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<document id="EA7BDFC1061D0A62687FB71F4FBF08B7" ID-CLB-Dataset="55716" ID-DOI="10.5852/ejt.2022.817.1763" ID-GBIF-Dataset="daf1b8a9-0042-4b39-8f9c-df87cb1603f5" ID-ISSN="2118-9773" ID-Zenodo-Dep="6518289" ID-ZooBank="22C5B528-F7D7-40F2-8877-C30FF6057157" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1651674041341" checkinUser="felipe" docAuthor="Tshernyshev, Sergei E." docDate="2022" docId="1E7A87E1505C5C17FDBC96BF36CF6BA1" docLanguage="en" docName="ejt-817_58-77.pdf" docOrigin="European Journal of Taxonomy 817" docSource="http://dx.doi.org/10.5852/ejt.2022.817.1763" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tonyattalus vanharteni Tshernyshev 2022, gen. et sp. nov." docType="treatment" docUuid="C239C42C-69A9-428A-BE8B-A8BAA64D1F9B" docUuidSource="ZooBank" docVersion="6" lastPageNumber="66" masterDocId="E243FF9950595C1FFF8D9279370A6F24" masterDocTitle="Soft-winged flower beetles (Coleoptera: Malachiidae) of the United Arab Emirates" masterLastPageNumber="77" masterPageNumber="58" pageNumber="63" updateTime="1699329578614" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="393A59069D93324EB2E0B20628B107EE">Soft-winged flower beetles (Coleoptera: Malachiidae) of the United Arab Emirates</mods:title>
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<mods:namePart id="B83802A2717CF5E02ECFC838484A1ED9">Tshernyshev, Sergei E.</mods:namePart>
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<mods:affiliation id="9B408159BFDB97F87355F00575C59355">Institute of Systematics and Ecology of Animals, Siberian Branch of the Russian Academy of Sciences, Frunze Street 11, 630091 Novosibirsk, Russia. Tomsk State University, Lenina prospekt 36, Tomsk 634050, Russia. Email: sch-sch @ mail. ru</mods:affiliation>
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<mods:title id="853EF5A0F1791442F3BE0BDAE8266A7A">European Journal of Taxonomy</mods:title>
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<mods:date id="B9E86D87CA6B6FE14ACEE9F7A1D13DE5">2022</mods:date>
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<mods:number id="48B4A79E856E58EBEE50383666A6B6D4">2022-04-29</mods:number>
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<mods:number id="0ACE2A632A51DBDCFE0A3F44C3657E6F">817</mods:number>
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<mods:classification id="668CE36D0074E7CD5891A72CA27E9D5A">journal article</mods:classification>
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<mods:identifier id="B7326FB57C8DE86341E41973FF2A16A8" type="DOI">10.5852/ejt.2022.817.1763</mods:identifier>
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<treatment id="1E7A87E1505C5C17FDBC96BF36CF6BA1" ID-DOI="http://doi.org/10.5281/zenodo.6521861" ID-GBIF-Taxon="195191935" ID-Zenodo-Dep="6521861" LSID="urn:lsid:zoobank.org:act:C239C42C-69A9-428A-BE8B-A8BAA64D1F9B" httpUri="http://treatment.plazi.org/id/1E7A87E1505C5C17FDBC96BF36CF6BA1" lastPageId="8" lastPageNumber="66" pageId="5" pageNumber="63">
<subSubSection id="DEC9657C505C5C1AFDBC96BF33096BC5" box="[561,1027,1222,1249]" pageId="5" pageNumber="63" type="nomenclature">
<paragraph id="966C36F7505C5C1AFDBC96BF33096BC5" blockId="5.[354,1234,1222,1319]" box="[561,1027,1222,1249]" pageId="5" pageNumber="63">
<heading id="CD24819B505C5C1AFDBC96BF33096BC5" box="[561,1027,1222,1249]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="63" reason="2">
<taxonomicName id="51D34D74505C5C1AFDBC96BF34416BC5" authority="Tshernyshev, 2022" authorityName="Tshernyshev" authorityYear="2022" box="[561,843,1222,1249]" class="Insecta" family="Melyridae" genus="Tonyattalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="species" species="vanharteni" status="gen. et sp. nov.">
<emphasis id="A4A7EAE5505C5C1AFDBC96BF34416BC5" bold="true" box="[561,843,1222,1249]" italics="true" pageId="5" pageNumber="63">Tonyattalus vanharteni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF94579E505C5C1AFCDE96BE33096BC5" box="[851,1027,1223,1249]" pageId="5" pageNumber="63" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DEC9657C505C5C1AFEEF9693345E6A03" pageId="5" pageNumber="63" type="description">
<paragraph id="966C36F7505C5C1AFEEF969333D86A20" blockId="5.[354,1234,1222,1319]" box="[354,1234,1258,1284]" pageId="5" pageNumber="63">
<uri id="E2423AF5505C5C1AFEEF969333D86A20" box="[354,1234,1258,1284]" pageId="5" pageNumber="63">
urn:lsid:zoobank.org:act:
<uuid id="E2750C22505C5C1AFD0F969333D86A20" box="[642,1234,1258,1284]" pageId="5" pageNumber="63">C239C42C-69A9-428A-BE8B-A8BAA64D1F9B</uuid>
</uri>
</paragraph>
<paragraph id="966C36F7505C5C1AFD529774345E6A03" blockId="5.[354,1234,1222,1319]" box="[735,852,1293,1319]" pageId="5" pageNumber="63">
<figureCitation id="0EE82A72505C5C1AFD529774345E6A03" box="[735,852,1293,1319]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="5" pageNumber="63">Figs 1–10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DEC9657C505C5C1AFF30973733C66AF9" pageId="5" pageNumber="63" type="diagnosis">
<paragraph id="966C36F7505C5C1AFF30973736C66A4C" blockId="5.[189,460,1358,1384]" box="[189,460,1358,1384]" pageId="5" pageNumber="63">
<heading id="CD24819B505C5C1AFF30973736C66A4C" bold="true" box="[189,460,1358,1384]" fontSize="11" level="3" pageId="5" pageNumber="63" reason="3">
<emphasis id="A4A7EAE5505C5C1AFF30973736C66A4C" bold="true" box="[189,460,1358,1384]" pageId="5" pageNumber="63">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="966C36F7505C5C1AFF30970433C66AF9" blockId="5.[189,1399,1405,1502]" pageId="5" pageNumber="63">
The new species is comparable with
<taxonomicName id="51D34D74505C5C1AFD0A970734E06AB3" authority="(Wittmer, 1980)" authorityName="Tshernyshev" authorityYear="2022" baseAuthorityName="Wittmer" baseAuthorityYear="1980" box="[647,1002,1405,1431]" class="Insecta" family="Melyridae" genus="Tonyattalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="species" species="buettikeri">
<emphasis id="A4A7EAE5505C5C1AFD0A970734116AB3" box="[647,795,1405,1431]" italics="true" pageId="5" pageNumber="63">T. buettikeri</emphasis>
(
<bibRefCitation id="F2424B06505C5C1AFCA3970434E86AB3" author="Wittmer W." box="[814,994,1405,1431]" pageId="5" pageNumber="63" pagination="114 - 118" refId="ref11519" refString="Wittmer W. 1980. Fauna of Saudi Arabia. Coleoptera: Fam. Drilidae, Malachiidae, Prionoceridae (2. Beitrag). Fauna of Saudi Arabia 2: 114 - 118." type="journal article" year="1980">Wittmer, 1980</bibRefCitation>
)
</taxonomicName>
, and differs in its yellow-brown colouration, the 3
<superScript id="61A69BBF505C5C1AFE0697D936936A8A" attach="left" box="[395,409,1440,1454]" fontSize="6" pageId="5" pageNumber="63">rd</superScript>
antennomere simple, lacking small stretched outer edge, 4
<superScript id="61A69BBF505C5C1AFBC697D933526A8A" attach="left" box="[1099,1112,1440,1454]" fontSize="6" pageId="5" pageNumber="63">th</superScript>
antennomere triangular, not dilated as the small flattened process at outer edge, apical antennomere 1.5 × shorter.
</paragraph>
</subSubSection>
<subSubSection id="DEC9657C505C5C1AFF30947C335A69B0" pageId="5" pageNumber="63" type="etymology">
<paragraph id="966C36F7505C5C1AFF30947C364F693B" blockId="5.[189,325,1541,1567]" box="[189,325,1541,1567]" pageId="5" pageNumber="63">
<heading id="CD24819B505C5C1AFF30947C364F693B" bold="true" box="[189,325,1541,1567]" fontSize="11" level="3" pageId="5" pageNumber="63" reason="3">
<emphasis id="A4A7EAE5505C5C1AFF30947C364F693B" bold="true" box="[189,325,1541,1567]" pageId="5" pageNumber="63">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="966C36F7505C5C1AFF30944A335A69B0" blockId="5.[189,1399,1587,1684]" pageId="5" pageNumber="63">
The new species, designated as the
<typeStatus id="49688855505C5C1AFDED944D3599696A" box="[608,659,1588,1614]" pageId="5" pageNumber="63">type</typeStatus>
species of the new genus ‘
<taxonomicName id="51D34D74505C5C1AFC5C944A33506969" authorityName="Tshernyshev" authorityYear="2022" box="[977,1114,1587,1613]" class="Insecta" family="Melyridae" genus="Tonyattalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="63" phylum="Arthropoda" rank="genus">
<emphasis id="A4A7EAE5505C5C1AFC5C944A33506969" box="[977,1114,1587,1613]" italics="true" pageId="5" pageNumber="63">Tonyattalus</emphasis>
</taxonomicName>
’, is named in honour of Antonius van Harten, enthusiastic organiser and participant of the
<collectingCountry id="EEC47667505C5C1AFC54942E331F6955" box="[985,1045,1623,1649]" name="United Arab Emirates" pageId="5" pageNumber="63">UAE</collectingCountry>
Insect Project, who collected series of soft-winged flower beetles in
<collectingCountry id="EEC47667505C5C1AFD099403348969B0" box="[644,899,1658,1684]" name="United Arab Emirates" pageId="5" pageNumber="63">United Arab Emirates</collectingCountry>
for the first time.
</paragraph>
</subSubSection>
<subSubSection id="DEC9657C505C5C18FF3094C234AE6C34" lastPageId="7" lastPageNumber="65" pageId="5" pageNumber="63" type="materials_examined">
<paragraph id="966C36F7505C5C1AFF3094C236A769F1" blockId="5.[189,429,1723,1749]" box="[189,429,1723,1749]" pageId="5" pageNumber="63">
<heading id="CD24819B505C5C1AFF3094C236A769F1" bold="true" box="[189,429,1723,1749]" fontSize="11" level="3" pageId="5" pageNumber="63" reason="3">
<emphasis id="A4A7EAE5505C5C1AFF3094C236A769F1" bold="true" box="[189,429,1723,1749]" pageId="5" pageNumber="63">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="966C36F7505C5C1AFF6794933450686E" blockId="5.[189,1398,1770,1867]" pageId="5" pageNumber="63">
<materialsCitation id="26BB3CAA505C5C1AFF6794933450686E" ID-GBIF-Occurrence="3762960328" collectingDate="2007-03-17" collectingDateMax="2007-03-24" collectingDateMin="2007-03-17" collectingMethod="light trap" collectionCode="ISEA" collectorName="A. van Harten" country="United Arab Emirates" latitude="25.17" location="Sharjah Desert Park" longLatPrecision="748" longitude="55.42" pageId="5" pageNumber="63" specimenCode="SCH_ISEA_000133" specimenCount="1" specimenCount-male="1" stateProvince="Ash Shariqah" typeStatus="holotype">
<emphasis id="A4A7EAE5505C5C1AFF67949336576820" bold="true" box="[234,349,1770,1796]" pageId="5" pageNumber="63">
<typeStatus id="49688855505C5C1AFF67949336576820" box="[234,349,1770,1796]" pageId="5" pageNumber="63" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="EEC47667505C5C1AFF30957735106803" box="[189,538,1805,1832]" name="United Arab Emirates" pageId="5" pageNumber="63">UNITED ARAB EMIRATES</collectingCountry>
•
<specimenCount id="80D5FD7E505C5C1AFDB895773544680C" box="[565,590,1806,1832]" pageId="5" pageNumber="63" type="male">♂</specimenCount>
;
<location id="930C602C505C5C1AFDD1957434476803" LSID="urn:lsid:plazi:treatment:1E7A87E1505C5C17FDBC96BF36CF6BA1:930C602C505C5C1AFDD1957434476803" box="[604,845,1805,1832]" country="United Arab Emirates" latitude="25.17" longLatPrecision="748" longitude="55.42" name="Sharjah Desert Park" pageId="5" pageNumber="63" stateProvince="Ash Shariqah">Sharjah Desert Park</location>
;
<geoCoordinate id="F3E75030505C5C1AFCD4957434CD680C" box="[857,967,1805,1832]" degrees="25.17" direction="north" orientation="latitude" pageId="5" pageNumber="63" precision="555" value="25.17">25.17° N</geoCoordinate>
,
<geoCoordinate id="F3E75030505C5C1AFC5995743335680C" box="[980,1087,1805,1832]" degrees="55.42" direction="east" orientation="longitude" pageId="5" pageNumber="63" precision="555" value="55.42">55.42° E</geoCoordinate>
;
<date id="E26D1037505C5C1AFBC0957432166803" box="[1101,1308,1805,1832]" pageId="5" pageNumber="63" value="2007-03-17" valueMax="2007-03-24" valueMin="2007-03-17">
<collectingDate id="F229E9DF505C5C1AFBC0957432166803" box="[1101,1308,1805,1832]" pageId="5" pageNumber="63" value="2007-03-17" valueMax="2007-03-24" valueMin="2007-03-17">17–24 Mar. 2007</collectingDate>
</date>
;
<collectorName id="3B265321505C5C1AFAAA95743607686F" pageId="5" pageNumber="63">A. van Harten</collectorName>
leg. (9043 UAE);
<collectingMethod id="4F924EE0505C5C1AFE659549355C686F" box="[488,598,1840,1867]" pageId="5" pageNumber="63">light trap</collectingMethod>
;
<specimenCode id="C6759E8C505C5C1AFDEF9549345F686E" box="[610,853,1840,1866]" pageId="5" pageNumber="63">SCH_ISEA_000133</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="966C36F7505C5C1AFF679508320E688B" blockId="5.[189,1284,1905,1967]" pageId="5" pageNumber="63">
<materialsCitation id="26BB3CAA505C5C1AFF679508320E688B" ID-GBIF-Occurrence="3762960324" collectingDate="2007-03-17" collectingDateMax="2007-03-24" collectingDateMin="2007-03-17" collectingMethod="light trap" collectionCode="ISEA" collectorName="A. van Harten" country="United Arab Emirates" latitude="25.17" location="UNITED ARAB EMIRATES" longLatPrecision="748" longitude="55.42" pageId="5" pageNumber="63" specimenCode="SCH_ISEA_000134" specimenCount="1" specimenCount-female="1" stateProvince="Ash Shariqah" typeStatus="allotype">
<emphasis id="A4A7EAE5505C5C1AFF679508365F68AF" bold="true" box="[234,341,1905,1931]" pageId="5" pageNumber="63">
<typeStatus id="49688855505C5C1AFF679508365F68AF" box="[234,341,1905,1931]" pageId="5" pageNumber="63" type="allotype">Allotype</typeStatus>
</emphasis>
<collectingCountry id="EEC47667505C5C1AFF3095EC3512688B" box="[189,536,1941,1967]" name="United Arab Emirates" pageId="5" pageNumber="63">UNITED ARAB EMIRATES</collectingCountry>
•
<specimenCount id="80D5FD7E505C5C1AFDBC95EC3542688B" box="[561,584,1941,1967]" pageId="5" pageNumber="63" type="female">♀</specimenCount>
; same collection data as for holotype;
<specimenCode id="C6759E8C505C5C1AFB8095EC320A688B" box="[1037,1280,1941,1967]" pageId="5" pageNumber="63">SCH_ISEA_000134</specimenCode>
.
</materialsCitation>
</paragraph>
<caption id="C2AC667F505F5C19FF309518347B68C2" ID-DOI="http://doi.org/10.5281/zenodo.6518293" ID-Zenodo-Dep="6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="6" pageNumber="64" startId="6.[189,242,1889,1915]" targetBox="[222,1352,267,1823]" targetPageId="6">
<paragraph id="966C36F7505F5C19FF309518347B68C2" blockId="6.[188,1399,1889,2022]" pageId="6" pageNumber="64">
<emphasis id="A4A7EAE5505F5C19FF30951836356858" bold="true" box="[189,319,1889,1916]" pageId="6" pageNumber="64">Figs 1–10.</emphasis>
<taxonomicName id="51D34D74505F5C19FEC7951B35556858" authority="Tshernyshev, 2022" authorityName="Tshernyshev" authorityYear="2022" box="[330,607,1890,1916]" class="Insecta" family="Melyridae" genus="Tonyattalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="64" phylum="Arthropoda" rank="species" species="vanharteni" status="gen. et sp. nov.">
<emphasis id="A4A7EAE5505F5C19FEC7951B35556858" box="[330,607,1890,1916]" italics="true" pageId="6" pageNumber="64">Tonyattalus vanharteni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF94579E505F5C19FDE7951B342D6858" box="[618,807,1890,1916]" pageId="6" pageNumber="64" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10).
<emphasis id="A4A7EAE5505F5C19FF7895FC362E68BB" bold="true" box="[245,292,1925,1951]" pageId="6" pageNumber="64">1, 9</emphasis>
. External appearance, dorsal view.
<emphasis id="A4A7EAE5505F5C19FD5C95FC340668BB" bold="true" box="[721,780,1925,1951]" pageId="6" pageNumber="64">2, 10</emphasis>
. External appearance, lateral view.
<emphasis id="A4A7EAE5505F5C19FB3195FC33C368BB" bold="true" box="[1212,1225,1925,1951]" pageId="6" pageNumber="64">3</emphasis>
. Left antenna.
<emphasis id="A4A7EAE5505F5C19FF3295D137C668E6" bold="true" box="[191,204,1960,1986]" pageId="6" pageNumber="64">4</emphasis>
. Head and pronotum, dorsal view.
<emphasis id="A4A7EAE5505F5C19FDEF95D1357B68E6" bold="true" box="[610,625,1960,1986]" pageId="6" pageNumber="64">5</emphasis>
. Left anterior tarsus.
<emphasis id="A4A7EAE5505F5C19FCE695D1347068E6" bold="true" box="[875,890,1960,1986]" pageId="6" pageNumber="64">6</emphasis>
. Pygidium.
<emphasis id="A4A7EAE5505F5C19FB8495D1331268E6" bold="true" box="[1033,1048,1960,1986]" pageId="6" pageNumber="64">7</emphasis>
. Ultimate abdominal ventrite.
<emphasis id="A4A7EAE5505F5C19FF3195B237C168C1" bold="true" box="[188,203,1995,2021]" pageId="6" pageNumber="64">8</emphasis>
. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="966C36F7505E5C18FF67937034AE6C34" blockId="7.[189,1399,265,784]" pageId="7" pageNumber="65">
<materialsCitation id="26BB3CAA505E5C18FF679370321E6E62" ID-GBIF-Occurrence="3762960305" collectingDate="2007-03-17" collectingDateMax="2007-03-24" collectingDateMin="2007-03-17" collectingMethod="light trap" collectionCode="ISEA" collectorName="A. van Harten" country="United Arab Emirates" latitude="25.17" location="UNITED ARAB EMIRATES" longLatPrecision="748" longitude="55.42" pageId="7" pageNumber="65" specimenCode="SCH_ISEA_000135" specimenCount="1" specimenCount-male="1" stateProvince="Ash Shariqah" typeStatus="paratype">
<emphasis id="A4A7EAE5505E5C18FF67937036636E07" bold="true" box="[234,361,265,291]" pageId="7" pageNumber="65">
<typeStatus id="49688855505E5C18FF67937036636E07" box="[234,361,265,291]" pageId="7" pageNumber="65" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="EEC47667505E5C18FF30935535126E62" box="[189,536,300,326]" name="United Arab Emirates" pageId="7" pageNumber="65">UNITED ARAB EMIRATES</collectingCountry>
•
<specimenCount id="80D5FD7E505E5C18FDBF9355356B6E62" box="[562,609,300,326]" pageId="7" pageNumber="65" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<specimenCode id="C6759E8C505E5C18FBA99352321E6E62" box="[1060,1300,299,326]" pageId="7" pageNumber="65">SCH_ISEA_000135</specimenCode>
</materialsCitation>
•
<materialsCitation id="26BB3CAA505E5C18FAA0935534AF6E4D" ID-GBIF-Occurrence="3762960318" collectingDate="2007-03-17" collectingDateMax="2007-03-24" collectingDateMin="2007-03-17" collectingMethod="light trap" collectionCode="ISEA" collectorName="A. van Harten" country="United Arab Emirates" latitude="25.17" location="Desert Park" longLatPrecision="748" longitude="55.42" pageId="7" pageNumber="65" specimenCode="SCH_ISEA_000136-139" specimenCount="4" specimenCount-female="4" stateProvince="Ash Shariqah" typeStatus="paratype">
<specimenCount id="80D5FD7E505E5C18FAA09355327A6E62" box="[1325,1392,300,326]" pageId="7" pageNumber="65" type="female">4 ♀♀</specimenCount>
, same collection data as for holotype;
<specimenCode id="C6759E8C505E5C18FDF7933634AF6E4D" box="[634,933,335,361]" pageId="7" pageNumber="65">SCH_ISEA_000136–139</specimenCode>
</materialsCitation>
•
<materialsCitation id="26BB3CAA505E5C18FC4D9336321A6EA8" ID-GBIF-Occurrence="3762960327" collectingDate="2006-04-01" collectingDateMax="2006-05-02" collectingDateMin="2006-04-01" collectingMethod="Malaise trap" collectionCode="ISEA" collectorName="A. Van Harten" country="United Arab Emirates" latitude="24.36" location="al-Ajban" longLatPrecision="751" longitude="55.01" pageId="7" pageNumber="65" specimenCode="SCH_ISEA_000140-141" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="80D5FD7E505E5C18FC4D9336330E6E4D" box="[960,1028,335,361]" pageId="7" pageNumber="65" type="female">2 ♀♀</specimenCount>
;
<location id="930C602C505E5C18FB99933633756E4D" LSID="urn:lsid:plazi:treatment:1E7A87E1505C5C17FDBC96BF36CF6BA1:930C602C505E5C18FB99933633756E4D" box="[1044,1151,335,361]" country="United Arab Emirates" latitude="24.36" longLatPrecision="751" longitude="55.01" name="al-Ajban" pageId="7" pageNumber="65">al-Ajban</location>
;
<geoCoordinate id="F3E75030505E5C18FB06933633F36E4D" box="[1163,1273,335,361]" degrees="24.36" direction="north" orientation="latitude" pageId="7" pageNumber="65" precision="555" value="24.36">24.36° N</geoCoordinate>
,
<geoCoordinate id="F3E75030505E5C18FA8B9336327B6E4D" box="[1286,1393,335,362]" degrees="55.01" direction="east" orientation="longitude" pageId="7" pageNumber="65" precision="555" value="55.01">55.01° E</geoCoordinate>
;
<date id="E26D1037505E5C18FF30930B36A36EA8" box="[189,425,370,397]" pageId="7" pageNumber="65" value="2006-04-01" valueMax="2006-05-02" valueMin="2006-04-01">
<collectingDate id="F229E9DF505E5C18FF30930B36A36EA8" box="[189,425,370,397]" pageId="7" pageNumber="65" value="2006-04-01" valueMax="2006-05-02" valueMin="2006-04-01">1 Apr.–2 May 2006</collectingDate>
</date>
;
<collectorName id="3B265321505E5C18FE38930B356A6EA9" box="[437,608,370,397]" pageId="7" pageNumber="65">A. Van Harten</collectorName>
leg. (5664 UAE);
<collectingMethod id="4F924EE0505E5C18FCCF930B34D36EA9" box="[834,985,370,397]" pageId="7" pageNumber="65">Malaise trap</collectingMethod>
;
<specimenCode id="C6759E8C505E5C18FC6B930B321A6EA8" box="[998,1296,370,396]" pageId="7" pageNumber="65">SCH_ISEA_000140–141</specimenCode>
</materialsCitation>
•
<materialsCitation id="26BB3CAA505E5C18FAA6930B36956EF7" ID-GBIF-Occurrence="3762960307" collectingDate="2007-05-06" collectingDateMax="2007-05-13" collectingDateMin="2007-05-06" collectingMethod="light trap" collectionCode="ISEA" collectorName="A. Van Harten" country="United Arab Emirates" latitude="25.48" location="Wadi Bih dam" longLatPrecision="748" longitude="56.04" pageId="7" pageNumber="65" specimenCode="SCH_ ISEA_000142-144" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="80D5FD7E505E5C18FAA6930B327B6EA9" box="[1323,1393,370,397]" pageId="7" pageNumber="65" type="male">3 ♂♂</specimenCount>
;
<location id="930C602C505E5C18FF3093EC36606E8B" LSID="urn:lsid:plazi:treatment:1E7A87E1505C5C17FDBC96BF36CF6BA1:930C602C505E5C18FF3093EC36606E8B" box="[189,362,405,431]" country="United Arab Emirates" latitude="25.48" longLatPrecision="748" longitude="56.04" name="Wadi Bih dam" pageId="7" pageNumber="65">Wadi Bih dam</location>
;
<geoCoordinate id="F3E75030505E5C18FEFB93EC36E96E94" box="[374,483,405,432]" degrees="25.48" direction="north" orientation="latitude" pageId="7" pageNumber="65" precision="555" value="25.48">25.48° N</geoCoordinate>
,
<geoCoordinate id="F3E75030505E5C18FE7D93EC35506E94" box="[496,602,405,432]" degrees="56.04" direction="east" orientation="longitude" pageId="7" pageNumber="65" precision="555" value="56.04">56.04° E</geoCoordinate>
;
<date id="E26D1037505E5C18FDEA93EC342E6E8B" box="[615,804,405,432]" pageId="7" pageNumber="65" value="2007-05-06" valueMax="2007-05-13" valueMin="2007-05-06">
<collectingDate id="F229E9DF505E5C18FDEA93EC342E6E8B" box="[615,804,405,432]" pageId="7" pageNumber="65" value="2007-05-06" valueMax="2007-05-13" valueMin="2007-05-06">6–13 May 2007</collectingDate>
</date>
;
<collectorName id="3B265321505E5C18FCA293EC34D26E94" box="[815,984,405,432]" pageId="7" pageNumber="65">A. Van Harten</collectorName>
leg. (9463 UAE);
<collectingMethod id="4F924EE0505E5C18FB3993EC32286E94" box="[1204,1314,405,432]" pageId="7" pageNumber="65">light trap</collectingMethod>
;
<specimenCode id="C6759E8C505E5C18FAA293EC36956EF7" pageId="7" pageNumber="65">SCH_ ISEA_000142–144</specimenCode>
</materialsCitation>
•
<materialsCitation id="26BB3CAA505E5C18FE3093C032076EF7" ID-GBIF-Occurrence="3762960322" box="[445,1293,440,467]" collectingDate="2007-05-06" collectingDateMax="2007-05-13" collectingDateMin="2007-05-06" collectingMethod="light trap" collectionCode="ISEA" collectorName="A. Van Harten" country="United Arab Emirates" latitude="25.48" location="Wadi Bih" longLatPrecision="748" longitude="56.04" pageId="7" pageNumber="65" specimenCode="SCH_ISEA_000145-146" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="80D5FD7E505E5C18FE3093C035086EF7" box="[445,514,441,467]" pageId="7" pageNumber="65" type="female">2 ♀♀</specimenCount>
; same collection data as for preceding;
<specimenCode id="C6759E8C505E5C18FC6E93C132076EF7" box="[995,1293,440,467]" pageId="7" pageNumber="65">SCH_ISEA_000145–146</specimenCode>
</materialsCitation>
•
<materialsCitation id="26BB3CAA505E5C18FAA693C0356E6D3E" ID-GBIF-Occurrence="3762960323" collectingDate="2008-04-30" collectingDateMax="2008-05-25" collectingDateMin="2008-04-30" collectingMethod="light trap" collectionCode="ISEA" collectorName="A. Van Harten" country="United Arab Emirates" latitude="25.17" location="Sharjah Desert Park" longLatPrecision="748" longitude="55.42" pageId="7" pageNumber="65" specimenCode="SCH_ISEA_000147-151" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Ash Shariqah" typeStatus="paratype">
<specimenCount id="80D5FD7E505E5C18FAA693C0327B6EF7" box="[1323,1393,441,467]" pageId="7" pageNumber="65" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="80D5FD7E505E5C18FF3093A537F56ED2" box="[189,255,476,502]" pageId="7" pageNumber="65" type="female">2 ♀♀</specimenCount>
;
<location id="930C602C505E5C18FE8093A536F06ED2" LSID="urn:lsid:plazi:treatment:1E7A87E1505C5C17FDBC96BF36CF6BA1:930C602C505E5C18FE8093A536F06ED2" box="[269,506,476,502]" country="United Arab Emirates" latitude="25.17" longLatPrecision="748" longitude="55.42" name="Sharjah Desert Park" pageId="7" pageNumber="65" stateProvince="Ash Shariqah">Sharjah Desert Park</location>
;
<geoCoordinate id="F3E75030505E5C18FD8893A2357B6ED2" box="[517,625,475,502]" degrees="25.17" direction="north" orientation="latitude" pageId="7" pageNumber="65" precision="555" value="25.17">25.17° N</geoCoordinate>
,
<geoCoordinate id="F3E75030505E5C18FDF093A235EC6ED2" box="[637,742,475,502]" degrees="55.42" direction="east" orientation="longitude" pageId="7" pageNumber="65" precision="555" value="55.42">55.42° E</geoCoordinate>
;
<date id="E26D1037505E5C18FD7F93A534FE6ED2" box="[754,1012,475,502]" pageId="7" pageNumber="65" value="2008-04-30" valueMax="2008-05-25" valueMin="2008-04-30">
<collectingDate id="F229E9DF505E5C18FD7F93A534FE6ED2" box="[754,1012,475,502]" pageId="7" pageNumber="65" value="2008-04-30" valueMax="2008-05-25" valueMin="2008-04-30">30 Apr.–25 May 2008</collectingDate>
</date>
;
<collectorName id="3B265321505E5C18FC7393A533AF6ED2" box="[1022,1189,476,502]" pageId="7" pageNumber="65">A. Van Harten</collectorName>
leg. (9535 UAE);
<collectingMethod id="4F924EE0505E5C18FF30938636266D3D" box="[189,300,511,537]" pageId="7" pageNumber="65">light trap</collectingMethod>
;
<specimenCode id="C6759E8C505E5C18FEB79386356E6D3E" box="[314,612,511,538]" pageId="7" pageNumber="65">SCH_ISEA_000147–151</specimenCode>
</materialsCitation>
•
<materialsCitation id="26BB3CAA505E5C18FD0D938633326D19" ID-GBIF-Occurrence="3762960321" collectingDate="2006-09-24" collectingDateMax="2006-10-22" collectingDateMin="2006-09-24" collectingMethod="water traps" collectionCode="ISEA" collectorName="A. Van Harten" country="United Arab Emirates" latitude="25.18" location="Wadi Maidaq" longLatPrecision="748" longitude="56.07" pageId="7" pageNumber="65" specimenCode="SCH_ISEA_000152-154" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="80D5FD7E505E5C18FD0D938635A56D3D" box="[640,687,511,537]" pageId="7" pageNumber="65" type="male">1 ♂</specimenCount>
,
<specimenCount id="80D5FD7E505E5C18FD30938634086D3D" box="[701,770,511,537]" pageId="7" pageNumber="65" type="female">2 ♀♀</specimenCount>
;
<location id="930C602C505E5C18FC9C938634BE6D3D" LSID="urn:lsid:plazi:treatment:1E7A87E1505C5C17FDBC96BF36CF6BA1:930C602C505E5C18FC9C938634BE6D3D" box="[785,948,511,537]" country="United Arab Emirates" latitude="25.18" longLatPrecision="748" longitude="56.07" name="Wadi Maidaq" pageId="7" pageNumber="65">Wadi Maidaq</location>
;
<geoCoordinate id="F3E75030505E5C18FC4F9386333A6D3D" box="[962,1072,511,538]" degrees="25.18" direction="north" orientation="latitude" pageId="7" pageNumber="65" precision="555" value="25.18">25.18° N</geoCoordinate>
,
<geoCoordinate id="F3E75030505E5C18FBB0938633A36D3D" box="[1085,1193,511,538]" degrees="56.07" direction="east" orientation="longitude" pageId="7" pageNumber="65" precision="555" value="56.07">56.07° E</geoCoordinate>
;
<date id="E26D1037505E5C18FB3A938637F16D18" pageId="7" pageNumber="65" value="2006-09-24" valueMax="2006-10-22" valueMin="2006-09-24">
<collectingDate id="F229E9DF505E5C18FB3A938637F16D18" pageId="7" pageNumber="65" value="2006-09-24" valueMax="2006-10-22" valueMin="2006-09-24">24 Sep.–22 Oct. 2006</collectingDate>
</date>
;
<collectorName id="3B265321505E5C18FE89905B36A06D19" box="[260,426,546,573]" pageId="7" pageNumber="65">A. Van Harten</collectorName>
leg. (8343 UAE);
<collectingMethod id="4F924EE0505E5C18FD0D905A340E6D19" box="[640,772,547,573]" pageId="7" pageNumber="65">water traps</collectingMethod>
;
<specimenCode id="C6759E8C505E5C18FC82905B33326D19" box="[783,1080,546,573]" pageId="7" pageNumber="65">SCH_ISEA_000152–154</specimenCode>
</materialsCitation>
•
<materialsCitation id="26BB3CAA505E5C18FBC2905B34AF6D44" ID-GBIF-Occurrence="3762960329" collectingDate="2006-09-24" collectingDateMax="2006-10-22" collectingDateMin="2006-09-24" collectingMethod="water traps" collectionCode="ISEA" collectorName="A. Van Harten" country="United Arab Emirates" latitude="25.18" location="Wadi Maidaq" longLatPrecision="748" longitude="56.07" pageId="7" pageNumber="65" specimenCode="SCH_ISEA_000155-156" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="80D5FD7E505E5C18FBC2905B33766D19" box="[1103,1148,546,573]" pageId="7" pageNumber="65" type="male">1 ♂</specimenCount>
,
<specimenCount id="80D5FD7E505E5C18FB0A905B33B96D19" box="[1159,1203,546,573]" pageId="7" pageNumber="65" type="female">1 ♀</specimenCount>
; same collection data as for preceding (9482 UAE);
<specimenCode id="C6759E8C505E5C18FDF6903C34AF6D44" box="[635,933,581,608]" pageId="7" pageNumber="65">SCH_ISEA_000155–156</specimenCode>
</materialsCitation>
•
<materialsCitation id="26BB3CAA505E5C18FC44903C327C6DA7" ID-GBIF-Occurrence="3762960310" collectingDate="2007-11-18" collectingDateMax="2007-12-15" collectingDateMin="2007-11-18" collectingMethod="pitfall traps" collectionCode="NHML" collectorName="A. Van Harten" country="United Arab Emirates" latitude="25.18" location="Wadi Maidaq" longLatPrecision="748" longitude="56.07" pageId="7" pageNumber="65" specimenCode="NHML_000157-160" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="80D5FD7E505E5C18FC44903C33196D44" box="[969,1043,581,608]" pageId="7" pageNumber="65" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="80D5FD7E505E5C18FBA8903C33676D44" box="[1061,1133,581,608]" pageId="7" pageNumber="65" type="female">2 ♀♀</specimenCount>
; same locality as for preceding;
<date id="E26D1037505E5C18FECC901035446DA7" box="[321,590,616,643]" pageId="7" pageNumber="65" value="2007-11-18" valueMax="2007-12-15" valueMin="2007-11-18">
<collectingDate id="F229E9DF505E5C18FECC901035446DA7" box="[321,590,616,643]" pageId="7" pageNumber="65" value="2007-11-18" valueMax="2007-12-15" valueMin="2007-11-18">18 Nov.–15 Dec. 2007</collectingDate>
</date>
;
<collectorName id="3B265321505E5C18FDD4901034086DA7" box="[601,770,617,643]" pageId="7" pageNumber="65">A. Van Harten</collectorName>
leg.; (9488 UAE);
<collectingMethod id="4F924EE0505E5C18FC65901133796DA7" box="[1000,1139,616,643]" pageId="7" pageNumber="65">pitfall traps</collectingMethod>
;
<specimenCode id="C6759E8C505E5C18FBF29011327C6DA7" box="[1151,1398,616,643]" pageId="7" pageNumber="65">NHML_000157–160</specimenCode>
</materialsCitation>
•
<materialsCitation id="26BB3CAA505E5C18FF4290F536A66DED" ID-GBIF-Occurrence="3762960330" collectingDate="2007-03" collectingDateMax="2007-04-31" collectingDateMin="2007-03" collectingMethod="water traps" collectionCode="ISEA" collectorName="C. Tourenq" country="United Arab Emirates" latitude="25.24" location="Wadi Wurayah" longLatPrecision="748" longitude="56.17" pageId="7" pageNumber="65" specimenCode="SCH_ISEA_000161" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="80D5FD7E505E5C18FF4290F537F76D82" box="[207,253,652,678]" pageId="7" pageNumber="65" type="female">1 ♀</specimenCount>
;
<location id="930C602C505E5C18FE8190F536B56D82" LSID="urn:lsid:plazi:treatment:1E7A87E1505C5C17FDBC96BF36CF6BA1:930C602C505E5C18FE8190F536B56D82" box="[268,447,652,678]" country="United Arab Emirates" latitude="25.24" longLatPrecision="748" longitude="56.17" name="Wadi Wurayah" pageId="7" pageNumber="65">Wadi Wurayah</location>
;
<geoCoordinate id="F3E75030505E5C18FE4690F235336D82" box="[459,569,651,678]" degrees="25.24" direction="north" orientation="latitude" pageId="7" pageNumber="65" precision="555" value="25.24">25.24° N</geoCoordinate>
,
<geoCoordinate id="F3E75030505E5C18FDCB90F235BB6D82" box="[582,689,651,678]" degrees="56.17" direction="east" orientation="longitude" pageId="7" pageNumber="65" precision="555" value="56.17">56.17° E</geoCoordinate>
;
<date id="E26D1037505E5C18FD3390F534776D82" box="[702,893,652,678]" pageId="7" pageNumber="65" value="2007-03" valueMax="2007-04-31" valueMin="2007-03-00">
<collectingDate id="F229E9DF505E5C18FD3390F534776D82" box="[702,893,652,678]" pageId="7" pageNumber="65" value="2007-03" valueMax="2007-04-31" valueMin="2007-03-00">Mar.–Apr. 2007</collectingDate>
</date>
;
<collectorName id="3B265321505E5C18FC0790F533076D82" box="[906,1037,652,678]" pageId="7" pageNumber="65">C. Tourenq</collectorName>
leg. (7482 UAE);
<collectingMethod id="4F924EE0505E5C18FB6190F532786D82" box="[1260,1394,652,678]" pageId="7" pageNumber="65">water traps</collectingMethod>
;
<specimenCode id="C6759E8C505E5C18FF3090D636A66DED" box="[189,428,687,713]" pageId="7" pageNumber="65">SCH_ISEA_000161</specimenCode>
</materialsCitation>
•
<materialsCitation id="26BB3CAA505E5C18FE4890D635636DC8" ID-GBIF-Occurrence="3762960331" collectingDate="2008-02-25" collectingDateMax="2007-04-31" collectingDateMin="2007-03" collectingMethod="hand-collected" collectionCode="ISEA" collectorName="A. Polaszek" country="United Arab Emirates" latitude="25.24" location="Wadi Wurayah" longLatPrecision="748" longitude="56.17" pageId="7" pageNumber="65" specimenCode="SCH_ISEA_000162" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="80D5FD7E505E5C18FE4890D636F86DED" box="[453,498,687,713]" pageId="7" pageNumber="65" type="female">1 ♀</specimenCount>
; same locality as for preceding;
<date id="E26D1037505E5C18FCE290D633066DED" box="[879,1036,687,714]" pageId="7" pageNumber="65" value="2008-02-25">
<collectingDate id="F229E9DF505E5C18FCE290D633066DED" box="[879,1036,687,714]" pageId="7" pageNumber="65" value="2008-02-25">25 Feb. 2008</collectingDate>
</date>
;
<collectorName id="3B265321505E5C18FB9A90D633A96DED" box="[1047,1187,687,713]" pageId="7" pageNumber="65">A. Polaszek</collectorName>
leg. (8807 UAE);
<collectingMethod id="4F924EE0505E5C18FF3090AB36646DC8" box="[189,366,722,748]" pageId="7" pageNumber="65">hand-collected</collectingMethod>
;
<specimenCode id="C6759E8C505E5C18FEF490AB35636DC8" box="[377,617,722,748]" pageId="7" pageNumber="65">SCH_ISEA_000162</specimenCode>
</materialsCitation>
•
<materialsCitation id="26BB3CAA505E5C18FD0F90AB34946C2B" ID-GBIF-Occurrence="3762960308" collectingDate="2006-09-24" collectingDateMax="2006-10-22" collectingDateMin="2006-09-24" collectingMethod="water traps" collectionCode="ISEA" collectorName="A. Van Harten" country="United Arab Emirates" latitude="25.09" location="Wadi Siji" longLatPrecision="749" longitude="56.02" pageId="7" pageNumber="65" specimenCode="SCH_ISEA_000163" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="80D5FD7E505E5C18FD0F90AB35BB6DC9" box="[642,689,722,749]" pageId="7" pageNumber="65" type="male">1 ♂</specimenCount>
;
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;
<geoCoordinate id="F3E75030505E5C18FCB490AB34AC6DC9" box="[825,934,722,749]" degrees="25.09" direction="north" orientation="latitude" pageId="7" pageNumber="65" precision="555" value="25.09">25.09° N</geoCoordinate>
,
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; 24
<date id="E26D1037505E5C18FBC090AB32456DC8" box="[1101,1359,722,748]" pageId="7" pageNumber="65" value="2006-09-24" valueMax="2006-10-22" valueMin="2006-09-24">
<collectingDate id="F229E9DF505E5C18FBC090AB32456DC8" box="[1101,1359,722,748]" pageId="7" pageNumber="65" value="2006-09-24" valueMax="2006-10-22" valueMin="2006-09-24">24 Sep.–22 Oct. 2006</collectingDate>
</date>
;
<collectorName id="3B265321505E5C18FAD790AB36486C34" pageId="7" pageNumber="65">A. Van Harten</collectorName>
leg. (8372 UAE);
<collectingMethod id="4F924EE0505E5C18FD90908F35A86C34" box="[541,674,758,784]" pageId="7" pageNumber="65">water traps</collectingMethod>
;
<specimenCode id="C6759E8C505E5C18FD23908C34946C2B" box="[686,926,757,783]" pageId="7" pageNumber="65">SCH_ISEA_000163</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DEC9657C505E5C17FF30914F34B06B32" lastPageId="8" lastPageNumber="66" pageId="7" pageNumber="65" type="description">
<paragraph id="966C36F7505E5C18FF30914F36456C74" blockId="7.[189,335,822,848]" box="[189,335,822,848]" pageId="7" pageNumber="65">
<heading id="CD24819B505E5C18FF30914F36456C74" bold="true" box="[189,335,822,848]" fontSize="11" level="3" pageId="7" pageNumber="65" reason="3">
<emphasis id="A4A7EAE5505E5C18FF30914F36456C74" bold="true" box="[189,335,822,848]" pageId="7" pageNumber="65">Description</emphasis>
</heading>
</paragraph>
<paragraph id="966C36F7505E5C18FF67911C35226C5B" blockId="7.[189,1399,869,931]" box="[234,552,869,895]" pageId="7" pageNumber="65">
<emphasis id="A4A7EAE5505E5C18FF67911C36AD6C5B" bold="true" box="[234,423,869,895]" pageId="7" pageNumber="65">
<typeStatus id="49688855505E5C18FF67911C36546C5B" box="[234,350,869,895]" pageId="7" pageNumber="65" type="holotype">Holotype</typeStatus>
, male
</emphasis>
(
<figureCitation id="0EE82A72505E5C18FE3A911C35156C5B" box="[439,543,869,895]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="7" pageNumber="65">Figs 1–2</figureCitation>
)
</paragraph>
<paragraph id="966C36F7505E5C18FF3091F0327D6C86" blockId="7.[189,1399,869,931]" box="[189,1399,904,931]" pageId="7" pageNumber="65">BODY. Parallel, slightly expanded posteriorly, with apical abdominal segments not covered by the elytra.</paragraph>
<paragraph id="966C36F7505E5C18FF3091B3354A6B69" blockId="7.[189,1401,970,1101]" pageId="7" pageNumber="65">COLOURATION. Yellow, except for meso- and metathoracic ventrites and two basal abdominal ventrites underside, traces of transverse fascia on head and distal side of pronotum, triangular spot on base of elytra narrowly continuing up into apical part, and narrow margins on apices of elytra brown, vesicles and thoracic mesepimera yellow.</paragraph>
<paragraph id="966C36F7505E5C18FF30960C33956A38" blockId="7.[189,1400,1141,1308]" pageId="7" pageNumber="65">
HEAD. Narrower as pronotum (
<figureCitation id="0EE82A72505E5C18FDBD960C35716BAB" box="[560,635,1141,1167]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="7" pageNumber="65">Fig. 4</figureCitation>
), slightly elongate, eyes small, weakly protruding over contour of head; frons and inter-ocular area flat and slightly impressed, glabrous, lacking depression or protuberances; clypeus slightly elongate, transverse, straight; labrum short and transverse; palpi short, ultimate palpomere thin, cylindrical, slightly narrowed and flattened apically. Surface of head dull, sparsely and finely punctured, evenly covered with white adpressed fine pubescence.
</paragraph>
<paragraph id="966C36F7505E5C18FF30973A327D692A" blockId="7.[189,1399,1347,1550]" pageId="7" pageNumber="65">
ANTENNAE. Flabellate, not long, reaching base of elytra; 1
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antennomere enlarged, sub-conic, 2
<superScript id="61A69BBF505E5C18FF41971F37D76A50" attach="left" box="[204,221,1382,1396]" fontSize="6" pageId="7" pageNumber="65">nd</superScript>
antennomere oval, ½ × as long as 3
<superScript id="61A69BBF505E5C18FD19971F35A86A50" attach="left" box="[660,674,1382,1396]" fontSize="6" pageId="7" pageNumber="65">rd</superScript>
antennomere, 3
<superScript id="61A69BBF505E5C18FCEF971F347A6A50" attach="left" box="[866,880,1382,1396]" fontSize="6" pageId="7" pageNumber="65">rd</superScript>
antennomere triangular, not wider than 4
<superScript id="61A69BBF505E5C18FAEF971F32656A50" attach="none" box="[1378,1391,1382,1396]" fontSize="6" pageId="7" pageNumber="65">th</superScript>
, 4
<superScript id="61A69BBF505E5C18FF4197F037D36AB3" attach="left" box="[204,217,1417,1431]" fontSize="6" pageId="7" pageNumber="65">th</superScript>
antennomere wide triangular, with somewhat dilated outer edge, 5
<superScript id="61A69BBF505E5C18FC6597F034FF6AB3" attach="left" box="[1000,1013,1417,1431]" fontSize="6" pageId="7" pageNumber="65">th</superScript>
with more strongly dilated outer edge, 6
<superScript id="61A69BBF505E5C18FE9A97D5362E6A9E" attach="none" box="[279,292,1452,1466]" fontSize="6" pageId="7" pageNumber="65">th</superScript>
–10
<superScript id="61A69BBF505E5C18FEDC97D536546A9E" attach="left" box="[337,350,1452,1466]" fontSize="6" pageId="7" pageNumber="65">th</superScript>
segments flabellate, cylindrical with long dilated outer edges, terminal antennomere distinctly longest, curved at basal third, evenly rounded at apex (
<figureCitation id="0EE82A72505E5C18FC2997A934E06ACE" box="[932,1002,1488,1514]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="7" pageNumber="65">Fig. 3</figureCitation>
), surface evenly covered with fine short, pale pubescence and erect long fine pale hairs on outer edges of dilated processes of antennomeres.
</paragraph>
<paragraph id="966C36F7505E5C18FF30944C348969B1" blockId="7.[189,1398,1589,1686]" pageId="7" pageNumber="65">
PRONOTUM (
<figureCitation id="0EE82A72505E5C18FEC7944C36E6696B" box="[330,492,1589,1615]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="7" pageNumber="65">Figs 2, 4, 10</figureCitation>
). Transverse, with rounded lateral sides, not sinuate or depressed at base, with straight anterior and posterior sides; margination thin and distinct, surface with smoothed sparse punctures and covered with short white adpressed fine hairs.
</paragraph>
<paragraph id="966C36F7505E5C18FF3094C4349569DE" blockId="7.[189,1399,1724,1786]" pageId="7" pageNumber="65">SCUTELLUM. Small, rectangular, narrow, slightly transverse, with slightly subparallel apex; surface sparsely punctured and covered with fine adpressed light hairs.</paragraph>
<paragraph id="966C36F7505E5C18FF30955B354E68A5" blockId="7.[189,1400,1825,1921]" pageId="7" pageNumber="65">
ELYTRA. Subparallel, weakly expanded posteriorly and not covering last segments of abdomen in both sexes (
<figureCitation id="0EE82A72505E5C18FE82953D36C2687A" box="[271,456,1860,1886]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="7" pageNumber="65">Figs 1–2, 9–10</figureCitation>
); humeri indistinct, small, protruding; surface densely punctured, covered with light dense adpressed short hairs.
</paragraph>
<paragraph id="966C36F7505E5C18FF3095D0354068E7" blockId="7.[189,586,1961,1987]" box="[189,586,1961,1987]" pageId="7" pageNumber="65">HIND WINGS. Normally developed.</paragraph>
<paragraph id="966C36F750515C17FF309370321D6EF7" blockId="8.[189,1398,265,467]" pageId="8" pageNumber="66">
LEGS. Thin and long; posterior femora reaching abdomen apex; anterior tarsi (
<figureCitation id="0EE82A7250515C17FBEF937033A36E00" box="[1122,1193,265,292]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="8" pageNumber="66">Fig. 5</figureCitation>
) five-segmented, compressed, with comb above 2
<superScript id="61A69BBF50515C17FDB1935235476E1D" attach="left" box="[572,589,299,313]" fontSize="6" pageId="8" pageNumber="66">nd</superScript>
segment, 2
<superScript id="61A69BBF50515C17FD5A935235E26E1D" attach="none" box="[727,744,299,313]" fontSize="6" pageId="8" pageNumber="66">nd</superScript>
–3
<superScript id="61A69BBF50515C17FC88935234196E1D" attach="left" box="[773,787,299,313]" fontSize="6" pageId="8" pageNumber="66">rd</superScript>
tarsomeres equal in length, 1
<superScript id="61A69BBF50515C17FBF89355338A6E1E" attach="left" box="[1141,1152,300,314]" fontSize="6" pageId="8" pageNumber="66">st</superScript>
1.2 × as long as 2
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, terminal tarsomere slightly narrower and depressed, longest, as long as 1
<superScript id="61A69BBF50515C17FBBD933633316E79" attach="none" box="[1072,1083,335,349]" fontSize="6" pageId="8" pageNumber="66">st</superScript>
–4
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tarsomeres in anterior legs, 1
<superScript id="61A69BBF50515C17FE83930B36136EA4" attach="none" box="[270,281,370,384]" fontSize="6" pageId="8" pageNumber="66">st</superScript>
–3
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in intermediate legs and 1
<superScript id="61A69BBF50515C17FD1D930B35916EA4" attach="none" box="[656,667,370,384]" fontSize="6" pageId="8" pageNumber="66">st</superScript>
–2
<superScript id="61A69BBF50515C17FD34930B35C06EA4" attach="left" box="[697,714,370,384]" fontSize="6" pageId="8" pageNumber="66">nd</superScript>
in posterior legs; claws thin, weakly curved and sharp, with dentate small lamellae at base; femora and tibia simple, not swollen or strongly curved, lacking indentation; surface indistinctly and covered with short and sparse adpressed light pubescence.
</paragraph>
<paragraph id="966C36F750515C17FF30938234276C2F" blockId="8.[187,1399,506,779]" pageId="8" pageNumber="66">
VENTRAL BODY. Surface weakly punctured, very sparsely covered with fine, depressed light pubescence; metathorax weakly swollen, narrow, transverse, lacking appendages or tuft of hairs, but with specific straight and edged distal side, which is slightly lamellate. Pygidium (
<figureCitation id="0EE82A7250515C17FC6B903833266D78" box="[998,1068,577,604]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="8" pageNumber="66">Fig. 6</figureCitation>
) undivided, evenly rounded and narrowed distally, with oval emargination in middle; 8
<superScript id="61A69BBF50515C17FCED901A34676D55" attach="left" box="[864,877,611,625]" fontSize="6" pageId="8" pageNumber="66">th</superScript>
ultimate abdominal ventrite (
<figureCitation id="0EE82A7250515C17FB4E901D320D6D5B" box="[1219,1287,612,639]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="8" pageNumber="66">Fig. 7</figureCitation>
) strongly narrowed, bilaciniate, but both lobes narrowly adjoined, evenly rounded distally, with round-triangular emargination in middle and sharp lateral processes; aedeagus (
<figureCitation id="0EE82A7250515C17FC3490D333096DE1" box="[953,1027,682,709]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="8" pageNumber="66">Fig. 8</figureCitation>
) elongate and strongly curved dorsally, typical of
<taxonomicName id="51D34D7450515C17FE2B90B735096DCC" authorityName="Abeille de Perrin" authorityYear="1890" box="[422,515,718,744]" class="Insecta" family="Melyridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="66" phylum="Arthropoda" rank="tribe" tribe="Attalini">Attalini</taxonomicName>
, with long sharpened bristle in inner sac in middle of distal part, tegmen wide, transverse, with thin short appendages (
<figureCitation id="0EE82A7250515C17FD5B908834166C2F" box="[726,796,753,779]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="8" pageNumber="66">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="966C36F750515C17FF30914A34A36C69" blockId="8.[189,937,818,845]" box="[189,937,818,845]" pageId="8" pageNumber="66">
MEASUREMENTS. Length
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, width (at elytral base)
<quantity id="512B9B1250515C17FCC5914B34AF6C69" box="[840,933,818,845]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="66" unit="mm" value="0.7">0.7 mm</quantity>
.
</paragraph>
<paragraph id="966C36F750515C17FF67910D364F6CAA" blockId="8.[189,1398,884,981]" box="[234,325,884,910]" pageId="8" pageNumber="66">
<emphasis id="A4A7EAE550515C17FF67910D364F6CAA" bold="true" box="[234,325,884,910]" pageId="8" pageNumber="66">Female</emphasis>
</paragraph>
<paragraph id="966C36F750515C17FF3091EE35266CF0" blockId="8.[189,1398,884,981]" pageId="8" pageNumber="66">
Differs from male by somewhat shorter and dentate antennae; anterior tarsi lacking comb on 2
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tarsomere; body slightly larger.
</paragraph>
<paragraph id="966C36F750515C17FF30918234B06B32" blockId="8.[189,954,1019,1046]" box="[189,954,1019,1046]" pageId="8" pageNumber="66">
Length
<quantity id="512B9B1250515C17FE9A918536A26B32" box="[279,424,1020,1046]" metricMagnitude="-3" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" pageId="8" pageNumber="66" unit="mm" value="2.3499999999999996" valueMax="2.4" valueMin="2.3">2.3–2.4 mm</quantity>
, width (at elytral base)
<quantity id="512B9B1250515C17FD369182342E6B32" box="[699,804,1019,1046]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="66" unit="mm" value="0.75">0.75 mm</quantity>
(
<figureCitation id="0EE82A7250515C17FCBE918234A76B32" box="[819,941,1019,1046]" captionStart="Figs 1–10" captionStartId="6.[189,242,1889,1915]" captionTargetBox="[222,1352,267,1823]" captionTargetId="figure-75@6.[219,1368,265,1849]" captionTargetPageId="6" captionText="Figs 1–10. Tonyattalus vanharteni gen. et sp. nov., holotype (SCH_ISEA_000133), ♂ (1–8), ♀ (9– 10). 1, 9. External appearance, dorsal view. 2, 10. External appearance, lateral view. 3. Left antenna. 4. Head and pronotum, dorsal view. 5. Left anterior tarsus. 6. Pygidium. 7. Ultimate abdominal ventrite. 8. Aedeagus, and tegmen, lateral view. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6518293" httpUri="https://zenodo.org/record/6518293/files/figure.png" pageId="8" pageNumber="66">Figs 9–10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DEC9657C50515C17FF30964436CF6BA1" box="[189,453,1085,1157]" pageId="8" pageNumber="66" type="distribution">
<paragraph id="966C36F750515C17FF30964436526B73" blockId="8.[189,344,1085,1111]" box="[189,344,1085,1111]" pageId="8" pageNumber="66">
<heading id="CD24819B50515C17FF30964436526B73" bold="true" box="[189,344,1085,1111]" fontSize="11" level="3" pageId="8" pageNumber="66" reason="3">
<emphasis id="A4A7EAE550515C17FF30964436526B73" bold="true" box="[189,344,1085,1111]" pageId="8" pageNumber="66">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="966C36F750515C17FF30961236CF6BA1" blockId="8.[189,453,1131,1157]" box="[189,453,1131,1157]" pageId="8" pageNumber="66">
<collectingCountry id="EEC4766750515C17FF30961236CA6BA1" box="[189,448,1131,1157]" name="United Arab Emirates" pageId="8" pageNumber="66">United Arab Emirates</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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