1151,481,495]" fontSize="6" pageId="9" pageNumber="180">3</superScript>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="64FD3113FF87FFB6FB3DFEAA4C87DCCD" box="[1168,1190,484,509]" italics="true" pageId="9" pageNumber="180">la</emphasis>
|
||
<superScript id="A1FC4049FF87FFB6FB06FEAF4C92DCDF" attach="left" box="[1195,1203,481,495]" fontSize="6" pageId="9" pageNumber="180">
|
||
<emphasis id="64FD3113FF87FFB6FB06FEAF4C92DCDF" box="[1195,1203,481,495]" italics="true" pageId="9" pageNumber="180">1</emphasis>
|
||
</superScript>
|
||
and
|
||
<emphasis id="64FD3113FF87FFB6FB43FEAA4D33DCDF" box="[1262,1298,481,509]" italics="true" pageId="9" pageNumber="180">
|
||
la
|
||
<superScript id="A1FC4049FF87FFB6FAA7FEAF4D33DCDF" attach="right" box="[1290,1298,481,495]" fontSize="6" pageId="9" pageNumber="180">2</superScript>
|
||
</emphasis>
|
||
not present. Opercular rows: two papillae at position of the missing preopercular canal (marked PC on
|
||
<figureCitation id="CEB2F184FF87FFB6FB29FD464CE6DF11" box="[1156,1223,520,545]" captionStart="FIGURE 2" captionStartId="4.[151,250,1022,1044]" captionTargetBox="[201,1426,297,947]" captionTargetId="figure@4.[161,1426,193,1001]" captionTargetPageId="4" captionText="FIGURE 2. Cerogobius petrophilus sp. nov. Cephalic sensory papillae, lateral view of paratype PMR VP4501, female, 15.36 + 3.21 mm, and dorsal view of holotype, SMF 35961, female, 16.32 + 3.54 mm. Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/2589830/files/figure.png" pageId="9" pageNumber="180">Fig. 2</figureCitation>
|
||
); transverse row
|
||
<emphasis id="64FD3113FF87FFB6FA2BFD474DBDDF10" box="[1414,1436,521,544]" italics="true" pageId="9" pageNumber="180">ot</emphasis>
|
||
(2–4); superior longitudinal row
|
||
<emphasis id="64FD3113FF87FFB6FDB4FD634A13DF74" box="[537,562,557,580]" italics="true" pageId="9" pageNumber="180">os</emphasis>
|
||
(1–2); and inferior longitudinal row
|
||
<emphasis id="64FD3113FF87FFB6FC41FD634C23DF74" box="[1004,1026,557,580]" italics="true" pageId="9" pageNumber="180">oi</emphasis>
|
||
(1) not visible in every specimen. Anterior dorsal rows: two papillae behind posterior upper edge of eye on the position of the missing anterior oculoscapular canal (marked OC on
|
||
<figureCitation id="CEB2F184FF87FFB6FD95FD3A4A5CDFBD" box="[568,637,628,653]" captionStart="FIGURE 2" captionStartId="4.[151,250,1022,1044]" captionTargetBox="[201,1426,297,947]" captionTargetId="figure@4.[161,1426,193,1001]" captionTargetPageId="4" captionText="FIGURE 2. Cerogobius petrophilus sp. nov. Cephalic sensory papillae, lateral view of paratype PMR VP4501, female, 15.36 + 3.21 mm, and dorsal view of holotype, SMF 35961, female, 16.32 + 3.54 mm. Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/2589830/files/figure.png" pageId="9" pageNumber="180">Fig. 2</figureCitation>
|
||
); transverse row
|
||
<emphasis id="64FD3113FF87FFB6FCEEFD3B4B70DFBC" box="[835,849,629,652]" italics="true" pageId="9" pageNumber="180">n</emphasis>
|
||
(2–4) behind eye, rows
|
||
<emphasis id="64FD3113FF87FFB6FBC4FD3B4C56DFBC" box="[1129,1143,629,652]" italics="true" pageId="9" pageNumber="180">o</emphasis>
|
||
and
|
||
<emphasis id="64FD3113FF87FFB6FB1DFD3B4CE5DFBC" box="[1200,1220,629,652]" italics="true" pageId="9" pageNumber="180">m</emphasis>
|
||
not present, row
|
||
<emphasis id="64FD3113FF87FFB6FA23FD3B4DBDDFBC" box="[1422,1436,629,652]" italics="true" pageId="9" pageNumber="180">g</emphasis>
|
||
longitudinal (3–5); row
|
||
<emphasis id="64FD3113FF87FFB6FE06FDD64998DF81" box="[427,441,664,689]" italics="true" pageId="9" pageNumber="180">h</emphasis>
|
||
(1–2) not visible in every specimen. Interorbital rows: two pairs of larger papillae are present in interorbital area anteriorly and posteriorly on the place of the missing anterior oculoscapular head canal (marked OC on
|
||
<figureCitation id="CEB2F184FF87FFB6FEE1FDAE49B1DFC9" box="[332,400,736,761]" captionStart="FIGURE 2" captionStartId="4.[151,250,1022,1044]" captionTargetBox="[201,1426,297,947]" captionTargetId="figure@4.[161,1426,193,1001]" captionTargetPageId="4" captionText="FIGURE 2. Cerogobius petrophilus sp. nov. Cephalic sensory papillae, lateral view of paratype PMR VP4501, female, 15.36 + 3.21 mm, and dorsal view of holotype, SMF 35961, female, 16.32 + 3.54 mm. Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/2589830/files/figure.png" pageId="9" pageNumber="180">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5636ED01FF87FFB6FF6AFC4A491DD90D" blockId="9.[151,1437,151,1877]" pageId="9" pageNumber="180">
|
||
<emphasis id="64FD3113FF87FFB6FF6AFC4A4A36DE2D" box="[199,535,772,797]" italics="true" pageId="9" pageNumber="180">Color of live and fresh female</emphasis>
|
||
(
|
||
<figureCitation id="CEB2F184FF87FFB6FD8BFC4A4AF9DE2D" box="[550,728,772,797]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[159,1427,373,1730]" captionTargetId="figure@5.[159,1427,373,1730]" captionTargetPageId="5" captionText="FIGURE 3. Cerogobius petrophilus sp. nov. A: paratype, SMF 35963, female, 14.81 + 2.97 mm, freshly collected specimen, Thuwal, Saudi Arabia. Holotype, SMF 35961, female, 16.32 + 3.54 mm, Thuwal, Saudi Arabia, B: freshly collected specimen; C: preserved specimen. Photos by E.M. Troyer (A & B), M. Kovačić (C)." httpUri="https://zenodo.org/record/2589832/files/figure.png" pageId="9" pageNumber="180">Fig. 3A and 3B</figureCitation>
|
||
). Head and body translucent i.e. showing limited transparence. Cheek, postorbital head, prepectoral and prepelvic areas, and anterior body to below origin of the second dorsal fin densely covered with variable-sized melanophores. Remaining body covered with few blackish speckles; row of eight, small, irregular black spots along base of second dorsal fin. Snout, tentacle, and orbits pale cream. First dorsal fin with translucent membranes, outer half of fin with many black dots, basal half with few dots on spines and membranes, about middle of first four spines and first interspinous membrane with whitish speckles. Second dorsal and caudal fins with translucent membranes and a row of alternating white and blackish speckles along spine and soft rays. Anal fin with whitish rays and translucent membranes. Pelvic fins densely dotted with numerous melanophores.
|
||
</paragraph>
|
||
<paragraph id="5636ED01FF87FFB6FF6AFB064CC4DB4D" blockId="9.[151,1437,151,1877]" pageId="9" pageNumber="180">
|
||
<emphasis id="64FD3113FF87FFB6FF6AFB0649CED951" box="[199,495,1096,1121]" italics="true" pageId="9" pageNumber="180">Color of preserved female</emphasis>
|
||
(
|
||
<figureCitation id="CEB2F184FF87FFB6FDADFB064A78D951" box="[512,601,1096,1121]" captionStart="FIGURE 3" captionStartId="5.[151,250,1751,1773]" captionTargetBox="[159,1427,373,1730]" captionTargetId="figure@5.[159,1427,373,1730]" captionTargetPageId="5" captionText="FIGURE 3. Cerogobius petrophilus sp. nov. A: paratype, SMF 35963, female, 14.81 + 2.97 mm, freshly collected specimen, Thuwal, Saudi Arabia. Holotype, SMF 35961, female, 16.32 + 3.54 mm, Thuwal, Saudi Arabia, B: freshly collected specimen; C: preserved specimen. Photos by E.M. Troyer (A & B), M. Kovačić (C)." httpUri="https://zenodo.org/record/2589832/files/figure.png" pageId="9" pageNumber="180">Fig. 3C</figureCitation>
|
||
). Coloration of preserved female similar to fresh specimen. Body opaque and whitish with dark brown pigmentation with large melanophores mostly on head and in anterior part of body and no other pigments visible. Head and body to below origin of the second dorsal fin densely pigmented with variable-sized melanophores with very poorly recognizable pattern. Snout weakly pigmented, dark vertical bar around posterior edge of eye and pigmented bar on check from eye to lips, followed by a pale band, containing scattered melanophores, that extends from nape over posterior cheek and angle of jaws to below it. Preopercle darker and less densely pigmented opercle and pectoral-fin base. Ventral side of head heavily pigmented with tiny melanophores except for pale area below each side angle of jaw. Breast in front of pelvic fin pale, belly pigmented. Eyes with bright whitish iris and blurred whitish pupil due to the alcohol fixation. The melanophores on the body behind the vertical through origin of the second dorsal are scattered, smaller and more obscure than anteriorly. Only visible patterns are dashed narrow line along lateral midline and dark spots at the origin of each dorsal and anal fin ray. First dorsal fin dotted with dark brown melanophores and with white dots restricted to anterior origin and at spines as oblique pattern of dots going downwards from spines I to spine VI; second dorsal fin with white spine and rays, and mostly translucent interradial membranes, melanophores, if present, rare at spine and some ray origins and also rarely scattered on rays; caudal and anal fin with transparent membranes. Pectoral and pelvic fins intensively dotted with dark brown melanophores, matching anterior body pigmentation (
|
||
<figureCitation id="CEB2F184FF87FFB6FB2DF92A4CF9DB4D" box="[1152,1240,1636,1661]" captionStart="FIGURE 1" captionStartId="3.[151,250,1806,1828]" captionTargetBox="[188,1398,190,1789]" captionTargetId="figure@3.[188,1398,190,1789]" captionTargetPageId="3" captionText="FIGURE 1. Cerogobius petrophilus sp. nov.: A: head with A—unpaired horn-like tentacle at snout midline in the level of nostrils, B—anterior nostril a short tube without process from rim, C—posterior nostril a short tube (paratype, PMR VP4501, female, 15.36 + 3.21 mm); B: upper lip covered with thin dermal flap, extending upwards over cheek and nearly reaching eye (holotype, SMF 35961, female, 16.32 + 3.54 mm); C: pelvic fins completely separated and without frenum (holotype, SMF 35961, female, 16.32 + 3.54 mm). Material reversibly stained; D: upper lip just broader than cheek depth and dermal flap covering only lip (paratype, SMF 35963, female, 14.81 + 2.97 mm), in B) and D) edge of dermal flap marked with arrow. Photos by M. Kovačić." httpUri="https://zenodo.org/record/2589828/files/figure.png" pageId="9" pageNumber="180">Fig. 1C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E93BE8AFF87FFB6FF6AF9C64C50DA65" pageId="9" pageNumber="180" type="materials_examined">
|
||
<paragraph id="5636ED01FF87FFB6FF6AF9C64C50DA65" blockId="9.[151,1437,151,1877]" pageId="9" pageNumber="180">
|
||
<emphasis id="64FD3113FF87FFB6FF6AF9C64B57DB90" box="[199,886,1672,1697]" italics="true" pageId="9" pageNumber="180">Alive coloration of non-collected material of unidentified sex</emphasis>
|
||
(
|
||
<figureCitation id="CEB2F184FF87FFB6FC2BF9C64BF2DB91" box="[902,979,1672,1697]" captionStart="FIGURE 5" captionStartId="10.[151,250,1141,1163]" captionTargetBox="[159,1427,193,1120]" captionTargetId="figure@10.[159,1427,193,1120]" captionTargetPageId="10" captionText="FIGURE 5. Cerogobius petrophilus sp. nov. A-C: alive individuals, Egypt, Marsa Alam, Marsa Shagra; D: typical habitat at All Fahal reef near Thuwal, Saudi Arabia. Photos B. Hazes (A-C) and E.M. Troyer (D)." httpUri="https://zenodo.org/record/2589836/files/figure.png" pageId="9" pageNumber="180">Figs. 5</figureCitation>
|
||
A-C). Coloration of head blackish (
|
||
<figureCitation id="CEB2F184FF87FFB6FAC9F9C648FDDBF5" captionStart="FIGURE 5" captionStartId="10.[151,250,1141,1163]" captionTargetBox="[159,1427,193,1120]" captionTargetId="figure@10.[159,1427,193,1120]" captionTargetPageId="10" captionText="FIGURE 5. Cerogobius petrophilus sp. nov. A-C: alive individuals, Egypt, Marsa Alam, Marsa Shagra; D: typical habitat at All Fahal reef near Thuwal, Saudi Arabia. Photos B. Hazes (A-C) and E.M. Troyer (D)." httpUri="https://zenodo.org/record/2589836/files/figure.png" pageId="9" pageNumber="180">Figs. 5A, B</figureCitation>
|
||
) or pale cream (
|
||
<figureCitation id="CEB2F184FF87FFB6FE31F9E249D7DBF5" box="[412,502,1708,1733]" captionStart="FIGURE 5" captionStartId="10.[151,250,1141,1163]" captionTargetBox="[159,1427,193,1120]" captionTargetId="figure@10.[159,1427,193,1120]" captionTargetPageId="10" captionText="FIGURE 5. Cerogobius petrophilus sp. nov. A-C: alive individuals, Egypt, Marsa Alam, Marsa Shagra; D: typical habitat at All Fahal reef near Thuwal, Saudi Arabia. Photos B. Hazes (A-C) and E.M. Troyer (D)." httpUri="https://zenodo.org/record/2589836/files/figure.png" pageId="9" pageNumber="180">Fig. 5C</figureCitation>
|
||
), covered with many irregular melanophores. In the specimen with blackish head (
|
||
<figureCitation id="CEB2F184FF87FFB6FF32F99E4909DBD9" box="[159,296,1744,1769]" captionStart="FIGURE 5" captionStartId="10.[151,250,1141,1163]" captionTargetBox="[159,1427,193,1120]" captionTargetId="figure@10.[159,1427,193,1120]" captionTargetPageId="10" captionText="FIGURE 5. Cerogobius petrophilus sp. nov. A-C: alive individuals, Egypt, Marsa Alam, Marsa Shagra; D: typical habitat at All Fahal reef near Thuwal, Saudi Arabia. Photos B. Hazes (A-C) and E.M. Troyer (D)." httpUri="https://zenodo.org/record/2589836/files/figure.png" pageId="9" pageNumber="180">Figs. 5A, B</figureCitation>
|
||
), snout, tentacle, orbit and jaws pale gray suffused with pale orange; postorbital head mainly black except for few small white spots on opercle and for an irregular, oblique, white band, containing scattered melanophores, that extends from nape to posterior chin. Individuals with blackish head are possibly males, considering their dark coloration compared to the
|
||
<typeStatus id="893253A3FF87FFB6FD6AF8734AD6DA65" box="[711,759,1853,1877]" pageId="9" pageNumber="180">type</typeStatus>
|
||
specimens which all are females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="02F6BD89FF84FFB5FF3AFB3B4C2FD99A" httpUri="https://zenodo.org/record/2589836/files/figure.png" pageId="10" pageNumber="181" startId="10.[151,250,1141,1163]" targetBox="[159,1427,193,1120]" targetPageId="10">
|
||
<paragraph id="5636ED01FF84FFB5FF3AFB3B4C2FD99A" blockId="10.[151,1436,1141,1194]" pageId="10" pageNumber="181">
|
||
<emphasis id="64FD3113FF84FFB5FF3AFB3B4935D9BB" bold="true" box="[151,276,1141,1163]" pageId="10" pageNumber="181">FIGURE 5.</emphasis>
|
||
<taxonomicName id="91899682FF84FFB5FEB6FB3B4A2AD9BB" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[283,523,1141,1163]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="181" phylum="Chordata" rank="species" species="petrophilus" status="sp. nov.">
|
||
<emphasis id="64FD3113FF84FFB5FEB6FB3B4A2AD9BB" box="[283,523,1141,1163]" italics="true" pageId="10" pageNumber="181">Cerogobius petrophilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FD3113FF84FFB5FDBFFB384AB9D9BB" bold="true" box="[530,664,1141,1163]" pageId="10" pageNumber="181">
|
||
<taxonomicNameLabel id="7FCE8C68FF84FFB5FDBFFB384A42D9BB" box="[530,611,1142,1163]" pageId="10" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A-C
|
||
</emphasis>
|
||
: alive individuals, Egypt, Marsa Alam, Marsa Shagra;
|
||
<emphasis id="64FD3113FF84FFB5FB7DFB384CC3D9BB" bold="true" box="[1232,1250,1142,1163]" pageId="10" pageNumber="181">D</emphasis>
|
||
: typical habitat at All Fahal reef near Thuwal, Saudi Arabia. Photos B. Hazes (A-C) and E.M. Troyer (D).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="1E93BE8AFF84FFB5FF6AFBB54B0FD8B0" pageId="10" pageNumber="181" type="etymology">
|
||
<paragraph id="5636ED01FF84FFB5FF6AFBB54B0FD8B0" blockId="10.[151,1437,1275,1876]" pageId="10" pageNumber="181">
|
||
<emphasis id="64FD3113FF84FFB5FF6AFBB54967D824" bold="true" box="[199,326,1275,1300]" pageId="10" pageNumber="181">Etymology</emphasis>
|
||
. The genus name
|
||
<taxonomicName id="91899682FF84FFB5FDB5FBB54ABBD824" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[536,666,1275,1300]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="173" phylum="Chordata" rank="genus" status="gen. nov.">
|
||
<emphasis id="64FD3113FF84FFB5FDB5FBB54ABBD824" box="[536,666,1275,1300]" italics="true" pageId="10" pageNumber="181">Cerogobius</emphasis>
|
||
</taxonomicName>
|
||
is formed from the Greek word
|
||
<emphasis id="64FD3113FF84FFB5FBA3FBB54C6AD824" box="[1038,1099,1275,1300]" italics="true" pageId="10" pageNumber="181">keras</emphasis>
|
||
(Latin =
|
||
<emphasis id="64FD3113FF84FFB5FB15FBB24CD5D823" box="[1208,1268,1276,1299]" italics="true" pageId="10" pageNumber="181">ceros</emphasis>
|
||
; horn) and the Latin word
|
||
<emphasis id="64FD3113FF84FFB5FEB3FA51497FD808" box="[286,350,1311,1336]" italics="true" pageId="10" pageNumber="181">gobio</emphasis>
|
||
(small fish or gudgeon), and is in reference to the horn-like tentacle on the head. The specific name
|
||
<taxonomicName id="91899682FF84FFB5FF4DFA0D497ED86C" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[224,351,1347,1372]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="181" phylum="Chordata" rank="species" species="petrophilus">
|
||
<emphasis id="64FD3113FF84FFB5FF4DFA0D497ED86C" box="[224,351,1347,1372]" italics="true" pageId="10" pageNumber="181">petrophilus</emphasis>
|
||
</taxonomicName>
|
||
is formed from the Latin words
|
||
<emphasis id="64FD3113FF84FFB5FD5DFA0A4B0AD86B" box="[752,811,1348,1371]" italics="true" pageId="10" pageNumber="181">petra</emphasis>
|
||
(rock) and
|
||
<emphasis id="64FD3113FF84FFB5FC10FA0D4C20D86C" box="[957,1025,1347,1372]" italics="true" pageId="10" pageNumber="181">philus</emphasis>
|
||
(having an affinity for), and is in reference to the rocky habitat where the species was found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E93BE8AFF84FFB5FF6AFAC54AA4DBAF" pageId="10" pageNumber="181" type="distribution">
|
||
<paragraph id="5636ED01FF84FFB5FF6AFAC549B6D8DC" blockId="10.[151,1437,1275,1876]" pageId="10" pageNumber="181">
|
||
<emphasis id="64FD3113FF84FFB5FF6AFAC54978D894" bold="true" box="[199,345,1419,1444]" pageId="10" pageNumber="181">Distribution</emphasis>
|
||
. At present only known from the northern part of the Red Sea, from collection taken at Thuwal in
|
||
<collectingCountry id="2E9EAD91FF84FFB5FF3AFAE1490CD8F8" box="[151,301,1455,1480]" name="Saudi Arabia" pageId="10" pageNumber="181">Saudi Arabia</collectingCountry>
|
||
, and SCUBA observations at the southern tip of Ras Mohammed national park and in Marsa Shagra, Marsa Alam, in
|
||
<collectingCountry id="2E9EAD91FF84FFB5FEE0FA9A49B5D8DC" box="[333,404,1492,1516]" name="Egypt" pageId="10" pageNumber="181">Egypt</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="5636ED01FF84FFB5FF6AFAB94AA4DBAF" blockId="10.[151,1437,1275,1876]" pageId="10" pageNumber="181">
|
||
<emphasis id="64FD3113FF84FFB5FF6AFAB94902DB20" bold="true" box="[199,291,1527,1552]" pageId="10" pageNumber="181">Habitat</emphasis>
|
||
. Collected on rubble-sand patches at the base of isolated coral blocks near the margin of Al Fahal reef in the central Red Sea, near Thuwal,
|
||
<collectingCountry id="2E9EAD91FF84FFB5FD95F9554AEDDB04" box="[568,716,1563,1588]" name="Saudi Arabia" pageId="10" pageNumber="181">Saudi Arabia</collectingCountry>
|
||
(
|
||
<figureCitation id="CEB2F184FF84FFB5FD71F9554B17DB04" box="[732,822,1563,1588]" captionStart="FIGURE 5" captionStartId="10.[151,250,1141,1163]" captionTargetBox="[159,1427,193,1120]" captionTargetId="figure@10.[159,1427,193,1120]" captionTargetPageId="10" captionText="FIGURE 5. Cerogobius petrophilus sp. nov. A-C: alive individuals, Egypt, Marsa Alam, Marsa Shagra; D: typical habitat at All Fahal reef near Thuwal, Saudi Arabia. Photos B. Hazes (A-C) and E.M. Troyer (D)." httpUri="https://zenodo.org/record/2589836/files/figure.png" pageId="10" pageNumber="181">Fig. 5D</figureCitation>
|
||
). The
|
||
<typeStatus id="893253A3FF84FFB5FCD3F9554BC1DB04" box="[894,992,1563,1588]" pageId="10" pageNumber="181" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="893253A3FF84FFB5FBBAF9524CA2DB04" box="[1047,1155,1564,1588]" pageId="10" pageNumber="181" type="paratype">paratypes</typeStatus>
|
||
were found living inside small holes in large pieces of dead coral rock covered with short algae at a depth of
|
||
<specimenCount id="408F2688FF84FFB5FBE9F9714C57DB67" box="[1092,1142,1599,1623]" pageId="10" pageNumber="181" type="generic">
|
||
<quantity id="917140E4FF84FFB5FBE9F9714C53DB67" box="[1092,1138,1599,1623]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="181" unit="m" value="8.0">8 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
Two other specimens, not examined in this study, were collected at a depth of
|
||
<quantity id="917140E4FF84FFB5FD48F92D4B3EDB4B" box="[741,799,1635,1660]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="181" unit="m" value="15.0">15 m</quantity>
|
||
<materialsCitation id="E6E1E75CFF84FFB5FC8AF92D4AA4DBAF" ID-GBIF-Occurrence="2012823386" collectingDate="2017-05" elevation="5" pageId="10" pageNumber="181" specimenCount="5">
|
||
in
|
||
<date id="2237CBC1FF84FFB5FCE8F92A4B99DB4B" box="[837,952,1635,1660]" pageId="10" pageNumber="181" value="2017-05">
|
||
<collectingDate id="32733229FF84FFB5FCE8F92A4B99DB4B" box="[837,952,1635,1660]" pageId="10" pageNumber="181" value="2017-05">May 2017</collectingDate>
|
||
</date>
|
||
at the same locality. Observations in other localities were made at depths of about
|
||
<specimenCount id="408F2688FF84FFB5FDFEF9C94AA4DBAF" box="[595,645,1671,1696]" pageId="10" pageNumber="181" type="generic">
|
||
<quantity id="917140E4FF84FFB5FDFEF9C94AA0DBAF" box="[595,641,1671,1696]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="181" unit="m" value="5.0">
|
||
<elevation id="DDA40A32FF84FFB5FDFEF9C94AA0DBAF" box="[595,641,1671,1696]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="181" unit="m" value="5.0">5 m</elevation>
|
||
</quantity>
|
||
.
|
||
</specimenCount>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="1E93BE8AFF84FFB1FF6AF9E54A77DDC3" lastPageId="14" lastPageNumber="185" pageId="10" pageNumber="181" type="description">
|
||
<paragraph id="5636ED01FF84FFB5FF6AF9E54B1CDA64" blockId="10.[151,1437,1275,1876]" pageId="10" pageNumber="181">
|
||
<emphasis id="64FD3113FF84FFB5FF6AF9E5491ADBF4" bold="true" box="[199,315,1707,1732]" pageId="10" pageNumber="181">Osteology</emphasis>
|
||
. In general, the osteology of
|
||
<taxonomicName id="91899682FF84FFB5FD22F9E54BB8DBF4" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[655,921,1707,1732]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="181" phylum="Chordata" rank="species" species="petrophilus">
|
||
<emphasis id="64FD3113FF84FFB5FD22F9E54BB8DBF4" box="[655,921,1707,1732]" italics="true" pageId="10" pageNumber="181">Cerogobius petrophilus</emphasis>
|
||
</taxonomicName>
|
||
is remarkably conservative in comparison to other gobies. The overall osteology shows a broad suite of characters that are shared across most generalized of species of the
|
||
<taxonomicName id="91899682FF84FFB5FE97F9BD4985DA3C" box="[314,420,1779,1804]" class="Actinopterygii" family="Gobiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="181" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
|
||
, especially species with a 32211-0 dorsal pterygiophore pattern and six spines in the first dorsal fin. Images of the main skeletal elements (
|
||
<figureCitation id="CEB2F184FF84FFB5FD7BF8594B07DA00" box="[726,806,1815,1840]" captionStart="FIGURE 6" captionStartId="11.[151,250,1390,1412]" captionTargetBox="[359,1247,202,1368]" captionTargetId="figure@11.[340,1247,180,1368]" captionTargetPageId="11" captionText="FIGURE 6. Cranium of Cerogobius petrophilus sp. nov., UW 158279. EO—exoccipital, EPO—epiotic, F- frontal, LE—lateral ethmoid, ME—median ethmoid, PTM—posttemporal, PTO—pterotic, SOC—supraoccipital, SPH—sphenotic." httpUri="https://zenodo.org/record/2589838/files/figure.png" pageId="10" pageNumber="181">Figs. 6</figureCitation>
|
||
–12) and generalized descriptions are provided here to serve as the basis of future comparative osteological studies.
|
||
</paragraph>
|
||
<caption id="02F6BD89FF85FFB4FF3AFA204CDDD892" httpUri="https://zenodo.org/record/2589838/files/figure.png" pageId="11" pageNumber="182" startId="11.[151,250,1390,1412]" targetBox="[359,1247,202,1368]" targetPageId="11">
|
||
<paragraph id="5636ED01FF85FFB4FF3AFA204CDDD892" blockId="11.[151,1435,1390,1442]" pageId="11" pageNumber="182">
|
||
<emphasis id="64FD3113FF85FFB4FF3AFA204933D8B3" bold="true" box="[151,274,1390,1412]" pageId="11" pageNumber="182">FIGURE 6.</emphasis>
|
||
Cranium of
|
||
<taxonomicName id="91899682FF85FFB4FE3CFA204A5ED8B4" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[401,639,1390,1412]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="182" phylum="Chordata" rank="species" species="petrophilus" status="sp. nov.">
|
||
<emphasis id="64FD3113FF85FFB4FE3CFA204A5ED8B4" box="[401,639,1390,1412]" italics="true" pageId="11" pageNumber="182">Cerogobius petrophilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FD3113FF85FFB4FD29FA204AF5D8B3" bold="true" box="[644,724,1390,1411]" pageId="11" pageNumber="182">
|
||
<taxonomicNameLabel id="7FCE8C68FF85FFB4FD29FA204AF5D8B3" box="[644,724,1390,1411]" pageId="11" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, UW 158279. EO—exoccipital, EPO—epiotic, F- frontal, LE—lateral ethmoid, ME—median ethmoid, PTM—posttemporal, PTO—pterotic, SOC—supraoccipital, SPH—sphenotic.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5636ED01FF85FFB4FF6AFA9E4B6FDBBE" blockId="11.[151,1436,1487,2012]" pageId="11" pageNumber="182">
|
||
<emphasis id="64FD3113FF85FFB4FF6AFA9E490AD8D7" box="[199,299,1488,1511]" italics="true" pageId="11" pageNumber="182">Cranium</emphasis>
|
||
(
|
||
<figureCitation id="CEB2F184FF85FFB4FE92FA8149ABD8D8" box="[319,394,1487,1512]" captionStart="FIGURE 6" captionStartId="11.[151,250,1390,1412]" captionTargetBox="[359,1247,202,1368]" captionTargetId="figure@11.[340,1247,180,1368]" captionTargetPageId="11" captionText="FIGURE 6. Cranium of Cerogobius petrophilus sp. nov., UW 158279. EO—exoccipital, EPO—epiotic, F- frontal, LE—lateral ethmoid, ME—median ethmoid, PTM—posttemporal, PTO—pterotic, SOC—supraoccipital, SPH—sphenotic." httpUri="https://zenodo.org/record/2589838/files/figure.png" pageId="11" pageNumber="182">Fig. 6</figureCitation>
|
||
). The roof of the cranium in
|
||
<taxonomicName id="91899682FF85FFB4FD5BFA9E4BBAD8D8" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[758,923,1487,1512]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="182" phylum="Chordata" rank="species" species="petrophilus">
|
||
<emphasis id="64FD3113FF85FFB4FD5BFA9E4BBAD8D8" box="[758,923,1487,1512]" italics="true" pageId="11" pageNumber="182">C. petrophilus</emphasis>
|
||
</taxonomicName>
|
||
is primarily comprised of the frontals and sphenotics anteriorly, and the supraoccipital, pterotic, epiotics posteriorly. The sphenotics have a prominent lateral flange that forms the posterolateral margin of the orbit, with the anterior orbit being formed from the lateral ethmoids. The supraoccipital bears a low dorsal crest on its posterior half, and is enclosed posteriorly by the epiotics, laterally by the pterotics, and anteriorly by the frontals. The pectoral girdle connects to the cranium dorsally via the posttemporal, which is attached to the epiotic.
|
||
</paragraph>
|
||
<paragraph id="5636ED01FF85FFB4FF6AF9D64C81DAEC" blockId="11.[151,1436,1487,2012]" pageId="11" pageNumber="182">
|
||
<emphasis id="64FD3113FF85FFB4FF6AF9D649F3DB9F" box="[199,466,1687,1712]" italics="true" pageId="11" pageNumber="182">Jaws and suspensorium</emphasis>
|
||
(
|
||
<figureCitation id="CEB2F184FF85FFB4FE4FF9D94A09DB9F" box="[482,552,1687,1712]" captionStart="FIGURE 7" captionStartId="12.[151,250,1090,1112]" captionTargetBox="[219,1374,193,1050]" captionTargetId="figure@12.[212,1374,193,1069]" captionTargetPageId="12" captionText="FIGURE 7. Suspensorium of Cerogobius petrophilus sp. nov., UW 158279. ART—articular, BR—branchiostegal ray, CHceratohyal, D—dentary, EH—epihyal, HYO—hyomandibular, IH—interhyal, MAX—maxilla, MT—metapterygoid, PALpalatine, PMX—premaxilla, POP—preopercle, PS—parasphenoid, PT—pterygoid (ectopterygoid), QU—quadrate, SYMsymplectic." httpUri="https://zenodo.org/record/2589840/files/figure.png" pageId="11" pageNumber="182">Fig. 7</figureCitation>
|
||
). The premaxilla is nearly as long as the maxilla, and bears a moderately long ascending process that is approximately as tall as half the horizontal length of the premaxilla. The dentary is stout and robust with a large coronoid process. A thin Meckel’s cartilage runs along the medial face of the dentary and continues posteriorly onto the articular. The anterior arm of the suspensorium is formed from the descending process of the palatine and ectopterygoid, the latter overlapping the former along approximately one-half of its length. The palatine is roughly T-shaped, with the anterior heads articulating with the lateral ethmoid and a lateral process on the anterior head of the maxilla. A wide metapterygoid abuts the symplectic along its posteroventral surface, and joins the quadrate via a cartilaginous connection anteroventrally. The symplectic connects to the hyomandibular via a thin layer of cartilage, unlike the connections between the metapterygoid and the hyomandibular, and between the hyomandibular and the preopercle, which are both to bone.
|
||
</paragraph>
|
||
<caption id="02F6BD89FF82FFB3FF3AFB0C492DD984" httpUri="https://zenodo.org/record/2589840/files/figure.png" pageId="12" pageNumber="183" startId="12.[151,250,1090,1112]" targetBox="[219,1374,193,1050]" targetPageId="12">
|
||
<paragraph id="5636ED01FF82FFB3FF3AFB0C492DD984" blockId="12.[151,1436,1090,1204]" pageId="12" pageNumber="183">
|
||
<emphasis id="64FD3113FF82FFB3FF3AFB0C4935D968" bold="true" box="[151,276,1090,1112]" pageId="12" pageNumber="183">FIGURE 7.</emphasis>
|
||
Suspensorium of
|
||
<taxonomicName id="91899682FF82FFB3FE62FB0C4A9FD968" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[463,702,1090,1112]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="183" phylum="Chordata" rank="species" species="petrophilus" status="sp. nov.">
|
||
<emphasis id="64FD3113FF82FFB3FE62FB0C4A9FD968" box="[463,702,1090,1112]" italics="true" pageId="12" pageNumber="183">Cerogobius petrophilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FD3113FF82FFB3FD68FB0D4B36D968" bold="true" box="[709,791,1091,1112]" pageId="12" pageNumber="183">
|
||
<taxonomicNameLabel id="7FCE8C68FF82FFB3FD68FB0D4B36D968" box="[709,791,1091,1112]" pageId="12" pageNumber="183" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, UW 158279. ART—articular, BR—branchiostegal ray, CHceratohyal, D—dentary, EH—epihyal, HYO—hyomandibular, IH—interhyal, MAX—maxilla, MT—metapterygoid, PALpalatine, PMX—premaxilla, POP—preopercle, PS—parasphenoid, PT—pterygoid (ectopterygoid), QU—quadrate, SYMsymplectic.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5636ED01FF82FFB3FF6AFA4B49A9D8E2" blockId="12.[151,1437,1285,1850]" pageId="12" pageNumber="183">
|
||
<emphasis id="64FD3113FF82FFB3FF6AFA4B496AD82E" box="[199,331,1285,1310]" italics="true" pageId="12" pageNumber="183">Hyoid arch</emphasis>
|
||
(
|
||
<figureCitation id="CEB2F184FF82FFB3FEF2FA4B498BD82E" box="[351,426,1285,1310]" captionStart="FIGURE 7" captionStartId="12.[151,250,1090,1112]" captionTargetBox="[219,1374,193,1050]" captionTargetId="figure@12.[212,1374,193,1069]" captionTargetPageId="12" captionText="FIGURE 7. Suspensorium of Cerogobius petrophilus sp. nov., UW 158279. ART—articular, BR—branchiostegal ray, CHceratohyal, D—dentary, EH—epihyal, HYO—hyomandibular, IH—interhyal, MAX—maxilla, MT—metapterygoid, PALpalatine, PMX—premaxilla, POP—preopercle, PS—parasphenoid, PT—pterygoid (ectopterygoid), QU—quadrate, SYMsymplectic." httpUri="https://zenodo.org/record/2589840/files/figure.png" pageId="12" pageNumber="183">Fig. 7</figureCitation>
|
||
). A relatively elongate, narrow interhyal serves as the main connection between the epihyal+ceratohyal complex and the hyomandibular. The branchiostegal rays are poorly ossified in the specimens examined here. The first branchiostegal ray originates from the thin, elongate anterior arm of the ceratohyal. The next three branchiostegal rays originate from the stout, deep, trapezoid-shaped posterior end of the ceratohyal, which itself is joined synchondrally do the epihyal. The last branchiostegal ray originates from the epihyal, which is triangular in shape.
|
||
</paragraph>
|
||
<paragraph id="5636ED01FF82FFB3FF6AFA934A51DA0A" blockId="12.[151,1437,1285,1850]" pageId="12" pageNumber="183">
|
||
<emphasis id="64FD3113FF82FFB3FF6AFA9349D1D8C6" box="[199,496,1501,1526]" italics="true" pageId="12" pageNumber="183">Pectoral and pelvic girdle</emphasis>
|
||
(
|
||
<figureCitation id="CEB2F184FF82FFB3FDACFA934A51D8C6" box="[513,624,1501,1526]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="13.[151,250,1149,1171]" captionStartId-1="14.[151,250,1249,1271]" captionTargetBox-0="[241,1345,193,1119]" captionTargetBox-1="[174,1413,293,1227]" captionTargetId-0="figure@13.[241,1345,193,1127]" captionTargetId-1="figure@14.[174,1413,293,1227]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 8. Pectoral girdle of Cerogobius petrophilus sp. nov., UW 158279. CL—cleithrum, COR—coracoid, PTMposttemporal, RAD—radials, SCA—scapula, SCL—supracleithrum." captionText-1="FIGURE 9. Pelvic girdle of Cerogobius petrophilus sp. nov., UW 158279. CL—cleithrum, PIC—pelvic intercleithral cartilage, PV—pelvic bone, VIC—ventral intercleithral cartilage." httpUri-0="https://zenodo.org/record/2589842/files/figure.png" httpUri-1="https://zenodo.org/record/2589844/files/figure.png" pageId="12" pageNumber="183">Figs. 8–9</figureCitation>
|
||
). The posttemporal has two well-developed anterior limbs that connect the pectoral girdle to the neurocranium via attachments to the epiotic (dorsal limb) and the intercalar (ventral limb). There are four pectoral radials that are poorly ossified and surrounded by a thin layer of cartilage. The dorsalmost radial is small and triangular, with the remaining radials being larger and square to dumbbell shaped. The scapula is small, unossified, and has a large foramen dorsally. The scapula synchondrally connects the cleithrum anteriorly, the coracoid ventrally, and the pectoral radials laterally. The cleithrum has a deeply forked dorsal surface, and is attached ligamentously to the supracleithrum. Ventrally, the cleithrum has two pointed processes. The elongate anterior process connects to the corresponding process on the cleithrum of the opposing side via the ventral intercleithral cartilage. The shorter posterior process on the ventral surface of the cleithrum connects to the pelvic bone via the pelvic intercleithral cartilage.
|
||
</paragraph>
|
||
<caption id="02F6BD89FF83FFB2FF3AFB334B71D982" httpUri="https://zenodo.org/record/2589842/files/figure.png" pageId="13" pageNumber="184" startId="13.[151,250,1149,1171]" targetBox="[241,1345,193,1119]" targetPageId="13">
|
||
<paragraph id="5636ED01FF83FFB2FF3AFB334B71D982" blockId="13.[151,1435,1149,1202]" pageId="13" pageNumber="184">
|
||
<emphasis id="64FD3113FF83FFB2FF3AFB334939D9A2" bold="true" box="[151,280,1149,1171]" pageId="13" pageNumber="184">FIGURE 8.</emphasis>
|
||
Pectoral girdle of
|
||
<taxonomicName id="91899682FF83FFB2FE41FB334AC1D9A3" authority="Kovačić & Bogorodsky & Troyer & Tornabene, 2019" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[492,736,1149,1171]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="184" phylum="Chordata" rank="species" species="petrophilus" status="sp. nov.">
|
||
<emphasis id="64FD3113FF83FFB2FE41FB334AC1D9A3" box="[492,736,1149,1171]" italics="true" pageId="13" pageNumber="184">Cerogobius petrophilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="64FD3113FF83FFB2FD41FB334B63D9A2" bold="true" box="[748,834,1149,1170]" pageId="13" pageNumber="184">
|
||
<taxonomicNameLabel id="7FCE8C68FF83FFB2FD41FB334B63D9A2" box="[748,834,1149,1170]" pageId="13" pageNumber="184" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, UW 158279. CL—cleithrum, COR—coracoid, PTMposttemporal, RAD—radials, SCA—scapula, SCL—supracleithrum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5636ED01FF83FFB2FF6AFBB94AAFDB48" blockId="13.[151,1437,1271,2016]" pageId="13" pageNumber="184">
|
||
<emphasis id="64FD3113FF83FFB2FF6AFBB94967D820" box="[199,326,1271,1296]" italics="true" pageId="13" pageNumber="184">Gill arches</emphasis>
|
||
(
|
||
<figureCitation id="CEB2F184FF83FFB2FEFBFBB94994D820" box="[342,437,1271,1296]" captionStart="FIGURE 10" captionStartId="15.[151,250,1140,1162]" captionTargetBox="[368,1163,175,1138]" captionTargetId="figure@15.[340,1247,175,1165]" captionTargetPageId="15" captionText="FIGURE 10. Ventral gill arches of Cerogobius petrophilus sp. nov., UW 158279. BB—basibranchial, BH—basihyal, CBceratobranchial, DH—dorsal hypohyal, HB—hypobranchial, LPT—lower pharyngeal toothplate, UH—urohyal, VH—ventral hypohyal." httpUri="https://zenodo.org/record/2589846/files/figure.png" pageId="13" pageNumber="184">Figs. 10</figureCitation>
|
||
–11). The primary elements of the ventral gill arches consist of a series of five pairs of ceratobranchials. Ceratobranchials 1–4 are slender and elongate, with ceratobranchials 1–3 each being bound to hypobranchials 1–3, respectively. Hypobranchials 1 and 2 are slender and elongate. Hypobranchial 3 is short and stout. Ceratobranchial 4 connects directly to basibranchial 4, which is small and unossified. Basibranchials 1–3 are elongate and well ossified. Ceratobranchial 5 forms a slender lower pharyngeal tooth plate, which is completely separate from the tooth plate on the opposing side. The urohyal is broadly spatulate with a cartilaginous anterior margin. The dorsal gill arches are composed primarily of four pairs of epibranchials. Epibranchial 1 has a deeply forked dorsal head, with the posterior fork connecting to the pharyngobranchial toothplate 2 via an elongate interarcual cartilage. Epibranchials 2 and 3 articulate with pharyngobranchial toothplates 2 and 3 respectively. Epibranchial 4 articulates with pharyngobranchial toothplate 4, which is largely fused to pharyngobranchial toothplate 3 to form a single toothed surface.
|
||
</paragraph>
|
||
<paragraph id="5636ED01FF83FFB2FF6AF9CD4B50DAD0" blockId="13.[151,1437,1271,2016]" pageId="13" pageNumber="184">
|
||
<emphasis id="64FD3113FF83FFB2FF6AF9CD49CADBAC" box="[199,491,1667,1692]" italics="true" pageId="13" pageNumber="184">Axial and caudal skeleton</emphasis>
|
||
(Fig. 12). There are 10 precaudal vertebrae and 17 caudal vertebrae. The dorsal-fin pterygiophore pattern is 3-22110. Two anal-fin pterygiophores insert anterior to the first haemal arch. There are ossified epineurals visible in association with vertebrae 1–8, with unossified elements visible on at least the next two vertebrae. There are no dorsal or ventral postcleithra visible. Several elements in the axial and caudal skeleton are not well ossified or did not take up alizarin red stain, but are nevertheless still present as hard, translucent elements. Most second dorsal-fin pterygiophores are unstained, as are the epural, hypural 5, parhypural, and the ventral half of the last haemal spine. The urostyle is fused to hypurals 3 + 4 (fused), which are separated by gap from hypurals 1 + 2 (fused). The dorsal and ventral procurrent cartilage each support three unsegmented procurrent caudal-fin rays. Single segmented fin-rays are associated with the epural, hypural 5, and the parahypural. Hypurals 1 + 2 and 3 + 4 are each associated with seven branched fin-rays.
|
||
</paragraph>
|
||
<paragraph id="5636ED01FF80FFB1FF6AFFD94A77DDC3" blockId="14.[151,1436,151,243]" pageId="14" pageNumber="185">
|
||
<emphasis id="64FD3113FF80FFB1FF6AFFD949D4DD80" bold="true" box="[199,501,151,176]" pageId="14" pageNumber="185">Genetics and phylogeny</emphasis>
|
||
. The molecular phylogenetic analysis supports a close relationship between
|
||
<taxonomicName id="91899682FF80FFB1FF3AFFF749B4DDE2" baseAuthorityName="Klunzinger" baseAuthorityYear="1871" box="[151,405,185,210]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="185" phylum="Chordata" rank="species" species="vulgaris">
|
||
<emphasis id="64FD3113FF80FFB1FF3AFFF749B4DDE2" box="[151,405,185,210]" italics="true" pageId="14" pageNumber="185">Hetereleotris vulgaris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="91899682FF80FFB1FE76FFF74ACADDE2" authorityName="Kovačić & Bogorodsky & Troyer & Tornabene" authorityYear="2019" box="[475,747,185,210]" class="Actinopterygii" family="Gobiidae" genus="Cerogobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Perciformes" pageId="14" pageNumber="185" phylum="Chordata" rank="species" species="petrophilus">
|
||
<emphasis id="64FD3113FF80FFB1FE76FFF74ACADDE2" box="[475,747,185,210]" italics="true" pageId="14" pageNumber="185">Cerogobius petrophilus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="CEB2F184FF80FFB1FCAFFFF74B6FDDE1" box="[770,846,185,210]" captionStart="FIGURE 4" captionStartId="7.[151,250,1946,1968]" captionTargetBox="[205,1292,205,1916]" captionTargetId="figure@7.[201,1389,193,1926]" captionTargetPageId="7" captionText="FIGURE 4. Bayesian inference molecular phylogeny of Gobiidae inferred from 5704 bp of combined nuclear genes and mtDNA. “Priolepis lineage” and “Gobiosomatini” refer to clades from Agorreta et al. (2013). Support values are posterior probabilities." httpUri="https://zenodo.org/record/2589834/files/figure.png" pageId="14" pageNumber="185">Fig. 4</figureCitation>
|
||
), matching the evidences from morphology (see Remarks for the genus and Discussion).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |