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<document ID-DOI="http://dx.doi.org/10.3897/dez.69.85584" ID-Pensoft-Pub="1860-1324-2-139" ID-Pensoft-UUID="03EFF5CBC2CF53CFA27D350B268657C3" ID-ZooBank="73B92B049B454237B81497A21DB59468" ModsDocID="1860-1324-69-2-139" checkinTime="1658439162963" checkinUser="pensoft" docAuthor="Kobor, Peter" docDate="2022" docId="B8A2FE46238E5AB58A32008A09AE9022" docLanguage="en" docName="DeutEntomolZeit 69(2): 139-150" docOrigin="Deutsche Entomologische Zeitschrift 69 (2)" docPubDate="2022-07-21" docSource="http://dx.doi.org/10.3897/dez.69.85584" docTitle="Umbrageocoris woodwardi Kóbor 2022, comb. nov." docType="treatment" docVersion="1" id="03EFF5CBC2CF53CFA27D350B268657C3" lastPageNumber="139" masterDocId="03EFF5CBC2CF53CFA27D350B268657C3" masterDocTitle="A redefinition of Umbrageocoris with new species and new combinations (Heteroptera, Lygaeoidea, Geocoridae)" masterLastPageNumber="150" masterPageNumber="139" pageNumber="139" updateTime="1658439162963" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A redefinition of Umbrageocoris with new species and new combinations (Heteroptera, Lygaeoidea, Geocoridae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kobor, Peter</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0266-7303</mods:nameIdentifier>
<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Eoetvoes Lorand Research Network, 15 Herman Otto street, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">kobor.peter@atk.hu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Deutsche Entomologische Zeitschrift</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-07-21</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>69</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>139</mods:start>
<mods:end>150</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/dez.69.85584</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/dez.69.85584</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-1324-2-139</mods:identifier>
<mods:identifier type="ZooBank">73B92B049B454237B81497A21DB59468</mods:identifier>
<mods:identifier type="Pensoft-UUID">03EFF5CBC2CF53CFA27D350B268657C3</mods:identifier>
</mods:mods>
<subSection pageId="0" pageNumber="139" type="results">
<treatment LSID="urn:lsid:plazi:treatment:B8A2FE46238E5AB58A32008A09AE9022" httpUri="http://treatment.plazi.org/id/B8A2FE46238E5AB58A32008A09AE9022" lastPageNumber="139" pageId="0" pageNumber="139">
<subSubSection pageId="0" pageNumber="139" type="nomenclature">
<paragraph pageId="0" pageNumber="139">
<taxonomicName LSID="B8A2FE46-238E-5AB5-8A32-008A09AE9022" authority="(Malipatil, 1994)" authorityName="Kóbor" authorityYear="2022" baseAuthorityName="Malipatil" baseAuthorityYear="1994" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Umbrageocoris woodwardi" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="woodwardi" status="comb. nov.">Umbrageocoris woodwardi (Malipatil, 1994)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="139">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="description">
<paragraph pageId="0" pageNumber="139">
<figureCitation captionStart="Figures 5–10" captionStartId="F2" captionText="Figures 5 - 10. Exoskeletal characters of Umbrageocoris: peritreme of metathoracic scent efferent apparatus of U. kondorosyi (5), U. maai maai (6), U. boonei sp. nov. (7), U. malipatili sp. nov. (8), U. elegantulus (9), and U. woodwardi (10) (images not to scale)." figureDoi="10.3897/dez.69.85584.figures5-10" httpUri="https://binary.pensoft.net/fig/718125" pageId="0" pageNumber="139">Figs 10</figureCitation>
<figureCitation captionStart="Figures 19–24" captionStartId="F5" captionText="Figures 19 - 24. Dorsal habitus of Umbrageocoris species: 19. U. kondorosyi (holotype, HNHM), 20. U. maai maai (holotype, BPBM), 21. U. maai timorensis (holotype, BPBM), 22. U. boonei sp. nov. (holotype, BPBM), 23. U. malipatili sp. nov. (holotype, MNHN), 24. U. woodwardi (SEMC) (scale bar = 1 mm for all images)." figureDoi="10.3897/dez.69.85584.figures19-24" httpUri="https://binary.pensoft.net/fig/718128" pageId="0" pageNumber="139">, 24</figureCitation>
<figureCitation captionStart="Figures 27–28" captionStartId="F7" captionText="Figures 27 - 28. Distribution map of Umbrageocoris species: 27. Indomalayan species (U. maai ssp. and U. boonei sp. nov.), 28. Australasian species (U. elegantulus, U. kondorosyi, U. malipatili sp. nov. and U. woodwardi)." figureDoi="10.3897/dez.69.85584.figures27-28" httpUri="https://binary.pensoft.net/fig/718130" pageId="0" pageNumber="139">, 28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="materials_examined">
<paragraph pageId="0" pageNumber="139">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="139">
<materialsCitation collectingDate="1959-10-07" collectingDateMax="1959-10-14" collectingDateMin="1959-10-07" collectorName="T. C. Maa" country="Guinea" elevation="2100" location="New" municipality="Moife" specimenCount="1">
2 f (BPBM): &quot;
<location LSID="urn:lsid:plazi:treatment:B8A2FE46238E5AB58A32008A09AE9022:9C6425CA518CFB90CE04A07D1DEC8B20" country="Guinea" municipality="Moife" name="New">New</location>
<collectingCountry name="Guinea-Bissau">Guinea</collectingCountry>
: NE. /
<collectingMunicipality>Moife</collectingMunicipality>
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.1" unit="m" value="2100.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.1" unit="m" value="2100.0">2100 m</elevation>
</quantity>
. /
<collectingDate value="1959-10-07" valueMax="1959-10-14" valueMin="1959-10-07">7-14. X. 1959</collectingDate>
// Swept in / clearing //
<collectorName>T. C. Maa</collectorName>
/ Collector / BISHOP&quot;; 1 f BPBM: &quot;NE. / Moife
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.1" unit="m" value="2100.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.1" unit="m" value="2100.0">2100 m</elevation>
</quantity>
. /
<locationDeviation location="Okapu">
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.5" unit="km" value="15.0">15km</quantity>
NW of Okapu
</locationDeviation>
/
<collectingDate value="1959-10-07" valueMax="1959-10-14" valueMin="1959-10-07">7-14. X. 1959</collectingDate>
//
<collectorName>T. C. Maa</collectorName>
/ Collector / BISHOP&quot;
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="139">
Additions and corrections to the description of
<bibRefCitation DOI="https://doi.org/10.1071/IT9940299" author="Malipatil, MB" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="139" pagination="299 - 327" refId="B10" refString="Malipatil, MB, 1994. Revision of Australian Geocoris Fallen and Stylogeocoris Montandon (Heteroptera: Lygaeidae: Geocorinae). Invertebrate Systematics 8 (2): 299 - 327, DOI: https://doi.org/10.1071/IT9940299" title="Revision of Australian Geocoris Fallen and Stylogeocoris Montandon (Heteroptera: Lygaeidae: Geocorinae)." url="https://doi.org/10.1071/IT9940299" volume="8" year="1994">Malipatil (1994)</bibRefCitation>
. Head. Median longitudinal furrow of vertex minute, hardly surpassing base of clypeus. Labial trough cup-shaped with suture short, hardly reaching middle of venter of head. Thorax. Peritreme bulbous, dorsal flange not indented.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="diagnosis">
<paragraph pageId="0" pageNumber="139">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="139">
<taxonomicName class="Insecta" genus="U." kingdom="Animalia" lsidName="U. woodwardi" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="woodwardi">
<emphasis italics="true" pageId="0" pageNumber="139">U. woodwardi</emphasis>
</taxonomicName>
resembles
<taxonomicName class="Insecta" genus="U." kingdom="Animalia" lsidName="U. elengantulus" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="elengantulus">
<emphasis italics="true" pageId="0" pageNumber="139">U. elengantulus</emphasis>
</taxonomicName>
in general aspect, but it can be distinguished from the latter species based on the combination of the following characters: vertex with dark marking only basally (with more extensive dark markings in
<taxonomicName class="Insecta" genus="U." kingdom="Animalia" lsidName="U. elegantulus" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis italics="true" pageId="0" pageNumber="139">U. elegantulus</emphasis>
</taxonomicName>
); dorsal flange of peritreme simple (indented in
<taxonomicName class="Insecta" genus="U." kingdom="Animalia" lsidName="U. elegantulus" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis italics="true" pageId="0" pageNumber="139">U. elegantulus</emphasis>
</taxonomicName>
).
</paragraph>
<caption doi="10.3897/dez.69.85584.figures25-26" httpUri="https://binary.pensoft.net/fig/718129" pageId="0" pageNumber="139" start="Figures 25–26" startId="F6">
<paragraph pageId="0" pageNumber="139">
<emphasis bold="true" pageId="0" pageNumber="139">Figures 25-26.</emphasis>
Lectotype of
<taxonomicName authorityName="Distant" authorityYear="1904" class="Insecta" family="Geocoridae" genus="Geocoris" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Geocoris elegantulus" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis italics="true" pageId="0" pageNumber="139">Geocoris elegantulus</emphasis>
</taxonomicName>
Distant, 1904 (BMNH):
<emphasis bold="true" pageId="0" pageNumber="139">25.</emphasis>
Dorsal habitus (scale bar = 1 mm),
<emphasis bold="true" pageId="0" pageNumber="139">26.</emphasis>
Labels (image not to scale).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="distribution">
<paragraph pageId="0" pageNumber="139">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="139">
Most of the published records of the species are from Western Australia, where the species inhabits Brigalow tropical savanna and Eastern Australian temperate forest habitats (
<bibRefCitation DOI="https://doi.org/10.1071/IT9940299" author="Malipatil, MB" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="139" pagination="299 - 327" refId="B10" refString="Malipatil, MB, 1994. Revision of Australian Geocoris Fallen and Stylogeocoris Montandon (Heteroptera: Lygaeidae: Geocorinae). Invertebrate Systematics 8 (2): 299 - 327, DOI: https://doi.org/10.1071/IT9940299" title="Revision of Australian Geocoris Fallen and Stylogeocoris Montandon (Heteroptera: Lygaeidae: Geocorinae)." url="https://doi.org/10.1071/IT9940299" volume="8" year="1994">Malipatil 1994</bibRefCitation>
). In the material deposited in the BPBM three specimens from the Central Range montane rain forests of New Guinea are present.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="139" type="remarks">
<paragraph pageId="0" pageNumber="139">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="139">
Based on the specimens available as well as the original description and illustrations provided by
<bibRefCitation DOI="https://doi.org/10.1071/IT9940299" author="Malipatil, MB" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="139" pagination="299 - 327" refId="B10" refString="Malipatil, MB, 1994. Revision of Australian Geocoris Fallen and Stylogeocoris Montandon (Heteroptera: Lygaeidae: Geocorinae). Invertebrate Systematics 8 (2): 299 - 327, DOI: https://doi.org/10.1071/IT9940299" title="Revision of Australian Geocoris Fallen and Stylogeocoris Montandon (Heteroptera: Lygaeidae: Geocorinae)." url="https://doi.org/10.1071/IT9940299" volume="8" year="1994">Malipatil (1994)</bibRefCitation>
the species possesses a combination of characters diagnostic of
<taxonomicName authorityName="Kobor" authorityYear="2019" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Umbrageocoris" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="139">Umbrageocoris</emphasis>
</taxonomicName>
, including the presence of a median longitudinal furrow on vertex, a cup- or Y-shaped labial trough, and the arrangement of sutures of abdominal tergites 4/5 and 5/6. The following new combination is accordingly proposed:
<taxonomicName authorityName="Kóbor" authorityYear="2022" baseAuthorityName="Malipatil" baseAuthorityYear="1994" class="Insecta" family="Geocoridae" genus="Umbrageocoris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Umbrageocoris woodwardi" order="Hemiptera" pageId="0" pageNumber="139" phylum="Arthropoda" rank="species" species="woodwardi">
<emphasis italics="true" pageId="0" pageNumber="139">Umbrageocoris woodwardi</emphasis>
</taxonomicName>
(Malipatil, 1994), comb. nov.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>
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