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<document id="EB15F12EEAD56E015B92EE07865860CF" ID-DOI="10.11646/zootaxa.3716.4.2" ID-GBIF-Dataset="a9b85d9e-a8fb-4ceb-9f20-65d2cd3c84df" ID-ISSN="1175-5326" ID-Zenodo-Dep="220316" ID-ZooBank="F1AD3425-314F-4B5F-AE7A-2D683135DBE6" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460682586901" checkinUser="plazi" docAuthor="Pyrcz, Tomasz W., Prieto, Carlos, Viloria, Angel L. &amp; Andrade, Gonzalo" docDate="2013" docId="03B29E2BFFE61870FF0FCCB7FE37F9CA" docLanguage="en" docName="zt03716p538.pdf" docOrigin="Zootaxa 3716 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pedaliodes adrianae Pyrcz &amp; Prieto, n. sp." docType="treatment" docVersion="8" lastPageNumber="531" masterDocId="FF8BE653FFE41873FF98C97BFFC1FF9A" masterDocTitle="New species of high elevation cloud forest butterflies of the genus Pedaliodes Butler from the northern Colombian Andes (Lepidoptera, Nymphalidae, Satyrinae)" masterLastPageNumber="538" masterPageNumber="528" pageNumber="530" updateTime="1698350629498" updateUser="plazi">
<mods:mods id="FA589EA41561F3ACEF9ACBEF57C7D41A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FA14A659CFB30770AFA965FD9D7E031D">
<mods:title id="B8D782FD5E0C4DDCD101D9E3EAC1735D">New species of high elevation cloud forest butterflies of the genus Pedaliodes Butler from the northern Colombian Andes (Lepidoptera, Nymphalidae, Satyrinae)</mods:title>
</mods:titleInfo>
<mods:name id="83FC961D8B842C1AE1B88503276507AC" type="personal">
<mods:role id="99FD73A4A08629023F9F8BF27981FD96">
<mods:roleTerm id="6381C857668793CD0F2DA9F0985ADC84">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1C7C6BD48C46C9A766531EF37781B2FF">Pyrcz, Tomasz W.</mods:namePart>
</mods:name>
<mods:name id="B5DDFCC30F3A8BB3325C05295E645AD7" type="personal">
<mods:role id="7FA1BC4887FEB7635B4BD10A0DC52105">
<mods:roleTerm id="9A463BFE58E45B21CA65417289D11A96">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="45FBD7E6C6F391AC44BD8F467CD6A3FA">Prieto, Carlos</mods:namePart>
</mods:name>
<mods:name id="E504E428F6FDC6B27E0A3EA2EBD2047D" type="personal">
<mods:role id="EE2FEC93A1A4077663148DA52348F3C7">
<mods:roleTerm id="1B4D6228125BCAA45E761F39582FCBAA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D70F992EAECD2A8D45962A8475D484A1">Viloria, Angel L.</mods:namePart>
</mods:name>
<mods:name id="4CC70897D53FE578AEFD29DBF70B37B7" type="personal">
<mods:role id="83D551AB59B997BCAE7CBE617609C4CA">
<mods:roleTerm id="9FCBB54E4B318BD656CA3FADD3BD6859">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B59D4250F2E72CB0EB7A3918748F804D">Andrade, Gonzalo</mods:namePart>
</mods:name>
<mods:typeOfResource id="A550369831B29EB96186E89F15C27B7C">text</mods:typeOfResource>
<mods:relatedItem id="0F0EC31F780344A4A7C2AD6F2C00F3C2" type="host">
<mods:titleInfo id="C990A10A96BDFCD433F3368A90B83EF2">
<mods:title id="373AAC59903CB4D07C732C1481545714">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="766506F86225884C59770042987CD692">
<mods:date id="B0AF1D2B79DF019CC4858EF676E1E953">2013</mods:date>
<mods:detail id="BC4E429A26BE39A50F9D3580464C7FEF" type="volume">
<mods:number id="D9BB8581E65385947921160C1EEAC679">3716</mods:number>
</mods:detail>
<mods:detail id="9735C3C7D86018FEF48C4CBB9C0BDC80" type="issue">
<mods:number id="CAED03F6016183251B73F6749AD01048">4</mods:number>
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<mods:start id="08BA68F020D45A1BF0EA3CFF73C4331B">528</mods:start>
<mods:end id="5B20F65C64F7FB6019D62CCD60041B80">538</mods:end>
</mods:extent>
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<mods:classification id="3D6A6FD8D96C59E216F1A51FDA1F4FC5">journal article</mods:classification>
<mods:identifier id="27A0766BFA4B6B6229009F1703731F67" type="DOI">10.11646/zootaxa.3716.4.2</mods:identifier>
<mods:identifier id="7023FF7D7624E990331C4535983F136D" type="GBIF-Dataset">a9b85d9e-a8fb-4ceb-9f20-65d2cd3c84df</mods:identifier>
<mods:identifier id="EB821E56EF2E31A026256723623F6B72" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6E3E82970ECCFEACC4C53149C6058676" type="Zenodo-Dep">220316</mods:identifier>
<mods:identifier id="CA9E3DEB7D8F8FDBB0B35C97C4D2879E" type="ZooBank">F1AD3425-314F-4B5F-AE7A-2D683135DBE6</mods:identifier>
</mods:mods>
<treatment id="03B29E2BFFE61870FF0FCCB7FE37F9CA" ID-DOI="http://doi.org/10.5281/zenodo.6151277" ID-GBIF-Taxon="119580931" ID-Zenodo-Dep="6151277" LSID="urn:lsid:plazi:treatment:03B29E2BFFE61870FF0FCCB7FE37F9CA" httpUri="http://treatment.plazi.org/id/03B29E2BFFE61870FF0FCCB7FE37F9CA" lastPageId="3" lastPageNumber="531" pageId="2" pageNumber="530">
<subSubSection id="C3017CB6FFE61871FF0FCCB7FEA7F99D" pageId="2" pageNumber="530" type="nomenclature">
<paragraph id="8BA42F3DFFE61871FF0FCCB7FD5EFA7C" blockId="2.[151,671,1484,1543]" box="[151,671,1484,1510]" pageId="2" pageNumber="530">
<heading id="D0EC9851FFE61871FF0FCCB7FD5EFA7C" bold="true" box="[151,671,1484,1510]" fontSize="11" level="1" pageId="2" pageNumber="530" reason="1">
<emphasis id="B96FF32FFFE61871FF0FCCB7FD5EFA7C" bold="true" box="[151,671,1484,1510]" pageId="2" pageNumber="530">
<taxonomicName id="4C1B54BEFFE61871FF0FCCB7FD8EFA7C" authority="Pyrcz &amp; Prieto" authorityName="Pyrcz &amp; Prieto" box="[151,591,1484,1510]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="adrianae" status="sp. nov.">
<emphasis id="B96FF32FFFE61871FF0FCCB7FE4AFA7C" bold="true" box="[151,395,1484,1510]" italics="true" pageId="2" pageNumber="530">Pedaliodes adrianae</emphasis>
Pyrcz &amp; Prieto
</taxonomicName>
,
<taxonomicNameLabel id="A25C4E54FFE61871FDC4CCB7FD5EFA7C" box="[604,671,1484,1510]" pageId="2" pageNumber="530" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA42F3DFFE61871FF0FCC95FEA7F99D" blockId="2.[151,671,1484,1543]" box="[151,358,1518,1543]" pageId="2" pageNumber="530">
(
<figureCitation id="132033B8FFE61871FF07CC95FECBF99D" box="[159,266,1518,1543]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1932,1954]" captionTargetBox="[262,1342,215,1896]" captionTargetId="figure@5.[221,1381,195,1934]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Pedaliodes adults. 1, P. haydoni male (holotype); 2, P. porcia male; 3, P. adrianae male (holotype); 4, P. adrianae female (paratype); 5, P. rodriguezi male (holotype); 6, P. ro dr i gu e z i female (paratype); 7, P. philinae male (holotype); 8, P. philinae female (paratype)." httpUri="https://zenodo.org/record/220317/files/figure.png" pageId="2" pageNumber="530">Figs. 3, 4</figureCitation>
,
<figureCitation id="132033B8FFE61871FE8ECC94FEF5F99D" box="[278,308,1519,1543]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1891,1913]" captionTargetBox="[247,1333,233,1790]" captionTargetId="figure@6.[243,1347,193,1849]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Male genitalia. Pedaliodes haydoni (paratype): 9 A, genitalia in lateral view; 9 B, aedeagus in lateral view; 9 C, aedeagus in dorsal view; Pedaliodes adrianae (paratype): 10 A, genitalia in lateral view; 10 B, aedeagus in lateral view; 10 C, aedeagus in dorsal view; Pedaliodes philinae (paratype): 11 A, genitalia in lateral view; 11 B, aedeagus in lateral view; 11 C, aedeagus in dorsal view; Pedaliodes rodriguezi (paratype): 12 A, genitalia in lateral view; 12 B, aedeagus in lateral view; 12 C, aedeagus in dorsal view." httpUri="https://zenodo.org/record/220318/files/figure.png" pageId="2" pageNumber="530">10</figureCitation>
,
<figureCitation id="132033B8FFE61871FED8CC94FE9CF99D" box="[320,349,1519,1543]" captionStart="FIGURES 13 – 15. 13" captionStartId="7.[151,264,1953,1975]" captionTargetBox="[378,1185,207,1911]" captionTargetId="figure@7.[214,1373,193,1932]" captionTargetPageId="7" captionText="FIGURES 13 – 15. 13, Pedaliodes adrianae (paratype) female genitalia; 14, Pedaliodes philinae (paratype) female genitalia; 15, Pedaliodes rodriguezi male (Cerro San Felix, Antioquia, Colombia, 2900 m)" httpUri="https://zenodo.org/record/220319/files/figure.png" pageId="2" pageNumber="530">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3017CB6FFE61871FF0FCF42FDBDF95A" pageId="2" pageNumber="530" type="materials_examined">
<paragraph id="8BA42F3DFFE61871FF0FCF42FDBDF95A" blockId="2.[151,1437,1590,2017]" pageId="2" pageNumber="530">
<emphasis id="B96FF32FFFE61871FF0FCF42FE4AF9C8" bold="true" box="[151,395,1593,1618]" pageId="2" pageNumber="530">Material examined.</emphasis>
<typeStatus id="54A0919FFFE61871FE04CF41FDF1F9C8" box="[412,560,1594,1618]" pageId="2" pageNumber="530" type="holotype">HOLOTYPE</typeStatus>
(male):
<collectingCountry id="F30C6FADFFE61871FD3CCF42FCD4F9C8" box="[676,789,1593,1618]" name="Colombia" pageId="2" pageNumber="530">Colombia</collectingCountry>
, Antioquia, Urrao, Páramo de Frontino,
<geoCoordinate id="EE2F49FAFFE61871FA81CF42FA56F9C8" box="[1305,1431,1590,1618]" direction="north" orientation="latitude" pageId="2" pageNumber="530" precision="15" value="6.4063888">6o24’23’’N</geoCoordinate>
,
<geoCoordinate id="EE2F49FAFFE61871FF0FCF1BFEE7F9E3" box="[151,294,1629,1657]" direction="west" orientation="longitude" pageId="2" pageNumber="530" precision="15" value="-76.104996">76o06’18’’W</geoCoordinate>
,
<quantity id="4CE382D8FFE61871FEABCF1BFE4AF9E2" box="[307,395,1632,1656]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="530" unit="m" value="2800.0">2800 m</quantity>
, 0 8.2012, C. Prieto leg., MZUJ (to be deposited in MHN-UN);
<typeStatus id="54A0919FFFE61871FBC5CF1BFB39F9E2" box="[1117,1272,1632,1656]" pageId="2" pageNumber="530" type="paratype">PARATYPES</typeStatus>
(
<specimenCount id="9D1DE4B4FFE61871FA93CF1BFA97F9E3" box="[1291,1366,1632,1657]" pageId="2" pageNumber="530" type="male">1 male</specimenCount>
and
<specimenCount id="9D1DE4B4FFE61871FA16CF1BFF28F907" pageId="2" pageNumber="530" type="female">1 female</specimenCount>
):
<specimenCount id="9D1DE4B4FFE61871FF63CFFEFE8AF907" box="[251,331,1668,1693]" pageId="2" pageNumber="530" type="male">1 male</specimenCount>
:
<collectingCountry id="F30C6FADFFE61871FECDCFFFFE07F907" box="[341,454,1668,1693]" name="Colombia" pageId="2" pageNumber="530">Colombia</collectingCountry>
, Antioquia, Urrao, Páramo de Frontino,
<quantity id="4CE382D8FFE61871FC14CFFEFC22F906" box="[908,995,1669,1693]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="530" unit="m" value="3100.0">3100 m</quantity>
,
<date id="FFA509FDFFE61871FC77CFFEFB9CF907" box="[1007,1117,1669,1693]" pageId="2" pageNumber="530" value="2012-01-07">07.I.2012</date>
, i804, C. Prieto leg., CPC;
<specimenCount id="9D1DE4B4FFE61871FA16CFFEFF29F95B" pageId="2" pageNumber="530" type="female">1 female</specimenCount>
: same data as the
<typeStatus id="54A0919FFFE61871FE37CFD3FDD0F95B" box="[431,529,1704,1729]" pageId="2" pageNumber="530" type="holotype">holotype</typeStatus>
(MZUJ).
</paragraph>
</subSubSection>
<subSubSection id="C3017CB6FFE61870FF5FCFB7FB49FF4E" lastPageId="3" lastPageNumber="531" pageId="2" pageNumber="530" type="diagnosis">
<paragraph id="8BA42F3DFFE61870FF5FCFB7FB49FF4E" blockId="2.[151,1437,1590,2017]" lastBlockId="3.[151,1437,151,1616]" lastPageId="3" lastPageNumber="531" pageId="2" pageNumber="530">
<emphasis id="B96FF32FFFE61871FF5FCFB7FE80F97F" bold="true" box="[199,321,1740,1765]" pageId="2" pageNumber="530">Diagnosis.</emphasis>
HWV pattern is similar to several species of the
<taxonomicName id="4C1B54BEFFE61871FC18CFB7FB8BF97E" box="[896,1098,1740,1764]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="phaea">
<emphasis id="B96FF32FFFE61871FC18CFB7FB8BF97E" box="[896,1098,1740,1764]" italics="true" pageId="2" pageNumber="530">Pedaliodes phaea</emphasis>
</taxonomicName>
group, all characterized by a wide postdiscal—submarginal lighter, yellowish area, which is nearly straight from vein M1 to anal margin. However, in contrast to these species, the wings of
<taxonomicName id="4C1B54BEFFE61871FD4DCE6EFC97F8B6" box="[725,854,1812,1836]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="adrianae">
<emphasis id="B96FF32FFFE61871FD4DCE6EFC97F8B6" box="[725,854,1812,1836]" italics="true" pageId="2" pageNumber="530">P. adrianae</emphasis>
</taxonomicName>
are dorsally dark brown, whereas in the
<taxonomicName id="4C1B54BEFFE61871FA87CE6FFF1AF8CA" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="phaea">
<emphasis id="B96FF32FFFE61871FA87CE6FFF1AF8CA" italics="true" pageId="2" pageNumber="530">Pedaliodes phaea</emphasis>
</taxonomicName>
and related species there are invariably conspicuous orange patches on both the FW and HW or restricted to HW. The only exceptions are
<taxonomicName id="4C1B54BEFFE61871FE76CE26FDA7F8EE" box="[494,614,1884,1908]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="haydoni">
<emphasis id="B96FF32FFFE61871FE76CE26FDA7F8EE" box="[494,614,1884,1908]" italics="true" pageId="2" pageNumber="530">P. haydoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1B54BEFFE61871FD39CE26FCDBF8EE" box="[673,794,1884,1908]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="baccara">
<emphasis id="B96FF32FFFE61871FD39CE26FCDBF8EE" box="[673,794,1884,1908]" italics="true" pageId="2" pageNumber="530">P. baccara</emphasis>
</taxonomicName>
, and the differences between
<taxonomicName id="4C1B54BEFFE61871FBEACE26FB35F8EE" box="[1138,1268,1884,1908]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="adrianae">
<emphasis id="B96FF32FFFE61871FBEACE26FB35F8EE" box="[1138,1268,1884,1908]" italics="true" pageId="2" pageNumber="530">P. adrianae</emphasis>
</taxonomicName>
and these two species were specified under
<taxonomicName id="4C1B54BEFFE61871FE7ACEFAFD99F802" box="[482,600,1920,1944]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="haydoni">
<emphasis id="B96FF32FFFE61871FE7ACEFAFD99F802" box="[482,600,1920,1944]" italics="true" pageId="2" pageNumber="530">P. haydoni</emphasis>
</taxonomicName>
. The female genitalia of
<taxonomicName id="4C1B54BEFFE61871FCEDCEFBFB9EF802" box="[885,1119,1920,1944]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="adrianae">
<emphasis id="B96FF32FFFE61871FCEDCEFBFB9EF802" box="[885,1119,1920,1944]" italics="true" pageId="2" pageNumber="530">Pedaliodes adrianae</emphasis>
</taxonomicName>
are similar in most respects to those of
<taxonomicName id="4C1B54BEFFE61871FE8ACEDFFBE9F827" authority="Lamas, Viloria &amp; Pyrcz, 2010 (Pyrcz et al. 2010)" authorityName="Lamas, Viloria &amp; Pyrcz, 2010 (Pyrcz et al." authorityYear="2010" box="[274,1064,1956,1981]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="acjanaco">
<emphasis id="B96FF32FFFE61871FE8ACEDFFE3DF826" box="[274,508,1956,1980]" italics="true" pageId="2" pageNumber="530">Pedaliodes acjanaco</emphasis>
Lamas, Viloria &amp; Pyrcz, 2010 (Pyrcz
<emphasis id="B96FF32FFFE61871FC3DCEDEFC17F826" box="[933,982,1956,1980]" italics="true" pageId="2" pageNumber="530">et al</emphasis>
. 2010)
</taxonomicName>
. The ductus bursae is similarly to other species of
<emphasis id="B96FF32FFFE61871FE18CEB3FD72F87A" box="[384,691,1992,2016]" italics="true" pageId="2" pageNumber="530">
<taxonomicName id="4C1B54BEFFE61871FE18CEB3FD92F87A" box="[384,595,1992,2016]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="species" species="sensu">
Pedaliodes
<taxonomicNameLabel id="A25C4E54FFE61871FD8DCEB2FD92F87A" box="[533,595,1993,2016]" pageId="2" pageNumber="530" sensu="stricto">sensu</taxonomicNameLabel>
</taxonomicName>
stricto
</emphasis>
, differing from
<taxonomicName id="4C1B54BEFFE61871FC01CEB3FA56F87B" authority="Miller, Miller &amp; Viloria" authorityName="Miller, Miller &amp; Viloria" box="[921,1431,1992,2017]" class="Insecta" family="Nymphalidae" genus="Neopedaliodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="530" phylum="Arthropoda" rank="genus">
<emphasis id="B96FF32FFFE61871FC01CEB3FBFEF87A" box="[921,1087,1992,2016]" italics="true" pageId="2" pageNumber="530">Neopedaliodes</emphasis>
Miller, Miller &amp; Viloria
</taxonomicName>
,
<taxonomicName id="4C1B54BEFFE71870FF0FC9E3FE02FF35" authority="Forster" authorityName="Forster" box="[151,451,152,176]" class="Insecta" family="Nymphalidae" genus="Corderopedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="531" phylum="Arthropoda" rank="genus">
<emphasis id="B96FF32FFFE71870FF0FC9E3FEACFF2A" box="[151,365,152,176]" italics="true" pageId="3" pageNumber="531">Corderopedaliodes</emphasis>
Forster
</taxonomicName>
and
<taxonomicName id="4C1B54BEFFE71870FE62C9E3FCDCFF35" authority="Forster" authorityName="Forster" box="[506,797,152,176]" class="Insecta" family="Nymphalidae" genus="Physcopedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="531" phylum="Arthropoda" rank="genus">
<emphasis id="B96FF32FFFE71870FE62C9E3FD05FF2A" box="[506,708,152,176]" italics="true" pageId="3" pageNumber="531">Physcopedaliodes</emphasis>
Forster
</taxonomicName>
, which are characterised by a strongly sclerotized ductus bursae, two-thirds the length of the corpus bursae, and slightly twisted (Pyrcz
<emphasis id="B96FF32FFFE71870FB99C9C6FBF3FF4E" box="[1025,1074,188,212]" italics="true" pageId="3" pageNumber="531">et al</emphasis>
. 2013
<emphasis id="B96FF32FFFE71870FBE0C9C7FB40FF49" box="[1144,1153,188,211]" italics="true" pageId="3" pageNumber="531">)</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C3017CB6FFE71870FF5FC9A4FC02FA5A" pageId="3" pageNumber="531" type="description">
<paragraph id="8BA42F3DFFE71870FF5FC9A4FF6FFB5F" blockId="3.[151,1437,151,1616]" pageId="3" pageNumber="531">
<emphasis id="B96FF32FFFE71870FF5FC9A4FE98FF62" bold="true" box="[199,345,223,248]" pageId="3" pageNumber="531">Description.</emphasis>
<emphasis id="B96FF32FFFE71870FEF8C99BFE5BFF62" box="[352,410,224,248]" italics="true" pageId="3" pageNumber="531">Male</emphasis>
(
<figureCitation id="132033B8FFE71870FE33C9A4FE31FF62" box="[427,496,223,248]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1932,1954]" captionTargetBox="[262,1342,215,1896]" captionTargetId="figure@5.[221,1381,195,1934]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Pedaliodes adults. 1, P. haydoni male (holotype); 2, P. porcia male; 3, P. adrianae male (holotype); 4, P. adrianae female (paratype); 5, P. rodriguezi male (holotype); 6, P. ro dr i gu e z i female (paratype); 7, P. philinae male (holotype); 8, P. philinae female (paratype)." httpUri="https://zenodo.org/record/220317/files/figure.png" pageId="3" pageNumber="531">Fig. 3</figureCitation>
). Head: Antennae slender, reaching nearly half length of the costa, naked, dorsally dark brown, ventrally orange brown, club formed gradually, slightly thicker than shaft, composed of 11 flagellomeres. Eyes chocolate brown, lustrous, covered with dense, black hair. Labial palpi two times as long as the head, covered ventrally with black and golden yellow hairy scales, laterally with brown and sandy yellow scales, and dorsally brown scales. Frons with a tuft of rather short, chocolate brown and golden yellow hair. Thorax: Dorsally black, covered with brown and golden brown hair, denser on patagium, tegulae and prothorax; ventrally black; legs brown, femora covered with long blackish hair, tibiae and tarsi with dense sandy yellow scales, tarsi with numerous blackish, short spines. FW (length:
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, n=2) with a subacute apex, and gently concave, giving the impression of being lightly undulated due to the intermittently dark brown and longer fringes at vein ends and shorter, chestnut in the interveins. FWD almost uniform glossy dark brown, a shade lighter in distal one-third; scent patch large, roughly
<quantity id="4CE382D8FFE71870FE1ACB33FE20FDC5" box="[386,481,584,608]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="531" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
wide, compact, extending in median area from base of M1 to anal margin, entering discal-cell. HW rounded, with undulating outer margin; fringes intermittently dark brown and milky white from apex to M3. HWD uniform dark brown from M3 to tornus; uniform dark brown, less glossy than the FW, slightly hairy in basal part and along anal margin. FWV dull medium brown, a shade lighter between postdiscal and submarginal line, a diffuse milky white costal streak along postdiscal line from costa to vein M3; outer margin auburn; a series of three or four minute white dots in subapical area from R4–R5 to M2–M3. HWV auburn, liberally speckled with a sandy yellow, ripple-like pattern; a wide sandy yellow band extending distally from postdiscal line to submarginal area, with distal part densely suffused with brown scales, with a sharp, and mostly straight inner edge from vein M2 to anal margin, and an irregular outer edge with two deep basal incision along veins M2 and M3, roughly parallel to outer margin; a row of four minute, oval submarginal whitish dots in M1– M2, M2–M3, M3–Cu1 and Cu1–Cu2, the former two slightly more prominent. Abdomen: Black, hairy, dorsally and laterally covered with black, ventrally covered with chestnut and sandy yellow scales. Male genitalia (
<figureCitation id="132033B8FFE71870FADACAAFFA52FC76" box="[1346,1427,980,1005]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1891,1913]" captionTargetBox="[247,1333,233,1790]" captionTargetId="figure@6.[243,1347,193,1849]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Male genitalia. Pedaliodes haydoni (paratype): 9 A, genitalia in lateral view; 9 B, aedeagus in lateral view; 9 C, aedeagus in dorsal view; Pedaliodes adrianae (paratype): 10 A, genitalia in lateral view; 10 B, aedeagus in lateral view; 10 C, aedeagus in dorsal view; Pedaliodes philinae (paratype): 11 A, genitalia in lateral view; 11 B, aedeagus in lateral view; 11 C, aedeagus in dorsal view; Pedaliodes rodriguezi (paratype): 12 A, genitalia in lateral view; 12 B, aedeagus in lateral view; 12 C, aedeagus in dorsal view." httpUri="https://zenodo.org/record/220318/files/figure.png" pageId="3" pageNumber="531">Fig. 10</figureCitation>
) with dorsum of tegumen flat, uncus slender, gently arched, as long as tegumen dorsum, subunci long and slender, approximately 2/3 the length of uncus, pedunculus small, valvae roughly the length of tegumen+uncus, with a short, sharp dorsal process aligned with the acute distal extremity; saccus wide and shallow, aedeagus as long as saccus+valve, massive, flattened dorso ventrally, somewhat humped in the middle and with a sharp apical extremity, proximal opening 1/3 the length of the entire aedeagus, approximately the width of the remaining part of it.
</paragraph>
<paragraph id="8BA42F3DFFE71870FF5FCDABFC02FA5A" blockId="3.[151,1437,151,1616]" pageId="3" pageNumber="531">
<emphasis id="B96FF32FFFE71870FF5FCDABFEDAFB72" box="[199,283,1232,1256]" italics="true" pageId="3" pageNumber="531">Female</emphasis>
(
<figureCitation id="132033B8FFE71870FEB1CDB4FEAFFB72" box="[297,366,1231,1256]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1932,1954]" captionTargetBox="[262,1342,215,1896]" captionTargetId="figure@5.[221,1381,195,1934]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Pedaliodes adults. 1, P. haydoni male (holotype); 2, P. porcia male; 3, P. adrianae male (holotype); 4, P. adrianae female (paratype); 5, P. rodriguezi male (holotype); 6, P. ro dr i gu e z i female (paratype); 7, P. philinae male (holotype); 8, P. philinae female (paratype)." httpUri="https://zenodo.org/record/220317/files/figure.png" pageId="3" pageNumber="531">Fig. 4</figureCitation>
). Head and Thorax: Similar to male except considerably paler on both upper and underside. FW length
<quantity id="4CE382D8FFE71870FF7FCD8FFEFDFA96" box="[231,316,1268,1292]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="531" unit="mm" value="27.0">27 mm</quantity>
. Abdomen: Female genitalia (
<figureCitation id="132033B8FFE71870FD00CD8FFD2DFA96" box="[664,748,1268,1293]" captionStart="FIGURES 13 – 15. 13" captionStartId="7.[151,264,1953,1975]" captionTargetBox="[378,1185,207,1911]" captionTargetId="figure@7.[214,1373,193,1932]" captionTargetPageId="7" captionText="FIGURES 13 – 15. 13, Pedaliodes adrianae (paratype) female genitalia; 14, Pedaliodes philinae (paratype) female genitalia; 15, Pedaliodes rodriguezi male (Cerro San Felix, Antioquia, Colombia, 2900 m)" httpUri="https://zenodo.org/record/220319/files/figure.png" pageId="3" pageNumber="531">Fig. 13</figureCitation>
) with papillae anales medium sized. Antrum enclosed by a wide basin strengthen from the inside by a strongly sclerotized slat-like lamella antevaginalis (a common feature of all examined species of
<emphasis id="B96FF32FFFE71870FE34CC47FD65FACE" box="[428,676,1340,1364]" italics="true" pageId="3" pageNumber="531">
<taxonomicName id="4C1B54BEFFE71870FE34CC47FDAEFACE" box="[428,623,1340,1364]" class="Insecta" family="Nymphalidae" genus="Pedaliodes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="531" phylum="Arthropoda" rank="species" species="sensu">
Pedaliodes
<taxonomicNameLabel id="A25C4E54FFE71870FDA9CC46FDAEFACE" box="[561,623,1341,1364]" pageId="3" pageNumber="531" sensu="lato">sensu</taxonomicNameLabel>
</taxonomicName>
lato
</emphasis>
). Bursa copulatrix oval, gradually narrowing toward posterior and gradually transforming into ductus bursae. Signa approximately half the length of corpus burse, parallel and wide, with a spiny surface. Ductus bursae wide, slightly sclerotized, two-thirds length of corpus bursae, slightly twisted. Ductus seminalis connecting to ductus bursae near its proximal opening.
</paragraph>
</subSubSection>
<subSubSection id="C3017CB6FFE71870FF5FCCB7FC3AF992" pageId="3" pageNumber="531" type="etymology">
<paragraph id="8BA42F3DFFE71870FF5FCCB7FC3AF992" blockId="3.[151,1437,151,1616]" pageId="3" pageNumber="531">
<emphasis id="B96FF32FFFE71870FF5FCCB7FE8FFA7F" bold="true" box="[199,334,1484,1509]" pageId="3" pageNumber="531">Etymology.</emphasis>
This species is named after Adriana Maria Collazos Porras, an enthusiastic nature lover who accompanied the second author in many of his field expeditions in
<collectingCountry id="F30C6FADFFE71870FC1DCC94FC36F992" box="[901,1015,1519,1544]" name="Colombia" pageId="3" pageNumber="531">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3017CB6FFE71870FF5FCF6FFE37F9CA" pageId="3" pageNumber="531" type="discussion">
<paragraph id="8BA42F3DFFE71870FF5FCF6FFE37F9CA" blockId="3.[151,1437,151,1616]" pageId="3" pageNumber="531">
<emphasis id="B96FF32FFFE71870FF5FCF6FFE8EF9B7" bold="true" box="[199,335,1556,1581]" pageId="3" pageNumber="531">Comments.</emphasis>
This species is known exclusively from the
<taxonomicName id="4C1B54BEFFE71870FCCDCF6FFBD5F9B7" authority="Frontino" authorityName="Frontino" box="[853,1044,1556,1581]" class="Insecta" family="Nymphalidae" genus="Paramo" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="531" phylum="Arthropoda" rank="genus">Páramo Frontino</taxonomicName>
massif in the northern part of the Colombian Western Cordillera.
</paragraph>
</subSubSection>
</treatment>
</document>
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