533 lines
78 KiB
XML
533 lines
78 KiB
XML
<document id="33609D72D5906931B3443B4E5C942203" ID-CLB-Dataset="2588" ID-DOI="10.11646/zootaxa.5091.3.2" ID-GBIF-Dataset="f6fc2fe0-6f9c-48cf-b252-e49db45b4a98" ID-ISSN="1175-5326" ID-Zenodo-Dep="5848281" ID-ZooBank="57DE1453-41B1-43B2-9B9E-B72BDA973208" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1642146879451" checkinUser="plazi" docAuthor="Durante, Antonio & Apinda-Legnouo, Emelie Arlette" docDate="2022" docId="03ACBD4BFFABFFF50D8FFEAF37C3F9F4" docLanguage="en" docName="zootaxa.5091.3.2.pdf" docOrigin="Zootaxa 5091 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pusiola unicolor Durante & Apinda-Legnouo 2022, new species" docType="treatment" docVersion="7" lastPageNumber="419" masterDocId="FF95C533FFA9FFF60D18FFAD337BFF81" masterDocTitle="Fifth contribution to the study of the Lithosiini of Gabon: two genera new for Gabon, with a description of two new species (Lepidoptera: Erebidae: Arctiinae)" masterLastPageNumber="428" masterPageNumber="416" pageNumber="418" updateTime="1699302283932" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BD9237657732059F27ADC256BC133819">Fifth contribution to the study of the Lithosiini of Gabon: two genera new for Gabon, with a description of two new species (Lepidoptera: Erebidae: Arctiinae)</mods:title>
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<mods:name id="7F936972E575A03C81E9083CAAD7C7C3" type="personal">
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<mods:roleTerm id="5AC40141D8278829E89009FFC1DDDB4F">Author</mods:roleTerm>
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<mods:namePart id="DC76783EF31EFA07A683ADB42F26984F">Durante, Antonio</mods:namePart>
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<mods:affiliation id="5AD252288136050662EDE4BFE7EDA20B">Museo di Storia naturale del Salento. S. P. Calimera-Borgagne, Km 1, 73021 Calimera, Italy.</mods:affiliation>
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<mods:namePart id="BD61D3B2DADEDA88B7007C8E70A1D19E">Apinda-Legnouo, Emelie Arlette</mods:namePart>
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<mods:affiliation id="3EB6A16CC33B9F0F2FDC644D790AD3FE">Institut de Recherches sur l’Écologie Tropicale, Libreville, Gabon.</mods:affiliation>
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<mods:typeOfResource id="483F890E56DA48981513BEB6588259D3">text</mods:typeOfResource>
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<mods:title id="B59D76C91A3736B7BDE6C859AD6E9326">Zootaxa</mods:title>
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<mods:part id="D78E718199743025708AB4925850EB31">
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<mods:date id="582568D3C57648499DBBAE0DEA3D27FE">2022</mods:date>
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<mods:number id="8556D399E60741683D37995D148CE11F">2022-01-14</mods:number>
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<mods:number id="8340580526202349CD42BEF10E7BBC18">5091</mods:number>
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<treatment id="03ACBD4BFFABFFF50D8FFEAF37C3F9F4" ID-DOI="http://doi.org/10.5281/zenodo.5851082" ID-GBIF-Taxon="192131082" ID-Zenodo-Dep="5851082" LSID="urn:lsid:plazi:treatment:03ACBD4BFFABFFF50D8FFEAF37C3F9F4" httpUri="http://treatment.plazi.org/id/03ACBD4BFFABFFF50D8FFEAF37C3F9F4" lastPageId="3" lastPageNumber="419" pageId="2" pageNumber="418">
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<subSubSection id="C31F5FD6FFABFFF40D8FFEAF311CFE9D" box="[151,615,258,285]" pageId="2" pageNumber="418" type="nomenclature">
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<paragraph id="8BBA0C5DFFABFFF40D8FFEAF311CFE9D" blockId="2.[151,615,258,321]" box="[151,615,258,285]" pageId="2" pageNumber="418">
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<heading id="D0F2BB31FFABFFF40D8FFEAF311CFE9D" bold="true" box="[151,615,258,285]" fontSize="11" level="1" pageId="2" pageNumber="418" reason="1">
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<emphasis id="B971D04FFFABFFF40D8FFEAF311CFE9D" bold="true" box="[151,615,258,285]" pageId="2" pageNumber="418">
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<taxonomicName id="4C0577DEFFABFFF40D8FFEAF32ABFE9C" authority="Durante" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[151,464,258,285]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="unicolor" status="sp. nov.">
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<emphasis id="B971D04FFFABFFF40D8FFEAF321AFE9C" bold="true" box="[151,353,258,285]" italics="true" pageId="2" pageNumber="418">Pusiola unicolor</emphasis>
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Durante
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</taxonomicName>
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<taxonomicNameLabel id="A2426D34FFABFFF40CCFFEAE311CFE9D" box="[471,615,258,285]" pageId="2" pageNumber="418" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C31F5FD6FFABFFF40D8FFE8A3229FEC0" box="[151,338,295,321]" pageId="2" pageNumber="418" type="description">
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<paragraph id="8BBA0C5DFFABFFF40D8FFE8A3229FEC0" blockId="2.[151,615,258,321]" box="[151,338,295,321]" pageId="2" pageNumber="418">
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(
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<figureCitation id="133E10D8FFABFFF40D87FE8A3278FEC0" box="[159,259,295,321]" captionStart="FIGURES 1–8" captionStartId="4.[151,264,1866,1890]" captionTargetBox="[170,1414,194,1826]" captionTargetId="figure-21@4.[160,1427,179,1841]" captionTargetPageId="4" captionText="FIGURES 1–8. Lithosiini spp.: adults. 1, Pusiola unicolor n. sp.—holotype male. Gabon, Ivindo National Park. 2, ditto—paratype, male. Gabon, Ivindo National Park. 3, ditto—paratype, female. Gabon, Ivindo National Park. 4, ditto—paratype, female. Gabon, Ivindo National Park. 5, Pusiola celida—male. Gabon, Mondah forest. 6, Novilema bifurcum n. sp.—holotype, male. Gabon, Ivindo National Park. 7, Novilema fluminale—male. Gabon, Ivindo National Park. 8, ditto—male. Gabon, Ivindo National Park." figureDoi="http://doi.org/10.5281/zenodo.5848283" httpUri="https://zenodo.org/record/5848283/files/figure.png" pageId="2" pageNumber="418">Figs 1–4</figureCitation>
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,
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<figureCitation id="133E10D8FFABFFF40C08FE8A325BFEC0" box="[272,288,295,321]" captionStart="FIGURE 9" captionStartId="5.[151,250,1889,1913]" captionTargetBox="[185,1397,197,1850]" captionTargetId="figure-27@5.[171,1416,181,1865]" captionTargetPageId="5" captionText="FIGURE 9. Pusiola unicolor n. sp.—A: central segment of the antennal flagellum, male; B: central segment of the antennal flagellum, female; C: dorsal pheromone gland; D: acrosternite; E: forewing venation; F: hindwing venation. A: anal vein; co: coremata; CuA: Anterior cubital vein; dp: dome-shaped projections; M: median vein; pg: dorsal pheromone glands; R: radial vein; Rs: radial sector vein; Sc: subcostal vein; VIII st: eighth sternum." figureDoi="http://doi.org/10.5281/zenodo.5848285" httpUri="https://zenodo.org/record/5848285/files/figure.png" pageId="2" pageNumber="418">9</figureCitation>
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,
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<figureCitation id="133E10D8FFABFFF40C34FE8A3232FEC0" box="[300,329,295,321]" captionStart="FIGURE 10" captionStartId="6.[151,250,1756,1780]" captionTargetBox="[231,1357,193,1720]" captionTargetId="figure-21@6.[231,1357,181,1732]" captionTargetPageId="6" captionText="FIGURE 10. Pusiola unicolor n. sp.—A–D: male genitalia. A: processus distalis plicae, enlarged; B: genitalia, aedeagus removed; C: aedeagus, vesica everted; D: granicula of the vesica, enlarged. E–H: female genitalia. E: genitalia; F: enlarged spicule of the antrum; G: enlarged spicule of the foremost part of the corpus bursae; H: enlarged spicule of the medio-caudal part of the corpus bursae. aa: anterior apophyses; ab: appendix bursae; ae: aedeagus; an: antrum; ap: posterior apophyses; av: ala valvae; cb: corpus bursae; cp: coecum penis; dl: distal lobus; fs: dorsal flat sclerotization of the aedeagus; gr: granicula; ju: juxta; pa: papillae anales; pav: distal process of the ala valvae; pd: processus distalis plicae; pe: pedunculi; pg: dorsal pheromone glands; sa: saccus; sv: supravalva; te: tegumen; un: uncus; ve: vesica; vi: vinculum; VII st: seventh sternum; VIII st: eighth sternum; vl: ventral lobus." figureDoi="http://doi.org/10.5281/zenodo.5848287" httpUri="https://zenodo.org/record/5848287/files/figure.png" pageId="2" pageNumber="418">10</figureCitation>
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)
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C31F5FD6FFABFFF40D8FFEC23138FD4C" pageId="2" pageNumber="418" type="materials_examined">
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<paragraph id="8BBA0C5DFFABFFF40D8FFEC232BAFE2C" blockId="2.[151,1437,366,2013]" pageId="2" pageNumber="418">
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<materialsCitation id="3B6D0600FFABFFF40D8FFEC232BAFE2C" ID-GBIF-Occurrence="3436732309" collectingDate="2017-11-20" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B971D04FFFABFFF40D8FFEC2327FFE08" bold="true" box="[151,260,367,393]" pageId="2" pageNumber="418">
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<typeStatus id="54BEB2FFFFABFFF40D8FFEC2327FFE08" box="[151,260,367,393]" pageId="2" pageNumber="418" type="holotype">Holotype</typeStatus>
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</emphasis>
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:
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<specimenCount id="9D03C7D4FFABFFF40C0AFEC23252FE08" box="[274,297,367,393]" pageId="2" pageNumber="418" type="male">♂</specimenCount>
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,
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<collectingCountry id="F3124CCDFFABFFF40C2DFEC232E0FE08" box="[309,411,367,393]" name="Gabon" pageId="2" pageNumber="418">GABON</collectingCountry>
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,
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<location id="8EDA5A86FFABFFF40CBEFEC23297FE08" LSID="urn:lsid:plazi:treatment:03ACBD4BFFABFFF50D8FFEAF37C3F9F4:8EDA5A86FFABFFF40CBEFEC23297FE08" box="[422,492,367,393]" country="Gabon" latitude="0.5119445" longLatPrecision="21" longitude="12.803612" name="Ipassa" pageId="2" pageNumber="418">Ipassa</location>
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(Makokou),
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</quantity>
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,
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<geoCoordinate id="EE316A9AFFABFFF40E4BFEC230A0FE08" box="[851,987,367,393]" degrees="12" direction="east" minutes="48" orientation="longitude" pageId="2" pageNumber="418" precision="15" seconds="13" value="12.803612">12°48’13”E</geoCoordinate>
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, 18/
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<date id="FFBB2A9DFFABFFF40913FEC237F4FE08" box="[1035,1167,367,393]" pageId="2" pageNumber="418" value="2017-11-20">
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<collectingDate id="EFFFD375FFABFFF40913FEC237F4FE08" box="[1035,1167,367,393]" pageId="2" pageNumber="418" value="2017-11-20">20-11-2017</collectingDate>
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</date>
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,
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<collectorName id="26F0698BFFABFFF40982FEC2378FFE08" box="[1178,1268,367,393]" pageId="2" pageNumber="418">Durante</collectorName>
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leg., gen. sl. n. 989
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<collectionCode id="ED149498FFABFFF40DD0FE3E3276FE2C" box="[200,269,403,429]" country="India" httpUri="http://grbio.org/cool/dqz8-sfkc" name="Madras Museum" pageId="2" pageNumber="418">MAD</collectionCode>
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; in coll.
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<collectionCode id="ED149498FFABFFF40C75FE3E32C6FE2C" box="[365,445,403,429]" pageId="2" pageNumber="418">MSNS</collectionCode>
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.
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</paragraph>
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<paragraph id="8BBA0C5DFFABFFF40DDFFE1A3138FD4C" blockId="2.[151,1437,366,2013]" pageId="2" pageNumber="418">
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<materialsCitation id="3B6D0600FFABFFF40DDFFE1A31B2FE50" ID-GBIF-Occurrence="3436732317" box="[199,713,439,465]" collectingDate="2017-11-20" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="7" specimenCount-male="7" typeStatus="paratype">
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<emphasis id="B971D04FFFABFFF40DDFFE1A3244FE50" bold="true" box="[199,319,439,465]" pageId="2" pageNumber="418">
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<typeStatus id="54BEB2FFFFABFFF40DDFFE1A3244FE50" box="[199,319,439,465]" pageId="2" pageNumber="418" type="paratype">Paratypes</typeStatus>
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</emphasis>
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:
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<specimenCount id="9D03C7D4FFABFFF40C57FE1A32E9FE50" box="[335,402,439,465]" pageId="2" pageNumber="418" type="male">7 ♂♂</specimenCount>
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, same data as the holotype
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</materialsCitation>
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;
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<materialsCitation id="3B6D0600FFABFFF40FCDFE1B3660FE50" ID-GBIF-Occurrence="3436732311" box="[725,1307,438,465]" collectingDate="2013-12-13" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="5" specimenCount-male="5" typeStatus="paratype">
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<specimenCount id="9D03C7D4FFABFFF40FCDFE1B306CFE50" box="[725,791,438,465]" pageId="2" pageNumber="418" type="male">5 ♂♂</specimenCount>
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, same locality as the holotype, 4/
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<date id="FFBB2A9DFFABFFF4098FFE1A3660FE50" box="[1175,1307,439,465]" pageId="2" pageNumber="418" value="2013-12-13">
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<collectingDate id="EFFFD375FFABFFF4098FFE1A3660FE50" box="[1175,1307,439,465]" pageId="2" pageNumber="418" value="2013-12-13">13-12-2013</collectingDate>
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</date>
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</materialsCitation>
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||
;
|
||
<materialsCitation id="3B6D0600FFABFFF40830FE1A313FFE74" ID-GBIF-Occurrence="3436732301" collectingDate="2015-03-24" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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||
<specimenCount id="9D03C7D4FFABFFF40830FE1A362EFE50" box="[1320,1365,439,465]" pageId="2" pageNumber="418" type="male">1 ♂</specimenCount>
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||
, same locality as the holotype, 14/
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<date id="FFBB2A9DFFABFFF40CD4FE77313FFE74" box="[460,580,474,501]" pageId="2" pageNumber="418" value="2015-03-24">
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<collectingDate id="EFFFD375FFABFFF40CD4FE77313FFE74" box="[460,580,474,501]" pageId="2" pageNumber="418" value="2015-03-24">24-3-2015</collectingDate>
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</date>
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</materialsCitation>
|
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;
|
||
<materialsCitation id="3B6D0600FFABFFF40F56FE7637E8FE74" ID-GBIF-Occurrence="3436732304" box="[590,1171,475,501]" collectingDate="2017-11-24" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
|
||
<specimenCount id="9D03C7D4FFABFFF40F56FE7631F4FE74" box="[590,655,475,501]" pageId="2" pageNumber="418" type="male">3 ♂♂</specimenCount>
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||
, same locality as the holotype, 19/
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||
<date id="FFBB2A9DFFABFFF40917FE7637E8FE74" box="[1039,1171,475,501]" pageId="2" pageNumber="418" value="2017-11-24">
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<collectingDate id="EFFFD375FFABFFF40917FE7637E8FE74" box="[1039,1171,475,501]" pageId="2" pageNumber="418" value="2017-11-24">24-11-2017</collectingDate>
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</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B6D0600FFABFFF40986FE7632A3FD98" ID-GBIF-Occurrence="3436732316" collectingDate="2017-11-22" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="8" specimenCount-male="8" typeStatus="paratype">
|
||
<specimenCount id="9D03C7D4FFABFFF40986FE7637A4FE74" box="[1182,1247,475,501]" pageId="2" pageNumber="418" type="male">8 ♂♂</specimenCount>
|
||
, same locality as the holotype, 21/
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||
<date id="FFBB2A9DFFABFFF40C4CFE5232A3FD98" box="[340,472,511,537]" pageId="2" pageNumber="418" value="2017-11-22">
|
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<collectingDate id="EFFFD375FFABFFF40C4CFE5232A3FD98" box="[340,472,511,537]" pageId="2" pageNumber="418" value="2017-11-22">22-11-2017</collectingDate>
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</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B6D0600FFABFFF40CFBFE52302BFD98" ID-GBIF-Occurrence="3436732308" box="[483,848,511,537]" collectingDate="2017-11-20" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
|
||
<specimenCount id="9D03C7D4FFABFFF40CFBFE523159FD98" box="[483,546,511,537]" pageId="2" pageNumber="418" type="female">3 ♀♀</specimenCount>
|
||
, same data as the holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B6D0600FFABFFF40E42FE5236FFFD98" ID-GBIF-Occurrence="3436732326" box="[858,1412,510,537]" collectingDate="2010-04-10" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
|
||
<specimenCount id="9D03C7D4FFABFFF40E42FE5230E2FD98" box="[858,921,511,537]" pageId="2" pageNumber="418" type="female">2 ♀♀</specimenCount>
|
||
, same locality as the holotype, 5/
|
||
<date id="FFBB2A9DFFABFFF40814FE5336FFFD98" box="[1292,1412,510,537]" pageId="2" pageNumber="418" value="2010-04-10">
|
||
<collectingDate id="EFFFD375FFABFFF40814FE5336FFFD98" box="[1292,1412,510,537]" pageId="2" pageNumber="418" value="2010-04-10">10-4-2010</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B6D0600FFABFFF40896FE5231AFFDBC" ID-GBIF-Occurrence="3436732320" collectingDate="2011-03-11" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="7" specimenCount-female="7" typeStatus="paratype">
|
||
<specimenCount id="9D03C7D4FFABFFF40896FE5233B9FDBC" pageId="2" pageNumber="418" type="female">7 ♀♀</specimenCount>
|
||
, same locality as the holotype, 19-2/
|
||
<date id="FFBB2A9DFFABFFF40F78FD8E31AFFDBC" box="[608,724,547,573]" pageId="2" pageNumber="418" value="2011-03-11">
|
||
<collectingDate id="EFFFD375FFABFFF40F78FD8E31AFFDBC" box="[608,724,547,573]" pageId="2" pageNumber="418" value="2011-03-11">11-3-2011</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B6D0600FFABFFF40FF9FD8E3672FDBC" ID-GBIF-Occurrence="3436732322" box="[737,1289,547,573]" collectingDate="2012-02-28" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D03C7D4FFABFFF40FF9FD8E3070FDBC" box="[737,779,547,573]" pageId="2" pageNumber="418" type="female">1 ♀</specimenCount>
|
||
, same locality as the holotype, 16/
|
||
<date id="FFBB2A9DFFABFFF4098AFD8E3672FDBC" box="[1170,1289,547,573]" pageId="2" pageNumber="418" value="2012-02-28">
|
||
<collectingDate id="EFFFD375FFABFFF4098AFD8E3672FDBC" box="[1170,1289,547,573]" pageId="2" pageNumber="418" value="2012-02-28">28-2-2012</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B6D0600FFABFFF4080DFD8E3131FDE0" ID-GBIF-Occurrence="3436732302" collectingDate="2013-12-13" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
|
||
<specimenCount id="9D03C7D4FFABFFF4080DFD8E362EFDBC" box="[1301,1365,547,573]" pageId="2" pageNumber="418" type="female">3 ♀♀</specimenCount>
|
||
, same locality as the holotype, 4/
|
||
<date id="FFBB2A9DFFABFFF40CDFFDEA3131FDE0" box="[455,586,583,609]" pageId="2" pageNumber="418" value="2013-12-13">
|
||
<collectingDate id="EFFFD375FFABFFF40CDFFDEA3131FDE0" box="[455,586,583,609]" pageId="2" pageNumber="418" value="2013-12-13">13-12-2013</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B6D0600FFABFFF40F4FFDEA37D1FDE0" ID-GBIF-Occurrence="3436732313" box="[599,1194,582,609]" collectingDate="2015-03-24" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="14" specimenCount-female="14" typeStatus="paratype">
|
||
<specimenCount id="9D03C7D4FFABFFF40F4FFDEA31DCFDE0" box="[599,679,583,609]" pageId="2" pageNumber="418" type="female">14 ♀♀</specimenCount>
|
||
, same locality as the holotype, 14/
|
||
<date id="FFBB2A9DFFABFFF4092BFDEB37D1FDE0" box="[1075,1194,582,609]" pageId="2" pageNumber="418" value="2015-03-24">
|
||
<collectingDate id="EFFFD375FFABFFF4092BFDEB37D1FDE0" box="[1075,1194,582,609]" pageId="2" pageNumber="418" value="2015-03-24">24-3-2015</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B6D0600FFABFFF409AFFDEA3282FD04" ID-GBIF-Occurrence="3436732321" collectingDate="2017-11-20" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
|
||
<specimenCount id="9D03C7D4FFABFFF409AFFDEA3783FDE0" box="[1207,1272,583,609]" pageId="2" pageNumber="418" type="female">3 ♀♀</specimenCount>
|
||
, same locality as the holotype, 18/
|
||
<date id="FFBB2A9DFFABFFF40C6DFDC63282FD04" box="[373,505,619,645]" pageId="2" pageNumber="418" value="2017-11-20">
|
||
<collectingDate id="EFFFD375FFABFFF40C6DFDC63282FD04" box="[373,505,619,645]" pageId="2" pageNumber="418" value="2017-11-20">20-11-2017</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B6D0600FFABFFF40F1EFDC7372FFD04" ID-GBIF-Occurrence="3436732312" box="[518,1108,618,645]" collectingDate="2017-11-24" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="5" specimenCount-female="5" typeStatus="paratype">
|
||
<specimenCount id="9D03C7D4FFABFFF40F1EFDC7313CFD04" box="[518,583,618,645]" pageId="2" pageNumber="418" type="female">5 ♀♀</specimenCount>
|
||
, same locality as the holotype, 19/
|
||
<date id="FFBB2A9DFFABFFF40EC9FDC6372FFD04" box="[977,1108,619,645]" pageId="2" pageNumber="418" value="2017-11-24">
|
||
<collectingDate id="EFFFD375FFABFFF40EC9FDC6372FFD04" box="[977,1108,619,645]" pageId="2" pageNumber="418" value="2017-11-24">24-11-2017</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B6D0600FFABFFF40979FDC632CAFD28" ID-GBIF-Occurrence="3436732307" collectingDate="2017-11-22" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="14" specimenCount-female="14" typeStatus="paratype">
|
||
<specimenCount id="9D03C7D4FFABFFF40979FDC637CBFD04" box="[1121,1200,619,645]" pageId="2" pageNumber="418" type="female">14 ♀♀</specimenCount>
|
||
, same locality as the holotype, 21/
|
||
<date id="FFBB2A9DFFABFFF40C35FD2232CAFD28" box="[301,433,655,681]" pageId="2" pageNumber="418" value="2017-11-22">
|
||
<collectingDate id="EFFFD375FFABFFF40C35FD2232CAFD28" box="[301,433,655,681]" pageId="2" pageNumber="418" value="2017-11-22">22-11-2017</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
; all Durante leg.
|
||
<materialsCitation id="3B6D0600FFABFFF40F6DFD2237CCFD28" ID-GBIF-Occurrence="3436732315" box="[629,1207,654,681]" collectingDate="2016-06-25" collectionCode="MAD, MSNS" collectorName="Durante" country="Gabon" elevation="500" latitude="0.5119445" location="Ipassa" longLatPrecision="21" longitude="12.803612" pageId="2" pageNumber="418" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
|
||
<specimenCount id="9D03C7D4FFABFFF40F6DFD2231CDFD28" box="[629,694,655,681]" pageId="2" pageNumber="418" type="female">4 ♀♀</specimenCount>
|
||
, same locality as the holotype, 14/
|
||
<date id="FFBB2A9DFFABFFF40958FD2337CCFD28" box="[1088,1207,654,681]" pageId="2" pageNumber="418" value="2016-06-25">
|
||
<collectingDate id="EFFFD375FFABFFF40958FD2337CCFD28" box="[1088,1207,654,681]" pageId="2" pageNumber="418" value="2016-06-25">25-6-2016</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
, Durante & Fasiello leg. All in the first author’s collection.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F5FD6FFABFFF40DDFFD7B32D8FB44" pageId="2" pageNumber="418" type="diagnosis">
|
||
<paragraph id="8BBA0C5DFFABFFF40DDFFD7B32CFFC48" blockId="2.[151,1437,366,2013]" pageId="2" pageNumber="418">
|
||
<emphasis id="B971D04FFFABFFF40DDFFD7B3241FD71" bold="true" box="[199,314,726,752]" pageId="2" pageNumber="418">Diagnosis</emphasis>
|
||
. The absence of a discal dot on the forewings distinguishes
|
||
<taxonomicName id="4C0577DEFFABFFF40EEDFD7A37C8FD70" authority="Durante & Apinda-Legnouo, 2022" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[1013,1203,727,753]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="unicolor" status="sp. nov.">
|
||
<emphasis id="B971D04FFFABFFF40EEDFD7A37C8FD70" box="[1013,1203,727,753]" italics="true" pageId="2" pageNumber="418">Pusiola unicolor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B971D04FFFABFFF409A4FD7A3786FD70" bold="true" box="[1212,1277,727,753]" pageId="2" pageNumber="418">
|
||
<taxonomicNameLabel id="A2426D34FFABFFF409A4FD7A3786FD70" box="[1212,1277,727,753]" pageId="2" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from
|
||
<taxonomicName id="4C0577DEFFABFFF4085FFD7A33A0FC94" baseAuthorityName="Bethune-Baker" baseAuthorityYear="1911" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="celida">
|
||
<emphasis id="B971D04FFFABFFF4085FFD7A33A0FC94" italics="true" pageId="2" pageNumber="418">Pusiola celida</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0577DEFFABFFF40C0BFD5631CCFC97" authority="(Wallengren, 1860)" baseAuthorityName="Wallengren" baseAuthorityYear="1860" box="[275,695,763,790]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="cinerella">
|
||
<emphasis id="B971D04FFFABFFF40C0BFD5632A8FC94" box="[275,467,763,789]" italics="true" pageId="2" pageNumber="418">Pusiola cinerella</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9471ACFFABFFF40CFAFD5631D4FC97" author="Wallengren, H. D. J." box="[482,687,763,790]" pageId="2" pageNumber="418" pagination="33 - 46" refId="ref8059" refString="Wallengren, H. D. J. (1860) Lepidopterologische Mittheilungen. Wiener entomologische Monatschrift, 4 (2), 33 - 46." type="journal article" year="1860">Wallengren, 1860</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; the pale shades of yellow (creamy, ochreous, ivory) or light grey of the majority of the congeneric species are diagnostic compared to the dark grey-brown colour of
|
||
<taxonomicName id="4C0577DEFFABFFF40807FCB236E7FCB8" authority="Durante & Apinda-Legnouo, 2022" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[1311,1436,799,825]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="unicolor" status="sp. nov.">
|
||
<emphasis id="B971D04FFFABFFF40807FCB236E7FCB8" box="[1311,1436,799,825]" italics="true" pageId="2" pageNumber="418">P. unicolor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B971D04FFFABFFF40D8FFCEE33ACFCDC" bold="true" box="[151,215,835,861]" pageId="2" pageNumber="418">
|
||
<taxonomicNameLabel id="A2426D34FFABFFF40D8FFCEE33ACFCDC" box="[151,215,835,861]" pageId="2" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
The only species that could require comparison of the genitalia are
|
||
<taxonomicName id="4C0577DEFFABFFF40EF9FCEE3635FCDF" authority="(Wallengren, 1860)" baseAuthorityName="Wallengren" baseAuthorityYear="1860" box="[993,1358,835,862]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B971D04FFFABFFF40EF9FCEE3710FCDC" box="[993,1131,835,861]" italics="true" pageId="2" pageNumber="418">P. flavicosta</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9471ACFFABFFF40963FCEE363DFCDF" author="Wallengren, H. D. J." box="[1147,1350,835,862]" pageId="2" pageNumber="418" pagination="33 - 46" refId="ref8059" refString="Wallengren, H. D. J. (1860) Lepidopterologische Mittheilungen. Wiener entomologische Monatschrift, 4 (2), 33 - 46." type="journal article" year="1860">Wallengren, 1860</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0577DEFFABFFF40890FCEE32B8FC00" authority="(Kiriakoff, 1958)" baseAuthorityName="Kiriakoff" baseAuthorityYear="1958" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="tinaeella">
|
||
<emphasis id="B971D04FFFABFFF40890FCEE3382FC00" italics="true" pageId="2" pageNumber="418">P. tinaeella</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9471ACFFABFFF40C1FFCCA32C0FC00" author="Kiriakoff, S. G." box="[263,443,870,897]" pageId="2" pageNumber="418" refId="ref7473" refString="Kiriakoff, S. G. (1958) Ruwenzori Expedition, 1952. Vol. I. 2 - 3. Trustees of the British Museum, London, 53 pp." type="book" year="1958">Kiriakoff, 1958</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The former however possesses a yellow line along the forewing’s costal margin, absent in
|
||
<taxonomicName id="4C0577DEFFABFFF40DADFC26324AFC24" authority="Durante & Apinda-Legnouo, 2022" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[181,305,907,933]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="unicolor" status="sp. nov.">
|
||
<emphasis id="B971D04FFFABFFF40DADFC26324AFC24" box="[181,305,907,933]" italics="true" pageId="2" pageNumber="418">P. unicolor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B971D04FFFABFFF40C22FC263201FC24" bold="true" box="[314,378,907,933]" pageId="2" pageNumber="418">
|
||
<taxonomicNameLabel id="A2426D34FFABFFF40C22FC263201FC24" box="[314,378,907,933]" pageId="2" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; the latter is generally paler than
|
||
<taxonomicName id="4C0577DEFFABFFF40FE4FC263020FC24" authority="Durante & Apinda-Legnouo, 2022" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[764,859,907,933]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="unicolor" status="sp. nov.">
|
||
<emphasis id="B971D04FFFABFFF40FE4FC263020FC24" box="[764,859,907,933]" italics="true" pageId="2" pageNumber="418">unicolor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B971D04FFFABFFF40E7BFC2630D8FC24" bold="true" box="[867,931,907,933]" pageId="2" pageNumber="418">
|
||
<taxonomicNameLabel id="A2426D34FFABFFF40E7BFC2630D8FC24" box="[867,931,907,933]" pageId="2" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
In any case, the following genital apparatus characters are diagnostic.
|
||
</paragraph>
|
||
<paragraph id="8BBA0C5DFFABFFF40DDFFC7E32D8FB44" blockId="2.[151,1437,366,2013]" pageId="2" pageNumber="418">
|
||
Comparing
|
||
<taxonomicName id="4C0577DEFFABFFF40C54FC7E32AEFC6C" baseAuthorityName="Wallengren" baseAuthorityYear="1860" box="[332,469,979,1005]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B971D04FFFABFFF40C54FC7E32AEFC6C" box="[332,469,979,1005]" italics="true" pageId="2" pageNumber="418">P. flavicosta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0577DEFFABFFF40F14FC7E31F2FC6C" baseAuthorityName="Kiriakoff" baseAuthorityYear="1958" box="[524,649,979,1005]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="tinaeella">
|
||
<emphasis id="B971D04FFFABFFF40F14FC7E31F2FC6C" box="[524,649,979,1005]" italics="true" pageId="2" pageNumber="418">P. tinaeella</emphasis>
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C0577DEFFABFFF40FD2FC7E303FFC6C" authority="Durante & Apinda-Legnouo, 2022" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[714,836,979,1005]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="unicolor" status="sp. nov.">
|
||
<emphasis id="B971D04FFFABFFF40FD2FC7E303FFC6C" box="[714,836,979,1005]" italics="true" pageId="2" pageNumber="418">P. unicolor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B971D04FFFABFFF40E53FC7E30F1FC6C" bold="true" box="[843,906,979,1005]" pageId="2" pageNumber="418">
|
||
<taxonomicNameLabel id="A2426D34FFABFFF40E53FC7E30F1FC6C" box="[843,906,979,1005]" pageId="2" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the differential characters are as follows: i) the valva and uncus of
|
||
<taxonomicName id="4C0577DEFFABFFF40C6AFC5A3296FB90" authority="Durante & Apinda-Legnouo, 2022" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[370,493,1015,1041]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="unicolor" status="sp. nov.">
|
||
<emphasis id="B971D04FFFABFFF40C6AFC5A3296FB90" box="[370,493,1015,1041]" italics="true" pageId="2" pageNumber="418">P. unicolor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B971D04FFFABFFF40CEDFC5A314FFB90" bold="true" box="[501,564,1015,1041]" pageId="2" pageNumber="418">
|
||
<taxonomicNameLabel id="A2426D34FFABFFF40CEDFC5A314FFB90" box="[501,564,1015,1041]" pageId="2" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are longer and more slender; ii) the juxta is L-shaped and protrudes caudally, while it is trapezoid and flat in
|
||
<taxonomicName id="4C0577DEFFABFFF40CE0FBB6311EFBB4" baseAuthorityName="Wallengren" baseAuthorityYear="1860" box="[504,613,1051,1077]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B971D04FFFABFFF40CE0FBB6311EFBB4" box="[504,613,1051,1077]" italics="true" pageId="2" pageNumber="418">flavicosta</emphasis>
|
||
</taxonomicName>
|
||
, and X-shaped and flat in
|
||
<taxonomicName id="4C0577DEFFABFFF40E89FBB63088FBB4" baseAuthorityName="Kiriakoff" baseAuthorityYear="1958" box="[913,1011,1051,1077]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="tinaeella">
|
||
<emphasis id="B971D04FFFABFFF40E89FBB63088FBB4" box="[913,1011,1051,1077]" italics="true" pageId="2" pageNumber="418">tinaeella</emphasis>
|
||
</taxonomicName>
|
||
; iii) the saccus is larger and u-shaped in
|
||
<taxonomicName id="4C0577DEFFABFFF40DAEFB923248FBD8" authority="Durante & Apinda-Legnouo, 2022" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[182,307,1087,1113]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="unicolor" status="sp. nov.">
|
||
<emphasis id="B971D04FFFABFFF40DAEFB923248FBD8" box="[182,307,1087,1113]" italics="true" pageId="2" pageNumber="418">P. unicolor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B971D04FFFABFFF40C24FB923206FBD8" bold="true" box="[316,381,1087,1113]" pageId="2" pageNumber="418">
|
||
<taxonomicNameLabel id="A2426D34FFABFFF40C24FB923206FBD8" box="[316,381,1087,1113]" pageId="2" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, with a larger median notch, while it is slender and bifid in
|
||
<taxonomicName id="4C0577DEFFABFFF4092FFB9237DFFBD8" baseAuthorityName="Wallengren" baseAuthorityYear="1860" box="[1079,1188,1087,1113]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B971D04FFFABFFF4092FFB9237DFFBD8" box="[1079,1188,1087,1113]" italics="true" pageId="2" pageNumber="418">flavicosta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0577DEFFABFFF409C7FB92363AFBD8" baseAuthorityName="Kiriakoff" baseAuthorityYear="1958" box="[1247,1345,1087,1113]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="tinaeella">
|
||
<emphasis id="B971D04FFFABFFF409C7FB92363AFBD8" box="[1247,1345,1087,1113]" italics="true" pageId="2" pageNumber="418">tinaeella</emphasis>
|
||
</taxonomicName>
|
||
; iv) the aedeagus of
|
||
<taxonomicName id="4C0577DEFFABFFF40C3BFBCE32E5FBFC" authority="Durante & Apinda-Legnouo, 2022" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[291,414,1123,1149]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="unicolor" status="sp. nov.">
|
||
<emphasis id="B971D04FFFABFFF40C3BFBCE32E5FBFC" box="[291,414,1123,1149]" italics="true" pageId="2" pageNumber="418">P. unicolor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B971D04FFFABFFF40CBDFBCE329FFBFC" bold="true" box="[421,484,1123,1149]" pageId="2" pageNumber="418">
|
||
<taxonomicNameLabel id="A2426D34FFABFFF40CBDFBCE329FFBFC" box="[421,484,1123,1149]" pageId="2" pageNumber="418" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
bears only granicula (graniculi in
|
||
<bibRefCitation id="EF9471ACFFABFFF40E7EFBCE3735FBFC" author="Birket-Smith, J." box="[870,1102,1122,1149]" pageId="2" pageNumber="418" pagination="1 - 161" refId="ref6365" refString="Birket-Smith, J. (1965) A revision of the West African Eilemic moths, based on the male genitalia (Lep., Arctiidae, Lithosinae; incl. gen. Crocosia, Eilema, Lithosia, Pelosia, Phryganopsis a. o.). Haile Sellassie I University. Papers from the Faculty of science, Series C (Zoology), 1, 1 - 161." type="journal article" year="1965">Birket-Smith (1965)</bibRefCitation>
|
||
; see also
|
||
<bibRefCitation id="EF9471ACFFABFFF409AFFBCE362BFBFC" author="Tuxen, S. L." box="[1207,1360,1123,1149]" pageId="2" pageNumber="418" refId="ref7922" refString="Tuxen, S. L. (1970) Taxonomist's Glossary of Genitalia in Insects. 2 nd Edition. Munksgaard, Copenhagen, 359 pp." type="book" year="1970">Tuxen (1970)</bibRefCitation>
|
||
on the neuter grammatical gender of this noun) on the vesica, while there are granicula and a large thorn-like cornutus in
|
||
<taxonomicName id="4C0577DEFFABFFF40D8FFB06327FFB44" baseAuthorityName="Wallengren" baseAuthorityYear="1860" box="[151,260,1195,1221]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B971D04FFFABFFF40D8FFB06327FFB44" box="[151,260,1195,1221]" italics="true" pageId="2" pageNumber="418">flavicosta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0577DEFFABFFF40C22FB0632E7FB44" baseAuthorityName="Kiriakoff" baseAuthorityYear="1958" box="[314,412,1195,1221]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="tinaeella">
|
||
<emphasis id="B971D04FFFABFFF40C22FB0632E7FB44" box="[314,412,1195,1221]" italics="true" pageId="2" pageNumber="418">tinaeella</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F5FD6FFABFFF50DDFFB6336EFFD70" lastPageId="3" lastPageNumber="419" pageId="2" pageNumber="418" type="description">
|
||
<paragraph id="8BBA0C5DFFABFFF40DDFFB63305DFB68" blockId="2.[151,1437,366,2013]" box="[199,806,1230,1257]" pageId="2" pageNumber="418">
|
||
<emphasis id="B971D04FFFABFFF40DDFFB63322AFB69" bold="true" box="[199,337,1230,1256]" pageId="2" pageNumber="418">Description</emphasis>
|
||
. Wingspan
|
||
<specimenCount id="9D03C7D4FFABFFF40CCFFB623297FB68" box="[471,492,1231,1257]" pageId="2" pageNumber="418" type="male">♂</specimenCount>
|
||
<quantity id="4CFDA1B8FFABFFF40CEBFB6331FFFB68" box="[499,644,1230,1257]" metricMagnitude="-2" metricUnit="m" metricValue="1.825" metricValueMax="1.9" metricValueMin="1.75" pageId="2" pageNumber="418" unit="mm" value="18.25" valueMax="19.0" valueMin="17.5">17.5–19 mm</quantity>
|
||
;
|
||
<specimenCount id="9D03C7D4FFABFFF40F97FB6231DFFB68" box="[655,676,1231,1257]" pageId="2" pageNumber="418" type="female">♀</specimenCount>
|
||
<quantity id="4CFDA1B8FFABFFF40FB3FB623059FB68" box="[683,802,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="2" pageNumber="418" unit="mm" value="20.0" valueMax="21.0" valueMin="19.0">19-21 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBA0C5DFFABFFF40DDFFB5E3167FAF8" blockId="2.[151,1437,366,2013]" pageId="2" pageNumber="418">
|
||
Head frons and vertex, notum including patagia and tegulae, and abdominal tergites all same colour as forewing upperside or slightly darker; clypeus always darker; antennae ciliated (cilia longer in the male,
|
||
<figureCitation id="133E10D8FFABFFF409E5FABA36F5FAB0" box="[1277,1422,1303,1329]" captionStart="FIGURE 9" captionStartId="5.[151,250,1889,1913]" captionTargetBox="[185,1397,197,1850]" captionTargetId="figure-27@5.[171,1416,181,1865]" captionTargetPageId="5" captionText="FIGURE 9. Pusiola unicolor n. sp.—A: central segment of the antennal flagellum, male; B: central segment of the antennal flagellum, female; C: dorsal pheromone gland; D: acrosternite; E: forewing venation; F: hindwing venation. A: anal vein; co: coremata; CuA: Anterior cubital vein; dp: dome-shaped projections; M: median vein; pg: dorsal pheromone glands; R: radial vein; Rs: radial sector vein; Sc: subcostal vein; VIII st: eighth sternum." figureDoi="http://doi.org/10.5281/zenodo.5848285" httpUri="https://zenodo.org/record/5848285/files/figure.png" pageId="2" pageNumber="418">Figs. 9A, 9B</figureCitation>
|
||
), dorsally same colour as the vertex, ventrally unpigmented; labial palpi porrect, not extending beyond the frons, as dark as the clypeus or even darker.
|
||
</paragraph>
|
||
<paragraph id="8BBA0C5DFFABFFF40DDFFA2E3253F9D0" blockId="2.[151,1437,366,2013]" pageId="2" pageNumber="418">Thorax. Thoracic sternites as dark as the forewings underside; legs same colour, with the dorsal face of the first two pairs darker; spurs formula 0-2-4. Forewing upperside ground colour grey-brown, with a faint, very small dark vaguely triangular mark half way along the costa; a few specimens slightly darkening towards the termen. Forewing underside slightly darker than the upperside, anal area paler. Hindwing upperside slightly darker than or same colour as the forewing upperside; costal area sometimes paler. Hindwing underside same colour as the upperside. Fringes concolorous.</paragraph>
|
||
<paragraph id="8BBA0C5DFFABFFF40DDFF9F6315AF8A8" blockId="2.[151,1437,366,2013]" pageId="2" pageNumber="418">
|
||
Abdomen. Abdominal sternites as dark as thoracic sternites; abdominal scales slightly darker along the pleurae; anal tuft paler; male with the eighth abdominal segment very short, about ¼ the length of the seventh, anteriorly forming shallow double-dimple-like coremata (
|
||
<figureCitation id="133E10D8FFABFFF40FB4F90E307DF93C" box="[684,774,1699,1725]" captionStart="FIGURE 9" captionStartId="5.[151,250,1889,1913]" captionTargetBox="[185,1397,197,1850]" captionTargetId="figure-27@5.[171,1416,181,1865]" captionTargetPageId="5" captionText="FIGURE 9. Pusiola unicolor n. sp.—A: central segment of the antennal flagellum, male; B: central segment of the antennal flagellum, female; C: dorsal pheromone gland; D: acrosternite; E: forewing venation; F: hindwing venation. A: anal vein; co: coremata; CuA: Anterior cubital vein; dp: dome-shaped projections; M: median vein; pg: dorsal pheromone glands; R: radial vein; Rs: radial sector vein; Sc: subcostal vein; VIII st: eighth sternum." figureDoi="http://doi.org/10.5281/zenodo.5848285" httpUri="https://zenodo.org/record/5848285/files/figure.png" pageId="2" pageNumber="418">Fig. 9D</figureCitation>
|
||
), and latero-posteriorly two dome-shaped projections (
|
||
<figureCitation id="133E10D8FFABFFF40877F90E33C6F960" captionStart="FIGURE 9" captionStartId="5.[151,250,1889,1913]" captionTargetBox="[185,1397,197,1850]" captionTargetId="figure-27@5.[171,1416,181,1865]" captionTargetPageId="5" captionText="FIGURE 9. Pusiola unicolor n. sp.—A: central segment of the antennal flagellum, male; B: central segment of the antennal flagellum, female; C: dorsal pheromone gland; D: acrosternite; E: forewing venation; F: hindwing venation. A: anal vein; co: coremata; CuA: Anterior cubital vein; dp: dome-shaped projections; M: median vein; pg: dorsal pheromone glands; R: radial vein; Rs: radial sector vein; Sc: subcostal vein; VIII st: eighth sternum." figureDoi="http://doi.org/10.5281/zenodo.5848285" httpUri="https://zenodo.org/record/5848285/files/figure.png" pageId="2" pageNumber="418">Fig. 9D</figureCitation>
|
||
); female with seventh abdominal segment slightly sclerotized, one and a half times longer than the sixth (
|
||
<figureCitation id="133E10D8FFABFFF40877F96A33BCF884" captionStart="FIGURE 10" captionStartId="6.[151,250,1756,1780]" captionTargetBox="[231,1357,193,1720]" captionTargetId="figure-21@6.[231,1357,181,1732]" captionTargetPageId="6" captionText="FIGURE 10. Pusiola unicolor n. sp.—A–D: male genitalia. A: processus distalis plicae, enlarged; B: genitalia, aedeagus removed; C: aedeagus, vesica everted; D: granicula of the vesica, enlarged. E–H: female genitalia. E: genitalia; F: enlarged spicule of the antrum; G: enlarged spicule of the foremost part of the corpus bursae; H: enlarged spicule of the medio-caudal part of the corpus bursae. aa: anterior apophyses; ab: appendix bursae; ae: aedeagus; an: antrum; ap: posterior apophyses; av: ala valvae; cb: corpus bursae; cp: coecum penis; dl: distal lobus; fs: dorsal flat sclerotization of the aedeagus; gr: granicula; ju: juxta; pa: papillae anales; pav: distal process of the ala valvae; pd: processus distalis plicae; pe: pedunculi; pg: dorsal pheromone glands; sa: saccus; sv: supravalva; te: tegumen; un: uncus; ve: vesica; vi: vinculum; VII st: seventh sternum; VIII st: eighth sternum; vl: ventral lobus." figureDoi="http://doi.org/10.5281/zenodo.5848287" httpUri="https://zenodo.org/record/5848287/files/figure.png" pageId="2" pageNumber="418">Fig. 10E</figureCitation>
|
||
), and eighth abdominal segment the same length as the sixth, slightly sclerotized, with a V-shaped membranous notch in the middle of the sternum.
|
||
</paragraph>
|
||
<paragraph id="8BBA0C5DFFABFFF50DDFF89F3644FEC0" blockId="2.[151,1437,366,2013]" lastBlockId="3.[151,1437,151,1653]" lastPageId="3" lastPageNumber="419" pageId="2" pageNumber="418">
|
||
<emphasis id="B971D04FFFABFFF40DDFF89F3246F8CD" bold="true" box="[199,317,1842,1868]" pageId="2" pageNumber="418">Genitalia.</emphasis>
|
||
<specimenCount id="9D03C7D4FFABFFF40C5DF89E3226F8CC" box="[325,349,1843,1869]" pageId="2" pageNumber="418" type="male">♂</specimenCount>
|
||
. Uncus pointed, long and slender, bent slightly backwards; tegumen pear-shaped in dorsal view, with fairly large pedunculi (sensu
|
||
<bibRefCitation id="EF9471ACFFABFFF40F3AF8FA31A1F8F0" author="Diakonoff, A." box="[546,730,1879,1905]" pageId="2" pageNumber="418" pagination="111 - 240" refId="ref6582" refString="Diakonoff, A. (1939) The genera of Indo-Malayan and Papuan Tortricidae. Zoologische Mededeelingen Rijksmuseum van Natuurlijke Historie te Leiden, 21, 111 - 240." type="journal article" year="1939">Diakonoff 1939</bibRefCitation>
|
||
); vinculum slender, U-shaped, but with the saccus notched at the ventro-cephalic end; juxta protruding caudally, more or less axe- or L-shaped with the handle pointed, dorsally directed; valvellae appearing as weak, not clearly delimitated sclerotizations, ventro-laterally with respect to the juxta. Valva slender compared to the typically stouter
|
||
<taxonomicName id="4C0577DEFFABFFF40FF2F86E303BF85C" authorityName="Wallengren" authorityYear="1863" box="[746,832,1987,2013]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971D04FFFABFFF40FF2F86E303BF85C" box="[746,832,1987,2013]" italics="true" pageId="2" pageNumber="418">Pusiola</emphasis>
|
||
</taxonomicName>
|
||
valvae (e.g.
|
||
<taxonomicName id="4C0577DEFFABFFF40ED4F86E3728F85C" baseAuthorityName="Wallengren" baseAuthorityYear="1860" box="[972,1107,1987,2013]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="418" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B971D04FFFABFFF40ED4F86E3728F85C" box="[972,1107,1987,2013]" italics="true" pageId="2" pageNumber="418">P. flavicosta</emphasis>
|
||
</taxonomicName>
|
||
); distal half of the supravalva flap-like, sub-oval, ending proximally with a curved processus distalis plicae (
|
||
<figureCitation id="133E10D8FFAAFFF5090DFF3A3704FF30" box="[1045,1151,151,177]" captionStart="FIGURE 10" captionStartId="6.[151,250,1756,1780]" captionTargetBox="[231,1357,193,1720]" captionTargetId="figure-21@6.[231,1357,181,1732]" captionTargetPageId="6" captionText="FIGURE 10. Pusiola unicolor n. sp.—A–D: male genitalia. A: processus distalis plicae, enlarged; B: genitalia, aedeagus removed; C: aedeagus, vesica everted; D: granicula of the vesica, enlarged. E–H: female genitalia. E: genitalia; F: enlarged spicule of the antrum; G: enlarged spicule of the foremost part of the corpus bursae; H: enlarged spicule of the medio-caudal part of the corpus bursae. aa: anterior apophyses; ab: appendix bursae; ae: aedeagus; an: antrum; ap: posterior apophyses; av: ala valvae; cb: corpus bursae; cp: coecum penis; dl: distal lobus; fs: dorsal flat sclerotization of the aedeagus; gr: granicula; ju: juxta; pa: papillae anales; pav: distal process of the ala valvae; pd: processus distalis plicae; pe: pedunculi; pg: dorsal pheromone glands; sa: saccus; sv: supravalva; te: tegumen; un: uncus; ve: vesica; vi: vinculum; VII st: seventh sternum; VIII st: eighth sternum; vl: ventral lobus." figureDoi="http://doi.org/10.5281/zenodo.5848287" httpUri="https://zenodo.org/record/5848287/files/figure.png" pageId="3" pageNumber="419">Fig. 10A</figureCitation>
|
||
); ala valvae with slender proximal portion (sacculus) and pointed distal process, long, slender and strongly sclerified, imperceptibly distally exceeding the supravalva. Aedeagus about three times longer than wide, tubular, with an undelimited flat sclerotization at the dorsal distal end, coecum penis sub-conical, ductus seminalis inserted dorsally, vesica with globular general appearance, and a distal and a ventral lobus covered in oval-shaped scobinate granicula (
|
||
<figureCitation id="133E10D8FFAAFFF509C9FE8A3648FEC0" box="[1233,1331,295,321]" captionStart="FIGURE 10" captionStartId="6.[151,250,1756,1780]" captionTargetBox="[231,1357,193,1720]" captionTargetId="figure-21@6.[231,1357,181,1732]" captionTargetPageId="6" captionText="FIGURE 10. Pusiola unicolor n. sp.—A–D: male genitalia. A: processus distalis plicae, enlarged; B: genitalia, aedeagus removed; C: aedeagus, vesica everted; D: granicula of the vesica, enlarged. E–H: female genitalia. E: genitalia; F: enlarged spicule of the antrum; G: enlarged spicule of the foremost part of the corpus bursae; H: enlarged spicule of the medio-caudal part of the corpus bursae. aa: anterior apophyses; ab: appendix bursae; ae: aedeagus; an: antrum; ap: posterior apophyses; av: ala valvae; cb: corpus bursae; cp: coecum penis; dl: distal lobus; fs: dorsal flat sclerotization of the aedeagus; gr: granicula; ju: juxta; pa: papillae anales; pav: distal process of the ala valvae; pd: processus distalis plicae; pe: pedunculi; pg: dorsal pheromone glands; sa: saccus; sv: supravalva; te: tegumen; un: uncus; ve: vesica; vi: vinculum; VII st: seventh sternum; VIII st: eighth sternum; vl: ventral lobus." figureDoi="http://doi.org/10.5281/zenodo.5848287" httpUri="https://zenodo.org/record/5848287/files/figure.png" pageId="3" pageNumber="419">Fig 10D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBA0C5DFFAAFFF50DDFFEE63290FD98" blockId="3.[151,1437,151,1653]" pageId="3" pageNumber="419">
|
||
Female. Caudal ventral margin of the seventh abdominal segment curly and dense (lamella antevaginalis,
|
||
<figureCitation id="133E10D8FFAAFFF50868FEE633B3FE08" captionStart="FIGURE 10" captionStartId="6.[151,250,1756,1780]" captionTargetBox="[231,1357,193,1720]" captionTargetId="figure-21@6.[231,1357,181,1732]" captionTargetPageId="6" captionText="FIGURE 10. Pusiola unicolor n. sp.—A–D: male genitalia. A: processus distalis plicae, enlarged; B: genitalia, aedeagus removed; C: aedeagus, vesica everted; D: granicula of the vesica, enlarged. E–H: female genitalia. E: genitalia; F: enlarged spicule of the antrum; G: enlarged spicule of the foremost part of the corpus bursae; H: enlarged spicule of the medio-caudal part of the corpus bursae. aa: anterior apophyses; ab: appendix bursae; ae: aedeagus; an: antrum; ap: posterior apophyses; av: ala valvae; cb: corpus bursae; cp: coecum penis; dl: distal lobus; fs: dorsal flat sclerotization of the aedeagus; gr: granicula; ju: juxta; pa: papillae anales; pav: distal process of the ala valvae; pd: processus distalis plicae; pe: pedunculi; pg: dorsal pheromone glands; sa: saccus; sv: supravalva; te: tegumen; un: uncus; ve: vesica; vi: vinculum; VII st: seventh sternum; VIII st: eighth sternum; vl: ventral lobus." figureDoi="http://doi.org/10.5281/zenodo.5848287" httpUri="https://zenodo.org/record/5848287/files/figure.png" pageId="3" pageNumber="419">Fig. 10E</figureCitation>
|
||
); eighth segment as long as the sixth, slightly sclerotized, with no lamella postvaginalis; papillae anales gently tapering caudally; posterior apophyses slightly longer and more slender than the anterior ones; anterior apophyses somewhat longer than the eighth segment; dorsal pheromone glands unpaired, with one caudal opening at the intersegmental membrane between A8 and A9, anteriorly bifurcated into two fairly narrow lobes not exceeding the posterior apophyses (
|
||
<figureCitation id="133E10D8FFAAFFF50C9EFE5232A5FD98" box="[390,478,511,537]" captionStart="FIGURE 9" captionStartId="5.[151,250,1889,1913]" captionTargetBox="[185,1397,197,1850]" captionTargetId="figure-27@5.[171,1416,181,1865]" captionTargetPageId="5" captionText="FIGURE 9. Pusiola unicolor n. sp.—A: central segment of the antennal flagellum, male; B: central segment of the antennal flagellum, female; C: dorsal pheromone gland; D: acrosternite; E: forewing venation; F: hindwing venation. A: anal vein; co: coremata; CuA: Anterior cubital vein; dp: dome-shaped projections; M: median vein; pg: dorsal pheromone glands; R: radial vein; Rs: radial sector vein; Sc: subcostal vein; VIII st: eighth sternum." figureDoi="http://doi.org/10.5281/zenodo.5848285" httpUri="https://zenodo.org/record/5848285/files/figure.png" pageId="3" pageNumber="419">Fig. 9C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BBA0C5DFFAAFFF50DDFFD8E36EFFD70" blockId="3.[151,1437,151,1653]" pageId="3" pageNumber="419">
|
||
Ostium bursae as an inconspicuous opening medially in the intersegmental membrane between A7 and A8; antrum thick and short, truncated cone-shaped, with many thin spicules in the wall (
|
||
<figureCitation id="133E10D8FFAAFFF50925FDEA37E2FDE0" box="[1085,1177,583,609]" captionStart="FIGURE 10" captionStartId="6.[151,250,1756,1780]" captionTargetBox="[231,1357,193,1720]" captionTargetId="figure-21@6.[231,1357,181,1732]" captionTargetPageId="6" captionText="FIGURE 10. Pusiola unicolor n. sp.—A–D: male genitalia. A: processus distalis plicae, enlarged; B: genitalia, aedeagus removed; C: aedeagus, vesica everted; D: granicula of the vesica, enlarged. E–H: female genitalia. E: genitalia; F: enlarged spicule of the antrum; G: enlarged spicule of the foremost part of the corpus bursae; H: enlarged spicule of the medio-caudal part of the corpus bursae. aa: anterior apophyses; ab: appendix bursae; ae: aedeagus; an: antrum; ap: posterior apophyses; av: ala valvae; cb: corpus bursae; cp: coecum penis; dl: distal lobus; fs: dorsal flat sclerotization of the aedeagus; gr: granicula; ju: juxta; pa: papillae anales; pav: distal process of the ala valvae; pd: processus distalis plicae; pe: pedunculi; pg: dorsal pheromone glands; sa: saccus; sv: supravalva; te: tegumen; un: uncus; ve: vesica; vi: vinculum; VII st: seventh sternum; VIII st: eighth sternum; vl: ventral lobus." figureDoi="http://doi.org/10.5281/zenodo.5848287" httpUri="https://zenodo.org/record/5848287/files/figure.png" pageId="3" pageNumber="419">Fig 10F</figureCitation>
|
||
); ductus bursae absent; corpus bursae oval, membranous, with a dense scobination of thorn-like spicules in the tissue of the wall, thicker than those of the antrum. Thorn-like spicules are of two sizes: thinner and denser along the caudal 2/3 of the corpus (
|
||
<figureCitation id="133E10D8FFAAFFF50D87FD1E327BFD4C" box="[159,256,691,717]" captionStart="FIGURE 10" captionStartId="6.[151,250,1756,1780]" captionTargetBox="[231,1357,193,1720]" captionTargetId="figure-21@6.[231,1357,181,1732]" captionTargetPageId="6" captionText="FIGURE 10. Pusiola unicolor n. sp.—A–D: male genitalia. A: processus distalis plicae, enlarged; B: genitalia, aedeagus removed; C: aedeagus, vesica everted; D: granicula of the vesica, enlarged. E–H: female genitalia. E: genitalia; F: enlarged spicule of the antrum; G: enlarged spicule of the foremost part of the corpus bursae; H: enlarged spicule of the medio-caudal part of the corpus bursae. aa: anterior apophyses; ab: appendix bursae; ae: aedeagus; an: antrum; ap: posterior apophyses; av: ala valvae; cb: corpus bursae; cp: coecum penis; dl: distal lobus; fs: dorsal flat sclerotization of the aedeagus; gr: granicula; ju: juxta; pa: papillae anales; pav: distal process of the ala valvae; pd: processus distalis plicae; pe: pedunculi; pg: dorsal pheromone glands; sa: saccus; sv: supravalva; te: tegumen; un: uncus; ve: vesica; vi: vinculum; VII st: seventh sternum; VIII st: eighth sternum; vl: ventral lobus." figureDoi="http://doi.org/10.5281/zenodo.5848287" httpUri="https://zenodo.org/record/5848287/files/figure.png" pageId="3" pageNumber="419">Fig 10H</figureCitation>
|
||
), stronger and more dispersed along the anterior 1/3 (
|
||
<figureCitation id="133E10D8FFAAFFF50E4EFD1E30C5FD4C" box="[854,958,691,717]" captionStart="FIGURE 10" captionStartId="6.[151,250,1756,1780]" captionTargetBox="[231,1357,193,1720]" captionTargetId="figure-21@6.[231,1357,181,1732]" captionTargetPageId="6" captionText="FIGURE 10. Pusiola unicolor n. sp.—A–D: male genitalia. A: processus distalis plicae, enlarged; B: genitalia, aedeagus removed; C: aedeagus, vesica everted; D: granicula of the vesica, enlarged. E–H: female genitalia. E: genitalia; F: enlarged spicule of the antrum; G: enlarged spicule of the foremost part of the corpus bursae; H: enlarged spicule of the medio-caudal part of the corpus bursae. aa: anterior apophyses; ab: appendix bursae; ae: aedeagus; an: antrum; ap: posterior apophyses; av: ala valvae; cb: corpus bursae; cp: coecum penis; dl: distal lobus; fs: dorsal flat sclerotization of the aedeagus; gr: granicula; ju: juxta; pa: papillae anales; pav: distal process of the ala valvae; pd: processus distalis plicae; pe: pedunculi; pg: dorsal pheromone glands; sa: saccus; sv: supravalva; te: tegumen; un: uncus; ve: vesica; vi: vinculum; VII st: seventh sternum; VIII st: eighth sternum; vl: ventral lobus." figureDoi="http://doi.org/10.5281/zenodo.5848287" httpUri="https://zenodo.org/record/5848287/files/figure.png" pageId="3" pageNumber="419">Fig. 10G</figureCitation>
|
||
); no signa; appendix bursae conical, membranous, caudally with respect to the corpus bursae, the ductus seminalis initiating at the posterior part (
|
||
<figureCitation id="133E10D8FFAAFFF5083AFD7A36FCFD70" box="[1314,1415,727,753]" captionStart="FIGURE 10" captionStartId="6.[151,250,1756,1780]" captionTargetBox="[231,1357,193,1720]" captionTargetId="figure-21@6.[231,1357,181,1732]" captionTargetPageId="6" captionText="FIGURE 10. Pusiola unicolor n. sp.—A–D: male genitalia. A: processus distalis plicae, enlarged; B: genitalia, aedeagus removed; C: aedeagus, vesica everted; D: granicula of the vesica, enlarged. E–H: female genitalia. E: genitalia; F: enlarged spicule of the antrum; G: enlarged spicule of the foremost part of the corpus bursae; H: enlarged spicule of the medio-caudal part of the corpus bursae. aa: anterior apophyses; ab: appendix bursae; ae: aedeagus; an: antrum; ap: posterior apophyses; av: ala valvae; cb: corpus bursae; cp: coecum penis; dl: distal lobus; fs: dorsal flat sclerotization of the aedeagus; gr: granicula; ju: juxta; pa: papillae anales; pav: distal process of the ala valvae; pd: processus distalis plicae; pe: pedunculi; pg: dorsal pheromone glands; sa: saccus; sv: supravalva; te: tegumen; un: uncus; ve: vesica; vi: vinculum; VII st: seventh sternum; VIII st: eighth sternum; vl: ventral lobus." figureDoi="http://doi.org/10.5281/zenodo.5848287" httpUri="https://zenodo.org/record/5848287/files/figure.png" pageId="3" pageNumber="419">Fig. 10E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F5FD6FFAAFFF50DDFFD5632FBFCB8" pageId="3" pageNumber="419" type="etymology">
|
||
<paragraph id="8BBA0C5DFFAAFFF50DDFFD5632FBFCB8" blockId="3.[151,1437,151,1653]" pageId="3" pageNumber="419">
|
||
<emphasis id="B971D04FFFAAFFF50DDFFD563233FC94" bold="true" box="[199,328,763,789]" pageId="3" pageNumber="419">Etymology</emphasis>
|
||
. The name
|
||
<taxonomicName id="4C0577DEFFAAFFF50CCBFD563149FC94" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[467,562,763,789]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="unicolor">
|
||
<emphasis id="B971D04FFFAAFFF50CCBFD563149FC94" box="[467,562,763,789]" italics="true" pageId="3" pageNumber="419">unicolor</emphasis>
|
||
</taxonomicName>
|
||
(meaning “of the same colour”) refers to the usually identical colour of the fore- and hindwings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31F5FD6FFAAFFF50DDFFCEE37C3F9F4" pageId="3" pageNumber="419" type="discussion">
|
||
<paragraph id="8BBA0C5DFFAAFFF50DDFFCEE37F6F9AC" blockId="3.[151,1437,151,1653]" pageId="3" pageNumber="419">
|
||
<emphasis id="B971D04FFFAAFFF50DDFFCEE324FFCDC" bold="true" box="[199,308,835,861]" pageId="3" pageNumber="419">Remarks</emphasis>
|
||
. In this species, the morphology of the male genitalia diverges in some way from the typical pattern of the genus (i.e. from the
|
||
<typeStatus id="54BEB2FFFFAAFFF50CD9FCCA328AFC00" box="[449,497,871,897]" pageId="3" pageNumber="419">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C0577DEFFAAFFF50F48FCCA31A2FC00" baseAuthorityName="Wallengren" baseAuthorityYear="1860" box="[592,729,871,897]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B971D04FFFAAFFF50F48FCCA31A2FC00" box="[592,729,871,897]" italics="true" pageId="3" pageNumber="419">P. flavicosta</emphasis>
|
||
</taxonomicName>
|
||
): in the new species presented above the uncus is slender, long and almost straight, whereas in most of
|
||
<taxonomicName id="4C0577DEFFAAFFF50F46FC263076FC24" box="[606,781,907,933]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B971D04FFFAAFFF50F46FC2631CFFC24" box="[606,692,907,933]" italics="true" pageId="3" pageNumber="419">Pusiola</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
it is stouter (i.e. shorter and less slender), and is caudally curved; the valvae are proportionally longer; the tegumen is twice as long as the vinculum (in other
|
||
<taxonomicName id="4C0577DEFFAAFFF509F1FC0236E3FC48" box="[1257,1432,943,969]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B971D04FFFAAFFF509F1FC023644FC48" box="[1257,1343,943,969]" italics="true" pageId="3" pageNumber="419">Pusiola</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, valvae are nearly of the same length); the juxta is projecting sharply backwards. However, other characters allow us to at least tentatively include this species in the genus
|
||
<taxonomicName id="4C0577DEFFAAFFF50E00FC5A3015FB90" authorityName="Wallengren" authorityYear="1863" box="[792,878,1015,1041]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971D04FFFAAFFF50E00FC5A3015FB90" box="[792,878,1015,1041]" italics="true" pageId="3" pageNumber="419">Pusiola</emphasis>
|
||
</taxonomicName>
|
||
, such as the overall size of the adult; shape of the wings (slightly broader than other
|
||
<taxonomicName id="4C0577DEFFAAFFF50F0CFBB631FDFBB4" authorityName="Birket-Smith" authorityYear="1965" box="[532,646,1051,1077]" class="Insecta" family="Erebidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="subTribe" subTribe="Lithosiina">Lithosiina</taxonomicName>
|
||
genera); the general structure of the genitalia (curved process of the ala valvae, presence of the processus distalis plicae, notched saccus); and the presence of a subapical spur at the distal end of the aedeagus (in
|
||
<taxonomicName id="4C0577DEFFAAFFF50CBBFBCE3165FBFC" authority="Durante & Apinda-Legnouo, 2022" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[419,542,1123,1149]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="unicolor" status="sp. nov.">
|
||
<emphasis id="B971D04FFFAAFFF50CBBFBCE3165FBFC" box="[419,542,1123,1149]" italics="true" pageId="3" pageNumber="419">P. unicolor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B971D04FFFAAFFF50F3DFBCE311FFBFC" bold="true" box="[549,612,1123,1149]" pageId="3" pageNumber="419">
|
||
<taxonomicNameLabel id="A2426D34FFAAFFF50F3DFBCE311FFBFC" box="[549,612,1123,1149]" pageId="3" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the subapical spur has the shape of a flat apical sclerotization).The wing venation fits Hampson’s description (1900) for the genus, with the only exception being the apical veins in the forewing: in
|
||
<taxonomicName id="4C0577DEFFAAFFF50DE2FB06321CFB44" baseAuthorityName="Wallengren" baseAuthorityYear="1860" box="[250,359,1195,1221]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="flavicosta">
|
||
<emphasis id="B971D04FFFAAFFF50DE2FB06321CFB44" box="[250,359,1195,1221]" italics="true" pageId="3" pageNumber="419">flavicosta</emphasis>
|
||
</taxonomicName>
|
||
is [Rs2(Rs3, Rs4)]M1, whereas in
|
||
<taxonomicName id="4C0577DEFFAAFFF50FF7FB063035FB44" authority="Durante & Apinda-Legnouo, 2022" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[751,846,1195,1221]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="unicolor" status="sp. nov.">
|
||
<emphasis id="B971D04FFFAAFFF50FF7FB063035FB44" box="[751,846,1195,1221]" italics="true" pageId="3" pageNumber="419">unicolor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B971D04FFFAAFFF50E4DFB0630E8FB44" bold="true" box="[853,915,1195,1221]" pageId="3" pageNumber="419">
|
||
<taxonomicNameLabel id="A2426D34FFAAFFF50E4DFB0630E8FB44" box="[853,915,1195,1221]" pageId="3" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
it is [Rs2, Rs3) Rs4]M1 (
|
||
<figureCitation id="133E10D8FFAAFFF509AAFB063673FB44" box="[1202,1288,1195,1221]" captionStart="FIGURE 9" captionStartId="5.[151,250,1889,1913]" captionTargetBox="[185,1397,197,1850]" captionTargetId="figure-27@5.[171,1416,181,1865]" captionTargetPageId="5" captionText="FIGURE 9. Pusiola unicolor n. sp.—A: central segment of the antennal flagellum, male; B: central segment of the antennal flagellum, female; C: dorsal pheromone gland; D: acrosternite; E: forewing venation; F: hindwing venation. A: anal vein; co: coremata; CuA: Anterior cubital vein; dp: dome-shaped projections; M: median vein; pg: dorsal pheromone glands; R: radial vein; Rs: radial sector vein; Sc: subcostal vein; VIII st: eighth sternum." figureDoi="http://doi.org/10.5281/zenodo.5848285" httpUri="https://zenodo.org/record/5848285/files/figure.png" pageId="3" pageNumber="419">Fig. 9E</figureCitation>
|
||
). In addition, it should be remarked that the new species does not fit well within any other genus in Africa (
|
||
<bibRefCitation id="EF9471ACFFAAFFF509DFFB6236ECFB68" author="De Toulgoet, H." box="[1223,1431,1231,1257]" pageId="3" pageNumber="419" pagination="69 - 108" refId="ref6547" refString="De Toulgoet, H. (1984) Liste recapitulative des Lepidopteres Arctiidae et Nolidae de Madagascar et de l'Archipel des Comores. Miscelanea entomologica, 50 (3), 69 - 108." type="journal article" year="1984">De Toulgoët 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9471ACFFAAFFF50D8FFB5E3252FA8C" author="Kruger, M." box="[151,297,1266,1293]" pageId="3" pageNumber="419" pagination="1 - 176" refId="ref7647" refString="Kruger, M. (2015) Generic classification of Afrotropical footman moths sensu stricto (Lepidoptera: Erebidae: Arctiinae: Lithosiini (partim )). Transvaal Museum Monograph, 15, 1 - 176." type="journal article" year="2015">Krüger 2015</bibRefCitation>
|
||
), or even in South-East Asia. Holloway (2001) for instance, indicates the cases of similarity, e.g.
|
||
<taxonomicName id="4C0577DEFFAAFFF50879FB5E3296FAB3" authority="(Walker, 1862)" baseAuthorityName="Walker" baseAuthorityYear="1862" class="Insecta" family="Arctiidae" genus="Lambula" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="fuliginosa">
|
||
<emphasis id="B971D04FFFAAFFF50879FB5E323BFAB0" italics="true" pageId="3" pageNumber="419">Lambula fuliginosa</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9471ACFFAAFFF50C55FABA329EFAB3" author="Walker, F." box="[333,485,1303,1330]" pageId="3" pageNumber="419" pagination="82 - 145" refId="ref7987" refString="Walker, F. (1862) Catalogue of the Heterocerous Lapidopterous Insects collected at Sarawak, in Borneo, by Mr A. R. Wallace, with Descriptions of New Species. Journal of the Proceeding of the Linnean Society of London, Zoology, 6, 82 - 145. https: // doi. org / 10.1111 / j. 1096 - 3642.1862. tb 00935. x" type="journal article" year="1862">Walker, 1862</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C0577DEFFAAFFF50F34FABA30B0FAB3" authority="(Walker, 1862)" baseAuthorityName="Walker" baseAuthorityYear="1862" box="[556,971,1303,1330]" class="Insecta" family="Arctiidae" genus="Euconosia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="xylinoides">
|
||
<emphasis id="B971D04FFFAAFFF50F34FABA3060FAB0" box="[556,795,1303,1329]" italics="true" pageId="3" pageNumber="419">Euconosia xylinoides</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9471ACFFAAFFF50E30FABA30B8FAB3" author="Walker, F." box="[808,963,1303,1330]" pageId="3" pageNumber="419" pagination="82 - 145" refId="ref7987" refString="Walker, F. (1862) Catalogue of the Heterocerous Lapidopterous Insects collected at Sarawak, in Borneo, by Mr A. R. Wallace, with Descriptions of New Species. Journal of the Proceeding of the Linnean Society of London, Zoology, 6, 82 - 145. https: // doi. org / 10.1111 / j. 1096 - 3642.1862. tb 00935. x" type="journal article" year="1862">Walker, 1862</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Neither of both species are considered to be related to
|
||
<taxonomicName id="4C0577DEFFAAFFF50C31FA9632F3FAD4" authority="Durante & Apinda-Legnouo, 2022" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[297,392,1339,1365]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="unicolor" status="sp. nov.">
|
||
<emphasis id="B971D04FFFAAFFF50C31FA9632F3FAD4" box="[297,392,1339,1365]" italics="true" pageId="3" pageNumber="419">unicolor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B971D04FFFAAFFF50C97FA9632B5FAD4" bold="true" box="[399,462,1339,1365]" pageId="3" pageNumber="419">
|
||
<taxonomicNameLabel id="A2426D34FFAAFFF50C97FA9632B5FAD4" box="[399,462,1339,1365]" pageId="3" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Despite a similar uncus and bilobed saccus in all three species mentioned above, there are significant diagnostic differences: males of
|
||
<taxonomicName id="4C0577DEFFAAFFF50FB0FAF230F9FAF8" baseAuthorityName="Walker" baseAuthorityYear="1862" box="[680,898,1375,1401]" class="Insecta" family="Arctiidae" genus="Lambula" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="fuliginosa">
|
||
<emphasis id="B971D04FFFAAFFF50FB0FAF230F9FAF8" box="[680,898,1375,1401]" italics="true" pageId="3" pageNumber="419">Lambula fuliginosa</emphasis>
|
||
</taxonomicName>
|
||
possess i) a folding on the costa of forewing, ii) a small tuft of scales on costal area of the hindwing (Holloway 2001: 298), iii) a process on the ala valvae bending over the supravalva, iv) socii present, v) a slender aedeagus and a slender cornutus. Similarly, despite the similar uncus and bilobed saccus,
|
||
<taxonomicName id="4C0577DEFFAAFFF50CDBFA6631C8FA64" baseAuthorityName="Walker" baseAuthorityYear="1862" box="[451,691,1483,1509]" class="Insecta" family="Arctiidae" genus="Euconosia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="xylinoides">
|
||
<emphasis id="B971D04FFFAAFFF50CDBFA6631C8FA64" box="[451,691,1483,1509]" italics="true" pageId="3" pageNumber="419">Euconosia xylinoides</emphasis>
|
||
</taxonomicName>
|
||
differs by: i) differently-shaped forewings, ii) distal margin of supravalva with dense hairs, iii) stout process on the ala valvae rises from a lower position than in
|
||
<taxonomicName id="4C0577DEFFAAFFF509DBFA423608F988" box="[1219,1395,1519,1545]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B971D04FFFAAFFF509DBFA423662F988" box="[1219,1305,1519,1545]" italics="true" pageId="3" pageNumber="419">Pusiola</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, iv) presence of slender aedeagus and vesica with at least one long lobus in
|
||
<taxonomicName id="4C0577DEFFAAFFF50EAFF9BE37F6F9AC" box="[951,1165,1555,1581]" class="Insecta" family="Arctiidae" genus="Euconosia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B971D04FFFAAFFF50EAFF9BE3756F9AD" box="[951,1069,1555,1580]" italics="true" pageId="3" pageNumber="419">Euconosia</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BBA0C5DFFAAFFF50DDFF99A37C3F9F4" blockId="3.[151,1437,151,1653]" pageId="3" pageNumber="419">
|
||
Ultimately, it should be stressed that the placement of
|
||
<taxonomicName id="4C0577DEFFAAFFF50E38F99A30E1F9D0" authority="Durante & Apinda-Legnouo, 2022" authorityName="Durante & Apinda-Legnouo" authorityYear="2022" box="[800,922,1591,1617]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="species" species="unicolor" status="sp. nov.">
|
||
<emphasis id="B971D04FFFAAFFF50E38F99A30E1F9D0" box="[800,922,1591,1617]" italics="true" pageId="3" pageNumber="419">P. unicolor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B971D04FFFAAFFF50EB8F99A30A5F9D0" bold="true" box="[928,990,1591,1617]" pageId="3" pageNumber="419">
|
||
<taxonomicNameLabel id="A2426D34FFAAFFF50EB8F99A30A5F9D0" box="[928,990,1591,1617]" pageId="3" pageNumber="419" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
into the genus
|
||
<taxonomicName id="4C0577DEFFAAFFF5099EF99A37A7F9D0" authorityName="Wallengren" authorityYear="1863" box="[1158,1244,1591,1617]" class="Insecta" family="Arctiidae" genus="Pusiola" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="419" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B971D04FFFAAFFF5099EF99A37A7F9D0" box="[1158,1244,1591,1617]" italics="true" pageId="3" pageNumber="419">Pusiola</emphasis>
|
||
</taxonomicName>
|
||
has to be considered somewhat doubtful at least until a robust and contemporary revision of the genus is done.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |