584 lines
71 KiB
XML
584 lines
71 KiB
XML
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<mods:title id="E929C2C747FD3A37DDC39FAD5AB050DF">Two newly recognized species of Hemidactylus (Squamata, Gekkonidae) from the Arabian Peninsula and Sinai, Egypt</mods:title>
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<mods:namePart id="4D56B8BB263FCC66581BE0E21623C743">Smid, Jiri</mods:namePart>
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<mods:namePart id="0063A5B15553B1DC1C4652563C565582">Moravec, Jiri</mods:namePart>
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<mods:namePart id="C0B9A4135B1FA56E7412215D38418175">Kratochvil, Lukas</mods:namePart>
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<mods:namePart id="B93B678C89A7A9C0751B146E8BADA8F6">Nasher, Abdul Karim</mods:namePart>
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<mods:namePart id="9D6A215E911D5F0CF17B451D69D91D8A">Busais, Salem M.</mods:namePart>
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<mods:namePart id="34A5F26CE2847C1D90192C55D1A767CE">Shobrak, Mohammed Y.</mods:namePart>
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<mods:date id="9A1930510BDF991A88D86942D5F458AB">2013</mods:date>
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<mods:number id="FE981D2083FF1B8146485C1E21C1D53F">355</mods:number>
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<mods:start id="F60ACCE41FB97759C11908740F94366B">79</mods:start>
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<mods:end id="2E7AF7526E117B0BA9DF0900107B28E3">107</mods:end>
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<mods:url id="F4357719AFAE2C2BDF87B1389DF5B1FC">http://dx.doi.org/10.3897/zookeys.355.6190</mods:url>
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<mods:classification id="C801E35C1FD91FAA41A4975EF5152659">journal article</mods:classification>
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<mods:identifier id="A6592B7161C61261A6D0AB30A47C1B8B" type="DOI">http://dx.doi.org/10.3897/zookeys.355.6190</mods:identifier>
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<treatment id="B82F5E163694DF25C620F560CF65E7CE" ID-GBIF-Taxon="152049795" LSID="urn:lsid:plazi:treatment:B82F5E163694DF25C620F560CF65E7CE" httpUri="http://treatment.plazi.org/id/B82F5E163694DF25C620F560CF65E7CE" lastPageId="12" lastPageNumber="90" pageId="5" pageNumber="84">
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<subSubSection id="17F488F8529DD0FBE15C0C415CAD792C" pageId="5" pageNumber="84" type="nomenclature">
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<paragraph id="2DC1306EE67361F30A27C495E9475E62" pageId="5" pageNumber="84">
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Redescription of
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<taxonomicName id="07277688ABE86E40D324ED1E320FA5B1" ID-CoL="6LQ2R" LSID="http://species-id.net/wiki/Hemidactylus_granosus" authority="Heyden, 1827" authorityName="Heyden" authorityYear="1827" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="5" pageNumber="84" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus Heyden, 1827</taxonomicName>
|
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Figs 4, 5
|
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</paragraph>
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</subSubSection>
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<subSubSection id="E7082C92FB61AA8DFAEDEF49BF79A299" pageId="5" pageNumber="84" type="reference_group">
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<paragraph id="32CF22465DF4374CDFB6B518282D2FD4" pageId="5" pageNumber="84">
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<taxonomicName id="5CE997C285D6A0A4CDA49621A40C1659" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="5" pageNumber="84" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
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Heyden, 1827: p. 17; tab. 5, fig. 1. Lectotype SMF 8723 designated by
|
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<bibRefCitation id="4CA333C86CAB9C5505488E13B06DE880" author="Mertens, R" journalOrPublisher="Senckenbergiana Biologica" pageId="19" pageNumber="98" pagination="1 - 106" title="Die herpetologische Sektion des Natur-Museums und Forschungs-Institutes Senckenberg in Frankfurt a. M. nebst einem Verzeichnis ihrer Typen." volume="48" year="1967">Mertens (1967)</bibRefCitation>
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; collected by E.
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<normalizedToken id="722447204CD1AC1DDE4B116F8829FE3A" originalValue="Rüppell">Rueppell</normalizedToken>
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1827.
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</paragraph>
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<paragraph id="1D746BA29166EAED6E45A98886D35D55" pageId="5" pageNumber="84">
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||
<taxonomicName id="C95F27C5C662B3AFEF6E2FF67A5EFC58" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus turcicus" pageId="5" pageNumber="84" phylum="Chordata" rank="species" species="turcicus">Hemidactylus turcicus</taxonomicName>
|
||
(Linnaeus, 1758) -
|
||
<bibRefCitation id="6C071242EF67D37E42145DEACFAA8ADD" author="Boettger, O" journalOrPublisher="Frankfurt am Main" pageId="18" pageNumber="97" title="Katalog der Reptilien-Sammlung im Museum der Senckenbergischen Naturforschenden Gesellschaft in Frankfurt am Main." year="1893">Boettger (1893</bibRefCitation>
|
||
: 29; part.);
|
||
<bibRefCitation id="B377AC1394AC0BE775886C9E942EC7FB" author="Anderson, J" journalOrPublisher="Bernard Quaritch, London" pageId="18" pageNumber="97" title="Zoology of Egypt. Vol. I. Reptilia and Batrachia." year="1898">Anderson (1898</bibRefCitation>
|
||
: 80; part.);
|
||
<bibRefCitation id="539E86366FC23F3897CFE3DC39FF4410" pageId="5" pageNumber="84">Salvador (1981</bibRefCitation>
|
||
: 84; part.);
|
||
<bibRefCitation id="91823B9C578026B13892733AFD1660D9" author="Baha El Din, S" journalOrPublisher="The American University in Cairo Press, Cairo - New York" pageId="18" pageNumber="97" title="A guide to the Reptiles and Amphibians of Egypt." year="2006">Baha El Din (2006</bibRefCitation>
|
||
: 66; part.).
|
||
</paragraph>
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||
<paragraph id="3B1F08F22FD92FC1BB37CFAC56FFC71B" pageId="5" pageNumber="84">
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||
<taxonomicName id="A1041F37C2A86D61AD3352630603DEE9" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus turcicus subsp. turcicus" pageId="5" pageNumber="84" phylum="Chordata" rank="subSpecies" species="turcicus" subSpecies="turcicus">Hemidactylus turcicus turcicus</taxonomicName>
|
||
(Linnaeus, 1758) -
|
||
<bibRefCitation id="669F0E26A676CA7C959756056CC01770" author="Loveridge, A" journalOrPublisher="Bulletin of The Museum of Comparative Zoology" pageId="19" pageNumber="98" pagination="1 - 469" title="Revision of the african lizards of the family Gekkonidae." volume="98" year="1947">Loveridge (1947</bibRefCitation>
|
||
: 143; part.);
|
||
<bibRefCitation id="3E3BE37F9829EA6BF2C28F384F3D0B64" author="Mertens, R" journalOrPublisher="Waldemar Kramer, Frankfurt am Main" pageId="19" pageNumber="98" title="Die Amphibien und Reptilien Europas." year="1960">Mertens and Wermuth (1960</bibRefCitation>
|
||
: 79; part.);
|
||
<bibRefCitation id="E5B4DC13ECB364885B56DD813269CEFD" author="Baha El Din, SM" journalOrPublisher="Zoology in the Middle East" pageId="18" pageNumber="97" pagination="27 - 34" title="An overview of Egyptian species of Hemidactylus (Gekkonidae), with the description of a new species from the high mountains of South Sinai." url="10.1080/09397140.2005.10638078" volume="34" year="2005">Baha El Din (2005</bibRefCitation>
|
||
: 19; part.);
|
||
<bibRefCitation id="F8E3C3B2FFAB47D4F7535175E316F482" author="Mertens, R" journalOrPublisher="Senckenbergiana Biologica" pageId="19" pageNumber="98" pagination="1 - 106" title="Die herpetologische Sektion des Natur-Museums und Forschungs-Institutes Senckenberg in Frankfurt a. M. nebst einem Verzeichnis ihrer Typen." volume="48" year="1967">Mertens (1967</bibRefCitation>
|
||
: 55).
|
||
</paragraph>
|
||
<paragraph id="6EB760E7C3E113B3C6776545E95F146E" pageId="5" pageNumber="84">
|
||
<taxonomicName id="D6058988DAE592B568BFA492ED8EE62D" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus verrucosus" pageId="5" pageNumber="84" phylum="Chordata" rank="species" species="verrucosus">Hemidactylus verrucosus</taxonomicName>
|
||
(Cuvier, 1829 [corr.
|
||
<taxonomicName id="A711BD1AD879447C5E2B40A0536A3660" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus verrucosus" pageId="5" pageNumber="84" phylum="Chordata" rank="species" species="verrucosus">Hemidactylus verrucosus</taxonomicName>
|
||
Gray, 1831]) -
|
||
<bibRefCitation id="3EC1620D970743AEF2AF4DD8910347C2" author="Rueppell, E" journalOrPublisher="Museum Senckenberg, Frankfurt am Main" pageId="19" pageNumber="98" pagination="293 - 316" title="Verzeichniss der in dem Museum der Senckenbergischen naturforschenden Gesellschaft aufgestellten Sammlungen. Dritte Abteilung: Amphibien." year="1845">
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<normalizedToken id="98801A59D9E095A608C0B861BAD8D395" originalValue="Rüppell">Rueppell</normalizedToken>
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(1845
|
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</bibRefCitation>
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: 300; part.).
|
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</paragraph>
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<paragraph id="588902C5C1FDD5054B58A79E05E66B9C" pageId="5" pageNumber="84">
|
||
<taxonomicName id="1C12277DFB240C6549A7B77E57D6DEC9" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus" pageId="5" pageNumber="84" phylum="Chordata" rank="genus">Hemidactylus</taxonomicName>
|
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sp. 1 -
|
||
<bibRefCitation id="1388A9FB3FE935143A209CA699E73855" author="Moravec, J" journalOrPublisher="Zootaxa" pageId="19" pageNumber="98" pagination="21 - 38" title="High genetic differentiation within the Hemidactylus turcicus complex (Reptilia: Gekkonidae) in the Levant, with comments on the phylogeny and systematics of the genus." volume="2894" year="2011">Moravec et al. (2011</bibRefCitation>
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: 24);
|
||
<bibRefCitation id="9FA3A01A739CE9FC830D70C1C319F6AC" author="Carranza, S" journalOrPublisher="Zootaxa" pageId="18" pageNumber="97" pagination="1 - 95" title="A review of the geckos of the genus Hemidactylus (Squamata: Gekkonidae) from Oman based on morphology, mitochondrial and nuclear data, with descriptions of eight new species." volume="3378" year="2012">Carranza and Arnold (2012</bibRefCitation>
|
||
: 17);
|
||
<bibRefCitation id="BE653A26339C0B07C872A7143964F85E" author="Smid, J" journalOrPublisher="PLoS ONE" pageId="19" pageNumber="98" title="Out of Arabia: A Complex Biogeographic History of Multiple Vicariance and Dispersal Events in the Gecko Genus Hemidactylus (Reptilia: Gekkonidae)." url="10.1371/journal.pone.0064018" volume="8" year="2013">
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<normalizedToken id="202C9306C057AE63072B84478312AFF9" originalValue="Šmíd">Smid</normalizedToken>
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et al. (2013
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</bibRefCitation>
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||
: 3).
|
||
</paragraph>
|
||
<paragraph id="398C320C5A85FC8D3A38C3007636ED76" pageId="5" pageNumber="84">
|
||
<taxonomicName id="91B1A85FD73FDDD3018DE2CF09660E65" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="5" pageNumber="84" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
Terra typica (
|
||
<bibRefCitation id="3B268C34BD15A3A50275DBD67F9BA587" author="Heyden, CHGv" editor="Rueppell, E" journalOrPublisher="Heinrich Ludwig Broenner, Frankfurt am Main" pageId="19" pageNumber="98" pagination="1 - 24" title="Reptilien." year="1827">Heyden 1827</bibRefCitation>
|
||
): "Egypten, Arabien, und Abyssinien".
|
||
</paragraph>
|
||
<paragraph id="017EEAC9B1B043FC668EE4EAF12A382F" pageId="5" pageNumber="84">
|
||
<taxonomicName id="A6D1C40C1E0F5FBDA6110C313E635C55" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="5" pageNumber="84" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
Terra typica restricta [by lectotype designation by
|
||
<bibRefCitation id="13AC40EAC76F178C03FAF5383E5387CE" author="Mertens, R" journalOrPublisher="Senckenbergiana Biologica" pageId="19" pageNumber="98" pagination="1 - 106" title="Die herpetologische Sektion des Natur-Museums und Forschungs-Institutes Senckenberg in Frankfurt a. M. nebst einem Verzeichnis ihrer Typen." volume="48" year="1967">Mertens (1967)</bibRefCitation>
|
||
]: "Arabia petraea" = Sinai, Egypt.
|
||
</paragraph>
|
||
</subSubSection>
|
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<subSubSection id="ACBE687EB528F8B1B26A9C6F6500661F" lastPageId="6" lastPageNumber="85" pageId="5" pageNumber="84" type="material examined">
|
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<paragraph id="A331B10788832F1314441CE8A60E81DF" pageId="5" pageNumber="84">Material examined.</paragraph>
|
||
<paragraph id="54BA09BEAB766941D0EFB57898E44EAD" lastPageId="6" lastPageNumber="85" pageId="5" pageNumber="84">
|
||
SMF 8723 (lectotype, adult male), Petr. Arabica [Arabia petraea], collected by E.
|
||
<normalizedToken id="B0803FA2A96873FF230A703888DCED69" originalValue="Rüppell">Rueppell</normalizedToken>
|
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in 1827 (MorphoBank M305565-M305594); NMP6V 70163/1 (adult female, MorphoBank M305520-M305528), NMP6V 70163/2 (adult male, MorphoBank M305529-M305542), NMP6V 70163/3-4 (adult females, MorphoBank M305543-M305554, M305555-M305564), Egypt, South Sinai governorate, Sharm el-Sheikh (
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<geoCoordinate id="A77597FCB8503974815226AE1AE841A7" direction="north" orientation="latitude" precision="55" value="27.885">27.885°N</geoCoordinate>
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,
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<geoCoordinate id="06B9FA86FEA611E6A657C86F0560D517" direction="east" orientation="longitude" precision="55" value="34.317">34.317°E</geoCoordinate>
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), ca. 30 m a.s.l., collected by R.
|
||
<normalizedToken id="4115D730F9AEA0EB02739B78F7EED867" originalValue="Kovář">Kovar</normalizedToken>
|
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and R.
|
||
<normalizedToken id="13641DA4EF42276C9D907B37252F8483" originalValue="Víta">Vita</normalizedToken>
|
||
in 1996; ZFMK 94084, ZFMK 94085 (adult females, MorphoBank M305744-M305760, M305761-M305775), Saudi Arabia, Tabuk province, Al Wajh (
|
||
<geoCoordinate id="D0C7151FCE0FC68FF71B28B0A8C4D304" direction="north" orientation="latitude" precision="5" value="26.2076">26.2076°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="ACF4C3E3AFA9B96AEBE692CB156C4B1D" direction="east" orientation="longitude" precision="5" value="36.4976">36.4976°E</geoCoordinate>
|
||
), 5 m a.s.l., 31. V. 2012; ZFMK 94086 (adult female, MorphoBank M305778-M305791), ZFMK 94088, ZFMK 94089 (adult males, M305793-M305799, M305807, M305822-M305827, M305828-M305841), Saudi Arabia, Tabuk province, 15 km S of Al Wajh (
|
||
<geoCoordinate id="A5323A095CBE2023A625A4F237D33178" direction="north" orientation="latitude" precision="5" value="26.1226">26.1226°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="52A2EC31020E50A3E0A9C5C921396107" direction="east" orientation="longitude" precision="5" value="36.5689">36.5689°E</geoCoordinate>
|
||
), 25 m a.s.l., 31. V. 2012; TUZC-R10 (adult female, MorphoBank M305728-M305743), Saudi Arabia, Hail province, 180 km N of Hail (
|
||
<geoCoordinate id="8ED7F08A1F078CB3C7317655F6B16BA3" direction="north" orientation="latitude" precision="5" value="26.8831">26.8831°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="3ECFF1CBE5F139B2EA143D3F0BADC4AE" direction="east" orientation="longitude" precision="5" value="40.0874">40.0874°E</geoCoordinate>
|
||
), 1020 m a.s.l., 30. V. 2012; IBES10183, TUZC-R11 (adult males, MorphoBank M305656-M305671, M305688-M305701), ZFMK 94090,IBES10344 (adult females, MorphoBank M305672-M305687, M305702-M305717), Saudi Arabia, Makkah province, 30 km NE of Alhawiyah (
|
||
<geoCoordinate id="0C9F12024212B15BC328286CECF71344" direction="north" orientation="latitude" precision="5" value="21.6244">21.6244°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="24D65A30E8C018AA80E221E26121B904" direction="east" orientation="longitude" precision="5" value="40.7094">40.7094°E</geoCoordinate>
|
||
), 1295 m a.s.l., 28. V. 2012; IBES10150, IBES10363 (adult males, MorphoBank M305615-M305628, M305643-M305655), ZFMK 94091 (adult female, MorphoBank M305629-M305642), Saudi Arabia, Makkah province, 20 km S of
|
||
<pageBreakToken id="E2B280A229AD6C1C971161837CC7B2D3" pageId="6" pageNumber="85" start="start">Ashayrah</pageBreakToken>
|
||
(
|
||
<geoCoordinate id="4C8B93C01A6EE9C60C9F9511A369DA6E" direction="north" orientation="latitude" precision="5" value="21.6022">21.6022°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E52CA8D2F8A71E0D8126215D5E0F75CF" direction="east" orientation="longitude" precision="5" value="40.6911">40.6911°E</geoCoordinate>
|
||
), 1316 m a.s.l., 28. V. 2012. All Saudi specimens were collected by M. Shobrak, S. Carranza and T. Wilms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="47AAFBCAAD8449ADD2C9380184A27422" pageId="6" pageNumber="85" type="referred material">
|
||
<paragraph id="84FBC6B833E6267A588C4C8D2D710E5B" pageId="6" pageNumber="85">Referred material.</paragraph>
|
||
<paragraph id="9C6AE8F3F304EF5A4F8A2864D4552763" pageId="6" pageNumber="85">
|
||
SMB 10660, Egypt, Suez governorate, Ayoun Musa (
|
||
<geoCoordinate id="B2F8E85495FF252A86645707292055C7" direction="north" orientation="latitude" precision="55" value="29.875">29.875°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="7B2CCAF824F81645474FBB6A92B8EC62" direction="east" orientation="longitude" precision="55" value="32.649">32.649°E</geoCoordinate>
|
||
), ca. 12 m a.s.l., collected by S. Baha El Din, date unknown; TUZC-R9, Saudi Arabia, Tabuk province, 72 km N of Umluj (
|
||
<geoCoordinate id="CEEC15F3EAE0C842F70A9189F5B263C1" direction="north" orientation="latitude" precision="55" value="25.614">25.614°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="97D97897DB96BE1C1A3323F1A3D2715C" direction="east" orientation="longitude" precision="5" value="36.9867">36.9867°E</geoCoordinate>
|
||
), 19 m a.s.l., 31. V. 2012; IBES10001, Saudi Arabia, Riyadh province, Al Ghat (
|
||
<geoCoordinate id="3B5F01C603541448096F8CEF285FB8D3" direction="north" orientation="latitude" precision="5" value="26.0545">26.0545°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="69B30A29235EDC3A621C3A1DA831CB5A" direction="east" orientation="longitude" precision="5" value="45.0003">45.0003°E</geoCoordinate>
|
||
), 776 m a.s.l., 29. V. 2012; ZFMK 94087, TUZC-R8, Saudi Arabia, Tabuk province, 15 km S of Al Wajh (
|
||
<geoCoordinate id="ED88704D26AE92EF5A49A840A43710B2" direction="north" orientation="latitude" precision="5" value="26.1226">26.1226°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="1178438853929F9B2CCF147AA36D26B9" direction="east" orientation="longitude" precision="5" value="36.5689">36.5689°E</geoCoordinate>
|
||
), 25 m a.s.l., 31. V. 2012; ZFMK 87236, Saudi Arabia, Makkah province, Taif National Wildlife Research Center (
|
||
<geoCoordinate id="70213AB65F3B075C12AF1FD18948848E" direction="north" orientation="latitude" precision="555" value="21.25">21.25°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="43F393CAC844C30D7E97A20B9D936E4A" direction="east" orientation="longitude" precision="555" value="40.96">40.96°E</geoCoordinate>
|
||
), 25. VI. 2007 by T. Wilms. These specimens were used for the molecular analyses only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E1EA71EEF32D62828F9E43B83DAC4274" lastPageId="7" lastPageNumber="86" pageId="6" pageNumber="85" type="status and nomenclature">
|
||
<paragraph id="C5CBEBFA375C17EAFD03EA253EB6AF6F" pageId="6" pageNumber="85">Status and nomenclature.</paragraph>
|
||
<paragraph id="9B2B86404906790E7A11E12222D8E612" lastPageId="7" lastPageNumber="86" pageId="6" pageNumber="85">
|
||
<bibRefCitation id="3E6F97C72C7D2C66B59A2217A3AC8391" author="Heyden, CHGv" editor="Rueppell, E" journalOrPublisher="Heinrich Ludwig Broenner, Frankfurt am Main" pageId="19" pageNumber="98" pagination="1 - 24" title="Reptilien." year="1827">Heyden (1827)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="2F7606C9FE57B48F316BD729762602ED" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="granosus">
|
||
<pageBreakToken id="EE6EA6E6A73B0B01A7605D3E10382E12" pageId="7" pageNumber="86" start="start">Hemidactylus</pageBreakToken>
|
||
granosus
|
||
</taxonomicName>
|
||
as a new species occurring in Egypt, Arabia and Abyssinia (Ethiopia and Eritrea). Although not explicitly mentioned by the author, the description was apparently based on four specimens collected by
|
||
<normalizedToken id="A8F9980B22305E42F429CA9A998A7906" originalValue="Rüppell">Rueppell</normalizedToken>
|
||
currently deposited in the Senckenberg Naturmuseum Frankfurt (collection numbers SMF 8723-8726). Heyden did not diagnose the new species against
|
||
<taxonomicName id="765BCE29549A8AD30EA3034D5DEB4D85" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus turcicus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="turcicus">Hemidactylus turcicus</taxonomicName>
|
||
(Linnaeus, 1758) and in respect to our
|
||
<normalizedToken id="C5791E8DE7CCF44F8560B92D743CB82E" originalValue="today’s">today's</normalizedToken>
|
||
knowledge on the morphological variation in
|
||
<taxonomicName id="AA567DEADB625897B62284062AE7E3AA" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus" pageId="7" pageNumber="86" phylum="Chordata" rank="genus">Hemidactylus</taxonomicName>
|
||
the description of
|
||
<taxonomicName id="CA7CBC7D69A1B444293C76CD45750099" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
is very general. Traditionally,
|
||
<taxonomicName id="4590FCB62C24E23AD39D35099FF27B8D" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus turcicus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="turcicus">Hemidactylus turcicus</taxonomicName>
|
||
has been considered a common species widely distributed across the Mediterranean and the Middle East. As the general diagnostic characters of
|
||
<taxonomicName id="7C57B043CCFE985A9953F0C5F5B59EE0" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
given by
|
||
<bibRefCitation id="4BA461D6F00D1C2AD80FF9AAB86C13A3" author="Heyden, CHGv" editor="Rueppell, E" journalOrPublisher="Heinrich Ludwig Broenner, Frankfurt am Main" pageId="19" pageNumber="98" pagination="1 - 24" title="Reptilien." year="1827">Heyden (1827)</bibRefCitation>
|
||
were also applicable to
|
||
<taxonomicName id="7215F42B126D7F1B86A66D464AE919C3" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus turcicus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="turcicus">Hemidactylus turcicus</taxonomicName>
|
||
at that time, the name
|
||
<taxonomicName id="268AE2D971E6A5D167BBCFADCE6859C5" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
Heyden, 1827 was considered its junior synonym (e.g.
|
||
<bibRefCitation id="744DA1FAE141079284A1496D0746DD18" author="Boulenger, GA" journalOrPublisher="London" pageId="18" pageNumber="97" title="Catalogue of the lizards in the British Museum (Natural History). Vol. I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae." volumeTitle="Trustees of the British Museum" year="1885">Boulenger 1885</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="31EE79ACAEA252BA657487D21E28F9F6" author="Loveridge, A" journalOrPublisher="Bulletin of The Museum of Comparative Zoology" pageId="19" pageNumber="98" pagination="1 - 469" title="Revision of the african lizards of the family Gekkonidae." volume="98" year="1947">Loveridge 1947</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="A7D47A5E00C9634BBB637C8442A23FDC" author="Mertens, R" journalOrPublisher="Waldemar Kramer, Frankfurt am Main" pageId="19" pageNumber="98" title="Die Amphibien und Reptilien Europas." year="1960">Mertens and Wermuth 1960</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="3B7E66F5504A388E722A79727443B199" author="Mertens, R" journalOrPublisher="Senckenbergiana Biologica" pageId="19" pageNumber="98" pagination="1 - 106" title="Die herpetologische Sektion des Natur-Museums und Forschungs-Institutes Senckenberg in Frankfurt a. M. nebst einem Verzeichnis ihrer Typen." volume="48" year="1967">Mertens 1967</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B482B1C8E1BFD6CA5DCC6617F720D9EA" pageId="7" pageNumber="86">Salvador 1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F032E96965E3420CF9710B1138CF87B1" author="Baha El Din, S" journalOrPublisher="The American University in Cairo Press, Cairo - New York" pageId="18" pageNumber="97" title="A guide to the Reptiles and Amphibians of Egypt." year="2006">Baha El Din 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1901324A24ADFF08EC87B10D2AD63EBA" pageId="7" pageNumber="86">
|
||
Recent examination (by
|
||
<normalizedToken id="1FA478728531C27E4F4A1E7899BD7DCD" originalValue="JŠ">JS</normalizedToken>
|
||
) of four specimens collected by
|
||
<normalizedToken id="9D6F8529AD20DBE62E158D7B5797C51C" originalValue="Rüppell">Rueppell</normalizedToken>
|
||
(SMF 8723-8726) has shown that one of them [SMF 8723 designated by
|
||
<bibRefCitation id="B695AA2B5858C7064B0B9002578EAA18" author="Mertens, R" journalOrPublisher="Senckenbergiana Biologica" pageId="19" pageNumber="98" pagination="1 - 106" title="Die herpetologische Sektion des Natur-Museums und Forschungs-Institutes Senckenberg in Frankfurt a. M. nebst einem Verzeichnis ihrer Typen." volume="48" year="1967">Mertens (1967)</bibRefCitation>
|
||
as lectotype of
|
||
<taxonomicName id="2B92D26C9D5D5BECC611CBC7367B9D17" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
; for description see below] corresponds morphologically to
|
||
<taxonomicName id="76135E229C69A9554175FAF885156FBA" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus" pageId="7" pageNumber="86" phylum="Chordata" rank="genus">Hemidactylus</taxonomicName>
|
||
sp. 1 from Sinai. The other three specimens from this series morphologically correspond to
|
||
<taxonomicName id="47DC010B533E9FC383C87534A90DC3D9" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus robustus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="robustus">Hemidactylus robustus</taxonomicName>
|
||
Heyden, 1827 (SMF 8725, 8726) and
|
||
<taxonomicName id="CB3F1C54A19076DCF56DE5A860F8FFCB" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus" pageId="7" pageNumber="86" phylum="Chordata" rank="genus">Hemidactylus</taxonomicName>
|
||
cf. granosus (SMF 8724), an animal superficially resembling
|
||
<taxonomicName id="A9B26630FECF43C0A3EEDEF2487F0BA4" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
but differing from the members of the '
|
||
<taxonomicName id="291EE9437DEC1077FE806D5CF4F31E09" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus saba" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="saba">Hemidactylus saba</taxonomicName>
|
||
species
|
||
<normalizedToken id="83D4E10ECDEC85F113659D920D1C9994" originalValue="group’">group'</normalizedToken>
|
||
in several important characters (see below). These findings lead to the conclusion that
|
||
<taxonomicName id="74DEF8E9840BE16C00BBE461AF61A59A" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
Heyden, 1827 is a valid taxon and needs to be resurrected from the synonymy of
|
||
<taxonomicName id="6772F743D72D55FFC08F649111149BED" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus turcicus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="turcicus">Hemidactylus turcicus</taxonomicName>
|
||
. In the light of current knowledge, the range of
|
||
<taxonomicName id="69EC0A27E9D1D12B9460D6C1078ACF1D" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus turcicus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="turcicus">Hemidactylus turcicus</taxonomicName>
|
||
does not include a large part of Egypt, being restricted mostly to northern Egypt including Sinai and its Red Sea coast. The species is also missing in Arabia (sensu lato) and Ethiopia (
|
||
<bibRefCitation id="3E041BD6887D573F465F599104E5EEED" author="Carranza, S" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="18" pageNumber="97" pagination="531 - 545" title="Systematics, biogeography and evolution of Hemidactylus geckos (Reptilia: Gekkonidae) elucidated using mitochondrial DNA sequences." url="10.1016/j.ympev.2005.07.012" volume="38" year="2006">Carranza and Arnold 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="C7F3C4934E4E8DAE0A145E4B61C26A83" author="Moravec, J" journalOrPublisher="Zootaxa" pageId="19" pageNumber="98" pagination="21 - 38" title="High genetic differentiation within the Hemidactylus turcicus complex (Reptilia: Gekkonidae) in the Levant, with comments on the phylogeny and systematics of the genus." volume="2894" year="2011">Moravec et al. 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="2CEEFA05856989EE291EF20B35966BB0" author="Rato, C" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="19" pageNumber="98" pagination="365 - 373" title="When selection deceives phylogeographic interpretation: The case of the Mediterranean house gecko, Hemidactylus turcicus (Linnaeus, 1758)." url="10.1016/j.ympev.2010.12.004" volume="58" year="2011">Rato et al. 2011</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="DB62A16D586A5C719C21F04A2A6D7CD3" author="Smid, J" journalOrPublisher="PLoS ONE" pageId="19" pageNumber="98" title="Out of Arabia: A Complex Biogeographic History of Multiple Vicariance and Dispersal Events in the Gecko Genus Hemidactylus (Reptilia: Gekkonidae)." url="10.1371/journal.pone.0064018" volume="8" year="2013">
|
||
<normalizedToken id="4360B808F4AC77D4FF6FAB755B3205D8" originalValue="Šmíd">Smid</normalizedToken>
|
||
et al. 2013
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="51E161BA79CA84AA04F01BAE1129A64B" pageId="7" pageNumber="86" type="diagnosis">
|
||
<paragraph id="B9C35262A62EBB32BEE75A393B32107F" pageId="7" pageNumber="86">Diagnosis.</paragraph>
|
||
<paragraph id="AA77641DA6B5BEF3FA7791C4DE98777A" pageId="7" pageNumber="86">
|
||
<taxonomicName id="AA8DA069B830127018ECE1D709A9FD1C" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
is a member of the '
|
||
<taxonomicName id="43759CEA7C5BD621BD5F37CBEB0226DB" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus saba" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="saba">Hemidactylus saba</taxonomicName>
|
||
species
|
||
<normalizedToken id="FACC5F569FBD0B4AA30FAEA34D696707" originalValue="group’">group'</normalizedToken>
|
||
within the Arabian radiation of the Arid clade as evidenced by the mtDNA and nDNA analyses. The species has the following combination of molecular and morphological characters: (1) Uncorrected genetic distance from
|
||
<taxonomicName id="BCEABF99D9FC2CDF47E57A467305141F" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus saba" pageId="7" pageNumber="86" phylum="Chordata" rank="species" species="saba">Hemidactylus saba</taxonomicName>
|
||
: 9.9-10.2% in 12S, 14.5-15.5% in cytb; from
|
||
<taxonomicName id="D51B30CB70FFD765AFE7C90D93D43A28" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus" pageId="7" pageNumber="86" phylum="Chordata" rank="genus">Hemidactylus</taxonomicName>
|
||
sp. 4: 10.2-12.3% in 12S, 11.2-13.5% in cytb; (2) small size, SVL 39.0-53.2 mm in males, 40.6-53.3 mm in females; (3) rather elongated head, head length 24-28% of SVL, head width 68-86% of head length, head depth 33-47% of head length; (4) tail length 107-130% of SVL; (5) uppermost nasals separated by a small shield in 89% of specimens; (6) large anterior postmentals in wide mutual contact, and always in contact with the 1st and 2nd lower labial; (7) 9-11 upper labials; (8) 7-9 lower labials; (9) 14-15 longitudinal rows of enlarged, subtriangular, distinctly keeled dorsal tubercles; (10) 7-8 lamellae under the 1st toe and 10-13 under the 4th toe; (11) ca. 6-8 tail segments bearing 6 pointed tubercles; (12) 4-7 preanal pores in males forming a continuous row on the left and right side; (13) subcaudals enlarged; (14) in life, dorsum pale buff with dark brown spots tending to form transverse bands or X-shaped markings, dark horizontal stripe in prefrontal and temporal region, tail with ca. 10-13 dark brown transverse bands, venter white.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A8098088781F3E7F2BC8E23A4B73293" lastPageId="9" lastPageNumber="88" pageId="7" pageNumber="86" type="description of the lectotype">
|
||
<paragraph id="4C6900B894D40EBB8B6A5D3FADF5696D" pageId="7" pageNumber="86">Description of the lectotype.</paragraph>
|
||
<paragraph id="273C83A3CF75F119A8A0DE9EA223B7D8" lastPageId="8" lastPageNumber="87" pageId="7" pageNumber="86">
|
||
SMF 8723, adult male [erroneously determined as female by
|
||
<bibRefCitation id="C16D924762F0DAE915E94217C9EB3849" author="Mertens, R" journalOrPublisher="Senckenbergiana Biologica" pageId="19" pageNumber="98" pagination="1 - 106" title="Die herpetologische Sektion des Natur-Museums und Forschungs-Institutes Senckenberg in Frankfurt a. M. nebst einem Verzeichnis ihrer Typen." volume="48" year="1967">Mertens (1967)</bibRefCitation>
|
||
]. Head and body moderately depressed (Fig. 4). Upper labials (10/10), lower labials (8/7). Nostril between rostral, three subequal nasals and in punctual contact with first upper labial. Uppermost nasals separated by a small inserted scale. Mental triangular, as long as wide. Anterior postmentals long, in a broad contact
|
||
<pageBreakToken id="3E20010432C514AE0D0AF798E741E159" pageId="8" pageNumber="87" start="start">with</pageBreakToken>
|
||
each other, both in contact with the 1st and 2nd lower labial reaching in about one fourth of the width of the 2nd labial. Second postmentals almost round, touching only the 2nd lower labial (Fig. 5). Two enlarged scales behind each second postmental, the lateral ones in contact with the 3rd lower labial. Eye moderate (E/HL=0.26). Head long, distinctly separated from body by a slender neck. Crescent-shaped ear opening. Interorbital region, crown of head and temporal area above the level of ear opening covered by round smooth tubercles. Dorsal region of the specimen is slightly scarred so it is not possible to count the enlarged tubercles on both sides precisely, but there are seven longitudinal rows of large, keeled and caudally pointed tubercles on the left side from which we infer there were originally 14 rows on both sides together. Lower arms, thighs and lower legs with prominent tubercles without keels. Tail original with 6 segments bearing 6 pointed tubercles, broken into three pieces, subcaudals enlarged from just after the hemipenial bulges. Lamellae under the 1st toe 7/7, lamellae under the 4th toe 11/11. Four preanal pores in a continuous row. No femoral pores or enlarged femoral scales. Colour (in alcohol) faded due to long fixation.
|
||
</paragraph>
|
||
<caption id="5A8C2D114F33067A649AA9E6D50BB423" pageId="8" pageNumber="87">
|
||
<paragraph id="5B06DB5480FF40FFD0CAB8BCA8498B46" pageId="8" pageNumber="87">
|
||
Figure 4. Male lectotype of
|
||
<taxonomicName id="9CCAAD02912E25B22B69AF35979FD766" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="8" pageNumber="87" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
(SMF 8723) from Sinai, Egypt. General habitus, lateral and ventral view of the head, precloacal region with preanal pores, right hind leg. Scale refers to the uppermost picture only.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="616F471D31B41AFEF80F73952E62D652" pageId="8" pageNumber="87">
|
||
<paragraph id="A91689BBAC12798F998A2D5B901111B8" pageId="8" pageNumber="87">
|
||
Figure 5. Schematic drawing of the chin region of the lectotype and a new specimen from Sinai of
|
||
<taxonomicName id="304A742ABAED9FB1ED731EAED1453269" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="8" pageNumber="87" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
, the holotype of
|
||
<taxonomicName id="E69392E63398D7BF42FFEE1AAB157539" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus ulii" pageId="8" pageNumber="87" phylum="Chordata" rank="species" species="ulii">Hemidactylus ulii</taxonomicName>
|
||
sp. n., and
|
||
<taxonomicName id="72C1145E648EFEACE2021DA70DEC4E6B" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus turcicus" pageId="8" pageNumber="87" phylum="Chordata" rank="species" species="turcicus">Hemidactylus turcicus</taxonomicName>
|
||
from Sinai.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="99E5E1944DE8179A9724AC3958D508B8" pageId="8" pageNumber="87">Measurements (in mm): SVL 51.5, HL 12.9, HW 9.8, HD 6.0, E 3.3, AG 23.7.</paragraph>
|
||
<paragraph id="9E02578D18BF398AFA491E523A396CE6" pageId="9" pageNumber="88">
|
||
<pageBreakToken id="E9747E967288841D64366B1C7F16BEF3" pageId="9" pageNumber="88" start="start">Paralectotype</pageBreakToken>
|
||
SMF 8724 differs from other individuals of
|
||
<taxonomicName id="9C06DC5AB15D0869C85B8ACE55A39DD3" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
in having relatively high head (HD 50% of HL), lower number of lower labials (6), uppermost nasals in wide contact, first postmentals in contact with 1st lower labials, and 2 preanal pores.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CEBE3DD5E643D2E5E7E1A360C86992C8" pageId="9" pageNumber="88" type="comparison">
|
||
<paragraph id="A35F160E425D836392DD109B76DB5D49" pageId="9" pageNumber="88">Comparison.</paragraph>
|
||
<paragraph id="749C4EAF74152026ECC6C472D04F487A" pageId="9" pageNumber="88">
|
||
<taxonomicName id="BC01A770E13A1105E910BCBA567D173F" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
can be distinguished from other member of the '
|
||
<taxonomicName id="828E014FE3116FA875088B2DAC6553E1" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus saba" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="saba">Hemidactylus saba</taxonomicName>
|
||
species
|
||
<normalizedToken id="794472D692B15E757F9DCC669DC41B27" originalValue="group’">group'</normalizedToken>
|
||
and from other congeners distributed in Sinai and the Red Sea coast by the following set of characters (see also Table 2).
|
||
</paragraph>
|
||
<caption id="D5044108A95132C071D6AD73528E63AA" pageId="9" pageNumber="88">
|
||
<paragraph id="710A32A5695C19D003DC2CC776B31041" pageId="9" pageNumber="88">
|
||
Table 2. Morphological comparison among members of the '
|
||
<taxonomicName id="2F8F1E05BD8600AD73D7550E67E4332C" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus saba" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="saba">Hemidactylus saba</taxonomicName>
|
||
species
|
||
<normalizedToken id="FDA2AAB4ECE93A54BC74EAC5492EBE94" originalValue="group’">group'</normalizedToken>
|
||
and with other
|
||
<taxonomicName id="22AA43AD5BD9534A07F25AC25C201190" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus" pageId="9" pageNumber="88" phylum="Chordata" rank="genus">Hemidactylus</taxonomicName>
|
||
species from Sinai and SW Yemen. The values are given as follows: sample size, mean
|
||
<normalizedToken id="CE8FC0674FBEC550E5DFF459AC5269A4" originalValue="±">+/-</normalizedToken>
|
||
standard deviation above, min. - max. value below.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D263CDF6063CE0679C525BABAA5C43E5" pageId="9" pageNumber="88">
|
||
<table id="0EADEDEF5B8743A1E13F11D7A6073794" pageId="9" pageNumber="88">
|
||
<tr id="280B6CB33EFC939E0B464B404DF76A39" pageId="9" pageNumber="88">
|
||
<th id="D6C1918DD5A860F73EEE4AE968C76EED" colspan="1" pageId="9" pageNumber="88" rowspan="2">Species / Character</th>
|
||
<th id="55DC836D79818A8BE2C578FA8C8F17BD" colspan="6" pageId="9" pageNumber="88" rowspan="1">
|
||
<taxonomicName id="30960F7DBEF846238C38DB09EC89CC54" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus saba" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="saba">Hemidactylus saba</taxonomicName>
|
||
species group
|
||
</th>
|
||
<th id="BAB2DB2015B76ACD3339C038947507FB" colspan="2" pageId="9" pageNumber="88" rowspan="2">
|
||
<taxonomicName id="05E26789A588A25C2221682FB8937987" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus robustus" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="robustus">Hemidactylus robustus</taxonomicName>
|
||
</th>
|
||
<th id="585405C3FFA70F497BCE1102ABF9ADAF" colspan="2" pageId="9" pageNumber="88" rowspan="2">
|
||
<taxonomicName id="66CE306F7EC7E53FE4728A88C1D4F4AF" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus turcicus" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="turcicus">Hemidactylus turcicus</taxonomicName>
|
||
</th>
|
||
<th id="C5E72A93170BD8F750F76B0F2F172565" colspan="2" pageId="9" pageNumber="88" rowspan="2">
|
||
<taxonomicName id="2D1CBF5D96AE2E77CEED7FB01704C99F" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus mindiae" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="mindiae">Hemidactylus mindiae</taxonomicName>
|
||
</th>
|
||
<th id="4DC425E3364A8210C1114B0F9BAE080D" colspan="2" pageId="9" pageNumber="88" rowspan="2">
|
||
<taxonomicName id="C3D31817D8C75D4B2588891ACD10F586" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus jumailiae" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="jumailiae">Hemidactylus jumailiae</taxonomicName>
|
||
</th>
|
||
<th id="CA059F7F821BFB98D9083ADA3C0AE02F" colspan="2" pageId="9" pageNumber="88" rowspan="2">
|
||
<taxonomicName id="BA9024E28A01AF2DCECB54B422E24416" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus yerburii subsp. yerburii" pageId="9" pageNumber="88" phylum="Chordata" rank="subSpecies" species="yerburii" subSpecies="yerburii">Hemidactylus yerburii yerburii</taxonomicName>
|
||
</th>
|
||
<th id="59ACBDD206F17EE6000360C785CBDD88" colspan="2" pageId="9" pageNumber="88" rowspan="2">
|
||
<taxonomicName id="0B5BF511CA5C9C319F320EB6137C351D" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus yerburii subsp. montanus" pageId="9" pageNumber="88" phylum="Chordata" rank="subSpecies" species="yerburii" subSpecies="montanus">Hemidactylus yerburii montanus</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="955AF4ACBB4B421DB12B6D3E24171623" pageId="9" pageNumber="88">
|
||
<th id="2567C0DD3BC2CF1BCC4F5041F8117656" colspan="2" pageId="9" pageNumber="88" rowspan="1">
|
||
<taxonomicName id="2A9AAED33DC67E965A878579E421E3E4" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
</th>
|
||
<th id="3BDDFA7524AC4C99102EFD2DCF608442" colspan="2" pageId="9" pageNumber="88" rowspan="1">
|
||
<taxonomicName id="04C1D133E613B84367E90A7DFB082D84" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus saba" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="saba">Hemidactylus saba</taxonomicName>
|
||
</th>
|
||
<th id="561AE4AA96E65FC2FD92C9153B89E566" colspan="2" pageId="9" pageNumber="88" rowspan="1">
|
||
<taxonomicName id="6A6456D4FFB67502C0EF5B445C08AA09" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus ulii" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="ulii">Hemidactylus ulii</taxonomicName>
|
||
sp. n.
|
||
</th>
|
||
</tr>
|
||
<tr id="51EB1C893AB81F20AB117A34C49FABEA" pageId="9" pageNumber="88">
|
||
<td id="515EC76ACD5E9CE2AC9232ED65A435D3" colspan="1" pageId="9" pageNumber="88" rowspan="1">stnd</td>
|
||
</tr>
|
||
<tr id="B65F8473459CD2E4411AF11F16006DA3" pageId="9" pageNumber="88">
|
||
<td id="98F86A34416CC0DBDA155266C2EA231E" colspan="1" pageId="9" pageNumber="88" rowspan="2">st</td>
|
||
</tr>
|
||
<tr id="94E0F75359C7754E41AC8612A6402F91" pageId="9" pageNumber="88">
|
||
<td id="936D72DEF58CAC6DF0F87547EF464F11" colspan="1" pageId="9" pageNumber="88" rowspan="2">th</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="3B3D4B77E95C06F28E5E707062DF0049" pageId="9" pageNumber="88">
|
||
From
|
||
<taxonomicName id="DA91D2500971061D092D5F05B83437DE" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus saba" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="saba">Hemidactylus saba</taxonomicName>
|
||
by having distinctly keeled dorsal tubercles (smooth in
|
||
<taxonomicName id="E160EEFA125D39C3075FAEA6B6DCF468" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus saba" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="saba">Hemidactylus saba</taxonomicName>
|
||
), and lower number of lamellae under the 1st toe (7-8 vs. 8-9).
|
||
</paragraph>
|
||
<paragraph id="4017A90B8DC64D6B9AA17E8A5B65FFFA" pageId="9" pageNumber="88">
|
||
From
|
||
<taxonomicName id="4016A4999814E18C717DC44287636C48" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus" pageId="9" pageNumber="88" phylum="Chordata" rank="genus">Hemidactylus</taxonomicName>
|
||
sp. 4 (described below) by its larger size (max. SVL 53.2 mm vs. 40.4 mm in males, 53.3 mm vs. 40.7 mm in females), in having more frequently separated uppermost nasals (100% vs. 60% of specimens), lower number of preanal pores in males (4-7 vs. 8), and higher number of lamellae under the 1st (7-8 vs. 5-6) and 4th (10-13 vs. 8-9) toe.
|
||
</paragraph>
|
||
<paragraph id="AADCEE81AC125DCEF9C666D101E89657" pageId="9" pageNumber="88">
|
||
From
|
||
<taxonomicName id="54E0B590D980296F292F8D12CD75BEC2" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus flaviviridis" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="flaviviridis">Hemidactylus flaviviridis</taxonomicName>
|
||
by its smaller size (max. SVL 53.2 mm in males and 53.3 mm in females vs. up to 90 mm [
|
||
<bibRefCitation id="6131CD700BA2F6AFE11C9C7D185A839E" author="Anderson, SC" journalOrPublisher="Society for the Study of Amphibians and Reptiles" pageId="18" pageNumber="97" title="The lizards of Iran." year="1999">Anderson (1999)</bibRefCitation>
|
||
; sexes not distinguished]), by the presence of enlarged dorsal tubercles, and the absence of femoral pores in males.
|
||
</paragraph>
|
||
<paragraph id="D2B5187544BA0E5A22A2A5384C91C8E0" pageId="9" pageNumber="88">
|
||
From
|
||
<taxonomicName id="4029353AF378C86087BD38790409DC6A" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus mindiae" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="mindiae">Hemidactylus mindiae</taxonomicName>
|
||
by the lower number of supralabials (9-11 vs. 10-12), by having anterior postmentals in wide contact (punctual in
|
||
<taxonomicName id="BFD10D10C7639E83B6BFE649E661E29B" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus mindiae" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="mindiae">Hemidactylus mindiae</taxonomicName>
|
||
) and keeled dorsal tubercles (smooth in
|
||
<taxonomicName id="5FDB2043A92F8619C4E6DF8BB522D881" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus mindiae" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="mindiae">Hemidactylus mindiae</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="C07BAD35BC0C7DE2BD3879640B61B894" pageId="9" pageNumber="88">
|
||
From
|
||
<taxonomicName id="A4B30F478CE87CF1C0818EF195DC8F0E" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus robustus" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="robustus">Hemidactylus robustus</taxonomicName>
|
||
by the larger size of males (max. SVL 53.2 mm vs. 43.7 mm), longer tail (tail length 53.0-64.8 mm vs. 40.9-48.7 mm), and lower number of preanal pores in males (4-7 vs. 5-8).
|
||
</paragraph>
|
||
<paragraph id="8ED6F29FD4A22FAB7391FE4FC88A8AD0" pageId="9" pageNumber="88">
|
||
From
|
||
<taxonomicName id="BDB939B672362FD8B0ADBDFC569E57DD" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus turcicus" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="turcicus">Hemidactylus turcicus</taxonomicName>
|
||
by its higher number of upper labials (9-11 vs. 7-10), in having anterior postmentals more frequently in contact with 2nd lower labial (100% vs. 12.1%), in having anterior postmentals in wide mutual contact behind the mental scale (contact punctual in 67% specimens of
|
||
<taxonomicName id="1FA1B1C981D6A4C8B12A478CF1FD506E" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus turcicus" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="turcicus">Hemidactylus turcicus</taxonomicName>
|
||
), and by the lower number of preanal pores in males (4-7 vs. 6-10).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FDBF30EB602788F244C54AA172C5192" lastPageId="10" lastPageNumber="89" pageId="9" pageNumber="88" type="variation">
|
||
<paragraph id="F40C09CA57878F51E8492B79EAAC4065" pageId="9" pageNumber="88">Variation.</paragraph>
|
||
<paragraph id="0E38C533917DADA65A8BF73E6CD08CE3" pageId="9" pageNumber="88">Specimens with intact tail vary in number of tail segments bearing 6 pointed tubercles (7-8). The original portion of the tail of the female NMP6V 70163/4 is very wide at the base, separated from cloacal region by a basal constriction. One specimen (IBES10212) is the only animal with 15 longitudinal rows of enlarged tubercles. Another one (IBES10284) has uppermost nasals in wide contact. Most striking is the variation in the number of preanal pores in males. Whereas the lectotype and the only male from Sinai (NMP6V 70163/2) have both 4 pores, all males from Saudi Arabia have 6-7 pores. There seems to be clinal variability in this character, males from NW of the known range (Fig. 6) possess only 4 preanal pores, all animals from the eastern Red Sea coast in Saudi Arabia have 6 pores and a single individual from the southern limit of the range has 7 pores.</paragraph>
|
||
<caption id="1878C92CEE2EFB0B7422EA6589043F3D" pageId="9" pageNumber="88">
|
||
<paragraph id="7F68BFCD2A014D73276A26DAC5865CD6" pageId="9" pageNumber="88">
|
||
Figure 6. Distribution map of
|
||
<taxonomicName id="504747C8C146AADDB3E6A9AE9897B1C5" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
,
|
||
<taxonomicName id="2FCA7C9DDE04AE0CE54BB71CA8204BF6" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus saba" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="saba">Hemidactylus saba</taxonomicName>
|
||
and
|
||
<taxonomicName id="EEA635A18AB6BB4B72C883D878C734D4" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus ulii" pageId="9" pageNumber="88" phylum="Chordata" rank="species" species="ulii">Hemidactylus ulii</taxonomicName>
|
||
sp. n. For the list of locality names and their corresponding numbers in the map see Table 1.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="90CA58DC4EFB3244F37618B651793691" lastPageId="10" lastPageNumber="89" pageId="9" pageNumber="88">
|
||
Coloration (in life) pale buff dorsally (Fig. 7). Conspicuous dark brown horizontal stripe in loreal and temporal area, terminated at the level of ear from where it continues in a series of dark patches on the neck. Four barely visible X-shaped markings on dorsum formed mainly by dark brown enlarged tubercles (first on nape, second across scapulae, third in lumbal region, and fourth just in front of the anterior insertion of hind limbs). Isolated dark brown stripe runs across body in the place of posterior insertion of hind
|
||
<pageBreakToken id="DC53DEBA9B3D09F21D09ADA7586EB9FD" pageId="10" pageNumber="89" start="start">limbs</pageBreakToken>
|
||
. Regenerated tails are uniformly buff from above. Dorsum, sides of chin, underside of front and hind limbs and underside of tail with faint stipple visible under magnification. Belly white. Tips of fingers and toes black behind insertion of terminal phalanges. Coloration is consistent among all specimens and varies only in distinctness of the markings.
|
||
</paragraph>
|
||
<caption id="C52C6391A84983A141A8F9062E9709F1" pageId="10" pageNumber="89">
|
||
<paragraph id="4A6B63F5AFDBE2024204303B8E8A894D" pageId="10" pageNumber="89">
|
||
Figure 7. Live specimens of
|
||
<taxonomicName id="9A5780ECA3235157689AB6B4B3DAA892" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="10" pageNumber="89" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
from Saudi Arabia. A IBES10344, 30 km NE of Alhawiyah (loc. number 8) B TUZC-R10, 180 km W of Hail (6) C ZFMK 94091, 20 km S of Ashayrah (9) D ZFMK 94086, 15 km S of Al Wajh (4).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="923168E9C4386DB3E333B82B8FE7A802" pageId="10" pageNumber="89">There is a very low variation in mtDNA between specimens from Sinai and Saudi Arabia (max. 1.3% in both 12S and cytb). All animals from Sinai share the same haplotypes in 12S and also cytb gene. All four nuclear loci studied show some degree of intraspecific variation (Fig. 3).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EE9F42A3C7811446CF8C2B4A11237362" lastPageId="12" lastPageNumber="91" pageId="10" pageNumber="89" type="distribution and ecology">
|
||
<paragraph id="C1DCB6BF62136CC7103EF99A01C72447" pageId="10" pageNumber="89">Distribution and ecology.</paragraph>
|
||
<paragraph id="048EF5486A0457EE5ECF49CD4300F7F8" lastPageId="11" lastPageNumber="90" pageId="10" pageNumber="89">
|
||
Eduard
|
||
<normalizedToken id="09CC7529BE315AB81C633F518C0DE53A" originalValue="Rüppell">Rueppell</normalizedToken>
|
||
collected the original series in 1827 when he began his marine biological studies of the Red Sea and travelled from Egypt to Eritrea. There is no specific information that he went to Arabia as well (
|
||
<bibRefCitation id="565B5408EDABAFFE50022956698DC5EC" author="Rueppell, E" journalOrPublisher="Heinrich Ludwig Broenner, Frankfurt am Main" pageId="19" pageNumber="98" title="Atlas zu der Reise im no ̈ rdlichen Afrika" year="1826 - 1828">
|
||
<normalizedToken id="0AA7318BBBC221C00EBCC82A96E5D584" originalValue="Rüppell">Rueppell</normalizedToken>
|
||
1826-1828
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="432D62A4E6E0086D3F50125900D443CE" author="Klausewitz, W" journalOrPublisher="Zoology in the Middle East" pageId="19" pageNumber="98" pagination="7 - 12" title="Frankfurt versus Berlin: The Red Sea explorers Wilhelm Hemprich, Christian Ehrenberg and Eduard Rueppell." url="10.1080/09397140.2002.10637935" volume="27" year="2002">Klausewitz 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A227919C6F24D992569C0AEFF967A424" author="Wagner, RK" journalOrPublisher="Papageien" pageId="19" pageNumber="98" pagination="244 - 249" title="Eduard Rueppell - Leben und Werk eines Forschungsreisenden." volume="7" year="2008">Wagner 2008</bibRefCitation>
|
||
); therefore the original distribution of
|
||
<taxonomicName id="E352C91399C64577271E5ABA08035C7C" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="11" pageNumber="90" phylum="Chordata" rank="species" species="granosus">
|
||
<pageBreakToken id="61038C7258F7E2B0887413D2F9FB203A" pageId="11" pageNumber="90" start="start">Hemidactylus</pageBreakToken>
|
||
granosus
|
||
</taxonomicName>
|
||
described as "Egypt, Arabia, and Abyssinia [Ethiopia and Eritrea]" by
|
||
<bibRefCitation id="09E687DC12D23A52B2FAEC05441C5CF0" author="Heyden, CHGv" editor="Rueppell, E" journalOrPublisher="Heinrich Ludwig Broenner, Frankfurt am Main" pageId="19" pageNumber="98" pagination="1 - 24" title="Reptilien." year="1827">Heyden (1827)</bibRefCitation>
|
||
was probably too general and incorrect. Because there were no other specimens assignable with certainty to
|
||
<taxonomicName id="F8CFA478CB5BE5F4CA452366510AC54E" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="11" pageNumber="90" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
apart from the four individuals collected in Sinai (SMF 8723-8726, for their current status see 'Status and
|
||
<normalizedToken id="9501F78A3E087635DD5F882C6D86527B" originalValue="nomenclature’">nomenclature'</normalizedToken>
|
||
section) (
|
||
<bibRefCitation id="CF31008B086F7ACE4CDD75355402B7A9" author="Boettger, O" journalOrPublisher="Frankfurt am Main" pageId="18" pageNumber="97" title="Katalog der Reptilien-Sammlung im Museum der Senckenbergischen Naturforschenden Gesellschaft in Frankfurt am Main." year="1893">Boettger 1893</bibRefCitation>
|
||
), one of which became the lectotype after
|
||
<bibRefCitation id="E74F686DD4AAAEE21E28501EA5218D0F" author="Mertens, R" journalOrPublisher="Senckenbergiana Biologica" pageId="19" pageNumber="98" pagination="1 - 106" title="Die herpetologische Sektion des Natur-Museums und Forschungs-Institutes Senckenberg in Frankfurt a. M. nebst einem Verzeichnis ihrer Typen." volume="48" year="1967">
|
||
<normalizedToken id="2C52FB8F141E43E91DDC8BB5C3A4FD12" originalValue="Mertens’">Mertens'</normalizedToken>
|
||
(1967)
|
||
</bibRefCitation>
|
||
designation, Sinai could be considered the only reliable locality for
|
||
<taxonomicName id="B3232D4C864265401F83D2C2EC151DF4" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="11" pageNumber="90" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
. Here,
|
||
<taxonomicName id="DCF276C22ED28E5E8D9D499C9E410973" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="11" pageNumber="90" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
is also confirmed from two coastal localities in south and west Sinai and from coastal and inland regions in western and central Saudi Arabia (Fig. 6). Nevertheless, a wider distribution of the species along the Red Sea coast can be expected. According to
|
||
<bibRefCitation id="FE9E6C45D91A097D96F52B3B955D8C21" author="Baha El Din, SM" journalOrPublisher="Zoology in the Middle East" pageId="18" pageNumber="97" pagination="27 - 34" title="An overview of Egyptian species of Hemidactylus (Gekkonidae), with the description of a new species from the high mountains of South Sinai." url="10.1080/09397140.2005.10638078" volume="34" year="2005">Baha El Din (2005)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="04A0D368F9BAF42AE098CFC7C384F374" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus" pageId="11" pageNumber="90" phylum="Chordata" rank="genus">Hemidactylus</taxonomicName>
|
||
geckos inhabiting the interior lowland of Sinai and the Eastern Desert in Egypt stand out in having notably coarse scalation. Interestingly, the areas with occurrence of animals with coarse scalation correspond with the presence of individuals with low numbers of preanal pores (
|
||
<bibRefCitation id="91C1FFEFB7D314D0EE1260E349B6FAC1" author="Baha El Din, SM" journalOrPublisher="Zoology in the Middle East" pageId="18" pageNumber="97" pagination="27 - 34" title="An overview of Egyptian species of Hemidactylus (Gekkonidae), with the description of a new species from the high mountains of South Sinai." url="10.1080/09397140.2005.10638078" volume="34" year="2005">Baha El Din 2005</bibRefCitation>
|
||
), which is typical for the Sinai populations of
|
||
<taxonomicName id="F88232B6F2AE4D5DA189BEE275D09EC7" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="11" pageNumber="90" phylum="Chordata" rank="species" species="granosus">Hemidactylus granosus</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9F33C470294442FA491B58486CF834DB" lastPageId="12" lastPageNumber="91" pageId="11" pageNumber="90">
|
||
In 1996, when the NMP specimens were collected, the locality in Sharm el-Sheikh was formed by a crop field supplied with drain water from nearby habitations. Geckos were found during the day under unused empty barrels and also inside buildings. Other species syntopic with
|
||
<taxonomicName id="E3CCB93AA8C9A7DDEE6330AC877AD298" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus granosus" pageId="12" pageNumber="91" phylum="Chordata" rank="species" species="granosus">
|
||
<pageBreakToken id="D46999F8A726D427DD3B93DFA31F532E" pageId="12" pageNumber="91" start="start">Hemidactylus</pageBreakToken>
|
||
granosus
|
||
</taxonomicName>
|
||
in Sharm el-Sheikh were:
|
||
<taxonomicName id="5188B0982E609ADB96AC3275F0F18742" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus turcicus" pageId="12" pageNumber="91" phylum="Chordata" rank="species" species="turcicus">Hemidactylus turcicus</taxonomicName>
|
||
,
|
||
<taxonomicName id="6EF30417FE0087E9A514A49B03795C63" class="Squamata" family="Scincidae" genus="Chalcides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chalcides ocellatus" pageId="12" pageNumber="91" phylum="Chordata" rank="species" species="ocellatus">Chalcides ocellatus</taxonomicName>
|
||
(
|
||
<normalizedToken id="443DFB28C45EFB6BA1C809AB59E23634" originalValue="Forskål">Forskal</normalizedToken>
|
||
, 1775),
|
||
<taxonomicName id="2860B164C767F44B0682903E226C090D" class="Squamata" family="Gekkonidae" genus="Stenodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenodactylus sthenodactylus" pageId="12" pageNumber="91" phylum="Chordata" rank="species" species="sthenodactylus">Stenodactylus sthenodactylus</taxonomicName>
|
||
(Lichtenstein, 1823), and
|
||
<taxonomicName id="BEC77E166A139D362245EF8C28CF8A4B" class="Squamata" family="Phyllodactylidae" genus="Ptyodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptyodactylus hasselquistii" pageId="12" pageNumber="91" phylum="Chordata" rank="species" species="hasselquistii">Ptyodactylus hasselquistii</taxonomicName>
|
||
(Donndorff, 1798) (R.
|
||
<normalizedToken id="F88A706C7F700D677201BA0938C6789F" originalValue="Víta">Vita</normalizedToken>
|
||
in litt, 2013). However, when visited again in 2010, the locality had changed dramatically (R.
|
||
<normalizedToken id="606ABC0B8787BB36C281D0A630A79014" originalValue="Víta">Vita</normalizedToken>
|
||
in litt, 2013). The whole area was under heavy development and the irrigation channels had disappeared. The current conditions at the place are unknown to us. In 2011 JM surveyed a neighbouring urban area east of this locality. It was covered by a mosaic of tourist resorts and abandoned ruderal plots. In dry anthropogenic habitats (e.g. rubbish dumps, road ditches, old walls and buildings, abandoned construction sites, natural but heavily disturbed open areas, etc.) dominated two very abundant gecko species.
|
||
<taxonomicName id="5367C1F714E2AB607EA934C9728FFFE3" class="Squamata" family="Phyllodactylidae" genus="Ptyodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptyodactylus hasselquistii" pageId="12" pageNumber="91" phylum="Chordata" rank="species" species="hasselquistii">Ptyodactylus hasselquistii</taxonomicName>
|
||
occupied primarily various vertical surfaces whereas
|
||
<taxonomicName id="95870459713F575CABF648739E8CB41A" class="Squamata" family="Gekkonidae" genus="Cyrtopodion" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyrtopodion scabrum" pageId="12" pageNumber="91" phylum="Chordata" rank="species" species="scabrum">Cyrtopodion scabrum</taxonomicName>
|
||
(Heyden, 1827) prevailed on the ground.
|
||
<taxonomicName id="97E3EA503D4B980419919EBE2A18DC0D" class="Squamata" family="Gekkonidae" genus="Tropiocolotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropiocolotes nattereri" pageId="12" pageNumber="91" phylum="Chordata" rank="species" species="nattereri">Tropiocolotes nattereri</taxonomicName>
|
||
Steindachner, 1901 was found in dry and relatively well-preserved natural places.
|
||
<taxonomicName id="22D052AFF9F287FECA62E1622AEA5D9C" class="Squamata" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hemidactylus turcicus" pageId="12" pageNumber="91" phylum="Chordata" rank="species" species="turcicus">Hemidactylus turcicus</taxonomicName>
|
||
was occasionally encountered in more humid artificial habitats in parks and hotel gardens. Specimens from Saudi Arabia were mostly collected during the day inside concrete tunnels under roads. In some of the tunnels they were syntopic with
|
||
<taxonomicName id="1817BCE891FC6F521D63EABFAC8827EC" class="Squamata" family="Phyllodactylidae" genus="Ptyodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptyodactylus hasselquistii" pageId="12" pageNumber="91" phylum="Chordata" rank="species" species="hasselquistii">Ptyodactylus hasselquistii</taxonomicName>
|
||
. One specimen was also collected on the walls of the Taif National Wildlife Research Centre, where it was also syntopic with
|
||
<taxonomicName id="BD8057B31F0E9E101F640F52761DEB76" class="Squamata" family="Phyllodactylidae" genus="Ptyodactylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptyodactylus hasselquistii" pageId="12" pageNumber="91" phylum="Chordata" rank="species" species="hasselquistii">Ptyodactylus hasselquistii</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |