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<mods:title id="E082401E2BB9F0A96F3C16ADA1BF9D42">Taxonomic revision and morphological cladistics analysis of the diatom genus Anorthoneis (Cocconeidaceae), with description of Anorthoneis arthus-bertrandii sp. nov.</mods:title>
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<taxonomicName id="4C05C2A1FFCC8369FF6442E78723A89E" authority="Hustedt 1955: 15" authorityName="Hustedt" authorityPageNumber="15" authorityYear="1955" box="[136,586,158,185]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Anorthoneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="16" pageNumber="217" phylum="Ochrophyta" rank="species" species="hummii">
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<emphasis id="B9716530FFCC8369FF6442E78411A89E" bold="true" box="[136,376,158,185]" italics="true" pageId="16" pageNumber="217">Anorthoneis hummii</emphasis>
<bibRefCitation id="EF94C4D3FFCC8369FE9342D98723A89E" author="Hustedt, F." box="[383,586,160,185]" pageId="16" pageNumber="217" pagination="5 - 67" refId="ref18183" refString="Hustedt, F. (1955) Marine littoral diatoms of Beaufort, North Carolina. Duke University Marine Station Bulletin 6: 5 - 67." type="journal article" year="1955">Hustedt 1955: 15</bibRefCitation>
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(
<figureCitation id="133EA5A7FFCC8369FDB542D987B0A89E" box="[601,729,160,185]" captionStart-0="FIGURES 4956" captionStart-1="FIGURES 5762" captionStart-2="FIGURES 63-67" captionStartId-0="16.[136,243,1776,1797]" captionStartId-1="17.[136,243,1432,1453]" captionStartId-2="18.[136,243,1627,1648]" captionTargetBox-0="[225,1366,281,1751]" captionTargetBox-1="[162,1430,201,1399]" captionTargetBox-2="[162,1430,436,1596]" captionTargetId-0="figure-53@16.[221,1366,266,1751]" captionTargetId-1="figure-14@17.[151,1436,190,1407]" captionTargetId-2="figure-133@18.[151,1436,425,1602]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionText-0="FIGURES 4956. Historical reports of Anorthoneis hummii by different authors. (49) Raphe-sternum and sternum valve drawings of A. hummii from Hustedt (1955) (holotype); (50) Raphe-sternum valve micrographs of A. hummii from Witkowski et al. (2000); (51, 52) Micrographs of A. hummii specimen H65158 from slide ZT1/87 (Holotype), material AM791 from Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven,Germany); (53) Micrograph of A. hummii specimen H65433 from slide ZT1/96 (Paralectotype), material AM795 from Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany); (54, 55) Micrographs of A. hummii specimen H65252 from slide ZT1/92 (isolectotype), material AM792 from Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany); (56) Micrograph of A. hummii specimen H65217a from slide ZT1/90 (lectotype), material AM792 from Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany). Magnification: Fig. 52: x1500. Scale bars: Figs 5156 = 10 μm." captionText-1="FIGURES 5762. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. (57) External sternum valve with asymmetrical hyaline central area and small roundish terminal areas (arrow); (58) Detail of sternum valve mantle terminating in a wide hyaline marginal area; (59) Detail of roundish/transapically elongated areolae externally occluded by thick perforated hymenes; (60) Detail of sternum valve mantle; (61) Internal view of areolae occluded by perforated hymens; (62) Internal sternum valve showing sternum and hyaline central area. Scale bars: Fig. 57 = 10 μm; Figs 62, 63 = 5 μm; Figs 58, 60 = 2 μm; Figs 59, 61 = 1 μm." captionText-2="FIGURES 63-67. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. 63. External view of raphe-sternum valve. (64) Detail of coaxial raphe central endings in external view lying in drop-shaped areas; (65) External view of distal raphe ending terminating in hyaline submarginal area; (66) Tilted external view (60°) of raphe-sternum mantle showing the opened valvocopula with smooth inner margin; (67) Tilted external view (60°) of raphe-sternum valve. Scale bars: Fig. 57 = 10 μm, Figs 62, 63, 67 = 5 μm; Figs 58, 60, 65 = 2 μm; Figs 59, 61, 64, 66 = 1 μm." figureDoi-0="http://doi.org/10.5281/zenodo.13720130" figureDoi-1="http://doi.org/10.5281/zenodo.13720132" figureDoi-2="http://doi.org/10.5281/zenodo.13720134" httpUri-0="https://zenodo.org/record/13720130/files/figure.png" httpUri-1="https://zenodo.org/record/13720132/files/figure.png" httpUri-2="https://zenodo.org/record/13720134/files/figure.png" pageId="16" pageNumber="217">Figs 4967</figureCitation>
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<paragraph id="8BBAB922FFCC8369FF6442BF8403A8D8" blockId="16.[136,1452,198,255]" pageId="16" pageNumber="217">
<emphasis id="B9716530FFCC8369FF6442BF8471A8FC" bold="true" box="[136,280,198,219]" pageId="16" pageNumber="217">References:—</emphasis>
<bibRefCitation id="EF94C4D3FFCC8369FEF442BF84C3A8FC" author="Hustedt, F." box="[280,426,198,219]" pageId="16" pageNumber="217" pagination="5 - 67" refId="ref18183" refString="Hustedt, F. (1955) Marine littoral diatoms of Beaufort, North Carolina. Duke University Marine Station Bulletin 6: 5 - 67." type="journal article" year="1955">Hustedt (1955)</bibRefCitation>
, p. 15, pl. 2 fig. 6, pl. 5 figs 1819; Simonsen (1987), p. 405, pl. 605, figs 19;
<bibRefCitation id="EF94C4D3FFCC8369FB4442BF80F9A8FC" author="Witkowski, A. &amp; Lange-Bertalot, H. &amp; Metzeltin, D." box="[1192,1424,198,219]" pageId="16" pageNumber="217" refId="ref20321" refString="Witkowski, A., Lange-Bertalot, H. &amp; Metzeltin, D. (2000) Diatom flora of marine coasts I. Iconographia Diatomologica. Vol. 7. Koeltz Scientific Books, 925 pp." type="book" year="2000">
Witkowski
<emphasis id="B9716530FFCC8369FAFB42BF8021A8FC" box="[1303,1352,198,219]" italics="true" pageId="16" pageNumber="217">et al.</emphasis>
(2000)
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, p. 97, pl. 42, fig. 22.
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<paragraph id="8BBAB922FFCC8369FF644489843BA002" blockId="16.[136,1452,1776,2085]" pageId="16" pageNumber="217">
<emphasis id="B9716530FFCC8369FF6444898454AF22" bold="true" box="[136,317,1776,1797]" pageId="16" pageNumber="217">FIGURES 4956.</emphasis>
Historical reports of
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<emphasis id="B9716530FFCC8369FDFF448987B1AF22" box="[531,728,1776,1797]" italics="true" pageId="16" pageNumber="217">Anorthoneis hummii</emphasis>
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by different authors. (49) Raphe-sternum and sternum valve drawings of
<taxonomicName id="4C05C2A1FFCC8369FF64456D8587AF0E" authorityName="Hustedt" authorityYear="1955" box="[136,238,1812,1833]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Anorthoneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="16" pageNumber="217" phylum="Ochrophyta" rank="species" species="hummii">
<emphasis id="B9716530FFCC8369FF64456D8587AF0E" box="[136,238,1812,1833]" italics="true" pageId="16" pageNumber="217">A. hummii</emphasis>
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from
<bibRefCitation id="EF94C4D3FFCC8369FEC2456D84A8AF0E" author="Hustedt, F." box="[302,449,1812,1833]" pageId="16" pageNumber="217" pagination="5 - 67" refId="ref18183" refString="Hustedt, F. (1955) Marine littoral diatoms of Beaufort, North Carolina. Duke University Marine Station Bulletin 6: 5 - 67." type="journal article" year="1955">Hustedt (1955)</bibRefCitation>
(holotype); (50) Raphe-sternum valve micrographs of
<taxonomicName id="4C05C2A1FFCC8369FC08456D8120AF0E" authorityName="Hustedt" authorityYear="1955" box="[996,1097,1812,1833]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Anorthoneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="16" pageNumber="217" phylum="Ochrophyta" rank="species" species="hummii">
<emphasis id="B9716530FFCC8369FC08456D8120AF0E" box="[996,1097,1812,1833]" italics="true" pageId="16" pageNumber="217">A. hummii</emphasis>
</taxonomicName>
from
<bibRefCitation id="EF94C4D3FFCC8369FB65456D8010AF0E" author="Witkowski, A. &amp; Lange-Bertalot, H. &amp; Metzeltin, D." box="[1161,1401,1812,1833]" pageId="16" pageNumber="217" refId="ref20321" refString="Witkowski, A., Lange-Bertalot, H. &amp; Metzeltin, D. (2000) Diatom flora of marine coasts I. Iconographia Diatomologica. Vol. 7. Koeltz Scientific Books, 925 pp." type="book" year="2000">
Witkowski
<emphasis id="B9716530FFCC8369FB16456D8047AF0E" box="[1274,1326,1812,1833]" italics="true" pageId="16" pageNumber="217">et al.</emphasis>
(2000)
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; (51, 52) Micrographs of
<taxonomicName id="4C05C2A1FFCC8369FEA1454184D8AF6A" authorityName="Hustedt" authorityYear="1955" box="[333,433,1848,1869]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Anorthoneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="16" pageNumber="217" phylum="Ochrophyta" rank="species" species="hummii">
<emphasis id="B9716530FFCC8369FEA1454184D8AF6A" box="[333,433,1848,1869]" italics="true" pageId="16" pageNumber="217">A. hummii</emphasis>
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specimen H65158 from slide ZT1/87 (Holotype), material AM791 from Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany); (53) Micrograph of
<taxonomicName id="4C05C2A1FFCC8369FCA0452586C4AF56" authorityName="Hustedt" authorityYear="1955" box="[844,941,1884,1905]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Anorthoneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="16" pageNumber="217" phylum="Ochrophyta" rank="species" species="hummii">
<emphasis id="B9716530FFCC8369FCA0452586C4AF56" box="[844,941,1884,1905]" italics="true" pageId="16" pageNumber="217">A. hummii</emphasis>
</taxonomicName>
specimen H65433 from slide ZT1/96 (Paralectotype), material AM795 from Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany); (54, 55) Micrographs of
<taxonomicName id="4C05C2A1FFCC8369FA7B45F985B9AF9E" authorityName="Hustedt" authorityYear="1955" class="Bacillariophyceae" family="Cocconeidaceae" genus="Anorthoneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="16" pageNumber="217" phylum="Ochrophyta" rank="species" species="hummii">
<emphasis id="B9716530FFCC8369FA7B45F985B9AF9E" italics="true" pageId="16" pageNumber="217">A. hummii</emphasis>
</taxonomicName>
specimen H65252 from slide ZT1/92 (isolectotype), material AM792 from Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany); (56) Micrograph of
<taxonomicName id="4C05C2A1FFCC8369FD5A45B18673AFFA" authorityName="Hustedt" authorityYear="1955" box="[694,794,1992,2013]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Anorthoneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="16" pageNumber="217" phylum="Ochrophyta" rank="species" species="hummii">
<emphasis id="B9716530FFCC8369FD5A45B18673AFFA" box="[694,794,1992,2013]" italics="true" pageId="16" pageNumber="217">A. hummii</emphasis>
</taxonomicName>
specimen H65217a from slide ZT1/90 (lectotype), material AM792 from Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany). Magnification: Fig. 52: x1500. Scale bars: Figs 5156 = 10 μm.
</paragraph>
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<paragraph id="8BBAB922FFCD8368FF6447E1861EAE46" blockId="17.[136,1452,1432,1633]" pageId="17" pageNumber="218">
<emphasis id="B9716530FFCD8368FF6447E18454AD8A" bold="true" box="[136,317,1432,1453]" pageId="17" pageNumber="218">FIGURES 5762.</emphasis>
<taxonomicName id="4C05C2A1FFCD8368FEA947E18762AD8A" authorityName="Hustedt" authorityYear="1955" box="[325,523,1432,1453]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Anorthoneis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="17" pageNumber="218" phylum="Ochrophyta" rank="species" species="hummii">
<emphasis id="B9716530FFCD8368FEA947E18762AD8A" box="[325,523,1432,1453]" italics="true" pageId="17" pageNumber="218">Anorthoneis hummii</emphasis>
</taxonomicName>
specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. (57) External sternum valve with asymmetrical hyaline central area and small roundish terminal areas (arrow); (58) Detail of sternum valve mantle terminating in a wide hyaline marginal area; (59) Detail of roundish/transapically elongated areolae externally occluded by thick perforated hymenes; (60) Detail of sternum valve mantle; (61) Internal view of areolae occluded by perforated hymens; (62) Internal sternum valve showing sternum and hyaline central area. Scale bars: Fig. 57 = 10 μm; Figs 62, 63 = 5 μm; Figs 58, 60 = 2 μm; Figs 59, 61 = 1 μm.
</paragraph>
</caption>
<subSubSection id="C31FEAA9FFCD836BFF6444E886DEA9B7" lastPageId="18" lastPageNumber="219" pageId="17" pageNumber="218" type="description">
<paragraph id="8BBAB922FFCD8368FF6444E88468AEE9" blockId="17.[136,1452,1681,1994]" pageId="17" pageNumber="218">
<emphasis id="B9716530FFCD8368FF6444E88473AE8D" bold="true" box="[136,282,1681,1706]" pageId="17" pageNumber="218">Description.</emphasis>
Valves outline varies from ovoid to roundish (
<figureCitation id="133EA5A7FFCD8368FCD744E886FCAE8D" box="[827,917,1681,1706]" captionStart="FIGURES 5762" captionStartId="17.[136,243,1432,1453]" captionTargetBox="[162,1430,201,1399]" captionTargetId="figure-14@17.[151,1436,190,1407]" captionTargetPageId="17" captionText="FIGURES 5762. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. (57) External sternum valve with asymmetrical hyaline central area and small roundish terminal areas (arrow); (58) Detail of sternum valve mantle terminating in a wide hyaline marginal area; (59) Detail of roundish/transapically elongated areolae externally occluded by thick perforated hymenes; (60) Detail of sternum valve mantle; (61) Internal view of areolae occluded by perforated hymens; (62) Internal sternum valve showing sternum and hyaline central area. Scale bars: Fig. 57 = 10 μm; Figs 62, 63 = 5 μm; Figs 58, 60 = 2 μm; Figs 59, 61 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720132" httpUri="https://zenodo.org/record/13720132/files/figure.png" pageId="17" pageNumber="218">Figs 57</figureCitation>
,
<figureCitation id="133EA5A7FFCD8368FC4844E886AAAE8D" box="[932,963,1681,1706]" captionStart="FIGURES 63-67" captionStartId="18.[136,243,1627,1648]" captionTargetBox="[162,1430,436,1596]" captionTargetId="figure-133@18.[151,1436,425,1602]" captionTargetPageId="18" captionText="FIGURES 63-67. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. 63. External view of raphe-sternum valve. (64) Detail of coaxial raphe central endings in external view lying in drop-shaped areas; (65) External view of distal raphe ending terminating in hyaline submarginal area; (66) Tilted external view (60°) of raphe-sternum mantle showing the opened valvocopula with smooth inner margin; (67) Tilted external view (60°) of raphe-sternum valve. Scale bars: Fig. 57 = 10 μm, Figs 62, 63, 67 = 5 μm; Figs 58, 60, 65 = 2 μm; Figs 59, 61, 64, 66 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720134" httpUri="https://zenodo.org/record/13720134/files/figure.png" pageId="17" pageNumber="218">63</figureCitation>
). Apical axis: 2741 μm; transapical axis: 2336 μm.
</paragraph>
<paragraph id="8BBAB922FFCD8368FF5144AE86FAAFED" blockId="17.[136,1452,1681,1994]" pageId="17" pageNumber="218">
<emphasis id="B9716530FFCD8368FF5144AE846CAED5" bold="true" box="[189,261,1751,1778]" pageId="17" pageNumber="218">
<emphasis id="B9716530FFCD8368FF5144AE85B6AED6" bold="true" box="[189,223,1751,1777]" italics="true" pageId="17" pageNumber="218">SV</emphasis>
:—
</emphasis>
Externally, valve face flat to convex, curving abruptly into a shallow, imperforate mantle (
<figureCitation id="133EA5A7FFCD8368FB1444A080C9AED5" box="[1272,1440,1753,1778]" captionStart="FIGURES 5762" captionStartId="17.[136,243,1432,1453]" captionTargetBox="[162,1430,201,1399]" captionTargetId="figure-14@17.[151,1436,190,1407]" captionTargetPageId="17" captionText="FIGURES 5762. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. (57) External sternum valve with asymmetrical hyaline central area and small roundish terminal areas (arrow); (58) Detail of sternum valve mantle terminating in a wide hyaline marginal area; (59) Detail of roundish/transapically elongated areolae externally occluded by thick perforated hymenes; (60) Detail of sternum valve mantle; (61) Internal view of areolae occluded by perforated hymens; (62) Internal sternum valve showing sternum and hyaline central area. Scale bars: Fig. 57 = 10 μm; Figs 62, 63 = 5 μm; Figs 58, 60 = 2 μm; Figs 59, 61 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720132" httpUri="https://zenodo.org/record/13720132/files/figure.png" pageId="17" pageNumber="218">Fig. 58 and 60</figureCitation>
). Sternum depressed, asymmetric, terminating in small roundish hyaline areas, close to the valve margin (
<figureCitation id="133EA5A7FFCD8368FAE94484803DAF31" box="[1285,1364,1789,1814]" captionStart="FIGURES 5762" captionStartId="17.[136,243,1432,1453]" captionTargetBox="[162,1430,201,1399]" captionTargetId="figure-14@17.[151,1436,190,1407]" captionTargetPageId="17" captionText="FIGURES 5762. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. (57) External sternum valve with asymmetrical hyaline central area and small roundish terminal areas (arrow); (58) Detail of sternum valve mantle terminating in a wide hyaline marginal area; (59) Detail of roundish/transapically elongated areolae externally occluded by thick perforated hymenes; (60) Detail of sternum valve mantle; (61) Internal view of areolae occluded by perforated hymens; (62) Internal sternum valve showing sternum and hyaline central area. Scale bars: Fig. 57 = 10 μm; Figs 62, 63 = 5 μm; Figs 58, 60 = 2 μm; Figs 59, 61 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720132" httpUri="https://zenodo.org/record/13720132/files/figure.png" pageId="17" pageNumber="218">Fig. 57</figureCitation>
arrow). Central area not clearly defined (
<figureCitation id="133EA5A7FFCD8368FE104558873BAF1D" box="[508,594,1825,1850]" captionStart="FIGURES 5762" captionStartId="17.[136,243,1432,1453]" captionTargetBox="[162,1430,201,1399]" captionTargetId="figure-14@17.[151,1436,190,1407]" captionTargetPageId="17" captionText="FIGURES 5762. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. (57) External sternum valve with asymmetrical hyaline central area and small roundish terminal areas (arrow); (58) Detail of sternum valve mantle terminating in a wide hyaline marginal area; (59) Detail of roundish/transapically elongated areolae externally occluded by thick perforated hymenes; (60) Detail of sternum valve mantle; (61) Internal view of areolae occluded by perforated hymens; (62) Internal sternum valve showing sternum and hyaline central area. Scale bars: Fig. 57 = 10 μm; Figs 62, 63 = 5 μm; Figs 58, 60 = 2 μm; Figs 59, 61 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720132" httpUri="https://zenodo.org/record/13720132/files/figure.png" pageId="17" pageNumber="218">Fig. 57</figureCitation>
). Transapical striae uniseriate on both the valve face and the mantle, from
<quantity id="4CFD14C7FFCD8368FA72455885F7AF79" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="17" pageNumber="218" unit="in" value="8.0">8 in</quantity>
10 μm close to the margin to
<quantity id="4CFD14C7FFCD8368FE1D453C8743AF79" box="[497,554,1861,1886]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="17" pageNumber="218" unit="in" value="10.0">10 in</quantity>
10 μm close to the sternum, composed of large, transapically elongated areolae (
<quantity id="4CFD14C7FFCD8368FF7D451085A4AFA5" box="[145,205,1897,1922]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="17" pageNumber="218" unit="in" value="10.0">10 in</quantity>
10 μm). Thick hymenes occluding externally the areolae perforated by round porelli in a circular to regular scatter (
<figureCitation id="133EA5A7FFCD8368FF0C45F4845AAF81" box="[224,307,1933,1958]" captionStart="FIGURES 5762" captionStartId="17.[136,243,1432,1453]" captionTargetBox="[162,1430,201,1399]" captionTargetId="figure-14@17.[151,1436,190,1407]" captionTargetPageId="17" captionText="FIGURES 5762. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. (57) External sternum valve with asymmetrical hyaline central area and small roundish terminal areas (arrow); (58) Detail of sternum valve mantle terminating in a wide hyaline marginal area; (59) Detail of roundish/transapically elongated areolae externally occluded by thick perforated hymenes; (60) Detail of sternum valve mantle; (61) Internal view of areolae occluded by perforated hymens; (62) Internal sternum valve showing sternum and hyaline central area. Scale bars: Fig. 57 = 10 μm; Figs 62, 63 = 5 μm; Figs 58, 60 = 2 μm; Figs 59, 61 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720132" httpUri="https://zenodo.org/record/13720132/files/figure.png" pageId="17" pageNumber="218">Fig. 59</figureCitation>
). Internally, valve face concave, with raised narrow sternum, expanding laterally into large asymmetric roundish central area (
<figureCitation id="133EA5A7FFCD8368FE6E45C884BEAFED" box="[386,471,1969,1994]" captionStart="FIGURES 5762" captionStartId="17.[136,243,1432,1453]" captionTargetBox="[162,1430,201,1399]" captionTargetId="figure-14@17.[151,1436,190,1407]" captionTargetPageId="17" captionText="FIGURES 5762. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. (57) External sternum valve with asymmetrical hyaline central area and small roundish terminal areas (arrow); (58) Detail of sternum valve mantle terminating in a wide hyaline marginal area; (59) Detail of roundish/transapically elongated areolae externally occluded by thick perforated hymenes; (60) Detail of sternum valve mantle; (61) Internal view of areolae occluded by perforated hymens; (62) Internal sternum valve showing sternum and hyaline central area. Scale bars: Fig. 57 = 10 μm; Figs 62, 63 = 5 μm; Figs 58, 60 = 2 μm; Figs 59, 61 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720132" httpUri="https://zenodo.org/record/13720132/files/figure.png" pageId="17" pageNumber="218">Fig. 62</figureCitation>
). Mantle virgae visibly raised (
<figureCitation id="133EA5A7FFCD8368FCDE45C886EFAFED" box="[818,902,1969,1994]" captionStart="FIGURES 5762" captionStartId="17.[136,243,1432,1453]" captionTargetBox="[162,1430,201,1399]" captionTargetId="figure-14@17.[151,1436,190,1407]" captionTargetPageId="17" captionText="FIGURES 5762. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. (57) External sternum valve with asymmetrical hyaline central area and small roundish terminal areas (arrow); (58) Detail of sternum valve mantle terminating in a wide hyaline marginal area; (59) Detail of roundish/transapically elongated areolae externally occluded by thick perforated hymenes; (60) Detail of sternum valve mantle; (61) Internal view of areolae occluded by perforated hymens; (62) Internal sternum valve showing sternum and hyaline central area. Scale bars: Fig. 57 = 10 μm; Figs 62, 63 = 5 μm; Figs 58, 60 = 2 μm; Figs 59, 61 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720132" httpUri="https://zenodo.org/record/13720132/files/figure.png" pageId="17" pageNumber="218">Fig. 61</figureCitation>
).
</paragraph>
<paragraph id="8BBAB922FFCE836BFF5142E786DEA9B7" blockId="18.[136,1452,158,400]" pageId="18" pageNumber="219">
<emphasis id="B9716530FFCE836BFF5142E78471A89F" bold="true" box="[189,280,158,184]" pageId="18" pageNumber="219">
<emphasis id="B9716530FFCE836BFF5142E7859BA89F" bold="true" box="[189,242,158,184]" italics="true" pageId="18" pageNumber="219">RSV</emphasis>
:—
</emphasis>
Externally, valve face almost flat to slightly concave, sloping gently into a shallow mantle (
<figureCitation id="133EA5A7FFCE836BFAAB42E680F6A89F" box="[1351,1439,159,184]" captionStart="FIGURES 63-67" captionStartId="18.[136,243,1627,1648]" captionTargetBox="[162,1430,436,1596]" captionTargetId="figure-133@18.[151,1436,425,1602]" captionTargetPageId="18" captionText="FIGURES 63-67. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. 63. External view of raphe-sternum valve. (64) Detail of coaxial raphe central endings in external view lying in drop-shaped areas; (65) External view of distal raphe ending terminating in hyaline submarginal area; (66) Tilted external view (60°) of raphe-sternum mantle showing the opened valvocopula with smooth inner margin; (67) Tilted external view (60°) of raphe-sternum valve. Scale bars: Fig. 57 = 10 μm, Figs 62, 63, 67 = 5 μm; Figs 58, 60, 65 = 2 μm; Figs 59, 61, 64, 66 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720134" httpUri="https://zenodo.org/record/13720134/files/figure.png" pageId="18" pageNumber="219">Fig. 67</figureCitation>
). Striae radial, uniseriate, becoming biseriate close to the valve margin,
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10 μm (
<figureCitation id="133EA5A7FFCE836BFB8542BA819DA8FB" box="[1129,1268,195,220]" captionStart="FIGURES 63-67" captionStartId="18.[136,243,1627,1648]" captionTargetBox="[162,1430,436,1596]" captionTargetId="figure-133@18.[151,1436,425,1602]" captionTargetPageId="18" captionText="FIGURES 63-67. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. 63. External view of raphe-sternum valve. (64) Detail of coaxial raphe central endings in external view lying in drop-shaped areas; (65) External view of distal raphe ending terminating in hyaline submarginal area; (66) Tilted external view (60°) of raphe-sternum mantle showing the opened valvocopula with smooth inner margin; (67) Tilted external view (60°) of raphe-sternum valve. Scale bars: Fig. 57 = 10 μm, Figs 62, 63, 67 = 5 μm; Figs 58, 60, 65 = 2 μm; Figs 59, 61, 64, 66 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720134" httpUri="https://zenodo.org/record/13720134/files/figure.png" pageId="18" pageNumber="219">Figs 65, 66</figureCitation>
). Areolae large, roundish (valve face) or transapically-elongated (mantle),
<quantity id="4CFD14C7FFCE836BFCF8429E8624A927" box="[788,845,231,256]" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="18" pageNumber="219" unit="in" value="30.0">30 in</quantity>
10 μm, occluded by hymenes ultrastructurally similar to those in the SV (
<figureCitation id="133EA5A7FFCE836BFE80437284AAA903" box="[364,451,267,292]" captionStart="FIGURES 63-67" captionStartId="18.[136,243,1627,1648]" captionTargetBox="[162,1430,436,1596]" captionTargetId="figure-133@18.[151,1436,425,1602]" captionTargetPageId="18" captionText="FIGURES 63-67. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. 63. External view of raphe-sternum valve. (64) Detail of coaxial raphe central endings in external view lying in drop-shaped areas; (65) External view of distal raphe ending terminating in hyaline submarginal area; (66) Tilted external view (60°) of raphe-sternum mantle showing the opened valvocopula with smooth inner margin; (67) Tilted external view (60°) of raphe-sternum valve. Scale bars: Fig. 57 = 10 μm, Figs 62, 63, 67 = 5 μm; Figs 58, 60, 65 = 2 μm; Figs 59, 61, 64, 66 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720134" httpUri="https://zenodo.org/record/13720134/files/figure.png" pageId="18" pageNumber="219">Fig. 66</figureCitation>
). Central area very narrow, not well discernible, flanked by imperforated small areolae (
<figureCitation id="133EA5A7FFCE836BFF7C4356858DA96F" box="[144,228,303,328]" captionStart="FIGURES 63-67" captionStartId="18.[136,243,1627,1648]" captionTargetBox="[162,1430,436,1596]" captionTargetId="figure-133@18.[151,1436,425,1602]" captionTargetPageId="18" captionText="FIGURES 63-67. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. 63. External view of raphe-sternum valve. (64) Detail of coaxial raphe central endings in external view lying in drop-shaped areas; (65) External view of distal raphe ending terminating in hyaline submarginal area; (66) Tilted external view (60°) of raphe-sternum mantle showing the opened valvocopula with smooth inner margin; (67) Tilted external view (60°) of raphe-sternum valve. Scale bars: Fig. 57 = 10 μm, Figs 62, 63, 67 = 5 μm; Figs 58, 60, 65 = 2 μm; Figs 59, 61, 64, 66 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720134" httpUri="https://zenodo.org/record/13720134/files/figure.png" pageId="18" pageNumber="219">Fig. 64</figureCitation>
). Proximal raphe endings expanded, pore-like, coaxial and lying in concave drop-shaped area (
<figureCitation id="133EA5A7FFCE836BFAEE4356803FA96F" box="[1282,1366,303,328]" captionStart="FIGURES 63-67" captionStartId="18.[136,243,1627,1648]" captionTargetBox="[162,1430,436,1596]" captionTargetId="figure-133@18.[151,1436,425,1602]" captionTargetPageId="18" captionText="FIGURES 63-67. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. 63. External view of raphe-sternum valve. (64) Detail of coaxial raphe central endings in external view lying in drop-shaped areas; (65) External view of distal raphe ending terminating in hyaline submarginal area; (66) Tilted external view (60°) of raphe-sternum mantle showing the opened valvocopula with smooth inner margin; (67) Tilted external view (60°) of raphe-sternum valve. Scale bars: Fig. 57 = 10 μm, Figs 62, 63, 67 = 5 μm; Figs 58, 60, 65 = 2 μm; Figs 59, 61, 64, 66 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720134" httpUri="https://zenodo.org/record/13720134/files/figure.png" pageId="18" pageNumber="219">Fig. 64</figureCitation>
). Distal raphe endings expanded, pore-like, hardly reaching hyaline submarginal areas (
<figureCitation id="133EA5A7FFCE836BFBEC432A813BA94B" box="[1024,1106,339,364]" captionStart="FIGURES 63-67" captionStartId="18.[136,243,1627,1648]" captionTargetBox="[162,1430,436,1596]" captionTargetId="figure-133@18.[151,1436,425,1602]" captionTargetPageId="18" captionText="FIGURES 63-67. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. 63. External view of raphe-sternum valve. (64) Detail of coaxial raphe central endings in external view lying in drop-shaped areas; (65) External view of distal raphe ending terminating in hyaline submarginal area; (66) Tilted external view (60°) of raphe-sternum mantle showing the opened valvocopula with smooth inner margin; (67) Tilted external view (60°) of raphe-sternum valve. Scale bars: Fig. 57 = 10 μm, Figs 62, 63, 67 = 5 μm; Figs 58, 60, 65 = 2 μm; Figs 59, 61, 64, 66 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720134" httpUri="https://zenodo.org/record/13720134/files/figure.png" pageId="18" pageNumber="219">Fig. 65</figureCitation>
, arrow), distant from the valve margin. Valvocopula open, with a smooth inner margin (
<figureCitation id="133EA5A7FFCE836BFCEC430E86C2A9B7" box="[768,939,375,400]" captionStart="FIGURES 63-67" captionStartId="18.[136,243,1627,1648]" captionTargetBox="[162,1430,436,1596]" captionTargetId="figure-133@18.[151,1436,425,1602]" captionTargetPageId="18" captionText="FIGURES 63-67. Anorthoneis hummii specimens from Hustedts cleaned materials AM791, AM792, Beaufort, North Carolina, USA (Hustedt Diatom Collection, Bremerhaven, Germany), SEM images. 63. External view of raphe-sternum valve. (64) Detail of coaxial raphe central endings in external view lying in drop-shaped areas; (65) External view of distal raphe ending terminating in hyaline submarginal area; (66) Tilted external view (60°) of raphe-sternum mantle showing the opened valvocopula with smooth inner margin; (67) Tilted external view (60°) of raphe-sternum valve. Scale bars: Fig. 57 = 10 μm, Figs 62, 63, 67 = 5 μm; Figs 58, 60, 65 = 2 μm; Figs 59, 61, 64, 66 = 1 μm." figureDoi="http://doi.org/10.5281/zenodo.13720134" httpUri="https://zenodo.org/record/13720134/files/figure.png" pageId="18" pageNumber="219">Figs 65 and 66</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>