treatments-xml/data/56/54/1E/56541E5AFFCABB5BFEC9072EFE422A0F.xml
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<document id="3EC5155A18F4FEC386B428269F58853A" ID-DOI="10.37520/aemnp.2022.010" ID-ISSN="1804-6487" ID-Zenodo-Dep="7161319" ID-ZooBank="D67D1D5C-7A1F-4FB7-93D8-DF39CC3F5FF1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665202576664" checkinUser="felipe" docAuthor="Tasaku, Yuto &amp; Maruyama, Munetoshi" docDate="2022" docId="56541E5AFFCABB5BFEC9072EFE422A0F" docLanguage="en" docName="ActaEntMusNatPra.62.1.137-144.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 62 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2022.010" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Eubadura hokkaidensis Tasaku &amp; Maruyama 2022, sp. nov." docType="treatment" docVersion="3" lastPageNumber="144" masterDocId="AA6D6622FFCCBB5CFFDE0435FFA9287A" masterDocTitle="Taxonomy of the intertidal athetine genus Eubadura stat. nov. (Coleoptera: Staphylinidae: Aleocharinae) from Japan, with descriptions of two new species" masterLastPageNumber="144" masterPageNumber="137" pageNumber="143" updateTime="1699365925587" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="67154281BC80EAF0A8DE2A8C316B4143">Taxonomy of the intertidal athetine genus Eubadura stat. nov. (Coleoptera: Staphylinidae: Aleocharinae) from Japan, with descriptions of two new species</mods:title>
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<mods:namePart id="30E4639557B7C4F35FD9DDBAA5A37366">Tasaku, Yuto</mods:namePart>
<mods:affiliation id="3D9103DF579B40A23D4F4D42D2DA7038">Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan</mods:affiliation>
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<mods:namePart id="63BC0EC6942C842D054A164B3BFCC3E3">Maruyama, Munetoshi</mods:namePart>
<mods:affiliation id="9B9858FEF0953565E7705302C76FC48E">The Kyushu University Museum, Hakozaki</mods:affiliation>
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<mods:typeOfResource id="B0EAB913BB43F83E6011C02727FFE7A9">text</mods:typeOfResource>
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<mods:title id="FB4941360E1208F7D87ECA4EDF98794E">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="DD5F4D42B704F37BCAFA623F288B9FF9">2022</mods:date>
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<mods:number id="2D1D03DBAE8538A786232A0CE0E24599">2022-07-12</mods:number>
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<treatment id="56541E5AFFCABB5BFEC9072EFE422A0F" ID-DOI="http://doi.org/10.5281/zenodo.7161323" ID-Zenodo-Dep="7161323" LSID="urn:lsid:plazi:treatment:56541E5AFFCABB5BFEC9072EFE422A0F" httpUri="http://treatment.plazi.org/id/56541E5AFFCABB5BFEC9072EFE422A0F" lastPageId="7" lastPageNumber="144" pageId="6" pageNumber="143">
<subSubSection id="96E7FCC7FFCABB5AFEC9072EFD242B37" pageId="6" pageNumber="143" type="nomenclature">
<paragraph id="DE42AF4CFFCABB5AFEC9072EFDD12B49" blockId="6.[279,632,795,819]" box="[279,632,795,819]" pageId="6" pageNumber="143">
<heading id="850A1820FFCABB5AFEC9072EFDD12B49" box="[279,632,795,819]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="143" reason="3">
<emphasis id="EC89735EFFCABB5AFEC9072EFDD12B49" bold="true" box="[279,632,795,819]" pageId="6" pageNumber="143">
<taxonomicName id="19FDD4CFFFCABB5AFEC9072EFDB42B49" ID-CoL="9GKW8" authority="Tasaku &amp; Maruyama, 2022" authorityName="Tasaku &amp; Maruyama" authorityYear="2022" box="[279,541,795,819]" class="Insecta" family="Staphylinidae" genus="Eubadura" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="143" phylum="Arthropoda" rank="species" species="hokkaidensis" status="sp. nov.">
<emphasis id="EC89735EFFCABB5AFEC9072EFDB42B49" bold="true" box="[279,541,795,819]" italics="true" pageId="6" pageNumber="143">Eubadura hokkaidensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7BACE25FFCABB5AFDFD0729FDD12B49" box="[547,632,796,819]" pageId="6" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DE42AF4CFFCABB5AFEDD070FFD242B37" blockId="6.[259,653,826,872]" box="[259,653,826,845]" pageId="6" pageNumber="143">[Japanese name: Kuro-umihime-hanekakushi]</paragraph>
</subSubSection>
<subSubSection id="96E7FCC7FFCABB5AFE5F0760FDA72B12" box="[385,526,853,872]" pageId="6" pageNumber="143" type="description">
<paragraph id="DE42AF4CFFCABB5AFE5F0760FDA72B12" blockId="6.[259,653,826,872]" box="[385,526,853,872]" pageId="6" pageNumber="143">
(
<figureCitation id="46C6B3C9FFCABB5AFE590760FE752B12" box="[391,476,853,872]" captionStart="Fig" captionStartId="1.[151,182,2034,2053]" captionTargetBox="[236,1358,199,1961]" captionTargetId="figure-15@1.[223,1364,189,1973]" captionTargetPageId="1" captionText="Fig. 1. Habitus of Eubadura spp. AC, G E. akiensis (Sawada, 1990); D, H E. yasukoae (Maruyama, Klimaszewski &amp; Gusarov, 2008); E, I E. onoi sp. nov.; F, J E. hokkaidensis sp. nov. Scale bars: 1.00 mm" figureDoi="http://doi.org/10.5281/zenodo.7161321" httpUri="https://zenodo.org/record/7161321/files/figure.png" pageId="6" pageNumber="143">Figs 1F, J</figureCitation>
;
<figureCitation id="46C6B3C9FFCABB5AFE380760FE5A2B12" box="[486,499,853,872]" captionStart="Fig" captionStartId="5.[152,183,2030,2049]" captionTargetBox="[243,1346,599,1996]" captionTargetId="figure-220@5.[230,1357,585,2011]" captionTargetPageId="5" captionText="Fig. 4. Eubadura hokkaidensis sp. nov. A male tergite VIII; B male sternite VIII; C female tergite VIII; D female sternite VIII; E spermatheca; F median lobe, ventral view; G median lobe, lateral view; H paramere. Scale bars: 0. 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7161329" httpUri="https://zenodo.org/record/7161329/files/figure.png" pageId="6" pageNumber="143">4</figureCitation>
;
<figureCitation id="46C6B3C9FFCABB5AFE220760FDAE2B12" box="[508,519,853,872]" captionStart="Fig" captionStartId="6.[150,181,2027,2046]" captionTargetBox="[203,1385,1095,2010]" captionTargetId="figure-481@6.[202,1385,1093,2010]" captionTargetPageId="6" captionText="Fig. 5. Known localities of Eubadura spp.Black circle E. akiensis (Sawada, 1990); black triangle E. yasukoae (Maruyama, Klimaszewski &amp; Gusarov, 2008); black square E. onoi sp. nov.; black star E. hokkaidensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7161331" httpUri="https://zenodo.org/record/7161331/files/figure.png" pageId="6" pageNumber="143">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="96E7FCC7FFCABB5AFF49074AFDDE2C48" pageId="6" pageNumber="143" type="materials_examined">
<paragraph id="DE42AF4CFFCABB5AFF49074AFDDE2C48" blockId="6.[151,761,895,1074]" pageId="6" pageNumber="143">
<emphasis id="EC89735EFFCABB5AFF49074AFEB52BE8" bold="true" box="[151,284,895,914]" pageId="6" pageNumber="143">Type material.</emphasis>
Hඈඅඈඍඒඉൾ:
<specimenCount id="C8FB64C5FFCABB5AFE58074AFE3F2BE8" box="[390,406,895,914]" pageId="6" pageNumber="143" type="male"></specimenCount>
,
<emphasis id="EC89735EFFCABB5AFE7E074AFDF02BE8" bold="true" box="[416,601,895,914]" pageId="6" pageNumber="143">
[
<collectingCountry id="A6EAEFDCFFCABB5AFE78074AFE582BE8" box="[422,497,895,914]" name="Japan" pageId="6" pageNumber="143">JAPAN</collectingCountry>
]:
<collectingRegion id="1C3961AEFFCABB5AFE21074AFDF02BE8" box="[511,601,895,914]" country="Japan" name="Hokkaido" pageId="6" pageNumber="143">Hokkaidô</collectingRegion>
</emphasis>
, Abashiri-shi, Komaba,
<date id="AA43898CFFCABB5AFF0E07ACFE9D2BD6" box="[208,308,921,940]" pageId="6" pageNumber="143" value="2021-06-20">
20
<collectionCode id="B8EC3789FFCABB5AFF3507ACFEA82BD6" box="[235,257,921,940]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="143">VI</collectionCode>
2021
</date>
, Y. Tasaku. (
<collectionCode id="B8EC3789FFCABB5AFE7E07ACFE7F2BD6" box="[416,470,921,940]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="143" type="Herbarium">KUM</collectionCode>
). Pൺඋൺඍඒඉൾඌ: 23 exs, same data as
<typeStatus id="014611EEFFCABB5AFF490781FF4B2BBD" box="[151,226,948,967]" pageId="6" pageNumber="143" type="holotype">holotype</typeStatus>
(
<collectionCode id="B8EC3789FFCABB5AFF320781FE882BBD" box="[236,289,948,967]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="143" type="Herbarium">KUM</collectionCode>
,
<collectionCode id="B8EC3789FFCABB5AFEF60781FEC12BBD" box="[296,360,948,967]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="6" pageNumber="143">NMPC</collectionCode>
); 5 exs, same data as
<typeStatus id="014611EEFFCABB5AFDC80781FDCA2BBD" box="[534,611,948,967]" pageId="6" pageNumber="143" type="holotype">holotype</typeStatus>
, but
<date id="AA43898CFFCABB5AFD570781FD5C2BBD" box="[649,757,948,967]" pageId="6" pageNumber="143" value="2020-09-16">16. IX. 2020</date>
, Y. Tasaku (
<collectionCode id="B8EC3789FFCABB5AFF2407FAFE992B98" box="[250,304,975,994]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="143" type="Herbarium">KUM</collectionCode>
); 3 exs, Ishikari-shi, Atsuta-ku, Betsukari, mouth of Atsuta-gawa riv.,
<date id="AA43898CFFCABB5AFEF507DCFE252B86" box="[299,396,1001,1020]" pageId="6" pageNumber="143" value="2021-06-05">
5.
<collectionCode id="B8EC3789FFCABB5AFEE107DCFEFC2B86" box="[319,341,1001,1020]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="143">VI</collectionCode>
. 2021
</date>
, Y.Tasaku (
<collectionCode id="B8EC3789FFCABB5AFE3307DCFD8A2B86" box="[493,547,1001,1020]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="143" type="Herbarium">KUM</collectionCode>
); 3 exs, Sôya, Esashi-tyô,
<date id="AA43898CFFCABB5AFF490031FEAA2C6D" box="[151,259,1028,1047]" pageId="6" pageNumber="143" value="2009-07-23">
23
<collectionCode id="B8EC3789FFCABB5AFF6C0031FF792C6D" box="[178,208,1028,1047]" pageId="6" pageNumber="143">VII</collectionCode>
2009
</date>
, M. Ôhara (
<collectionCode id="B8EC3789FFCABB5AFEB20031FE0B2C6D" box="[364,418,1028,1047]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="143" type="Herbarium">KUM</collectionCode>
); 2 exs, Nemuro, Rausu-tyô, Azabu,
<date id="AA43898CFFCABB5AFD3C0031FF492C48" pageId="6" pageNumber="143" value="2009-09-19">19 IX 2009</date>
, M. Ôhara, H. Yamamoto &amp; M. Furuta (
<collectionCode id="B8EC3789FFCABB5AFDE9002AFDC42C48" box="[567,621,1055,1074]" country="Ghana" lsid="urn:lsid:biocol.org:col:13924" name="Resource Management Support Center" pageId="6" pageNumber="143" type="Herbarium">KUM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="96E7FCC7FFCABB5BFCE2048BFDC729CF" lastPageId="7" lastPageNumber="144" pageId="6" pageNumber="143" type="description">
<paragraph id="DE42AF4CFFCABB5AFCE2048BFB462A8F" blockId="6.[828,1437,190,1077]" pageId="6" pageNumber="143">
<emphasis id="EC89735EFFCABB5AFCE2048BFC6F28AF" bold="true" box="[828,966,190,213]" pageId="6" pageNumber="143">Description.</emphasis>
Head, pronotum, elytra and abdomen almost black; antennae, mouthparts and legs blackish brown and yellowish brown. Head (
<figureCitation id="46C6B3C9FFCABB5AFB9D04CBFB38296F" box="[1091,1169,254,277]" captionStart="Fig" captionStartId="1.[151,182,2034,2053]" captionTargetBox="[236,1358,199,1961]" captionTargetId="figure-15@1.[223,1364,189,1973]" captionTargetPageId="1" captionText="Fig. 1. Habitus of Eubadura spp. AC, G E. akiensis (Sawada, 1990); D, H E. yasukoae (Maruyama, Klimaszewski &amp; Gusarov, 2008); E, I E. onoi sp. nov.; F, J E. hokkaidensis sp. nov. Scale bars: 1.00 mm" figureDoi="http://doi.org/10.5281/zenodo.7161321" httpUri="https://zenodo.org/record/7161321/files/figure.png" pageId="6" pageNumber="143">Fig. 1F</figureCitation>
) as long as wide. Pronotum slightly transverse, about 1.1 times as wide as long, narrowed posteriorly; postero-lateral corner rounded; surface finely punctured, densely covered with setae, with 7 macrosetae along lateral margin. Scutellum with integument smooth and sparsely covered with setae. Elytra (
<figureCitation id="46C6B3C9FFCABB5AFC9A058BFC3B29AF" box="[836,914,446,469]" captionStart="Fig" captionStartId="1.[151,182,2034,2053]" captionTargetBox="[236,1358,199,1961]" captionTargetId="figure-15@1.[223,1364,189,1973]" captionTargetPageId="1" captionText="Fig. 1. Habitus of Eubadura spp. AC, G E. akiensis (Sawada, 1990); D, H E. yasukoae (Maruyama, Klimaszewski &amp; Gusarov, 2008); E, I E. onoi sp. nov.; F, J E. hokkaidensis sp. nov. Scale bars: 1.00 mm" figureDoi="http://doi.org/10.5281/zenodo.7161321" httpUri="https://zenodo.org/record/7161321/files/figure.png" pageId="6" pageNumber="143">Fig. 1F</figureCitation>
) as wide as pronotum; surface finely punctured, densely covered with setae, with 23 small macrosetae laterally. Hind wings reduced, about 1/3 as long as elytra. Abdomen oblong, widest around segments VI and VII; surface smooth; tergites IIVII densely covered with setae. Antennae (
<figureCitation id="46C6B3C9FFCABB5AFC6F066BFC562A0F" box="[945,1023,606,629]" captionStart="Fig" captionStartId="1.[151,182,2034,2053]" captionTargetBox="[236,1358,199,1961]" captionTargetId="figure-15@1.[223,1364,189,1973]" captionTargetPageId="1" captionText="Fig. 1. Habitus of Eubadura spp. AC, G E. akiensis (Sawada, 1990); D, H E. yasukoae (Maruyama, Klimaszewski &amp; Gusarov, 2008); E, I E. onoi sp. nov.; F, J E. hokkaidensis sp. nov. Scale bars: 1.00 mm" figureDoi="http://doi.org/10.5281/zenodo.7161321" httpUri="https://zenodo.org/record/7161321/files/figure.png" pageId="6" pageNumber="143">Fig. 1F</figureCitation>
) almost as long as head and pronotum combined; antennomere I long; antennomere II shorter than I; antennomere III slightly shorter than II; antennomeres IVX moniliform, each almost as long as wide; antennomere XI long, almost twice as long as X.
</paragraph>
<paragraph id="DE42AF4CFFCABB5AFC8406C8FBDF2B8F" blockId="6.[828,1437,190,1077]" pageId="6" pageNumber="143">
<emphasis id="EC89735EFFCABB5AFC8406C8FC332B6F" bold="true" box="[858,922,765,789]" italics="true" pageId="6" pageNumber="143">Male.</emphasis>
Male tergite VIII (
<figureCitation id="46C6B3C9FFCABB5AFBA406CBFB7C2B6F" box="[1146,1237,766,789]" captionStart="Fig" captionStartId="5.[152,183,2030,2049]" captionTargetBox="[243,1346,599,1996]" captionTargetId="figure-220@5.[230,1357,585,2011]" captionTargetPageId="5" captionText="Fig. 4. Eubadura hokkaidensis sp. nov. A male tergite VIII; B male sternite VIII; C female tergite VIII; D female sternite VIII; E spermatheca; F median lobe, ventral view; G median lobe, lateral view; H paramere. Scale bars: 0. 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7161329" httpUri="https://zenodo.org/record/7161329/files/figure.png" pageId="6" pageNumber="143">Fig. 4A</figureCitation>
) densely covered with setae, with 4 pairs of macrosetae. Male sternite VIII (
<figureCitation id="46C6B3C9FFCABB5AFC9A070BFC302B2F" box="[836,921,830,853]" captionStart="Fig" captionStartId="5.[152,183,2030,2049]" captionTargetBox="[243,1346,599,1996]" captionTargetId="figure-220@5.[230,1357,585,2011]" captionTargetPageId="5" captionText="Fig. 4. Eubadura hokkaidensis sp. nov. A male tergite VIII; B male sternite VIII; C female tergite VIII; D female sternite VIII; E spermatheca; F median lobe, ventral view; G median lobe, lateral view; H paramere. Scale bars: 0. 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7161329" httpUri="https://zenodo.org/record/7161329/files/figure.png" pageId="6" pageNumber="143">Fig. 4B</figureCitation>
) rounded on posterior margin, with 8 pairs of macrosetae. Median lobe of aedeagus as in
<figureCitation id="46C6B3C9FFCABB5AFAD7076BFAD42B0F" box="[1289,1405,862,885]" captionStart="Fig" captionStartId="5.[152,183,2030,2049]" captionTargetBox="[243,1346,599,1996]" captionTargetId="figure-220@5.[230,1357,585,2011]" captionTargetPageId="5" captionText="Fig. 4. Eubadura hokkaidensis sp. nov. A male tergite VIII; B male sternite VIII; C female tergite VIII; D female sternite VIII; E spermatheca; F median lobe, ventral view; G median lobe, lateral view; H paramere. Scale bars: 0. 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7161329" httpUri="https://zenodo.org/record/7161329/files/figure.png" pageId="6" pageNumber="143">Figs 4FG</figureCitation>
; in lateral view (
<figureCitation id="46C6B3C9FFCABB5AFC19074BFBB32BEF" box="[967,1050,894,917]" captionStart="Fig" captionStartId="5.[152,183,2030,2049]" captionTargetBox="[243,1346,599,1996]" captionTargetId="figure-220@5.[230,1357,585,2011]" captionTargetPageId="5" captionText="Fig. 4. Eubadura hokkaidensis sp. nov. A male tergite VIII; B male sternite VIII; C female tergite VIII; D female sternite VIII; E spermatheca; F median lobe, ventral view; G median lobe, lateral view; H paramere. Scale bars: 0. 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7161329" httpUri="https://zenodo.org/record/7161329/files/figure.png" pageId="6" pageNumber="143">Fig. 4G</figureCitation>
) apical lobe bent paramerally, much narrowed subapically, sinuate abparamerally and acutely narrowed apically. Paramere (
<figureCitation id="46C6B3C9FFCABB5AFBA6078BFB632BAF" box="[1144,1226,958,981]" captionStart="Fig" captionStartId="5.[152,183,2030,2049]" captionTargetBox="[243,1346,599,1996]" captionTargetId="figure-220@5.[230,1357,585,2011]" captionTargetPageId="5" captionText="Fig. 4. Eubadura hokkaidensis sp. nov. A male tergite VIII; B male sternite VIII; C female tergite VIII; D female sternite VIII; E spermatheca; F median lobe, ventral view; G median lobe, lateral view; H paramere. Scale bars: 0. 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7161329" httpUri="https://zenodo.org/record/7161329/files/figure.png" pageId="6" pageNumber="143">Fig. 4H</figureCitation>
) with apical lobe of paramerite narrowed slightly.
</paragraph>
<paragraph id="DE42AF4CFFCABB5BFC8407C8FD6B292F" blockId="6.[828,1437,190,1077]" lastBlockId="7.[151,760,190,629]" lastPageId="7" lastPageNumber="144" pageId="6" pageNumber="143">
<emphasis id="EC89735EFFCABB5AFC8407C8FC1A2C6F" bold="true" box="[858,947,1021,1045]" italics="true" pageId="6" pageNumber="143">Female.</emphasis>
Female tergite VIII (
<figureCitation id="46C6B3C9FFCABB5AFB4C07CBFB4A2C6F" box="[1170,1251,1022,1045]" captionStart="Fig" captionStartId="5.[152,183,2030,2049]" captionTargetBox="[243,1346,599,1996]" captionTargetId="figure-220@5.[230,1357,585,2011]" captionTargetPageId="5" captionText="Fig. 4. Eubadura hokkaidensis sp. nov. A male tergite VIII; B male sternite VIII; C female tergite VIII; D female sternite VIII; E spermatheca; F median lobe, ventral view; G median lobe, lateral view; H paramere. Scale bars: 0. 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7161329" httpUri="https://zenodo.org/record/7161329/files/figure.png" pageId="6" pageNumber="143">Fig. 4C</figureCitation>
) densely covered with setae, with 4 macrosetae on each side; female sternite VIII (
<figureCitation id="46C6B3C9FFCBBB5BFF0C048BFE8D28AF" box="[210,292,190,213]" captionStart="Fig" captionStartId="5.[152,183,2030,2049]" captionTargetBox="[243,1346,599,1996]" captionTargetId="figure-220@5.[230,1357,585,2011]" captionTargetPageId="5" captionText="Fig. 4. Eubadura hokkaidensis sp. nov. A male tergite VIII; B male sternite VIII; C female tergite VIII; D female sternite VIII; E spermatheca; F median lobe, ventral view; G median lobe, lateral view; H paramere. Scale bars: 0. 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7161329" httpUri="https://zenodo.org/record/7161329/files/figure.png" pageId="7" pageNumber="144">Fig. 4D</figureCitation>
) rounded on posterior margin, with 6 macrosetae on each side. Spermatheca (
<figureCitation id="46C6B3C9FFCBBB5BFE3304EBFD92288F" box="[493,571,222,245]" captionStart="Fig" captionStartId="5.[152,183,2030,2049]" captionTargetBox="[243,1346,599,1996]" captionTargetId="figure-220@5.[230,1357,585,2011]" captionTargetPageId="5" captionText="Fig. 4. Eubadura hokkaidensis sp. nov. A male tergite VIII; B male sternite VIII; C female tergite VIII; D female sternite VIII; E spermatheca; F median lobe, ventral view; G median lobe, lateral view; H paramere. Scale bars: 0. 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7161329" httpUri="https://zenodo.org/record/7161329/files/figure.png" pageId="7" pageNumber="144">Fig. 4E</figureCitation>
) with basal part of stem once curved at apical and twice curved around base; capsule dilated apically, about 1/4 as long as stem, densely covered with wrinkles on internal side of apical 1/3.
</paragraph>
<caption id="8A82FFC4FFCABB5AFF4803DEFCB32062" ID-DOI="http://doi.org/10.5281/zenodo.7161331" ID-Zenodo-Dep="7161331" httpUri="https://zenodo.org/record/7161331/files/figure.png" pageId="6" pageNumber="143" startId="6.[150,181,2027,2046]" targetBox="[203,1385,1095,2010]" targetPageId="6">
<paragraph id="DE42AF4CFFCABB5AFF4803DEFCB32062" blockId="6.[150,1437,2027,2072]" pageId="6" pageNumber="143">
Fig. 5. Known localities of
<taxonomicName id="19FDD4CFFFCABB5AFEA503DEFE502F84" box="[379,505,2027,2046]" class="Insecta" family="Staphylinidae" genus="Eubadura" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="143" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EC89735EFFCABB5AFEA503DEFE792F84" box="[379,464,2027,2046]" italics="true" pageId="6" pageNumber="143">Eubadura</emphasis>
spp.
</taxonomicName>
Black circle
<taxonomicName id="19FDD4CFFFCABB5AFDAD03DEFCF32F84" authority="(Sawada, 1990)" authorityName="Tasaku &amp; Maruyama" authorityYear="2022" baseAuthorityName="Sawada" baseAuthorityYear="1990" box="[627,858,2027,2046]" class="Insecta" family="Staphylinidae" genus="Eubadura" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="143" phylum="Arthropoda" rank="species" species="akiensis">
<emphasis id="EC89735EFFCABB5AFDAD03DEFD662F84" box="[627,719,2027,2046]" italics="true" pageId="6" pageNumber="143">E. akiensis</emphasis>
(
<bibRefCitation id="BA6CD2BDFFCABB5AFD0703DEFCFD2F84" author="SAWADA K." box="[729,852,2027,2046]" pageId="6" pageNumber="143" pagination="273 - 307" refId="ref5181" refString="SAWADA K. 1990: New Species of Aleocharinae from Japan II (Coleoptera: Staphylinidae). Contributions from the Biological Laboratory, Kyoto University 27 (3): 273 - 307." type="journal article" year="1990">Sawada, 1990</bibRefCitation>
)
</taxonomicName>
; black triangle
<taxonomicName id="19FDD4CFFFCABB5AFC3603DEFF622062" authority="(Maruyama, Klimaszewski &amp; Gusarov, 2008)" baseAuthorityName="Maruyama, Klimaszewski &amp; Gusarov" baseAuthorityYear="2008" class="Insecta" family="Staphylinidae" genus="Osakatheta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="143" phylum="Arthropoda" rank="species" species="yasukoae">
<emphasis id="EC89735EFFCABB5AFC3603DEFBE42F84" box="[1000,1101,2027,2046]" italics="true" pageId="6" pageNumber="143">E. yasukoae</emphasis>
(
<bibRefCitation id="BA6CD2BDFFCABB5AFB8903DEFF6C2062" author="MARUYAMA M. &amp; KLIMASZEWSKI J. &amp; GUSAROV V." pageId="6" pageNumber="143" pagination="39 - 50" refId="ref5066" refString="MARUYAMA M., KLIMASZEWSKI J. &amp; GUSAROV V. 2008: Osakatheta yasukoae, a new intertidal genus and species of athetine rove beetles (Coleoptera, Staphylinidae, Aleocharinae) from Japan. Zootaxa 1683: 39 - 50." type="journal article" year="2008">Maruyama, Klimaszewski &amp; Gusarov, 2008</bibRefCitation>
)
</taxonomicName>
; black square
<taxonomicName id="19FDD4CFFFCABB5AFE8B0C30FE3B2062" authority="Tasaku &amp; Maruyama, 2022" authorityName="Tasaku &amp; Maruyama" authorityYear="2022" box="[341,402,2053,2072]" class="Insecta" family="Staphylinidae" genus="Eubadura" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="143" phylum="Arthropoda" rank="species" species="onoi" status="sp. nov.">
<emphasis id="EC89735EFFCABB5AFE8B0C30FE3B2062" box="[341,402,2053,2072]" italics="true" pageId="6" pageNumber="143">E. onoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7BACE25FFCABB5AFE460C30FE742062" box="[408,477,2053,2072]" pageId="6" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
; black star
<taxonomicName id="19FDD4CFFFCABB5AFD920C30FD7B2062" authority="Tasaku &amp; Maruyama, 2022" authorityName="Tasaku &amp; Maruyama" authorityYear="2022" box="[588,722,2053,2072]" class="Insecta" family="Staphylinidae" genus="Eubadura" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="143" phylum="Arthropoda" rank="species" species="hokkaidensis" status="sp. nov.">
<emphasis id="EC89735EFFCABB5AFD920C30FD7B2062" box="[588,722,2053,2072]" italics="true" pageId="6" pageNumber="143">E. hokkaidensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7BACE25FFCABB5AFD090C30FCB32062" box="[727,794,2053,2072]" pageId="6" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="DE42AF4CFFCBBB5BFF6B056BFDC729CF" blockId="7.[151,760,190,629]" pageId="7" pageNumber="144">
<emphasis id="EC89735EFFCBBB5BFF6B056BFEFF290F" bold="true" box="[181,342,350,373]" italics="true" pageId="7" pageNumber="144">Measurements</emphasis>
(mm). Body length ca. 1.902.20; fore body length ca. 1.01.1; head width 0.360.38; pronotal length 0.320.38; pronotal width 0.380.40.
</paragraph>
</subSubSection>
<subSubSection id="96E7FCC7FFCBBB5BFF49058BFD66298F" pageId="7" pageNumber="144" type="diagnosis">
<paragraph id="DE42AF4CFFCBBB5BFF49058BFD66298F" blockId="7.[151,760,190,629]" pageId="7" pageNumber="144">
<emphasis id="EC89735EFFCBBB5BFF49058BFE2629AF" bold="true" box="[151,399,446,469]" pageId="7" pageNumber="144">Differential diagnosis.</emphasis>
This species can be distinguished from other
<taxonomicName id="19FDD4CFFFCBBB5BFED105EBFE64298F" box="[271,461,478,501]" pageId="7" pageNumber="144">
<emphasis id="EC89735EFFCBBB5BFED105EBFED3298F" box="[271,378,478,501]" italics="true" pageId="7" pageNumber="144">Eubadura</emphasis>
species
</taxonomicName>
by its black body color.
</paragraph>
</subSubSection>
<subSubSection id="96E7FCC7FFCBBB5BFF4905CBFD672A6F" box="[151,718,510,533]" pageId="7" pageNumber="144" type="etymology">
<paragraph id="DE42AF4CFFCBBB5BFF4905CBFD672A6F" blockId="7.[151,760,190,629]" box="[151,718,510,533]" pageId="7" pageNumber="144">
<emphasis id="EC89735EFFCBBB5BFF4905CBFEB12A6F" bold="true" box="[151,280,510,533]" pageId="7" pageNumber="144">Etymology.</emphasis>
Named after the
<typeStatus id="014611EEFFCBBB5BFE0E05CBFE572A6F" box="[464,510,510,533]" pageId="7" pageNumber="144">type</typeStatus>
locality. Adjective.
</paragraph>
</subSubSection>
<subSubSection id="96E7FCC7FFCBBB5BFF49062BFDF12A2F" pageId="7" pageNumber="144" type="biology_ecology">
<paragraph id="DE42AF4CFFCBBB5BFF49062BFDF12A2F" blockId="7.[151,760,190,629]" pageId="7" pageNumber="144">
<emphasis id="EC89735EFFCBBB5BFF49062BFF592A4F" bold="true" box="[151,240,542,565]" pageId="7" pageNumber="144">Biology.</emphasis>
The beetles were found under supratidal driftwood or seaweeds on sandy and gravel beaches.
</paragraph>
</subSubSection>
<subSubSection id="96E7FCC7FFCBBB5BFF49066BFE422A0F" box="[151,491,606,629]" pageId="7" pageNumber="144" type="distribution">
<paragraph id="DE42AF4CFFCBBB5BFF49066BFE422A0F" blockId="7.[151,760,190,629]" box="[151,491,606,629]" pageId="7" pageNumber="144">
<emphasis id="EC89735EFFCBBB5BFF49066BFE832A0F" bold="true" box="[151,298,606,629]" pageId="7" pageNumber="144">Distribution.</emphasis>
<collectingCountry id="A6EAEFDCFFCBBB5BFEEE066BFED82A0F" box="[304,369,606,629]" name="Japan" pageId="7" pageNumber="144">Japan</collectingCountry>
:
<collectingRegion id="1C3961AEFFCBBB5BFEA5066BFE412A0F" box="[379,488,606,629]" country="Japan" name="Hokkaido" pageId="7" pageNumber="144">Hokkaidô</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>