treatments-xml/data/03/B5/EB/03B5EB57AE1FFFDD99C47F5405871A9E.xml
2024-06-21 12:22:17 +02:00

388 lines
50 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="391E56CAC701D0223E14003A97C41A89" ID-CLB-Dataset="9529" ID-DOI="10.11646/zootaxa.4881.3.7" ID-GBIF-Dataset="b7622904-4534-4fc0-aebd-d4766d680eb9" ID-ISSN="1175-5326" ID-Zenodo-Dep="4283960" ID-ZooBank="003285F9-AD4C-4B33-BCFD-E15821BF26C6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1605981939210" checkinUser="plazi" docAuthor="Tian, Mingyi &amp; He, Li" docDate="2020" docId="03B5EB57AE1FFFDD99C47F5405871A9E" docLanguage="en" docName="zootaxa.4881.3.7.pdf" docOrigin="Zootaxa 4881 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Huaius) hanwang Tian &amp; He 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="555" masterDocId="FF8C932FAE18FFD799537B4F01291F5F" masterDocTitle="New species and new record of cavernicolous ground beetles from Sichuan Province, China (Coleoptera: Carabidae: Pterostichini and Platynini)" masterLastPageNumber="558" masterPageNumber="545" pageNumber="552" updateTime="1698846498561" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="9F1BE08A7DC80BC3CC246E62EB3C3D79" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BF167F3DA53DC3A71373CFD649CAFF5A">
<mods:title id="95BB940ED30580088EF501BE4C17C15E">New species and new record of cavernicolous ground beetles from Sichuan Province, China (Coleoptera: Carabidae: Pterostichini and Platynini)</mods:title>
</mods:titleInfo>
<mods:name id="89DFBE86D7B8C8FAA9D3141F5D496997" type="personal">
<mods:role id="FF137E2356A53A3DC552CABAF0555E7C">
<mods:roleTerm id="469E736A48F6298B0A0576D3409ACCF4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C9652D8D2EF284F8600FCDDC8C69B4FB">Tian, Mingyi</mods:namePart>
<mods:affiliation id="EBCC71DC405ECD89D614FC6FD242900F">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642,</mods:affiliation>
</mods:name>
<mods:name id="83DA90883677C23949D2FF31B6BFD8E3" type="personal">
<mods:role id="B92A88827AFBC81CD87C1D853E6477F9">
<mods:roleTerm id="CD6E7267A13C5C5778513C2773CF1D26">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9B9EE8CC0F40E5132007A025E804DA20">He, Li</mods:namePart>
<mods:affiliation id="9308EA60A65506B00C08A4C037CCE6A1">Sichuan Cave Exploration Team, No. 66, 5 Shuangcheng Road, Chenghua District, Chengdu, 610051, China https: // orcid. org / 0000 - 0002 - 4597 - 3442</mods:affiliation>
</mods:name>
<mods:typeOfResource id="92D400AD0BA63AD20498D92A666152B2">text</mods:typeOfResource>
<mods:relatedItem id="51D24C76EAC3C717035DF536EAE5902B" type="host">
<mods:titleInfo id="ED8376F82D13598839EC1667C359288D">
<mods:title id="D2BF6D8AAAAD8112463FC0F7BD14A8A6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="71713B654A1CF0A8555132BA67EAC7B6">
<mods:date id="14404B8FDDEBFBCC0F33B009D68A140D">2020</mods:date>
<mods:detail id="C424DB35501D03A46B51950463C3FB44" type="pubDate">
<mods:number id="57396AE4D4E19BDBF6D61FDB31ACD891">2020-11-20</mods:number>
</mods:detail>
<mods:detail id="A9518B7CEF1E06F1AA481B02FCECCFE2" type="volume">
<mods:number id="841F79CA0AB749BE26D8C8682B9CA0B3">4881</mods:number>
</mods:detail>
<mods:detail id="EBF19A1151947E4616FFB112B47A47CB" type="issue">
<mods:number id="5D520B0B29DD3E24796446CE98653B62">3</mods:number>
</mods:detail>
<mods:extent id="0716361D7CECF529D02997CC2D644949" unit="page">
<mods:start id="33A178BC7FE5ACF98A04AE4567ACA320">545</mods:start>
<mods:end id="E36BC15437F56ABC7DE29895A26070BB">558</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="121040215DB38833FCEC806F2702552B">journal article</mods:classification>
<mods:identifier id="3E721ACC941789C8D21AA9FED7DAAF23" type="CLB-Dataset">9529</mods:identifier>
<mods:identifier id="95C467BF2103D36381CAAE20EEBD01A5" type="DOI">10.11646/zootaxa.4881.3.7</mods:identifier>
<mods:identifier id="906C9480765F461A3EF6C51DB6FEDE2C" type="GBIF-Dataset">b7622904-4534-4fc0-aebd-d4766d680eb9</mods:identifier>
<mods:identifier id="E72DC088E438410F56FC81788673DDC4" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="85FB846BBD61B8124824ECBCFED8B2C2" type="Zenodo-Dep">4283960</mods:identifier>
<mods:identifier id="89611332118E12CA5242926ACDFC210B" type="ZooBank">003285F9-AD4C-4B33-BCFD-E15821BF26C6</mods:identifier>
</mods:mods>
<treatment id="03B5EB57AE1FFFDD99C47F5405871A9E" ID-DOI="http://doi.org/10.5281/zenodo.4328763" ID-GBIF-Taxon="170510555" ID-Zenodo-Dep="4328763" LSID="urn:lsid:plazi:treatment:03B5EB57AE1FFFDD99C47F5405871A9E" httpUri="http://treatment.plazi.org/id/03B5EB57AE1FFFDD99C47F5405871A9E" lastPageId="10" lastPageNumber="555" pageId="7" pageNumber="552">
<subSubSection id="C30609CAAE1FFFD099C47F5400D71B04" pageId="7" pageNumber="552" type="nomenclature">
<paragraph id="8BA35A41AE1FFFD099C47F5403571B69" blockId="7.[151,638,1051,1150]" box="[151,638,1051,1078]" pageId="7" pageNumber="552">
<heading id="D0EBED2DAE1FFFD099C47F5403571B69" bold="true" box="[151,638,1051,1078]" fontSize="11" level="1" pageId="7" pageNumber="552" reason="1">
<emphasis id="B9688653AE1FFFD099C47F5403571B69" bold="true" box="[151,638,1051,1078]" pageId="7" pageNumber="552">
<taxonomicName id="4C1C21C2AE1FFFD099C47F5403301B69" ID-CoL="8M6X4" authority="Tian &amp; He, 2020" authorityName="Tian &amp; He" authorityYear="2020" box="[151,537,1051,1078]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="hanwang" status="sp. nov." subGenus="Huaius">
<emphasis id="B9688653AE1FFFD099C47F5403301B69" bold="true" box="[151,537,1051,1078]" italics="true" pageId="7" pageNumber="552">Pterostichus (Huaius) hanwang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B3B28AE1FFFD09B727F5303571B69" box="[545,638,1052,1078]" pageId="7" pageNumber="552" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA35A41AE1FFFD099C47F0F00D71B04" blockId="7.[151,638,1051,1150]" box="[151,510,1088,1115]" pageId="7" pageNumber="552">(Chinese name: ỡŦúãṃm)</paragraph>
</subSubSection>
<subSubSection id="C30609CAAE1FFFD099C47F2B00241B21" box="[151,269,1124,1150]" pageId="7" pageNumber="552" type="description">
<paragraph id="8BA35A41AE1FFFD099C47F2B00241B21" blockId="7.[151,638,1051,1150]" box="[151,269,1124,1150]" pageId="7" pageNumber="552">
<figureCitation id="132746C4AE1FFFD099C47F2B00241B21" box="[151,269,1124,1150]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStartId-0="8.[151,250,1996,2020]" captionStartId-1="9.[151,250,1877,1901]" captionStartId-2="10.[151,250,837,861]" captionStartId-3="11.[151,250,1944,1968]" captionStartId-4="12.[151,250,1685,1709]" captionTargetBox-1="[153,1429,198,1844]" captionTargetBox-2="[160,1432,240,807]" captionTargetBox-3="[269,1309,189,1898]" captionTargetBox-4="[151,1435,181,1661]" captionTargetId-0="figure-20@8.[338,1248,181,1973]" captionTargetId-1="figure-19@9.[151,1435,181,1853]" captionTargetId-2="figure-53@10.[151,1435,237,813]" captionTargetId-3="figure-19@11.[260,1328,181,1920]" captionTargetId-4="figure-20@12.[151,1435,181,1661]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="11" captionTargetPageId-4="12" captionText-0="FIGURE 6. Habitus of Pterostichus (Huaius) hanwang sp. nov., male, holotype." captionText-1="FIGURE 7. Head and pronotum (A, B) and labium (C, D) of Pterostichus (Huaius) species. A and C Pt. (H.) hanwang sp. nov.; B and D Pt. (H.) yuae Tian et Chen, 2019" captionText-2="FIGURE 8. Male genitalia of Pterostichus (Huaius) hanwang sp. nov. A median lobe and parameres, lateral view B apical lobe, dorsal view." captionText-3="FIGURE 9. Female genitalia of Pterostichus (Huaius) hanwang sp. nov. co common oviduct; bc bursa copulatrix; sp spermatheca; sd ductus of spermatheca; sg spermathecal gland; sgd ductus of the spermathecal gland; gscx gonosubcoxite; gcx gonocoxite." captionText-4="FIGURE 10. Cave Hanwang Dong, the type locality of Pterostichus (Huaius) hanwang, sp. nov. A, entrance; B, C, spectacular scenery inside the cave; D, the moist and gravelly areas in cave, showing the habitat where the beetles were collected; E, a running individual of P. hanwang, sp. nov. in cave; F, the original state of the beetles when the stone was turned over; G, two individuals of P. hanwang, sp. nov. were collected in a test tube." figureDoi-0="http://doi.org/10.5281/zenodo.4289215" figureDoi-1="http://doi.org/10.5281/zenodo.4283970" figureDoi-2="http://doi.org/10.5281/zenodo.4283972" figureDoi-3="http://doi.org/10.5281/zenodo.4283974" figureDoi-4="http://doi.org/10.5281/zenodo.4283976" httpUri-0="https://zenodo.org/record/4289215/files/figure.png" httpUri-1="https://zenodo.org/record/4283970/files/figure.png" httpUri-2="https://zenodo.org/record/4283972/files/figure.png" httpUri-3="https://zenodo.org/record/4283974/files/figure.png" httpUri-4="https://zenodo.org/record/4283976/files/figure.png" pageId="7" pageNumber="552">Figs. 610</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30609CAAE1FFFD099C47FE303611AC1" pageId="7" pageNumber="552" type="materials_examined">
<paragraph id="8BA35A41AE1FFFD099C47FE303611AC1" blockId="7.[151,1437,1196,2014]" pageId="7" pageNumber="552">
<emphasis id="B9688653AE1FFFD099C47FE3002F1B99" bold="true" box="[151,262,1196,1222]" pageId="7" pageNumber="552">Material.</emphasis>
<materialsCitation id="3B74501CAE1FFFD098437FE304B01BB5" ID-GBIF-Occurrence="2985238307" collectingDate="2018-09-22" collectionCode="SCAU" collectorName="Li He" county="cave Hanwang Dong" elevation="1210" latitude="32.577816" location="Zengjia" longLatPrecision="1" longitude="106.10703" municipality="Zhongbai" pageId="7" pageNumber="552" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">
<typeStatus id="54A7E4E3AE1FFFD098437FE300541B99" box="[272,381,1196,1222]" pageId="7" pageNumber="552" type="holotype">Holotype</typeStatus>
: male,
<collectingCounty id="62C222CDAE1FFFD098817FE203EC1B98" box="[466,709,1197,1223]" pageId="7" pageNumber="552">cave Hanwang Dong</collectingCounty>
,
<collectingMunicipality id="6BC7C03BAE1FFFD09B877FE3026B1B99" box="[724,834,1196,1222]" pageId="7" pageNumber="552">Zhongbai</collectingMunicipality>
,
<location id="8EC30C9AAE1FFFD09A037FE302801B99" LSID="urn:lsid:plazi:treatment:03B5EB57AE1FFFDD99C47F5405871A9E:8EC30C9AAE1FFFD09A037FE302801B99" box="[848,937,1196,1222]" county="cave Hanwang Dong" latitude="32.577816" longLatPrecision="1" longitude="106.10703" municipality="Zhongbai" name="Zengjia" pageId="7" pageNumber="552" stateProvince="Sichuan">Zengjia</location>
,
<location id="8EC30C9AAE1FFFD09AE47FE305361B99" LSID="urn:lsid:plazi:treatment:03B5EB57AE1FFFDD99C47F5405871A9E:8EC30C9AAE1FFFD09AE47FE305361B99" box="[951,1055,1196,1222]" county="cave Hanwang Dong" latitude="32.577816" longLatPrecision="1" longitude="106.10703" municipality="Zhongbai" name="Chaotian" pageId="7" pageNumber="552" stateProvince="Sichuan">Chaotian</location>
,
<location id="8EC30C9AAE1FFFD09D7E7FE305981B99" LSID="urn:lsid:plazi:treatment:03B5EB57AE1FFFDD99C47F5405871A9E:8EC30C9AAE1FFFD09D7E7FE305981B99" box="[1069,1201,1196,1222]" county="cave Hanwang Dong" latitude="32.577816" longLatPrecision="1" longitude="106.10703" municipality="Zhongbai" name="Guangyuan" pageId="7" pageNumber="552" stateProvince="Sichuan">Guangyuan</location>
,
<collectingRegion id="49D894A3AE1FFFD09DEC7FE304301B99" box="[1215,1305,1196,1222]" country="China" name="Sichuan" pageId="7" pageNumber="552">Sichuan</collectingRegion>
(HnjờỮ KḿfikṽǎẍüϕffifflỡŦḆ),
<geoCoordinate id="EE283C86AE1FFFD09B6A7F9E03CD1BB3" box="[569,740,1233,1260]" degrees="32" direction="north" minutes="34" orientation="latitude" pageId="7" pageNumber="552" precision="1" seconds="40.14" value="32.577816">32°3440.14″N</geoCoordinate>
,
<geoCoordinate id="EE283C86AE1FFFD09BA17F9E02B31BB3" box="[754,922,1233,1260]" degrees="106" direction="east" minutes="6" orientation="longitude" pageId="7" pageNumber="552" precision="1" seconds="25.32" value="106.10703">106°625.32″E</geoCoordinate>
,
<quantity id="4CE4F7A4AE1FFFD09AFB7F9F05281BB4" box="[936,1025,1232,1259]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="7" pageNumber="552" unit="m" value="1210.0">
<elevation id="0031BD72AE1FFFD09AFB7F9F05281BB4" box="[936,1025,1232,1259]" metricMagnitude="3" metricUnit="m" metricValue="1.21" pageId="7" pageNumber="552" unit="m" value="1210.0">1210 m</elevation>
</quantity>
,
<date id="FFA27C81AE1FFFD09D437F9F05BF1BB5" box="[1040,1174,1232,1258]" pageId="7" pageNumber="552" value="2018-09-22">
<collectingDate id="EFE68569AE1FFFD09D437F9F05BF1BB5" box="[1040,1174,1232,1258]" pageId="7" pageNumber="552" value="2018-09-22">IX-22-2018</collectingDate>
</date>
, leg.
<collectorName id="26E93F97AE1FFFD09D847F9F04341BB4" box="[1239,1309,1232,1259]" pageId="7" pageNumber="552">Li He</collectorName>
, in
<collectionCode id="ED0DC284AE1FFFD09C187F9F04B01BB5" box="[1355,1433,1232,1258]" pageId="7" pageNumber="552">SCAU</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B74501CAE1FFFD099C47FBA020D1A51" ID-GBIF-Occurrence="2985238313" box="[151,804,1268,1295]" collectingDate="2018-09-22" collectionCode="SCAU" collectorName="Li He" county="cave Hanwang Dong" elevation="1210" latitude="32.577816" location="Zengjia" longLatPrecision="1" longitude="106.10703" municipality="Zhongbai" pageId="7" pageNumber="552" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="paratype">
<typeStatus id="54A7E4E3AE1FFFD099C47FBA002D1A50" box="[151,260,1269,1295]" pageId="7" pageNumber="552" type="paratype">Paratypes</typeStatus>
(12):
<specimenCount id="9D1A91C8AE1FFFD098147FBB00BC1A51" box="[327,405,1268,1294]" pageId="7" pageNumber="552" type="male">1 male</specimenCount>
,
<specimenCount id="9D1A91C8AE1FFFD098F37FBB032A1A51" box="[416,515,1268,1294]" pageId="7" pageNumber="552" type="female">1 female</specimenCount>
,
<emphasis id="B9688653AE1FFFD09B5D7FBB036D1A51" box="[526,580,1268,1294]" italics="true" pageId="7" pageNumber="552">idem</emphasis>
<typeStatus id="54A7E4E3AE1FFFD09B197FBB03861A51" box="[586,687,1268,1294]" pageId="7" pageNumber="552" type="holotype">holotype</typeStatus>
, in
<collectionCode id="ED0DC284AE1FFFD09B857FBB020D1A51" box="[726,804,1268,1294]" pageId="7" pageNumber="552">SCAU</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B74501CAE1FFFD09A7C7FBB05A21A51" ID-GBIF-Occurrence="2985238314" box="[815,1163,1268,1294]" collectingDate="2018-09-22" collectionCode="SCAU" collectorName="Li He" county="cave Hanwang Dong" elevation="1210" latitude="32.577816" location="Zengjia" longLatPrecision="1" longitude="106.10703" municipality="Zhongbai" pageId="7" pageNumber="552" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<specimenCount id="9D1A91C8AE1FFFD09A7C7FBB02551A51" box="[815,892,1268,1294]" pageId="7" pageNumber="552" type="male">1 male</specimenCount>
,
<emphasis id="B9688653AE1FFFD09AD47FBB02941A51" box="[903,957,1268,1294]" italics="true" pageId="7" pageNumber="552">idem</emphasis>
<typeStatus id="54A7E4E3AE1FFFD09A977FBB05001A51" box="[964,1065,1268,1294]" pageId="7" pageNumber="552" type="holotype">holotype</typeStatus>
, in CHL
</materialsCitation>
;
<materialsCitation id="3B74501CAE1FFFD09DC57FBB00D71A6D" ID-GBIF-Occurrence="2985238304" collectingDate="2018-04-05" collectionCode="SCAU" collectorName="Li He" county="cave Hanwang Dong" elevation="1210" latitude="32.577816" location="Zengjia" longLatPrecision="1" longitude="106.10703" municipality="Zhongbai" pageId="7" pageNumber="552" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9D1A91C8AE1FFFD09DC57FBB05CA1A51" box="[1174,1251,1268,1294]" pageId="7" pageNumber="552" type="male">1 male</specimenCount>
, same collection data except
<date id="FFA27C81AE1FFFD098737E5700BE1A6C" box="[288,407,1304,1331]" pageId="7" pageNumber="552" value="2018-04-05">
<collectingDate id="EFE68569AE1FFFD098737E5700BE1A6C" box="[288,407,1304,1331]" pageId="7" pageNumber="552" value="2018-04-05">IV-5-2018</collectingDate>
</date>
, in CHL
</materialsCitation>
;
<materialsCitation id="3B74501CAE1FFFD09B597E5700A21A09" ID-GBIF-Occurrence="2985238310" collectingDate="2020-06-06" collectionCode="SCAU" collectorName="Li He, Yuan Li &amp; Yimei Wen" county="cave Hanwang Dong" elevation="1210" latitude="32.577816" location="Zengjia" longLatPrecision="1" longitude="106.10703" municipality="Zhongbai" pageId="7" pageNumber="552" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9D1A91C8AE1FFFD09B597E5703731A6D" box="[522,602,1304,1330]" pageId="7" pageNumber="552" type="male">1 male</specimenCount>
,
<specimenCount id="9D1A91C8AE1FFFD09B347E5703E51A6D" box="[615,716,1304,1330]" pageId="7" pageNumber="552" type="female">1 female</specimenCount>
, same collection data except
<date id="FFA27C81AE1FFFD09D497E5705BD1A6D" box="[1050,1172,1304,1330]" pageId="7" pageNumber="552" value="2020-06-06">
<collectingDate id="EFE68569AE1FFFD09D497E5705BD1A6D" box="[1050,1172,1304,1330]" pageId="7" pageNumber="552" value="2020-06-06">VI-6-2020</collectingDate>
</date>
, leg.
<collectorName id="26E93F97AE1FFFD09D817E57043E1A6C" box="[1234,1303,1304,1331]" pageId="7" pageNumber="552">Li He</collectorName>
,
<collectorName id="26E93F97AE1FFFD09C707E5604571A6D" box="[1315,1406,1304,1331]" pageId="7" pageNumber="552">Yuan Li</collectorName>
&amp;
<collectorName id="26E93F97AE1FFFD099C47E73003E1A08" box="[151,279,1340,1367]" pageId="7" pageNumber="552">Yimei Wen</collectorName>
, in
<collectionCode id="ED0DC284AE1FFFD0986E7E7300A21A09" box="[317,395,1340,1366]" pageId="7" pageNumber="552">SCAU</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B74501CAE1FFFD098C57E7301D91A25" ID-GBIF-Occurrence="2985238305" collectingDate="2020-06-06" collectionCode="SCAU" collectorName="Li He, Yuan Li &amp; Yimei Wen" county="cave Hanwang Dong" elevation="1210" latitude="32.577816" location="Zengjia" longLatPrecision="1" longitude="106.10703" municipality="Zhongbai" pageId="7" pageNumber="552" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9D1A91C8AE1FFFD098C57E7300C71A09" box="[406,494,1340,1366]" pageId="7" pageNumber="552" type="male">3 males</specimenCount>
,
<specimenCount id="9D1A91C8AE1FFFD098AA7E7303751A09" box="[505,604,1340,1366]" pageId="7" pageNumber="552" type="female">1 female</specimenCount>
, same collection data except
<date id="FFA27C81AE1FFFD09ACC7E7305311A09" box="[927,1048,1340,1366]" pageId="7" pageNumber="552" value="2020-06-06">
<collectingDate id="EFE68569AE1FFFD09ACC7E7305311A09" box="[927,1048,1340,1366]" pageId="7" pageNumber="552" value="2020-06-06">VI-6-2020</collectingDate>
</date>
, leg.
<collectorName id="26E93F97AE1FFFD09D017E7305BD1A08" box="[1106,1172,1340,1367]" pageId="7" pageNumber="552">Li He</collectorName>
,
<collectorName id="26E93F97AE1FFFD09DCD7E7205DE1A09" box="[1182,1271,1340,1367]" pageId="7" pageNumber="552">Yuan Li</collectorName>
&amp;
<collectorName id="26E93F97AE1FFFD09C4B7E7304B01A08" box="[1304,1433,1340,1367]" pageId="7" pageNumber="552">Yimei Wen</collectorName>
, in CHL
</materialsCitation>
;
<materialsCitation id="3B74501CAE1FFFD099A97E2F01FA1AC1" ID-GBIF-Occurrence="2985238302" collectingDate="2020-06-06" collectionCode="SCAU" collectorName="Li He, Yuan Li &amp; Yimei Wen" county="cave Hanwang Dong" elevation="1210" latitude="32.577816" location="Zengjia" longLatPrecision="1" longitude="106.10703" municipality="Zhongbai" pageId="7" pageNumber="552" specimenCount="2" specimenCount-male="2" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9D1A91C8AE1FFFD099A97E2F007A1A25" box="[250,339,1376,1402]" pageId="7" pageNumber="552" type="male">2 males</specimenCount>
, same collection data except
<date id="FFA27C81AE1FFFD09BCB7E2F02381A25" box="[664,785,1376,1402]" pageId="7" pageNumber="552" value="2020-06-06">
<collectingDate id="EFE68569AE1FFFD09BCB7E2F02381A25" box="[664,785,1376,1402]" pageId="7" pageNumber="552" value="2020-06-06">VI-6-2020</collectingDate>
</date>
, leg.
<collectorName id="26E93F97AE1FFFD09A1F7E2F02A71A24" box="[844,910,1376,1403]" pageId="7" pageNumber="552">Li He</collectorName>
,
<collectorName id="26E93F97AE1FFFD09ACA7E2E02DB1A25" box="[921,1010,1376,1403]" pageId="7" pageNumber="552">Yuan Li</collectorName>
&amp;
<collectorName id="26E93F97AE1FFFD09D477E2F05BC1A24" box="[1044,1173,1376,1403]" pageId="7" pageNumber="552">Yimei Wen</collectorName>
, one in CCZ and one in CYL
</materialsCitation>
.
<materialsCitation id="3B74501CAE1FFFD0998E7ECB036D1AC1" ID-GBIF-Occurrence="2985238303" box="[221,580,1412,1439]" collectionCode="SCAU" pageId="7" pageNumber="552" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="paratype">
<specimenCount id="9D1A91C8AE1FFFD0998E7ECB00021AC1" box="[221,299,1412,1438]" pageId="7" pageNumber="552" type="male">1 male</specimenCount>
, spare in ethanol,
<collectionCode id="ED0DC284AE1FFFD098A57ECB036D1AC1" box="[502,580,1412,1438]" pageId="7" pageNumber="552">SCAU</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30609CAAE1FFFD099947EE702B31955" pageId="7" pageNumber="552" type="diagnosis">
<paragraph id="8BA35A41AE1FFFD099947EE702B31955" blockId="7.[151,1437,1196,2014]" pageId="7" pageNumber="552">
<emphasis id="B9688653AE1FFFD099947EE7006B1A9D" bold="true" box="[199,322,1448,1474]" pageId="7" pageNumber="552">Diagnosis.</emphasis>
Similar to
<taxonomicName id="4C1C21C2AE1FFFD098907EE6025E1A9D" authority="Tian and Chen, 2019" authorityName="Tian and Chen" authorityYear="2019" box="[451,887,1448,1474]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="552" phylum="Arthropoda" rank="species" species="yuae" subGenus="Huaius">
<emphasis id="B9688653AE1FFFD098907EE603A81A9D" box="[451,641,1448,1474]" italics="true" pageId="7" pageNumber="552">P. (Huaius) yuae</emphasis>
<bibRefCitation id="EF8D27B0AE1FFFD09BDA7EE7025E1A9D" author="Tian, M. Y. &amp; Huang, S. B. &amp; Chen, M. Z. &amp; Ding, K. J." box="[649,887,1448,1474]" pageId="7" pageNumber="552" pagination="1 - 16" refId="ref6051" refString="Tian, M. Y., Huang, S. B., Chen, M. Z. &amp; Ding, K. J. (2019) Remarkable cave-adapted ground beetles of the tribe Pterostichini from China: a new subgenus and three new species (Coleoptera: Carabidae). Annales de la Societe entomologique de France, New Series, 55 (1), 1 - 16. https: // doi. org / 10.1080 / 00379271.2018.1546554" type="journal article" year="2019">Tian and Chen, 2019</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation id="EF8D27B0AE1FFFD09AF57EE705761A9D" author="Tian, M. Y. &amp; Huang, S. B. &amp; Chen, M. Z. &amp; Ding, K. J." box="[934,1119,1448,1475]" pageId="7" pageNumber="552" pagination="1 - 16" refId="ref6051" refString="Tian, M. Y., Huang, S. B., Chen, M. Z. &amp; Ding, K. J. (2019) Remarkable cave-adapted ground beetles of the tribe Pterostichini from China: a new subgenus and three new species (Coleoptera: Carabidae). Annales de la Societe entomologique de France, New Series, 55 (1), 1 - 16. https: // doi. org / 10.1080 / 00379271.2018.1546554" type="journal article" year="2019">
Tian
<emphasis id="B9688653AE1FFFD09AB27EE6053D1A9D" box="[993,1044,1448,1474]" italics="true" pageId="7" pageNumber="552">et al</emphasis>
. 2019
</bibRefCitation>
), but slightly stouter. Upper surface smooth and glabrous, except pronotum punctate in and near the basal foveae; body strongly shiny, light brown, with antennae, mouthpart palpi and tarsi light reddish brown.
</paragraph>
</subSubSection>
<subSubSection id="C30609CAAE1FFFDD99947D5B052E1B4E" lastPageId="10" lastPageNumber="555" pageId="7" pageNumber="552" type="description">
<paragraph id="8BA35A41AE1FFFD099947D5B051B1971" blockId="7.[151,1437,1196,2014]" box="[199,1074,1556,1583]" pageId="7" pageNumber="552">
<emphasis id="B9688653AE1FFFD099947D5B00701971" bold="true" box="[199,345,1556,1582]" pageId="7" pageNumber="552">Description.</emphasis>
Length:
<quantity id="4CE4F7A4AE1FFFD098ED7D5B034A1970" box="[446,611,1556,1583]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.25" metricValueMin="1.15" pageId="7" pageNumber="552" unit="mm" value="12.0" valueMax="12.5" valueMin="11.5">11.512.5 mm</quantity>
; width:
<quantity id="4CE4F7A4AE1FFFD09BEE7D5B026E1970" box="[701,839,1556,1583]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.2" metricValueMin="3.1" pageId="7" pageNumber="552" unit="mm" value="3.15" valueMax="3.2" valueMin="3.1">3.13.2 mm</quantity>
. Habitus as in
<figureCitation id="132746C4AE1FFFD09AB97D5B05061971" box="[1002,1071,1556,1582]" captionStart="FIGURE 6" captionStartId="8.[151,250,1996,2020]" captionTargetId="figure-20@8.[338,1248,181,1973]" captionTargetPageId="8" captionText="FIGURE 6. Habitus of Pterostichus (Huaius) hanwang sp. nov., male, holotype." figureDoi="http://doi.org/10.5281/zenodo.4289215" httpUri="https://zenodo.org/record/4289215/files/figure.png" pageId="7" pageNumber="552">Fig. 6</figureCitation>
.
</paragraph>
<paragraph id="8BA35A41AE1FFFD099947D77056F19C5" blockId="7.[151,1437,1196,2014]" pageId="7" pageNumber="552">
Head slightly widened, HLm/LW = 1.361.39, HLl/LW = 1.101.18; neck short, anterior and posterior supraorbital setiferous pores at about middle and about apical 5/6 of head respectively; clypeus bisetose; mentum tooth short, bluntly bifid at apex; mentum bisetose, submentum quadrisetose (
<figureCitation id="132746C4AE1FFFD09AED7DCF051019C5" box="[958,1081,1664,1690]" captionStart="FIGURE 7" captionStartId="9.[151,250,1877,1901]" captionTargetBox="[153,1429,198,1844]" captionTargetId="figure-19@9.[151,1435,181,1853]" captionTargetPageId="9" captionText="FIGURE 7. Head and pronotum (A, B) and labium (C, D) of Pterostichus (Huaius) species. A and C Pt. (H.) hanwang sp. nov.; B and D Pt. (H.) yuae Tian et Chen, 2019" figureDoi="http://doi.org/10.5281/zenodo.4283970" httpUri="https://zenodo.org/record/4283970/files/figure.png" pageId="7" pageNumber="552">Fig. 7A, C</figureCitation>
).
</paragraph>
<paragraph id="8BA35A41AE1FFFD099947DEA05921859" blockId="7.[151,1437,1196,2014]" pageId="7" pageNumber="552">
Pronotum quadrate, transverse, PL/PW = 0.860.92; lateral margins evidently not sinuate before hind angles (but convergent and relatively straight); PL/HLm = 0.740.76; PW/HW = 1.191.23; widest at about 1/5 of pronotum from front; fore angles porrect, widely rounded, hind angles bluntly rectangular (
<figureCitation id="132746C4AE1FFFD09D077DA305871859" box="[1108,1198,1772,1798]" captionStart="FIGURE 7" captionStartId="9.[151,250,1877,1901]" captionTargetBox="[153,1429,198,1844]" captionTargetId="figure-19@9.[151,1435,181,1853]" captionTargetPageId="9" captionText="FIGURE 7. Head and pronotum (A, B) and labium (C, D) of Pterostichus (Huaius) species. A and C Pt. (H.) hanwang sp. nov.; B and D Pt. (H.) yuae Tian et Chen, 2019" figureDoi="http://doi.org/10.5281/zenodo.4283970" httpUri="https://zenodo.org/record/4283970/files/figure.png" pageId="7" pageNumber="552">Fig. 7A</figureCitation>
).
</paragraph>
<paragraph id="8BA35A41AE1FFFD099947C5F051F182D" blockId="7.[151,1437,1196,2014]" pageId="7" pageNumber="552">Elytra much longer than wide, EL/EW = 1.85, EW/PW = 1.261.28; widest at about 2/3 from base; umbilicate marginal series composed of 1517 pores on each elytron; two dorsal pores present on interval 3, all close to stria 2, at about middle and 3/4 of elytra respectively from base; scutellar striole absent.</paragraph>
<paragraph id="8BA35A41AE1FFFD099947C3300F61881" blockId="7.[151,1437,1196,2014]" pageId="7" pageNumber="552">Hind coxa trisetose; each femur in all legs with two setae posterio-ventrally. Prosternal process blunt at apex; metepisternum with outer upper angle acute; ventrites smooth, ventrites IVVI bisetose, ventrite VII bisetose in male, quadrisetose in female.</paragraph>
<caption id="DF630AC9AE10FFDF99C47C8302E418BA" ID-DOI="http://doi.org/10.5281/zenodo.4289215" ID-Zenodo-Dep="4289215" box="[151,973,1996,2021]" httpUri="https://zenodo.org/record/4289215/files/figure.png" pageId="8" pageNumber="553" targetBox="[357,1232,202,1960]" targetPageId="8">
<paragraph id="8BA35A41AE10FFDF99C47C8302E418BA" blockId="8.[151,973,1996,2021]" box="[151,973,1996,2021]" pageId="8" pageNumber="553">
<emphasis id="B9688653AE10FFDF99C47C83003A18BA" bold="true" box="[151,275,1996,2021]" pageId="8" pageNumber="553">FIGURE 6.</emphasis>
Habitus of
<taxonomicName id="4C1C21C2AE10FFDF98D97C8203E218BA" authority="Tian &amp; He, 2020" authorityName="Tian &amp; He" authorityYear="2020" box="[394,715,1997,2021]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="553" phylum="Arthropoda" rank="species" species="hanwang" status="sp. nov." subGenus="Huaius">
<emphasis id="B9688653AE10FFDF98D97C8203E218BA" box="[394,715,1997,2021]" italics="true" pageId="8" pageNumber="553">Pterostichus (Huaius) hanwang</emphasis>
</taxonomicName>
<emphasis id="B9688653AE10FFDF9B817C82020B18BA" bold="true" box="[722,802,1997,2021]" pageId="8" pageNumber="553">
<taxonomicNameLabel id="A25B3B28AE10FFDF9B817C82020B18BA" box="[722,802,1997,2021]" pageId="8" pageNumber="553" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, holotype.
</paragraph>
</caption>
<caption id="DF630AC9AE11FFDE99C47C1A031418CE" ID-DOI="http://doi.org/10.5281/zenodo.4283970" ID-Zenodo-Dep="4283970" httpUri="https://zenodo.org/record/4283970/files/figure.png" pageId="9" pageNumber="554" startId="9.[151,250,1877,1901]" targetBox="[153,1429,198,1844]" targetPageId="9">
<paragraph id="8BA35A41AE11FFDE99C47C1A031418CE" blockId="9.[151,1436,1877,1937]" pageId="9" pageNumber="554">
<emphasis id="B9688653AE11FFDE99C47C1A003B1832" bold="true" box="[151,274,1877,1901]" pageId="9" pageNumber="554">FIGURE 7.</emphasis>
Head and pronotum (
<emphasis id="B9688653AE11FFDE98BE7C1A03351832" bold="true" box="[493,540,1877,1901]" pageId="9" pageNumber="554">A, B</emphasis>
) and labium (
<emphasis id="B9688653AE11FFDE9BFB7C1A03F01832" bold="true" box="[680,729,1877,1901]" pageId="9" pageNumber="554">C, D</emphasis>
) of
<taxonomicName id="4C1C21C2AE11FFDE9A527C1A02F71832" authority="" authorityName="Tian and Huang" authorityYear="2019" box="[769,990,1877,1901]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="subGenus" subGenus="Huaius">
<emphasis id="B9688653AE11FFDE9A527C1A02F71832" box="[769,990,1877,1901]" italics="true" pageId="9" pageNumber="554">Pterostichus (Huaius)</emphasis>
</taxonomicName>
species.
<emphasis id="B9688653AE11FFDE9D6A7C1A05631832" bold="true" box="[1081,1098,1877,1901]" pageId="9" pageNumber="554">A</emphasis>
and
<emphasis id="B9688653AE11FFDE9D297C1A05A51832" bold="true" box="[1146,1164,1877,1901]" pageId="9" pageNumber="554">C</emphasis>
<taxonomicName id="4C1C21C2AE11FFDE9DC27C1904691832" authority="Tian &amp; He, 2020" authorityName="Tian &amp; He" authorityYear="2020" box="[1169,1344,1877,1901]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="hanwang" status="sp. nov." subGenus="Huaius">
<emphasis id="B9688653AE11FFDE9DC27C1904691832" box="[1169,1344,1877,1901]" italics="true" pageId="9" pageNumber="554">Pt. (H.) hanwang</emphasis>
</taxonomicName>
<emphasis id="B9688653AE11FFDE9C167C1A04BC1832" bold="true" box="[1349,1429,1877,1901]" pageId="9" pageNumber="554">
<taxonomicNameLabel id="A25B3B28AE11FFDE9C167C1A04BC1832" box="[1349,1429,1877,1901]" pageId="9" pageNumber="554" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B9688653AE11FFDE99C47C36018118CE" bold="true" box="[151,168,1913,1937]" pageId="9" pageNumber="554">B</emphasis>
and
<emphasis id="B9688653AE11FFDE998A7C3601C218CE" bold="true" box="[217,235,1913,1937]" pageId="9" pageNumber="554">D</emphasis>
<taxonomicName id="4C1C21C2AE11FFDE99A17C35031418CE" authority="Tian et Chen, 2019" authorityName="Tian et Chen" authorityYear="2019" box="[242,573,1913,1937]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="554" phylum="Arthropoda" rank="species" species="yuae" subGenus="Huaius">
<emphasis id="B9688653AE11FFDE99A17C35005C18CE" box="[242,373,1914,1937]" italics="true" pageId="9" pageNumber="554">Pt. (H.) yuae</emphasis>
<bibRefCitation id="EF8D27B0AE11FFDE98287C36031418CE" author="Tian, M. Y. &amp; Huang, S. B. &amp; Chen, M. Z. &amp; Ding, K. J." box="[379,573,1913,1937]" pageId="9" pageNumber="554" pagination="1 - 16" refId="ref6051" refString="Tian, M. Y., Huang, S. B., Chen, M. Z. &amp; Ding, K. J. (2019) Remarkable cave-adapted ground beetles of the tribe Pterostichini from China: a new subgenus and three new species (Coleoptera: Carabidae). Annales de la Societe entomologique de France, New Series, 55 (1), 1 - 16. https: // doi. org / 10.1080 / 00379271.2018.1546554" type="journal article" year="2019">Tian et Chen, 2019</bibRefCitation>
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BA35A41AE12FFDD99947BD802181F8A" blockId="10.[151,1436,151,213]" pageId="10" pageNumber="555">
Male genitalia (
<figureCitation id="132746C4AE12FFDD982A7BD800971FEE" box="[377,446,151,177]" captionStart="FIGURE 8" captionStartId="10.[151,250,837,861]" captionTargetBox="[160,1432,240,807]" captionTargetId="figure-53@10.[151,1435,237,813]" captionTargetPageId="10" captionText="FIGURE 8. Male genitalia of Pterostichus (Huaius) hanwang sp. nov. A median lobe and parameres, lateral view B apical lobe, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4283972" httpUri="https://zenodo.org/record/4283972/files/figure.png" pageId="10" pageNumber="555">Fig. 8</figureCitation>
): Similar to
<taxonomicName id="4C1C21C2AE12FFDD9B1E7BD802A21FEE" authority="Tian and Ding, 2019" authorityName="Tian and Ding" authorityYear="2019" box="[589,907,151,177]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="555" phylum="Arthropoda" rank="species" species="caoi">
<emphasis id="B9688653AE12FFDD9B1E7BD803B31FEF" box="[589,666,151,176]" italics="true" pageId="10" pageNumber="555">P. caoi</emphasis>
<bibRefCitation id="EF8D27B0AE12FFDD9BF27BD802A21FEE" author="Tian, M. Y. &amp; Huang, S. B. &amp; Chen, M. Z. &amp; Ding, K. J." box="[673,907,151,177]" pageId="10" pageNumber="555" pagination="1 - 16" refId="ref6051" refString="Tian, M. Y., Huang, S. B., Chen, M. Z. &amp; Ding, K. J. (2019) Remarkable cave-adapted ground beetles of the tribe Pterostichini from China: a new subgenus and three new species (Coleoptera: Carabidae). Annales de la Societe entomologique de France, New Series, 55 (1), 1 - 16. https: // doi. org / 10.1080 / 00379271.2018.1546554" type="journal article" year="2019">Tian and Ding, 2019</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation id="EF8D27B0AE12FFDD9AEA7BD805591FEE" author="Tian, M. Y. &amp; Huang, S. B. &amp; Chen, M. Z. &amp; Ding, K. J." box="[953,1136,151,177]" pageId="10" pageNumber="555" pagination="1 - 16" refId="ref6051" refString="Tian, M. Y., Huang, S. B., Chen, M. Z. &amp; Ding, K. J. (2019) Remarkable cave-adapted ground beetles of the tribe Pterostichini from China: a new subgenus and three new species (Coleoptera: Carabidae). Annales de la Societe entomologique de France, New Series, 55 (1), 1 - 16. https: // doi. org / 10.1080 / 00379271.2018.1546554" type="journal article" year="2019">
Tian
<emphasis id="B9688653AE12FFDD9AA07BD8050C1FEE" box="[1011,1061,151,177]" italics="true" pageId="10" pageNumber="555">et al</emphasis>
. 2019
</bibRefCitation>
), but strongly bisinuate on ventral side, and apex sharp instead of blunt in lateral view.
</paragraph>
<caption id="DF630AC9AE12FFDD99C4780A00621CDE" ID-DOI="http://doi.org/10.5281/zenodo.4283972" ID-Zenodo-Dep="4283972" httpUri="https://zenodo.org/record/4283972/files/figure.png" pageId="10" pageNumber="555" startId="10.[151,250,837,861]" targetBox="[160,1432,240,807]" targetPageId="10">
<paragraph id="8BA35A41AE12FFDD99C4780A00621CDE" blockId="10.[151,1436,837,897]" pageId="10" pageNumber="555">
<emphasis id="B9688653AE12FFDD99C4780A003C1C02" bold="true" box="[151,277,837,861]" pageId="10" pageNumber="555">FIGURE 8.</emphasis>
Male genitalia of
<taxonomicName id="4C1C21C2AE12FFDD9884780A02341C02" authority="Tian &amp; He, 2020" authorityName="Tian &amp; He" authorityYear="2020" box="[471,797,837,861]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="555" phylum="Arthropoda" rank="species" species="hanwang" status="sp. nov." subGenus="Huaius">
<emphasis id="B9688653AE12FFDD9884780A02341C02" box="[471,797,837,861]" italics="true" pageId="10" pageNumber="555">Pterostichus (Huaius) hanwang</emphasis>
</taxonomicName>
<emphasis id="B9688653AE12FFDD9A76780A02BB1C02" bold="true" box="[805,914,837,861]" pageId="10" pageNumber="555">
<taxonomicNameLabel id="A25B3B28AE12FFDD9A76780A02511C02" box="[805,888,837,861]" pageId="10" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
median lobe and parameres, lateral view
<emphasis id="B9688653AE12FFDD9C15780A047E1C02" bold="true" box="[1350,1367,837,861]" pageId="10" pageNumber="555">B</emphasis>
apical lobe, dorsal view.
</paragraph>
</caption>
<paragraph id="8BA35A41AE12FFDD999478E0052E1B4E" blockId="10.[151,1437,943,1473]" pageId="10" pageNumber="555">
Female genitalia (
<figureCitation id="132746C4AE12FFDD98DC78E000FD1C96" box="[399,468,943,969]" captionStart="FIGURE 9" captionStartId="11.[151,250,1944,1968]" captionTargetBox="[269,1309,189,1898]" captionTargetId="figure-19@11.[260,1328,181,1920]" captionTargetPageId="11" captionText="FIGURE 9. Female genitalia of Pterostichus (Huaius) hanwang sp. nov. co common oviduct; bc bursa copulatrix; sp spermatheca; sd ductus of spermatheca; sg spermathecal gland; sgd ductus of the spermathecal gland; gscx gonosubcoxite; gcx gonocoxite." figureDoi="http://doi.org/10.5281/zenodo.4283974" httpUri="https://zenodo.org/record/4283974/files/figure.png" pageId="10" pageNumber="555">Fig. 9</figureCitation>
): Gonosubcoxite with four setae along outer margin; gonocoxite faintly bent outwards, with longer ensiform setae on outer margin; bursa copulatrix wide, simply saccate, evidently folded; oviduct situated in middle position, spermathecal gland large and long, spermatheca short.
</paragraph>
</subSubSection>
<subSubSection id="C30609CAAE12FFDD99947F5403611B22" pageId="10" pageNumber="555" type="discussion">
<paragraph id="8BA35A41AE12FFDD99947F5403611B22" blockId="10.[151,1437,943,1473]" pageId="10" pageNumber="555">
<emphasis id="B9688653AE12FFDD99947F5400121B6A" bold="true" box="[199,315,1051,1077]" pageId="10" pageNumber="555">Remarks.</emphasis>
Similar to
<taxonomicName id="4C1C21C2AE12FFDD98EE7F5302021B6A" authority="Tian and Chen" authorityName="Tian and Chen" authorityYear="2019" box="[445,811,1051,1078]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="555" phylum="Arthropoda" rank="species" species="yuae" subGenus="Huaius">
<emphasis id="B9688653AE12FFDD98EE7F5303551B6A" box="[445,636,1052,1078]" italics="true" pageId="10" pageNumber="555">P. (Huaius) yuae</emphasis>
Tian and Chen
</taxonomicName>
(in
<bibRefCitation id="EF8D27B0AE12FFDD9A087F54053D1B6A" author="Tian, M. Y. &amp; Huang, S. B. &amp; Chen, M. Z. &amp; Ding, K. J." box="[859,1044,1051,1078]" pageId="10" pageNumber="555" pagination="1 - 16" refId="ref6051" refString="Tian, M. Y., Huang, S. B., Chen, M. Z. &amp; Ding, K. J. (2019) Remarkable cave-adapted ground beetles of the tribe Pterostichini from China: a new subgenus and three new species (Coleoptera: Carabidae). Annales de la Societe entomologique de France, New Series, 55 (1), 1 - 16. https: // doi. org / 10.1080 / 00379271.2018.1546554" type="journal article" year="2019">
Tian
<emphasis id="B9688653AE12FFDD9AC57F5302E11B6A" box="[918,968,1051,1077]" italics="true" pageId="10" pageNumber="555">et al</emphasis>
. 2019
</bibRefCitation>
), but median tooth bluntly bifid at apex, instead sharply bifid (
<figureCitation id="132746C4AE12FFDD98807F7003661B05" box="[467,591,1087,1114]" captionStart="FIGURE 7" captionStartId="9.[151,250,1877,1901]" captionTargetBox="[153,1429,198,1844]" captionTargetId="figure-19@9.[151,1435,181,1853]" captionTargetPageId="9" captionText="FIGURE 7. Head and pronotum (A, B) and labium (C, D) of Pterostichus (Huaius) species. A and C Pt. (H.) hanwang sp. nov.; B and D Pt. (H.) yuae Tian et Chen, 2019" figureDoi="http://doi.org/10.5281/zenodo.4283970" httpUri="https://zenodo.org/record/4283970/files/figure.png" pageId="10" pageNumber="555">Fig. 7C, D</figureCitation>
); pronotum wider, lateral margins not sinuate before hind angles (
<figureCitation id="132746C4AE12FFDD9C6E7F7001851B21" captionStart="FIGURE 7" captionStartId="9.[151,250,1877,1901]" captionTargetBox="[153,1429,198,1844]" captionTargetId="figure-19@9.[151,1435,181,1853]" captionTargetPageId="9" captionText="FIGURE 7. Head and pronotum (A, B) and labium (C, D) of Pterostichus (Huaius) species. A and C Pt. (H.) hanwang sp. nov.; B and D Pt. (H.) yuae Tian et Chen, 2019" figureDoi="http://doi.org/10.5281/zenodo.4283970" httpUri="https://zenodo.org/record/4283970/files/figure.png" pageId="10" pageNumber="555">Fig. 7A, B</figureCitation>
); and elytra without scutellar striole.
</paragraph>
</subSubSection>
<subSubSection id="C30609CAAE12FFDD99947FC803551BFE" box="[199,636,1159,1186]" pageId="10" pageNumber="555" type="etymology">
<paragraph id="8BA35A41AE12FFDD99947FC803551BFE" blockId="10.[151,1437,943,1473]" box="[199,636,1159,1186]" pageId="10" pageNumber="555">
<emphasis id="B9688653AE12FFDD99947FC800671BFE" bold="true" box="[199,334,1159,1185]" pageId="10" pageNumber="555">Etymology.</emphasis>
Refers to the
<typeStatus id="54A7E4E3AE12FFDD98B87FC703321BFD" box="[491,539,1160,1186]" pageId="10" pageNumber="555">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C30609CAAE12FFDD99947FE405871A9E" pageId="10" pageNumber="555" type="distribution">
<paragraph id="8BA35A41AE12FFDD99947FE4040C1B9A" blockId="10.[151,1437,943,1473]" box="[199,1317,1195,1222]" pageId="10" pageNumber="555">
<emphasis id="B9688653AE12FFDD99947FE400481B9A" bold="true" box="[199,353,1195,1221]" pageId="10" pageNumber="555">Distribution.</emphasis>
<collectingCountry id="F30B1AD1AE12FFDD98347FE400831B9A" box="[359,426,1195,1221]" name="China" pageId="10" pageNumber="555">China</collectingCountry>
(
<collectingRegion id="49D894A3AE12FFDD98EA7FE403331B99" box="[441,538,1195,1222]" country="China" name="Sichuan" pageId="10" pageNumber="555">Sichuan</collectingRegion>
). Known only from the cave Hanwang Dong in Guangyuan (
<figureCitation id="132746C4AE12FFDD9D977FE4043E1B9A" box="[1220,1303,1195,1221]" captionStart="FIGURE 10" captionStartId="12.[151,250,1685,1709]" captionTargetBox="[151,1435,181,1661]" captionTargetId="figure-20@12.[151,1435,181,1661]" captionTargetPageId="12" captionText="FIGURE 10. Cave Hanwang Dong, the type locality of Pterostichus (Huaius) hanwang, sp. nov. A, entrance; B, C, spectacular scenery inside the cave; D, the moist and gravelly areas in cave, showing the habitat where the beetles were collected; E, a running individual of P. hanwang, sp. nov. in cave; F, the original state of the beetles when the stone was turned over; G, two individuals of P. hanwang, sp. nov. were collected in a test tube." figureDoi="http://doi.org/10.5281/zenodo.4283976" httpUri="https://zenodo.org/record/4283976/files/figure.png" pageId="10" pageNumber="555">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8BA35A41AE12FFDD99947F8000B91AC2" blockId="10.[151,1437,943,1473]" pageId="10" pageNumber="555">
The cave Hanwang Dong is located at the base of a limestone mountain, with a large entrance (
<figureCitation id="132746C4AE12FFDD9DBF7F80047B1BB6" box="[1260,1362,1231,1257]" captionStart="FIGURE 10" captionStartId="12.[151,250,1685,1709]" captionTargetBox="[151,1435,181,1661]" captionTargetId="figure-20@12.[151,1435,181,1661]" captionTargetPageId="12" captionText="FIGURE 10. Cave Hanwang Dong, the type locality of Pterostichus (Huaius) hanwang, sp. nov. A, entrance; B, C, spectacular scenery inside the cave; D, the moist and gravelly areas in cave, showing the habitat where the beetles were collected; E, a running individual of P. hanwang, sp. nov. in cave; F, the original state of the beetles when the stone was turned over; G, two individuals of P. hanwang, sp. nov. were collected in a test tube." figureDoi="http://doi.org/10.5281/zenodo.4283976" httpUri="https://zenodo.org/record/4283976/files/figure.png" pageId="10" pageNumber="555">Fig. 10A</figureCitation>
) and a large main passage. The main passage extends inward along a stream. It is a beautiful cave with spectacular scenery inside (
<figureCitation id="132746C4AE12FFDD98447E5800891A6E" box="[279,416,1303,1329]" captionStart="FIGURE 10" captionStartId="12.[151,250,1685,1709]" captionTargetBox="[151,1435,181,1661]" captionTargetId="figure-20@12.[151,1435,181,1661]" captionTargetPageId="12" captionText="FIGURE 10. Cave Hanwang Dong, the type locality of Pterostichus (Huaius) hanwang, sp. nov. A, entrance; B, C, spectacular scenery inside the cave; D, the moist and gravelly areas in cave, showing the habitat where the beetles were collected; E, a running individual of P. hanwang, sp. nov. in cave; F, the original state of the beetles when the stone was turned over; G, two individuals of P. hanwang, sp. nov. were collected in a test tube." figureDoi="http://doi.org/10.5281/zenodo.4283976" httpUri="https://zenodo.org/record/4283976/files/figure.png" pageId="10" pageNumber="555">Fig. 10B, C</figureCitation>
). Individuals of
<taxonomicName id="4C1C21C2AE12FFDD9B087E5703F61A6E" authority="Tian &amp; He, 2020" authorityName="Tian &amp; He" authorityYear="2020" box="[603,735,1303,1329]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="555" phylum="Arthropoda" rank="species" species="hanwang" status="sp. nov.">
<emphasis id="B9688653AE12FFDD9B087E5703F61A6E" box="[603,735,1303,1329]" italics="true" pageId="10" pageNumber="555">P. hanwang</emphasis>
</taxonomicName>
<emphasis id="B9688653AE12FFDD9BB47E58026B1A6E" bold="true" box="[743,834,1303,1329]" pageId="10" pageNumber="555">
<taxonomicNameLabel id="A25B3B28AE12FFDD9BB47E58026B1A6E" box="[743,834,1303,1329]" pageId="10" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were collected under stone in the moist and gravelly areas about
<quantity id="4CE4F7A4AE12FFDD984F7E7400B31A09" box="[284,410,1339,1366]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="10" pageNumber="555" unit="m" value="250.0" valueMax="400.0" valueMin="100.0">100400 m</quantity>
from the entrance (
<figureCitation id="132746C4AE12FFDD9B207E7403EB1A0A" box="[627,706,1339,1365]" captionStart="FIGURE 10" captionStartId="12.[151,250,1685,1709]" captionTargetBox="[151,1435,181,1661]" captionTargetId="figure-20@12.[151,1435,181,1661]" captionTargetPageId="12" captionText="FIGURE 10. Cave Hanwang Dong, the type locality of Pterostichus (Huaius) hanwang, sp. nov. A, entrance; B, C, spectacular scenery inside the cave; D, the moist and gravelly areas in cave, showing the habitat where the beetles were collected; E, a running individual of P. hanwang, sp. nov. in cave; F, the original state of the beetles when the stone was turned over; G, two individuals of P. hanwang, sp. nov. were collected in a test tube." figureDoi="http://doi.org/10.5281/zenodo.4283976" httpUri="https://zenodo.org/record/4283976/files/figure.png" pageId="10" pageNumber="555">Fig. 10</figureCitation>
DG). The cave holds a rich fauna. Apart from
<taxonomicName id="4C1C21C2AE12FFDD9D9C7E7304781A0A" authorityName="Tian &amp; He" authorityYear="2020" box="[1231,1361,1339,1365]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="555" phylum="Arthropoda" rank="species" species="hanwang">
<emphasis id="B9688653AE12FFDD9D9C7E7304781A0A" box="[1231,1361,1339,1365]" italics="true" pageId="10" pageNumber="555">P. hanwang</emphasis>
</taxonomicName>
, many other cave animals are also living in this cave, such as three trechines, bats, millipedes, spiders, moths, diplurans, amphipods and snails.
</paragraph>
<paragraph id="8BA35A41AE12FFDD99947EE705871A9E" blockId="10.[151,1437,943,1473]" box="[199,1198,1447,1473]" pageId="10" pageNumber="555">
<taxonomicName id="4C1C21C2AE12FFDD99947EE700631A9E" authority="Tian &amp; He, 2020" authorityName="Tian &amp; He" authorityYear="2020" box="[199,330,1447,1473]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="555" phylum="Arthropoda" rank="species" species="hanwang" status="sp. nov.">
<emphasis id="B9688653AE12FFDD99947EE700631A9E" box="[199,330,1447,1473]" italics="true" pageId="10" pageNumber="555">P. hanwang</emphasis>
</taxonomicName>
<emphasis id="B9688653AE12FFDD98037EE800831A9E" bold="true" box="[336,426,1447,1473]" pageId="10" pageNumber="555">
<taxonomicNameLabel id="A25B3B28AE12FFDD98037EE800831A9E" box="[336,426,1447,1473]" pageId="10" pageNumber="555" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the first troglobitic pterostichine recorded from
<collectingRegion id="49D894A3AE12FFDD9AB17EE805831A9E" box="[994,1194,1447,1473]" country="China" name="Sichuan" pageId="10" pageNumber="555">Sichuan Province</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>