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<mods:title id="3CC8DB56E3B9A3192BE7C373E0892D96">A taxonomic study on Epedanidae from Thailand including functional aspects of male genital morphology (Opiliones, Laniatores)</mods:title>
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<mods:namePart id="44449F41E95AE11FC4D8AF76D41A7D37">Zhang, Chao</mods:namePart>
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<treatment id="9000C3A3306A5DB2B9BF6DBB8F758C25" ID-GBIF-Taxon="162267203" LSID="urn:lsid:plazi:treatment:9000C3A3306A5DB2B9BF6DBB8F758C25" httpUri="http://treatment.plazi.org/id/9000C3A3306A5DB2B9BF6DBB8F758C25" lastPageNumber="25" pageId="0" pageNumber="25" scope_family="Epedanidae" scope_order="Opiliones" scope_subOrder="Laniatores">
<subSubSection id="EBC451E3B12FB6AB90556D6F70731342" pageId="0" pageNumber="25" type="nomenclature">
<paragraph id="628C07DB2F8F7975E8A42B946DE3325A" pageId="0" pageNumber="25">
<taxonomicName id="5CB3FAE60EDD1868A546C054D175E509" ID-CoL="7PTZQ" LSID="9000C3A3-306A-5DB2-B9BF-6DBB8F758C25" authority="Pocock, 1903" authorityName="Pocock" authorityYear="1903" class="Arachnida" family="Epedanidae" genus="Plistobunus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plistobunus" order="Opiliones" pageId="0" pageNumber="25" phylum="Arthropoda" rank="genus">Plistobunus Pocock, 1903</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="9F61204A6E1F9894A5395397D9651C11" pageId="0" pageNumber="25" type="reference_group">
<paragraph id="82828324BF1D911A089C0521F19795ED" pageId="0" pageNumber="25">
<taxonomicName id="4B137C7ABB343CE565A508A63D314953" authorityName="Pocock" authorityYear="1903" class="Arachnida" family="Epedanidae" genus="Plistobunus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plistobunus" order="Opiliones" pageId="0" pageNumber="25" phylum="Arthropoda" rank="genus">Plistobunus</taxonomicName>
<bibRefCitation id="EB440E582E667CDF2CB80D346EAFA470" DOI="https://doi.org/10.1080/00222930308678797" author="Pocock, RI" journalOrPublisher="The Annals and Magazine of Natural History" pageId="0" pageNumber="25" pagination="433 - 450" refId="B11" refString="Pocock, RI, 1903. Fifteen new species and two new genera of tropical southern Opiliones. The Annals and Magazine of Natural History 11 (65): 433 - 450, DOI: https://doi.org/10.1080/00222930308678797" title="Fifteen new species and two new genera of tropical southern Opiliones." url="https://doi.org/10.1080/00222930308678797" volume="11" year="1903">Pocock 1903</bibRefCitation>
: 447;
<bibRefCitation id="AB7A294C1B63377385ABE20D8D547206" author="Roewer, CF" journalOrPublisher="A, Original-Arbeiten" pageId="0" pageNumber="25" pagination="1 - 242" refId="B12" refString="Roewer, CF, 1912. Die Familien der Assamiiden und Phalangodiden der Opiliones - Laniatores. (= Assamiden, Dampetriden, Phalangodiden, Epedaniden, Biantiden, Zalmoxiden, Samoiden, Palpipediden anderer Autoren). Archiv fuer Naturgeschichte, Berlin, Abt. A, Original-Arbeiten 78 (3): 1 - 242" title="Die Familien der Assamiiden und Phalangodiden der Opiliones - Laniatores. (= Assamiden, Dampetriden, Phalangodiden, Epedaniden, Biantiden, Zalmoxiden, Samoiden, Palpipediden anderer Autoren). Archiv fuer Naturgeschichte, Berlin, Abt." volume="78" year="1912">Roewer 1912</bibRefCitation>
: 232;
<bibRefCitation id="7B85F40B2DB986EBBDF8410239F1846E" author="Roewer, CF" journalOrPublisher="Gustav Fischer, Jena" pageId="0" pageNumber="25" refId="B14" refString="Roewer, CF, 1923. Die Weberknechte der Erde. Systematische Bearbeitung der bisher bekannten Opiliones. Gustav Fischer, Jena" title="Die Weberknechte der Erde. Systematische Bearbeitung der bisher bekannten Opiliones." year="1923">1923</bibRefCitation>
: 207;
<bibRefCitation id="4E05C8AA69460FA665C072C465103423" author="Roewer, CF" journalOrPublisher="Veroeffentlichungen aus dem Deutschen Kolonial- und Uebersee-Museum in Bremen" pageId="0" pageNumber="25" pagination="81 - 169" refId="B16" refString="Roewer, CF, 1938. Ueber Acrobuninae, Epedaninae und Sarasinicinae. Weitere Weberknechte IX. (9. Ergaenzung der &quot;Weberknechte der Erde&quot; 1923). Veroeffentlichungen aus dem Deutschen Kolonial- und Uebersee-Museum in Bremen 2 (2): 81 - 169" title="Ueber Acrobuninae, Epedaninae und Sarasinicinae. Weitere Weberknechte IX. (9. Ergaenzung der &quot; Weberknechte der Erde &quot; 1923)." volume="2" year="1938">1938</bibRefCitation>
: 124;
<bibRefCitation id="A7DFB154A77E7951EB705E928C112BEE" DOI="https://doi.org/10.3897/zookeys.112.1110" author="Lian, WG" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="39 - 52" refId="B8" refString="Lian, WG, Zhang, C, Zhang, F, 2011. Review of the genus Plistobunus Pocock, 1903, with description of a new species from Hainan Island, China (Opiliones, Laniatores, Epedanidae). ZooKeys 112: 39 - 52, DOI: https://doi.org/10.3897/zookeys.112.1110" title="Review of the genus Plistobunus Pocock, 1903, with description of a new species from Hainan Island, China (Opiliones, Laniatores, Epedanidae)." url="https://doi.org/10.3897/zookeys.112.1110" volume="112" year="2011">Lian et al. 2011</bibRefCitation>
: 41.
</paragraph>
</subSubSection>
<subSubSection id="60024D4FE1FB8B4DC8BE308052F5EDD0" pageId="0" pageNumber="25" type="type species">
<paragraph id="8BA02FA854723844027C5DF5F41B5719" pageId="0" pageNumber="25">Type species.</paragraph>
<paragraph id="A70E89CAAEFC10A577586DD8237C3395" pageId="0" pageNumber="25">
<taxonomicName id="26DBEA8DBEFC024985B384AF0E192803" authorityName="Pocock" authorityYear="1903" class="Arachnida" family="Epedanidae" genus="Plistobunus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plistobunus rapax" order="Opiliones" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="rapax">
<emphasis id="FCFC2C8804D425D63EA366206A2765B5" italics="true" pageId="0" pageNumber="25">Plistobunus rapax</emphasis>
</taxonomicName>
Pocock, 1903, by original designation.
</paragraph>
</subSubSection>
<subSubSection id="4F5F5E10A9924F693EFF2C89EBE0E7E8" pageId="0" pageNumber="25" type="extended diagnosis">
<paragraph id="2ABE3896C59EF309744047B0E7BACCC7" pageId="0" pageNumber="25">Extended diagnosis.</paragraph>
<paragraph id="E387CEC4A70C6AEDC4095E2DAFB865CB" pageId="0" pageNumber="25">
Medium-sized epedanines (2.40-3.57 mm) with a long median spine on the ocularium. Prosoma with a row of 4-6 setiferous tubercles on each side of the frontal margin. Areae II and IV with or without long spines. The basichelicerite of chelicerae elongate and armed with numerous tubercles, dorsally with three in males and with two in female (
<bibRefCitation id="377EDC88A3FD1216531B6DE3BD22B403" DOI="https://doi.org/10.3897/zookeys.112.1110" author="Lian, WG" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="39 - 52" refId="B8" refString="Lian, WG, Zhang, C, Zhang, F, 2011. Review of the genus Plistobunus Pocock, 1903, with description of a new species from Hainan Island, China (Opiliones, Laniatores, Epedanidae). ZooKeys 112: 39 - 52, DOI: https://doi.org/10.3897/zookeys.112.1110" title="Review of the genus Plistobunus Pocock, 1903, with description of a new species from Hainan Island, China (Opiliones, Laniatores, Epedanidae)." url="https://doi.org/10.3897/zookeys.112.1110" volume="112" year="2011">Lian et al. 2011</bibRefCitation>
: 42, figs 2, 3; 44: 9, 10; 46: 18, 19. Figs
<figureCitation id="2C9B26A77107A138501D57A53B8804DF" captionStart="Figures 32–39" captionStartId="F5" captionText="Figures 32 - 39. Plistobunus jaegeri sp. nov., male holotype 32 body, lateral view 33 left chelicera, medial view 34 same, lateral view 35 left basichelicerite, dorsal view 36 left cheliceral hand, frontal view 37 left cheliceral fingers, frontal view 38 left pedipalp, medial view 39 same, lateral view. Scale bars: 1 mm (32 - 36, 38, 39), 0.5 mm (37)." figureDoi="10.3897/zookeys.915.47626.figures32-39" httpUri="https://binary.pensoft.net/fig/383025" pageId="0" pageNumber="25">33-35</figureCitation>
,
<figureCitation id="07755B3293E0809594A616AADA7E1536" captionStart="Figures 40–54" captionStartId="F6" captionText="Figures 40 - 54. Plistobunus jaegeri sp. nov. male (Figs 40 - 44, holotype) and female (Figs 45 - 54, paratype) 40, 45 body, dorsal view 41 right leg I, retrolateral view 42 - 44, 46 femur of right legs, retrolateral view 42 leg II 43 leg III 44 leg IV 46 leg I 47 left chelicera, medial view 48 same, lateral view 49 left basichelicerite, dorsal view 50 left cheliceral hand, frontal view 51 left cheliceral fingers, frontal view 52 left pedipalp, medial view 53 ovipositor, ventral view 54 same, dorsal view. Scale bars: 1 mm (40 - 46, 52), 0.5 mm (47 - 51), 0.25 mm (53, 54)." figureDoi="10.3897/zookeys.915.47626.figures40-54" httpUri="https://binary.pensoft.net/fig/383026" pageId="0" pageNumber="25">47-49</figureCitation>
). Pedipalpal femur of male ventrally with 9-13 seta-tipped tubercles, and distally with one or two tubercles on medial side. Pedipalpal patella with at least three seta-tipped tubercles. Distitarsus of leg I with two segments. The distal part of pars basalis of penis widened and with a stereoscopic ventral plate. Stylus columnar. Capsula interna ventrally and capsula externa dorsally surround the stylus.
</paragraph>
</subSubSection>
<subSubSection id="C4B3A7ED6EE73922A35F001FD6474D18" pageId="0" pageNumber="25" type="included species">
<paragraph id="44FF0C02D57367AC8E39D335FBF8BF50" pageId="0" pageNumber="25">Included species.</paragraph>
<paragraph id="2BF3027C6C1586C6F4F53E6BB4D66D72" pageId="0" pageNumber="25">
<taxonomicName id="19FE821F3DC0ECB77C754BE5788D8704" authorityName="Pocock" authorityYear="1903" class="Arachnida" family="Epedanidae" genus="Plistobunus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plistobunus rapax" order="Opiliones" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="rapax">
<emphasis id="2BF3837C9337DEB65971A30813FAAFDB" italics="true" pageId="0" pageNumber="25">Plistobunus rapax</emphasis>
</taxonomicName>
Pocock, 1903,
<taxonomicName id="75B6D56CD068815EF5ACCEBAC29FF508" class="Arachnida" family="Epedanidae" genus="Plistobunus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plistobunus columnarius" order="Opiliones" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="columnarius">
<emphasis id="517501F0D6336F60362D93ABC3011239" italics="true" pageId="0" pageNumber="25">Plistobunus columnarius</emphasis>
</taxonomicName>
<bibRefCitation id="8791C97D132C390E1324E71BD2EEB349" DOI="https://doi.org/10.3897/zookeys.112.1110" author="Lian, WG" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="39 - 52" refId="B8" refString="Lian, WG, Zhang, C, Zhang, F, 2011. Review of the genus Plistobunus Pocock, 1903, with description of a new species from Hainan Island, China (Opiliones, Laniatores, Epedanidae). ZooKeys 112: 39 - 52, DOI: https://doi.org/10.3897/zookeys.112.1110" title="Review of the genus Plistobunus Pocock, 1903, with description of a new species from Hainan Island, China (Opiliones, Laniatores, Epedanidae)." url="https://doi.org/10.3897/zookeys.112.1110" volume="112" year="2011">Lian et al., 2011</bibRefCitation>
, and
<taxonomicName id="B03E519261763CD0A6AB5CF74EF2292D" authorityName="Zhang &amp; Martens" authorityYear="2020" class="Arachnida" family="Epedanidae" genus="Plistobunus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plistobunus jaegeri" order="Opiliones" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="jaegeri">
<emphasis id="7DD4DDCB3DA989D3DDD7E93773F1441D" italics="true" pageId="0" pageNumber="25">Plistobunus jaegeri</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="565DBF611930497BA237A6C406E3BE7F" pageId="0" pageNumber="25" type="distribution">
<paragraph id="CB73CE755DD343BA3675E4D7A93E76A9" pageId="0" pageNumber="25">Distribution.</paragraph>
<paragraph id="35D0288173C6BFACF22A61BA9C0C31B5" pageId="0" pageNumber="25">China (Hong Kong and Hainan Island), Thailand (Ubon Ratchathani).</paragraph>
</subSubSection>
<subSubSection id="6F77554C3255FD5DAFF23AF9503B52C3" pageId="0" pageNumber="25" type="sexual dimorphism">
<paragraph id="40B33BDB537B470B497336056549266D" pageId="0" pageNumber="25">Sexual dimorphism.</paragraph>
<paragraph id="518B2BCC4ABAD52D6D3CD145A0BE2E9C" pageId="0" pageNumber="25">The species of this genus possess conspicuous sexual dimorphism in the armature of the appendages. The tubercles or spines in male on the basichelicerite of chelicerae, and those of the femur of pedipalps and legs are much more robust or conspicuous than those of in female; the shape of body is very similar in both sexes.</paragraph>
<paragraph id="20BF936911BD037C7FFE74C8EB21A7AA" pageId="0" pageNumber="25">
The numbers of seta-tipped tubercles distally on medial side of pedipalpal femur gives an impression of the differences between the species. Male specimens of
<taxonomicName id="EFF678B856BBD82873928256A9F819EB" lsidName="P. rapax" pageId="0" pageNumber="25" rank="species" species="rapax">
<emphasis id="8337F54F1A1343151F937082CE871308" italics="true" pageId="0" pageNumber="25">P. rapax</emphasis>
</taxonomicName>
and
<taxonomicName id="125B56A2E0C62606DEF96E326F4B7F88" lsidName="P. columnarius" pageId="0" pageNumber="25" rank="species" species="columnarius">
<emphasis id="96B6B313C73EB3C31978BE6FC12769B5" italics="true" pageId="0" pageNumber="25">P. columnarius</emphasis>
</taxonomicName>
are described with two (
<bibRefCitation id="7DECCED7CA07A74DE47F3D80226B1DB8" DOI="https://doi.org/10.3897/zookeys.112.1110" author="Lian, WG" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="39 - 52" refId="B8" refString="Lian, WG, Zhang, C, Zhang, F, 2011. Review of the genus Plistobunus Pocock, 1903, with description of a new species from Hainan Island, China (Opiliones, Laniatores, Epedanidae). ZooKeys 112: 39 - 52, DOI: https://doi.org/10.3897/zookeys.112.1110" title="Review of the genus Plistobunus Pocock, 1903, with description of a new species from Hainan Island, China (Opiliones, Laniatores, Epedanidae)." url="https://doi.org/10.3897/zookeys.112.1110" volume="112" year="2011">Lian et al. 2011</bibRefCitation>
: 42, fig. 6; 44, fig. 14) and those of
<taxonomicName id="C2158D9EE45FEB677D93B7B1320EF3CA" lsidName="P. jaegeri" pageId="0" pageNumber="25" rank="species" species="jaegeri">
<emphasis id="8F87D9BB19FE2B10F25F2EAF2AB17C1A" italics="true" pageId="0" pageNumber="25">P. jaegeri</emphasis>
</taxonomicName>
sp. nov. with only one such tubercle (Fig.
<figureCitation id="FBE922F369372984CBC947E715397183" captionStart="Figures 32–39" captionStartId="F5" captionText="Figures 32 - 39. Plistobunus jaegeri sp. nov., male holotype 32 body, lateral view 33 left chelicera, medial view 34 same, lateral view 35 left basichelicerite, dorsal view 36 left cheliceral hand, frontal view 37 left cheliceral fingers, frontal view 38 left pedipalp, medial view 39 same, lateral view. Scale bars: 1 mm (32 - 36, 38, 39), 0.5 mm (37)." figureDoi="10.3897/zookeys.915.47626.figures32-39" httpUri="https://binary.pensoft.net/fig/383025" pageId="0" pageNumber="25">38</figureCitation>
). Females of
<taxonomicName id="35D7AEE6D3E6BC2BE98281F045921ADB" lsidName="P. columnarius" pageId="0" pageNumber="25" rank="species" species="columnarius">
<emphasis id="AF9970596E01CB5A09332975A71CD1D1" italics="true" pageId="0" pageNumber="25">P. columnarius</emphasis>
</taxonomicName>
(
<bibRefCitation id="F4F86F4FC965F8CB85DB5D18B2275F9D" DOI="https://doi.org/10.3897/zookeys.112.1110" author="Lian, WG" journalOrPublisher="ZooKeys" pageId="0" pageNumber="25" pagination="39 - 52" refId="B8" refString="Lian, WG, Zhang, C, Zhang, F, 2011. Review of the genus Plistobunus Pocock, 1903, with description of a new species from Hainan Island, China (Opiliones, Laniatores, Epedanidae). ZooKeys 112: 39 - 52, DOI: https://doi.org/10.3897/zookeys.112.1110" title="Review of the genus Plistobunus Pocock, 1903, with description of a new species from Hainan Island, China (Opiliones, Laniatores, Epedanidae)." url="https://doi.org/10.3897/zookeys.112.1110" volume="112" year="2011">Lian et al. 2011</bibRefCitation>
: 46, fig. 22) and
<taxonomicName id="68FB8A159C7A80E76BC2347D5ACC0BB7" lsidName="P. jaegeri" pageId="0" pageNumber="25" rank="species" species="jaegeri">
<emphasis id="D488547F8C999FF7FA016A4EDC7D3749" italics="true" pageId="0" pageNumber="25">P. jaegeri</emphasis>
</taxonomicName>
sp. nov. (Fig.
<figureCitation id="44306A99E3494EEE0375866456CE4AEA" captionStart="Figures 40–54" captionStartId="F6" captionText="Figures 40 - 54. Plistobunus jaegeri sp. nov. male (Figs 40 - 44, holotype) and female (Figs 45 - 54, paratype) 40, 45 body, dorsal view 41 right leg I, retrolateral view 42 - 44, 46 femur of right legs, retrolateral view 42 leg II 43 leg III 44 leg IV 46 leg I 47 left chelicera, medial view 48 same, lateral view 49 left basichelicerite, dorsal view 50 left cheliceral hand, frontal view 51 left cheliceral fingers, frontal view 52 left pedipalp, medial view 53 ovipositor, ventral view 54 same, dorsal view. Scale bars: 1 mm (40 - 46, 52), 0.5 mm (47 - 51), 0.25 mm (53, 54)." figureDoi="10.3897/zookeys.915.47626.figures40-54" httpUri="https://binary.pensoft.net/fig/383026" pageId="0" pageNumber="25">52</figureCitation>
) possess no tubercles of this size or position (female of
<taxonomicName id="0ACCEC56966BC09FAC89C3E31F916F46" lsidName="P. rapax" pageId="0" pageNumber="25" rank="species" species="rapax">
<emphasis id="96FE52CD39E85250E4BC23F9D343BD29" italics="true" pageId="0" pageNumber="25">P. rapax</emphasis>
</taxonomicName>
unknown).
</paragraph>
</subSubSection>
</treatment>
</document>
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