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<document id="FCBEEE885F93D6878528D22AEBB592A5" ID-CLB-Dataset="235405" ID-DOI="10.11646/zootaxa.5256.5.2" ID-GBIF-Dataset="1830c929-4423-42b4-a4ae-c60c525a2f70" ID-ISSN="1175-5326" ID-Zenodo-Dep="7758834" ID-ZooBank="5278C906-086F-4B62-ADC0-75E8F481D95D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1679468506190" checkinUser="plazi" docAuthor="Park, Kyu-Tek &amp; Heppner, John B." docDate="2023" docId="9F59CE71EB238F4BFF6C943FB87FD7DC" docLanguage="en" docName="zootaxa.5256.5.2.pdf" docOrigin="Zootaxa 5256 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.5.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thubana spiniosa Park &amp; Heppner 2023, sp. nov." docType="treatment" docUuid="9EB1F313-055D-47BD-8FEB-B1486FCFE63D" docUuidSource="ZooBank" docVersion="7" lastPageNumber="441" masterDocId="6360B609EB268F4CFFFB9001BB49D60D" masterDocTitle="Review of the genera Thubana Walker and Torodora Meyrick (Lepidoptera: Lecithoceridae) from Sumatra, Indonesia" masterLastPageNumber="456" masterPageNumber="434" pageNumber="439" updateTime="1699477203389" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="5CB2F60E3372DE851CFBFB0C145BD8CF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="7F9A697D8D60E0BBB988FEFAB9E378A8">Review of the genera Thubana Walker and Torodora Meyrick (Lepidoptera: Lecithoceridae) from Sumatra, Indonesia</mods:title>
</mods:titleInfo>
<mods:name id="ADAF0BEFA98B8C4E2B4DFCDD60DE2D6F" type="personal">
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<mods:namePart id="C0B81782E704E42176F538BFEC96BEAB">Park, Kyu-Tek</mods:namePart>
<mods:affiliation id="641FF0C36CC3BFF5FCC663746856CC60">Bioresource and Environmental Center, Incheon National University, Incheon, 22012 Korea.</mods:affiliation>
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</mods:role>
<mods:namePart id="CF33BA59EE504115C3D8F7E8C9AD4FE5">Heppner, John B.</mods:namePart>
<mods:affiliation id="73CFFBD6AFAD46682152BED31281C275">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, USA. jheppner @ flmnh. ufI. edu.</mods:affiliation>
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<mods:title id="26614BAC44FB1033011A3098922EC01C">Zootaxa</mods:title>
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<mods:part id="B561D831E7F7DCA15535176EA1353652">
<mods:date id="8DABFE978CBDC25032F5542E1DC5B283">2023</mods:date>
<mods:detail id="3A6B8BECBBA8FDE99EAC30A70A84945C" type="pubDate">
<mods:number id="352454EC974EA459EB91B621DDE461D1">2023-03-22</mods:number>
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<mods:detail id="D5C7B2BB8EC286312A4CBA8AF8012304" type="volume">
<mods:number id="ED9141DCB34864DEE1F499F22BA90B2B">5256</mods:number>
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<mods:number id="E6E066274269CB10EFF0BF6B7699BA7A">5</mods:number>
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<mods:url id="4A766A394B10777C8EBEBFEE7F17386C">http://dx.doi.org/10.11646/zootaxa.5256.5.2</mods:url>
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<mods:classification id="0C192DAB19A38E0F2EBDCE09F6DD4F59">journal article</mods:classification>
<mods:identifier id="FE645350EE515ED7A7D50C1956F7E26D" type="CLB-Dataset">235405</mods:identifier>
<mods:identifier id="22D0E0A74250D831C614E7F143BB014B" type="DOI">10.11646/zootaxa.5256.5.2</mods:identifier>
<mods:identifier id="5947ECFF1B92E4BD1D3D768C05192B26" type="GBIF-Dataset">1830c929-4423-42b4-a4ae-c60c525a2f70</mods:identifier>
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<treatment id="9F59CE71EB238F4BFF6C943FB87FD7DC" ID-DOI="http://doi.org/10.5281/zenodo.7761087" ID-GBIF-Taxon="209464676" ID-Zenodo-Dep="7761087" LSID="urn:lsid:zoobank.org:act:9EB1F313-055D-47BD-8FEB-B1486FCFE63D" httpUri="http://treatment.plazi.org/id/9F59CE71EB238F4BFF6C943FB87FD7DC" lastPageId="7" lastPageNumber="441" pageId="5" pageNumber="439">
<subSubSection id="5FEA2CECEB238F49FF6C943FBF52D270" pageId="5" pageNumber="439" type="nomenclature">
<paragraph id="174F7F67EB238F49FF6C943FB9FFD255" blockId="5.[151,1051,1086,1184]" box="[151,694,1086,1113]" pageId="5" pageNumber="439">
<heading id="4C07C80BEB238F49FF6C943FB9FFD255" bold="true" box="[151,694,1086,1113]" fontSize="11" level="1" pageId="5" pageNumber="439" reason="1">
<emphasis id="2584A375EB238F49FF6C943FB9FFD255" bold="true" box="[151,694,1086,1113]" pageId="5" pageNumber="439">
<taxonomicName id="D0F004E4EB238F49FF6C943FB902D255" ID-CoL="B7R95" authority="Park &amp; Heppner" authorityName="Park &amp; Heppner" authorityYear="2023" box="[151,587,1086,1113]" class="Insecta" family="Lecithoceridae" genus="Thubana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="439" phylum="Arthropoda" rank="species" species="spiniosa" status="sp. nov.">
<emphasis id="2584A375EB238F49FF6C943FBA39D255" bold="true" box="[151,368,1086,1113]" italics="true" pageId="5" pageNumber="439">Thubana spiniosa</emphasis>
Park &amp; Heppner
</taxonomicName>
,
<taxonomicNameLabel id="3EB71E0EEB238F49FDA3943FB9FFD255" box="[600,694,1086,1112]" pageId="5" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="174F7F67EB238F49FF6C9463BF52D270" blockId="5.[151,1051,1086,1184]" box="[151,1051,1122,1149]" pageId="5" pageNumber="439">
LSID:
<uri id="63617365EB238F49FF1F9463BF52D270" box="[228,1051,1122,1149]" pageId="5" pageNumber="439">
urn:lsid:zoobank.org:act:
<uuid id="635645B2EB238F49FE0C9463BF52D270" box="[503,1051,1122,1149]" pageId="5" pageNumber="439">9EB1F313-055D-47BD-8FEB-B1486FCFE63D</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="5FEA2CECEB238F49FF6C9487BBA2D2AD" box="[151,235,1158,1184]" pageId="5" pageNumber="439" type="description">
<paragraph id="174F7F67EB238F49FF6C9487BBA2D2AD" blockId="5.[151,1051,1086,1184]" box="[151,235,1158,1184]" pageId="5" pageNumber="439">
(
<figureCitation id="8FCB63E2EB238F49FF649487BBAAD2AD" box="[159,227,1158,1184]" captionStart="FIGURE 4" captionStartId="6.[151,250,1824,1848]" captionTargetBox="[161,1429,192,1784]" captionTargetId="figure-20@6.[151,1436,181,1800]" captionTargetPageId="6" captionText="FIGURE 4. Thubana spiniosa Park &amp; Heppner, sp. nov.: A, adult, holotype; B, head, dorsal view; C, head with labial palpus; D, male genitalia, slide no. JBH-4199; E, ditto, aedeagus; F, abdominal segments VII-VIII. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7758842" httpUri="https://zenodo.org/record/7758842/files/figure.png" pageId="5" pageNumber="439">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5FEA2CECEB238F49FF6C94CFB9F4D301" pageId="5" pageNumber="439" type="materials_examined">
<paragraph id="174F7F67EB238F49FF6C94CFB9F4D301" blockId="5.[151,1437,1230,2019]" pageId="5" pageNumber="439">
<emphasis id="2584A375EB238F49FF6C94CFBA00D2E5" bold="true" box="[151,329,1230,1256]" pageId="5" pageNumber="439">Type specimen</emphasis>
.
<materialsCitation id="A798753AEB238F49FEA194CFB9F4D301" ID-GBIF-Occurrence="4063375309" collectingDate="1992-08-24" collectingDateMax="1992-08-26" collectingDateMin="1992-08-24" collectorName="J. B. Heppner &amp; E. W. Diehl" elevation="1200" location="Pematang Siantar" pageId="5" pageNumber="439" specimenCount="1" stateProvince="Sumatera Utara" typeStatus="holotype">
<emphasis id="2584A375EB238F49FEA194CFBA88D2E5" box="[346,449,1230,1256]" italics="true" pageId="5" pageNumber="439">
<typeStatus id="C84BC1C5EB238F49FEA194CFBA88D2E5" box="[346,449,1230,1256]" pageId="5" pageNumber="439" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectorName id="BA051AB1EB238F49FE2994CFB97ED2E4" box="[466,567,1230,1257]" pageId="5" pageNumber="439">Male, N.</collectorName>
<collectingRegion id="D534B185EB238F49FDBB94CFB9E9D2E5" box="[576,672,1230,1256]" country="Indonesia" name="Sumatera Utara" pageId="5" pageNumber="439">Sumatra</collectingRegion>
,
<quantity id="D008D282EB238F49FD5594CFB9BED2E5" box="[686,759,1230,1256]" metricMagnitude="4" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="439" unit="km" value="43.0">43 km</quantity>
SW
<location id="122F29BCEB238F49FCC894CEB8B5D2E5" LSID="urn:lsid:plazi:treatment:9F59CE71EB238F4BFF6C943FB87FD7DC:122F29BCEB238F49FCC894CEB8B5D2E5" box="[819,1020,1230,1257]" name="Pematang Siantar" pageId="5" pageNumber="439" stateProvince="Sumatera Utara">Pematang Siantar</location>
, ca.
<quantity id="D008D282EB238F49FBC994CFBFDAD2E4" box="[1074,1171,1230,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="439" unit="m" value="1200.0">
<elevation id="9CDD9854EB238F49FBC994CFBFDAD2E4" box="[1074,1171,1230,1257]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="439" unit="m" value="1200.0">1,200 m</elevation>
</quantity>
,
<date id="634E59A7EB238F49FB5A94CFBE2AD2E5" box="[1185,1379,1230,1256]" pageId="5" pageNumber="439" value="1992-08-24" valueMax="1992-08-26" valueMin="1992-08-24">
<collectingDate id="730AA04FEB238F49FB5A94CFBE2AD2E5" box="[1185,1379,1230,1256]" pageId="5" pageNumber="439" value="1992-08-24" valueMax="1992-08-26" valueMin="1992-08-24">24–26 Aug 1992</collectingDate>
</date>
,
<collectorName id="BA051AB1EB238F49FA8A94CEBBBED300" pageId="5" pageNumber="439">J.B. Heppner</collectorName>
&amp;
<collectorName id="BA051AB1EB238F49FEE094F2BAD3D301" box="[283,410,1266,1293]" pageId="5" pageNumber="439">E.W. Diehl</collectorName>
, gen. slide no. JBH-4149.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5FEA2CECEB238F49FF3C9517B978D005" pageId="5" pageNumber="439" type="diagnosis">
<paragraph id="174F7F67EB238F49FF3C9517B978D005" blockId="5.[151,1437,1230,2019]" pageId="5" pageNumber="439">
<emphasis id="2584A375EB238F49FF3C9517BA08D33C" bold="true" box="[199,321,1302,1329]" pageId="5" pageNumber="439">Diagnosis.</emphasis>
The new species is superficially similar to
<taxonomicName id="D0F004E4EB238F49FCD69516BFECD33C" authority="Park &amp; Abang, 2005" authorityName="Park &amp; Abang" authorityYear="2005" box="[813,1189,1302,1329]" class="Insecta" family="Lecithoceridae" genus="Thubana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="439" phylum="Arthropoda" rank="species" species="ochracea">
<emphasis id="2584A375EB238F49FCD69516B8F8D33D" box="[813,945,1302,1328]" italics="true" pageId="5" pageNumber="439">T. ochracea</emphasis>
<bibRefCitation id="73610296EB238F49FC429517BFECD33C" author="Park, K. - T. &amp; Sugisima, K. &amp; Abang, F." box="[953,1189,1302,1329]" pageId="5" pageNumber="439" pagination="133 - 141" refId="ref8631" refString="Park, K. - T., Sugisima, K. &amp; Abang, F. (2005) Genus Thubana in the Malaysian Borneo (Lepidoptera, Lecithoceridae) with descriptions of seven new species. Journal of Asia Pacific Entomology, 8 (2), 133 - 141. https: // doi. org / 10.1016 / S 1226 - 8615 (08) 60083 - 7" type="journal article" year="2005">Park &amp; Abang, 2005</bibRefCitation>
</taxonomicName>
, which was described from
<collectingCountry id="6FE73FF7EB238F49FF2D953BBA70D359" box="[214,313,1338,1364]" name="Malaysia" pageId="5" pageNumber="439">Sarawak</collectingCountry>
,
<collectingCountry id="6FE73FF7EB238F49FEBD953BBAE4D359" box="[326,429,1338,1364]" name="Malaysia" pageId="5" pageNumber="439">Malaysia</collectingCountry>
(
<typeStatus id="C84BC1C5EB238F49FE47953BB969D359" box="[444,544,1338,1364]" pageId="5" pageNumber="439" type="holotype">holotype</typeStatus>
in KNA), especially the dark-brown ground color of both wings, but it can be distinguished by the more or less elliptical costal patch and the termen of both wings being very oblique, and the fringe with white scales narrow only between R
<subScript id="8B747D22EB238F49FD539595B9F9D3AF" attach="left" box="[680,688,1428,1442]" fontSize="6" pageId="5" pageNumber="439">5</subScript>
and M
<subScript id="8B747D22EB238F49FD009594B84AD3AE" attach="left" box="[763,771,1429,1443]" fontSize="6" pageId="5" pageNumber="439">2</subScript>
, whereas in
<taxonomicName id="D0F004E4EB238F49FC769582BF59D391" authorityName="Park &amp; Abang" authorityYear="2005" box="[909,1040,1410,1436]" class="Insecta" family="Lecithoceridae" genus="Thubana" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="439" phylum="Arthropoda" rank="species" species="ochracea">
<emphasis id="2584A375EB238F49FC769582BF59D391" box="[909,1040,1410,1436]" italics="true" pageId="5" pageNumber="439">T. ochracea</emphasis>
</taxonomicName>
the costal patch is quadrate, the termen is less oblique with the white fringe from beyond apex to middle of termen. The male genitalia differ from each other by the cucullus having a row of strong spines along the ventral margin, and the juxta with a large, triangular median plate (illustrated in
<figureCitation id="8FCB63E2EB238F49FE3095EFB96CD005" box="[459,549,1518,1544]" captionStart="FIGURE 4" captionStartId="6.[151,250,1824,1848]" captionTargetBox="[161,1429,192,1784]" captionTargetId="figure-20@6.[151,1436,181,1800]" captionTargetPageId="6" captionText="FIGURE 4. Thubana spiniosa Park &amp; Heppner, sp. nov.: A, adult, holotype; B, head, dorsal view; C, head with labial palpus; D, male genitalia, slide no. JBH-4199; E, ditto, aedeagus; F, abdominal segments VII-VIII. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7758842" httpUri="https://zenodo.org/record/7758842/files/figure.png" pageId="5" pageNumber="439">Fig. 4D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5FEA2CECEB238F4BFF3C9613BAD8D768" lastPageId="7" lastPageNumber="441" pageId="5" pageNumber="439" type="description">
<paragraph id="174F7F67EB238F49FF3C9613B854D020" blockId="5.[151,1437,1230,2019]" box="[199,797,1554,1581]" pageId="5" pageNumber="439">
<emphasis id="2584A375EB238F49FF3C9613BA18D021" bold="true" box="[199,337,1554,1580]" pageId="5" pageNumber="439">Description</emphasis>
. Male (
<figureCitation id="8FCB63E2EB238F49FE539613B96FD021" box="[424,550,1554,1580]" captionStart="FIGURE 4" captionStartId="6.[151,250,1824,1848]" captionTargetBox="[161,1429,192,1784]" captionTargetId="figure-20@6.[151,1436,181,1800]" captionTargetPageId="6" captionText="FIGURE 4. Thubana spiniosa Park &amp; Heppner, sp. nov.: A, adult, holotype; B, head, dorsal view; C, head with labial palpus; D, male genitalia, slide no. JBH-4199; E, ditto, aedeagus; F, abdominal segments VII-VIII. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7758842" httpUri="https://zenodo.org/record/7758842/files/figure.png" pageId="5" pageNumber="439">Figs 4A, B</figureCitation>
). Wingspan 19.0 mm.
</paragraph>
<paragraph id="174F7F67EB238F49FF3C9637BABAD0ED" blockId="5.[151,1437,1230,2019]" pageId="5" pageNumber="439">
<emphasis id="2584A375EB238F49FF3C9637BA4DD05D" box="[199,260,1590,1616]" italics="true" pageId="5" pageNumber="439">Head</emphasis>
: frons shiny creamy white; vertex same color, mixed with brownish scales posteriorly, with orange-white erect scales laterally. Antenna longer than forewing, about 1.3 times; scape elongated, slightly broadened distally, brownish dorsally; flagellum yellowish white, with very weak annulations. Second segment of labial palpus (
<figureCitation id="8FCB63E2EB238F49FA94967FBBF0D0B1" captionStart="FIGURE 4" captionStartId="6.[151,250,1824,1848]" captionTargetBox="[161,1429,192,1784]" captionTargetId="figure-20@6.[151,1436,181,1800]" captionTargetPageId="6" captionText="FIGURE 4. Thubana spiniosa Park &amp; Heppner, sp. nov.: A, adult, holotype; B, head, dorsal view; C, head with labial palpus; D, male genitalia, slide no. JBH-4199; E, ditto, aedeagus; F, abdominal segments VII-VIII. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7758842" httpUri="https://zenodo.org/record/7758842/files/figure.png" pageId="5" pageNumber="439">Fig. 4C</figureCitation>
) thickened, strongly arched and upturned, mustard brown on outer surface; 3rd segment slender, longer than 2nd segment, dark brown ventrally.
</paragraph>
<paragraph id="174F7F67EB238F49FF3C96EBBA6DD1D0" blockId="5.[151,1437,1230,2019]" pageId="5" pageNumber="439">
<emphasis id="2584A375EB238F49FF3C96EBBA51D109" box="[199,280,1770,1796]" italics="true" pageId="5" pageNumber="439">Thorax</emphasis>
: tegula and thorax dark mustard brown, with some yellowish-white, long, erect scales laterally. Hind tibia with yellowish-white rough scales mixing with brownish scales dorsally. Forewing ground color dark mustard brown; costa nearly straight before 3/4; costal patch more or less elliptical, yellowish-white, just beyond middle, with three small, yellowish-white, short streaks before apex; apex obtuse; termen oblique; fringe with narrow, white basal line, with white scales narrowly between R
<subScript id="8B747D22EB238F49FD3B978DB981D197" attach="left" box="[704,712,1932,1946]" fontSize="6" pageId="5" pageNumber="439">5</subScript>
and M
<subScript id="8B747D22EB238F49FCEE978CB854D196" attach="left" box="[789,797,1933,1947]" fontSize="6" pageId="5" pageNumber="439">2</subScript>
. Hindwing ground color same as that of forewing, without any markings; apex sharply produced; termen very oblique, slightly sinuate; fringe concolorous, but few white scales at M
<subScript id="8B747D22EB238F49FEEE97D4BA54D1EE" attach="left" box="[277,285,2005,2019]" fontSize="6" pageId="5" pageNumber="439">1</subScript>
.
</paragraph>
<caption id="438F2FEFEB208F4AFF6C9721BFBFD156" ID-DOI="http://doi.org/10.5281/zenodo.7758842" ID-Zenodo-Dep="7758842" httpUri="https://zenodo.org/record/7758842/files/figure.png" pageId="6" pageNumber="440" startId="6.[151,250,1824,1848]" targetBox="[161,1429,192,1784]" targetPageId="6">
<paragraph id="174F7F67EB208F4AFF6C9721BFBFD156" blockId="6.[151,1436,1824,1884]" pageId="6" pageNumber="440">
<emphasis id="2584A375EB208F4AFF6C9721BA44D135" bold="true" box="[151,269,1824,1848]" pageId="6" pageNumber="440">FIGURE 4</emphasis>
.
<taxonomicName id="D0F004E4EB208F4AFEE19721B934D13A" authority="Park &amp; Heppner" authorityName="Park &amp; Heppner" authorityYear="2023" box="[282,637,1824,1848]" class="Insecta" family="Lecithoceridae" genus="Thubana" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="440" phylum="Arthropoda" rank="species" species="spiniosa" status="sp. nov.">
<emphasis id="2584A375EB208F4AFEE19721BA86D13A" box="[282,463,1824,1848]" italics="true" pageId="6" pageNumber="440">Thubana spiniosa</emphasis>
Park &amp; Heppner
</taxonomicName>
,
<emphasis id="2584A375EB208F4AFD719721B992D135" bold="true" box="[650,731,1824,1848]" pageId="6" pageNumber="440">
<taxonomicNameLabel id="3EB71E0EEB208F4AFD719721B992D135" box="[650,731,1824,1848]" pageId="6" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, adult, holotype; B, head, dorsal view; C, head with labial palpus; D, male genitalia, slide no. JBH-4199; E, ditto, aedeagus; F, abdominal segments VII-VIII. Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="174F7F67EB208F4AFF3C978BBA9ED1C5" blockId="6.[151,1436,1930,2029]" pageId="6" pageNumber="440">
<emphasis id="2584A375EB208F4AFF3C978BBA78D1A9" box="[199,305,1930,1956]" italics="true" pageId="6" pageNumber="440">Abdomen</emphasis>
: mustard brown on dorsal surface; spinous zones on tergites broadly developed; abdominal segments VI-VIII as shown in
<figureCitation id="8FCB63E2EB208F4AFE7B97AFBA9AD1C5" box="[384,467,1966,1992]" captionStart="FIGURE 4" captionStartId="6.[151,250,1824,1848]" captionTargetBox="[161,1429,192,1784]" captionTargetId="figure-20@6.[151,1436,181,1800]" captionTargetPageId="6" captionText="FIGURE 4. Thubana spiniosa Park &amp; Heppner, sp. nov.: A, adult, holotype; B, head, dorsal view; C, head with labial palpus; D, male genitalia, slide no. JBH-4199; E, ditto, aedeagus; F, abdominal segments VII-VIII. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7758842" httpUri="https://zenodo.org/record/7758842/files/figure.png" pageId="6" pageNumber="440">
Fig.
<specimenCount id="01F6B4EEEB208F4AFE4F97AFBA9AD1C5" box="[436,467,1966,1992]" pageId="6" pageNumber="440" type="generic">4F</specimenCount>
</figureCitation>
.
</paragraph>
<paragraph id="174F7F67EB208F4BFF3C97D3B9AFD74C" blockId="6.[151,1436,1930,2029]" lastBlockId="7.[151,1436,151,465]" lastPageId="7" lastPageNumber="441" pageId="6" pageNumber="440">
<emphasis id="2584A375EB208F4AFF3C97D3BA27D1E1" box="[199,366,2002,2028]" italics="true" pageId="6" pageNumber="440">Male genitalia</emphasis>
(
<figureCitation id="8FCB63E2EB208F4AFE8597D3B948D1E1" box="[382,513,2002,2028]" captionStart="FIGURE 4" captionStartId="6.[151,250,1824,1848]" captionTargetBox="[161,1429,192,1784]" captionTargetId="figure-20@6.[151,1436,181,1800]" captionTargetPageId="6" captionText="FIGURE 4. Thubana spiniosa Park &amp; Heppner, sp. nov.: A, adult, holotype; B, head, dorsal view; C, head with labial palpus; D, male genitalia, slide no. JBH-4199; E, ditto, aedeagus; F, abdominal segments VII-VIII. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7758842" httpUri="https://zenodo.org/record/7758842/files/figure.png" pageId="6" pageNumber="440">Figs 4C, D</figureCitation>
): Uncus slender, nearly parallel sided. Mesial process of gnathos relatively long, strongly bent beyond 2/3. Tegumen deeply concave on anterior margin Valva broad basally, narrowed at conjunction with cucullus; costa deeply concave before cucullus; cucullus thumb-like, densely setose, with a row of strong spines along outer margin; apex rounded. Juxta broad, short, with large, triangular mesial plate. Vinculum narrow, banded. Aedeagus shorter than valva, broad basally, with several conical spines in apical part dorsally; cornuti consisting of several short, sclerotized bars as in
<figureCitation id="8FCB63E2EB218F4BFD689126B9ABD74C" box="[659,738,295,321]" captionStart="FIGURE 4" captionStartId="6.[151,250,1824,1848]" captionTargetBox="[161,1429,192,1784]" captionTargetId="figure-20@6.[151,1436,181,1800]" captionTargetPageId="6" captionText="FIGURE 4. Thubana spiniosa Park &amp; Heppner, sp. nov.: A, adult, holotype; B, head, dorsal view; C, head with labial palpus; D, male genitalia, slide no. JBH-4199; E, ditto, aedeagus; F, abdominal segments VII-VIII. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7758842" httpUri="https://zenodo.org/record/7758842/files/figure.png" pageId="7" pageNumber="441">Fig 4E</figureCitation>
.
</paragraph>
<paragraph id="174F7F67EB218F4BFF3C914ABAD8D768" blockId="7.[151,1436,151,465]" box="[199,401,331,357]" pageId="7" pageNumber="441">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="5FEA2CECEB218F4BFF3C916FB93CD784" box="[199,629,366,393]" pageId="7" pageNumber="441" type="distribution">
<paragraph id="174F7F67EB218F4BFF3C916FB93CD784" blockId="7.[151,1436,151,465]" box="[199,629,366,393]" pageId="7" pageNumber="441">
<emphasis id="2584A375EB218F4BFF3C916FBA29D784" bold="true" box="[199,352,366,393]" pageId="7" pageNumber="441">Distribution.</emphasis>
<collectingCountry id="6FE73FF7EB218F4BFE9C916EBA9DD784" box="[359,468,367,393]" name="Indonesia" pageId="7" pageNumber="441">Indonesia</collectingCountry>
(N.
<collectingRegion id="D534B185EB218F4BFDFC916EB923D784" box="[519,618,367,393]" country="Indonesia" name="Sumatera Utara" pageId="7" pageNumber="441">Sumatra</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="5FEA2CECEB218F4BFF3C9192B87FD7DC" pageId="7" pageNumber="441" type="etymology">
<paragraph id="174F7F67EB218F4BFF3C9192B87FD7DC" blockId="7.[151,1436,151,465]" pageId="7" pageNumber="441">
<emphasis id="2584A375EB218F4BFF3C9192BA06D7A0" bold="true" box="[199,335,403,429]" pageId="7" pageNumber="441">Etymology.</emphasis>
The specific name is derived from the Latin,
<emphasis id="2584A375EB218F4BFCBD9192B8CAD7A1" box="[838,899,403,428]" italics="true" pageId="7" pageNumber="441">spina</emphasis>
(= spine) with Latin suffix, -
<emphasis id="2584A375EB218F4BFB399192BFA0D7A1" box="[1218,1257,403,428]" italics="true" pageId="7" pageNumber="441">osa</emphasis>
, referring to the spines along the outer margin of valva in the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>
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