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<document ID-DOI="http://doi.org/10.5281/zenodo.6485242" ID-GBIF-Dataset="3dfc64e7-8c23-456d-9678-1bb0858ac470" ID-GBIF-Taxon="150645364" ID-Zenodo-Dep="6485242" checkinTime="1545431109525" checkinUser="PlaziZenodoSync" docAuthor="Jan van Tol &amp; André Günther" docDate="2018" docId="0395A91CAC52FFA1951EFF52FD5EFC2E" docLanguage="en" docName="10.5281zenodo.1481114_vanTol_Guenther_2018.pdf" docOrigin="Odonatologica 47 (3)" docStyle="DocumentStyle{}" docTitle="Rhinocypha frontalis subsp. sulselensis Jan van Tol &amp; André Günther 2018, ssp. nov." docType="treatment" docVersion="4" lastPageId="39" lastPageNumber="338" masterDocId="FFACD164AC71FF869443FFAFFF05FFBC" masterDocTitle="The Odonata of Sulawesi and adjacent islands. Part 8. Revision of the genus Rhinocypha Rambur, 1842 (Chlorocyphidae)" masterLastPageNumber="386" masterPageNumber="299" originalUpdateDomain="Boston" originalUpdateTime="1544520335069" originalUpdateUser="PlaziZenodoSync" pageId="35" pageNumber="334" updateTime="1650928538772" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The Odonata of Sulawesi and adjacent islands. Part 8. Revision of the genus Rhinocypha Rambur, 1842 (Chlorocyphidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jan van Tol</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>André Günther</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Odonatologica</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-12-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>47</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>299</mods:start>
<mods:end>386</mods:end>
</mods:extent>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.1481114</mods:identifier>
<mods:identifier type="GBIF-Dataset">3dfc64e7-8c23-456d-9678-1bb0858ac470</mods:identifier>
<mods:identifier type="Zenodo-Dep">1481114</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6485242" ID-GBIF-Taxon="150645364" ID-Zenodo-Dep="6485242" LSID="urn:lsid:plazi:treatment:0395A91CAC52FFA1951EFF52FD5EFC2E" httpUri="http://treatment.plazi.org/id/0395A91CAC52FFA1951EFF52FD5EFC2E" lastPageId="39" lastPageNumber="338" pageId="35" pageNumber="334">
<subSubSection box="[349,845,253,281]" pageId="35" pageNumber="334" type="nomenclature">
<paragraph blockId="35.[159,1042,253,724]" box="[349,845,253,281]" pageId="35" pageNumber="334">
<heading bold="true" box="[349,845,253,281]" centered="true" fontSize="11" level="2" pageId="35" pageNumber="334" reason="4">
<emphasis bold="true" box="[349,845,253,281]" pageId="35" pageNumber="334">
<taxonomicName authority="Jan van Tol &amp; André Günther, 2018" authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" box="[349,734,253,281]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="334" phylum="Arthropoda" rank="subSpecies" species="frontalis" status="ssp. nov." subSpecies="sulselensis">
<emphasis bold="true" box="[349,734,253,281]" italics="true" pageId="35" pageNumber="334">Rhinocypha frontalis sulselensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[741,845,253,281]" pageId="35" pageNumber="334" rank="subSpecies">
<taxonomicName box="[741,841,253,281]" pageId="35" pageNumber="334" rank="subSpecies" subSpecies="nov">ssp. nov</taxonomicName>
.
</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[482,713,287,315]" pageId="35" pageNumber="334" type="description">
<paragraph blockId="35.[159,1042,253,724]" box="[482,713,287,315]" pageId="35" pageNumber="334">
<heading box="[482,713,287,315]" centered="true" fontSize="11" level="4" pageId="35" pageNumber="334" reason="4">
(
<figureCitation box="[488,577,287,315]" captionStart="Figure 34" captionStartId="37.[159,227,1103,1128]" captionTargetBox="[157,1036,529,1062]" captionTargetId="figure@37.[157,1036,529,1062]" captionTargetPageId="37" captionText="Figure 34. Rhinocypha frontalis sulselensis ssp. nov. Male, holotype. Sulawesi Selatan, Borong Rapoa, 2023-viii-1949 [JvT 22887]. Photo: JvT" httpUri="https://zenodo.org/record/2200463/files/figure.png" pageId="35" pageNumber="334">Figs 34</figureCitation>
,
<figureCitation box="[587,621,287,315]" captionStart="Figure 35" captionStartId="37.[651,719,1433,1458]" captionTargetBox="[169,619,1216,1538]" captionTargetId="figure@37.[156,627,1200,1576]" captionTargetPageId="37" captionText="Figure 35. Rhinocypha frontalis sulselensis ssp. nov. Male, head, dorsal view. Locality as in Fig. 34. Artwork by JvT" httpUri="https://zenodo.org/record/2200465/files/figure.png" pageId="35" pageNumber="334">35</figureCitation>
,
<figureCitation box="[630,664,287,315]" captionStart="Figures 3637" captionStartId="38.[159,238,1170,1195]" captionTargetBox="[469,1036,782,1539]" captionTargetPageId="38" captionText="Figures 3637. Rhinocypha frontalis subspecies. Wings, female. 36 — Subspecies frontalis. Sulawesi Utara, surroundings Tondano, 09-vi-1940 [JvT 22866]; 37 — subspecies sulselensis. Sulawesi Selatan, Boeloedoea, vi-1935 [JvT 22891]. Both with same magnification. Photos: JvT" httpUri="https://zenodo.org/record/2200471/files/figure.png" pageId="35" pageNumber="334">37</figureCitation>
, 84)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="334" type="materials_examined">
<paragraph blockId="35.[159,1042,253,724]" box="[159,360,333,359]" pageId="35" pageNumber="334">
<heading bold="true" box="[159,360,333,359]" fontSize="11" level="3" pageId="35" pageNumber="334" reason="6">
<emphasis bold="true" box="[159,360,333,359]" pageId="35" pageNumber="334">Material studied</emphasis>
</heading>
</paragraph>
<paragraph blockId="35.[159,1042,253,724]" pageId="35" pageNumber="334">
<materialsCitation ID-GBIF-Occurrence="1949138302" collectingDate="1949-08-20" collectingDateMax="1949-08-23" collectingDateMin="1949-08-20" location="Leg. A. Diakonoff" municipality="Borong Rapoa" pageId="35" pageNumber="334" specimenCode="JvT 22887" specimenCount="801" specimenCount-male="801" typeStatus="holotype">
<typeStatus box="[159,273,371,397]" pageId="35" pageNumber="334">
<emphasis bold="true" box="[159,273,371,397]" pageId="35" pageNumber="334">Holotype</emphasis>
</typeStatus>
<specimenCount box="[280,302,371,397]" pageId="35" pageNumber="334" type="male"></specimenCount>
.
<emphasis bold="true" box="[316,329,371,397]" pageId="35" pageNumber="334">»</emphasis>
<collectingMunicipality box="[330,503,371,397]" pageId="35" pageNumber="334">Borong Rapoa</collectingMunicipality>
.
<specimenCount box="[513,594,371,397]" pageId="35" pageNumber="334" type="male">
<quantity box="[513,590,371,397]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="35" pageNumber="334" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
<collectingDate box="[601,802,371,397]" pageId="35" pageNumber="334" value="1949-08-20" valueMax="1949-08-23" valueMin="1949-08-20">
<date box="[601,802,371,397]" pageId="35" pageNumber="334" value="1949-08-20" valueMax="1949-08-23" valueMin="1949-08-20">2023 Aug. 1949</date>
</collectingDate>
.
<location LSID="urn:lsid:plazi:treatment:0395A91CAC52FFA1951EFF52FD5EFC2E:8EE34ED1AC52FFA5976EFEDCFB07FE31" box="[813,1026,371,397]" municipality="Borong Rapoa" name="Leg. A. Diakonoff" pageId="35" pageNumber="334">Leg. A. Diakonoff</location>
« (
<specimenCode box="[167,290,408,434]" pageId="35" pageNumber="334">JvT 22887</specimenCode>
; in RMNH) (
<figureCitation box="[444,529,408,434]" captionStart="Figure 34" captionStartId="37.[159,227,1103,1128]" captionTargetBox="[157,1036,529,1062]" captionTargetId="figure@37.[157,1036,529,1062]" captionTargetPageId="37" captionText="Figure 34. Rhinocypha frontalis sulselensis ssp. nov. Male, holotype. Sulawesi Selatan, Borong Rapoa, 2023-viii-1949 [JvT 22887]. Photo: JvT" httpUri="https://zenodo.org/record/2200463/files/figure.png" pageId="35" pageNumber="334">Fig. 34</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph blockId="35.[159,1042,253,724]" box="[159,384,445,472]" pageId="35" pageNumber="334">
<emphasis bold="true" box="[159,279,446,472]" pageId="35" pageNumber="334">Paratypes</emphasis>
(7♂ 4♀)
</paragraph>
<paragraph blockId="35.[159,1042,253,724]" box="[159,341,485,509]" pageId="35" pageNumber="334">
<emphasis bold="true" box="[159,341,485,509]" pageId="35" pageNumber="334">Sulawesi Selatan</emphasis>
</paragraph>
<paragraph blockId="35.[159,1042,253,724]" pageId="35" pageNumber="334">1♀, Boeloedoea (Barroe). 670 m. June 1935. Leg. C. Veen; 1♀, Boeloedoea (Bar- roe). Brooklet in forest. CdR XXXIII. 27 Apr. 1941. Leg. L. Coomans de Ruiter; 6♂ 1♀, Borong Rapoa. 800 m. 2023 Aug. 1949 [some handwritten labels with erroneous year 1948]. Leg. A. Diakonoff (same data as holotype); 1♀, Mangkoso, stream emerging from forest. Alt. at least 400 m. July 2013. Leg. R.A. Dow; 1♂, 20 km E of Padaelo: mountain stream crossing Jl. Poros Barru-Soppeng (4.51082°S, 119.78648°E). Alt. 698 m. 20 July 2013. Leg. H. Cahilog.</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="334" type="etymology">
<paragraph blockId="35.[159,1036,754,855]" box="[159,292,754,780]" pageId="35" pageNumber="334">
<heading bold="true" box="[159,292,754,780]" fontSize="11" level="3" pageId="35" pageNumber="334" reason="6">
<emphasis bold="true" box="[159,292,754,780]" pageId="35" pageNumber="334">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="35.[159,1036,754,855]" pageId="35" pageNumber="334">
<emphasis box="[159,279,791,817]" italics="true" pageId="35" pageNumber="334">Sulselensis</emphasis>
, an adjective based on the frequently used abbreviation SulSel, for the province of Sulawesi Selatan, or South Sulawesi.
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="334" type="diagnosis">
<paragraph blockId="35.[159,1036,887,1549]" box="[159,281,887,913]" pageId="35" pageNumber="334">
<heading bold="true" box="[159,281,887,913]" fontSize="11" level="3" pageId="35" pageNumber="334" reason="6">
<emphasis bold="true" box="[159,281,887,913]" pageId="35" pageNumber="334">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="35.[159,1036,887,1549]" pageId="35" pageNumber="334">
Largely agrees with specimens assigned to the nominotypical subspecies of
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[159,287,962,988]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="334" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[159,287,962,988]" italics="true" pageId="35" pageNumber="334">R. frontalis</emphasis>
</taxonomicName>
, but is distinguished by its significantly larger size, and a more extensive black area in the wing space between the Costa and Subcosta,
<emphasis box="[995,1029,1000,1025]" italics="true" pageId="35" pageNumber="334">i.e.</emphasis>
, from 8 Ax anterior to the nodus in the fore wing and 6 Ax anterior to the nodus in the hind wing (
<figureCitation box="[446,530,1074,1100]" captionStart="Figure 34" captionStartId="37.[159,227,1103,1128]" captionTargetBox="[157,1036,529,1062]" captionTargetId="figure@37.[157,1036,529,1062]" captionTargetPageId="37" captionText="Figure 34. Rhinocypha frontalis sulselensis ssp. nov. Male, holotype. Sulawesi Selatan, Borong Rapoa, 2023-viii-1949 [JvT 22887]. Photo: JvT" httpUri="https://zenodo.org/record/2200463/files/figure.png" pageId="35" pageNumber="334">Fig. 34</figureCitation>
, compare with
<figureCitation box="[709,779,1074,1100]" captionStart="Figure 3" captionStartId="14.[159,227,1021,1046]" captionTargetBox="[234,961,653,982]" captionTargetId="figure@14.[234,961,653,982]" captionTargetPageId="14" captionText="Figure 3. Rhinocypha frontalis, male. Stream Sg. Tumpa kecil, North Sulawesi, Bogani-Nani-Wartabone NP. Photo: AG (10-viii-2011)" httpUri="https://zenodo.org/record/2200381/files/figure.png" pageId="35" pageNumber="334">Figs 3</figureCitation>
,
<figureCitation box="[792,824,1074,1100]" captionStart="Figure 24" captionStartId="25.[159,227,1494,1519]" captionTargetBox="[355,994,1033,1455]" captionTargetId="figure@25.[200,994,1033,1455]" captionTargetPageId="25" captionText="Figure 24. Rhinocypha f. frontalis. Male, lectotype. [NHMUK 012500438]. Photo: B. Price, Natural History Museum, London" httpUri="https://zenodo.org/record/2200443/files/figure.png" pageId="35" pageNumber="334">24</figureCitation>
); opaque marking in wings wedge-shaped, and base reaching to distal side of q. The female is distinctly different from the nominotypical subspecies, particularly in the wing marking, although the nominotypical subspecies shows significant variation in these characters as well. In
<taxonomicName authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" box="[622,873,1223,1249]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="334" phylum="Arthropoda" rank="subSpecies" species="frontalis" subSpecies="sulselensis">
<emphasis box="[622,873,1223,1249]" italics="true" pageId="35" pageNumber="334">R. frontalis sulselensis</emphasis>
</taxonomicName>
the wings are nearly fully brown (
<figureCitation box="[393,478,1261,1287]" captionStart="Figures 3637" captionStartId="38.[159,238,1170,1195]" captionTargetBox="[469,1036,782,1539]" captionTargetPageId="38" captionText="Figures 3637. Rhinocypha frontalis subspecies. Wings, female. 36 — Subspecies frontalis. Sulawesi Utara, surroundings Tondano, 09-vi-1940 [JvT 22866]; 37 — subspecies sulselensis. Sulawesi Selatan, Boeloedoea, vi-1935 [JvT 22891]. Both with same magnification. Photos: JvT" httpUri="https://zenodo.org/record/2200471/files/figure.png" pageId="35" pageNumber="334">Fig. 37</figureCitation>
, compare with
<figureCitation box="[663,747,1261,1288]" captionStart="Figures 3637" captionStartId="38.[159,238,1170,1195]" captionTargetBox="[469,1036,782,1539]" captionTargetPageId="38" captionText="Figures 3637. Rhinocypha frontalis subspecies. Wings, female. 36 — Subspecies frontalis. Sulawesi Utara, surroundings Tondano, 09-vi-1940 [JvT 22866]; 37 — subspecies sulselensis. Sulawesi Selatan, Boeloedoea, vi-1935 [JvT 22891]. Both with same magnification. Photos: JvT" httpUri="https://zenodo.org/record/2200471/files/figure.png" pageId="35" pageNumber="334">Fig. 36</figureCitation>
): the base is very darkly enfumed, approximately to the level of the nodus, merging to the opaque part of the wing distal to nodus; apex of wings distal to middle of pterostigma paler, the tip opaque white. The females are thus rather similar to sympatric females of
<taxonomicName box="[362,523,1410,1436]" pageId="35" pageNumber="334">
<emphasis box="[362,523,1410,1436]" italics="true" pageId="35" pageNumber="334">R. monochroa</emphasis>
</taxonomicName>
(
<figureCitation box="[538,610,1410,1437]" captionStart="Figure 6" captionStartId="15.[828,896,675,700]" captionText="Figure 6. Rhinocypha monochroa, female. Stream Sg. Pattunuang Asue, Southwest Sulawesi. Photo: AG (01- -viii-2011)" pageId="35" pageNumber="334">Fig. 6</figureCitation>
). However, the latter species usually with fore wing transparent distal to
<emphasis box="[590,615,1448,1473]" italics="true" pageId="35" pageNumber="334">ca</emphasis>
14 Px anterior to pterostigma, and also has a fully (brownish) black frons; its pronotum has an oblong spot on the median lobe; the lateral lobe has only a small spot.
</paragraph>
</subSubSection>
<subSubSection lastPageId="37" lastPageNumber="336" pageId="36" pageNumber="335" type="description">
<paragraph blockId="36.[159,1036,253,541]" box="[159,305,253,279]" pageId="36" pageNumber="335">
<heading bold="true" box="[159,305,253,279]" fontSize="11" level="3" pageId="36" pageNumber="335" reason="6">
<emphasis bold="true" box="[159,305,253,279]" pageId="36" pageNumber="335">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="36.[159,1036,253,541]" box="[159,481,291,317]" pageId="36" pageNumber="335">
<emphasis bold="true" box="[159,220,291,317]" pageId="36" pageNumber="335">Male</emphasis>
(holotype, JvT 22887)
</paragraph>
<paragraph blockId="36.[159,1036,253,541]" pageId="36" pageNumber="335">Head — Labium, labrum and anteclypeus glossy brownish black; mandi- bles blue, except for a narrow black stripe bordering labrum; dorsal side of head velvet black; frons with a paired blue spot, large and squarish, tapering abaxiad; very small creamish spots behind lateral ocelli; spots on post-ocu- lar lobe sub-oval, larger than those on vertex; gena with large blue marking, somewhat extending along eye.</paragraph>
<paragraph blockId="36.[159,1037,589,952]" pageId="36" pageNumber="335">Thorax — Pronotum velvet black; anterior lobe with relative large oval, creamy, paired spot; lateral lobe fully covered by large creamy marking, and an oblong creamy stripe along margin below lateral lobe.</paragraph>
<paragraph blockId="36.[159,1037,589,952]" pageId="36" pageNumber="335">Synthorax. Mesopreepisternum creamish. Synthorax black, with pale markings as follows: a short pale stripe on dorsal carina in mesothoracic triangle; ante-humeral stripe about ⅔ the length of mesepisternum, taper- ing posteriad; mesepimeron posteriorly with short and narrow stripe; stripe over lower side of synthorax from mesokatepisternum to posterior corner of metepimeron, dorso-posterior projection on metepisternum not reaching posterior margin; coxa of all legs with narrow stripe along posterior margin.</paragraph>
<paragraph blockId="36.[159,842,998,1029]" box="[159,842,998,1029]" pageId="36" pageNumber="335">Legs —Black; tibiae of mid and hind legs white in distal ⁴/₅.</paragraph>
<paragraph blockId="36.[159,1035,1075,1213]" pageId="36" pageNumber="335">Wings — Base of wings transparent, somewhat enfumed; both wings with opaque brown band distal to Ax8 in costal and subcostal space, wedge- shaped and reaching to first fork in Rs; quadrangle transparent; pterostigma of fore and hind wings dark brown.</paragraph>
<paragraph blockId="36.[159,1036,1261,1325]" pageId="36" pageNumber="335">Abdomen — Dorsum black, with lateral blue markings similar to nomino- typical form.</paragraph>
<paragraph blockId="36.[159,1037,1374,1549]" box="[159,247,1374,1400]" pageId="36" pageNumber="335">
<heading bold="true" box="[159,247,1374,1400]" fontSize="11" level="3" pageId="36" pageNumber="335" reason="6">
<emphasis bold="true" box="[159,247,1374,1400]" pageId="36" pageNumber="335">Female</emphasis>
</heading>
</paragraph>
<paragraph blockId="36.[159,1037,1374,1549]" pageId="36" pageNumber="335">
Larger than nominotypical form; coloration very similar to that form, except for the coloration of the wings: fully brown, with base enfumed very dark, subtly merging towards the fully opaque distal part of the wing, distal to
<emphasis box="[1011,1036,1486,1511]" italics="true" pageId="36" pageNumber="335">ca</emphasis>
level of nodus; apex distal to middle of pterostigma paler, the tip opaque white.
</paragraph>
<paragraph blockId="37.[159,1036,253,504]" box="[159,431,253,280]" pageId="37" pageNumber="336">
<heading bold="true" box="[159,431,253,280]" fontSize="11" level="3" pageId="37" pageNumber="336" reason="6">
<emphasis bold="true" box="[159,431,253,280]" pageId="37" pageNumber="336">Variation in paratypes</emphasis>
</heading>
</paragraph>
<paragraph blockId="37.[159,1036,253,504]" pageId="37" pageNumber="336">Measurements of male specimens [mm] — Hind wing length length (n =4) x= 27.5 (27.027.5), hind wing width (n = 4) x =7.0 (6.57.0); abdomen (in- cluding appendages) x =22.5 (22.022.5).</paragraph>
<paragraph blockId="37.[159,1036,253,504]" pageId="37" pageNumber="336">Measurements of female specimens [mm] —Hind wing length (n =3) x= 30.0 (29.031.0); hind wing width (n = 3) x =6.5 (6.57.0); abdomen (in- cluding appendages) (n = 3) x =21.5 (20.022.5).</paragraph>
</subSubSection>
<caption httpUri="https://zenodo.org/record/2200463/files/figure.png" pageId="37" pageNumber="336" startId="37.[159,227,1103,1128]" targetBox="[157,1036,529,1062]" targetPageId="37">
<paragraph blockId="37.[159,1036,1102,1157]" pageId="37" pageNumber="336">
<emphasis bold="true" box="[159,268,1103,1128]" pageId="37" pageNumber="336">Figure 34.</emphasis>
<taxonomicName authority="Jan van Tol &amp; André Günther, 2018" authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" box="[275,617,1102,1126]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="336" phylum="Arthropoda" rank="subSpecies" species="frontalis" status="ssp. nov." subSpecies="sulselensis">
<taxonomicName box="[275,497,1102,1126]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="336" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis box="[275,617,1102,1126]" italics="true" pageId="37" pageNumber="336">
<taxonomicName box="[275,401,1102,1126]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="336" phylum="Arthropoda" rank="genus">Rhinocypha</taxonomicName>
<taxonomicName box="[408,617,1102,1126]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="336" phylum="Arthropoda" rank="subSpecies" species="frontalis" status="ssp. nov." subSpecies="sulselensis">
<taxonomicName box="[408,497,1102,1126]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="336" phylum="Arthropoda" rank="species" species="frontalis">frontalis</taxonomicName>
<taxonomicName box="[504,617,1102,1126]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="336" phylum="Arthropoda" rank="subSpecies" species="frontalis" status="ssp. nov." subSpecies="sulselensis">sulselensis</taxonomicName>
</taxonomicName>
</emphasis>
</taxonomicName>
</taxonomicName>
<taxonomicNameLabel box="[623,717,1102,1127]" pageId="37" pageNumber="336" rank="subSpecies">
<taxonomicName box="[623,714,1102,1127]" pageId="37" pageNumber="336" rank="subSpecies" subSpecies="nov">ssp. nov</taxonomicName>
.
</taxonomicNameLabel>
Male, holotype. Sulawesi Selatan, Borong Rapoa, 2023-viii-1949 [JvT 22887]. Photo: JvT
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/2200465/files/figure.png" pageId="37" pageNumber="336" startId="37.[651,719,1433,1458]" targetBox="[169,619,1216,1538]" targetPageId="37">
<paragraph blockId="37.[651,1037,1433,1546]" pageId="37" pageNumber="336">
<emphasis bold="true" box="[651,775,1433,1458]" pageId="37" pageNumber="336">Figure 35.</emphasis>
<taxonomicName authority="Jan van Tol &amp; André Günther, 2018" authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="336" phylum="Arthropoda" rank="subSpecies" species="frontalis" status="ssp. nov." subSpecies="sulselensis">
<taxonomicName box="[798,1035,1433,1457]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="336" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis italics="true" pageId="37" pageNumber="336">
<taxonomicName box="[798,924,1433,1457]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="336" phylum="Arthropoda" rank="genus">Rhinocypha</taxonomicName>
<taxonomicName class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="336" phylum="Arthropoda" rank="subSpecies" species="frontalis" status="ssp. nov." subSpecies="sulselensis">
<taxonomicName box="[946,1035,1433,1457]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="37" pageNumber="336" phylum="Arthropoda" rank="species" species="frontalis">frontalis</taxonomicName>
sulselensis
</taxonomicName>
</emphasis>
</taxonomicName>
</taxonomicName>
<taxonomicNameLabel box="[779,883,1462,1487]" pageId="37" pageNumber="336" rank="subSpecies">
<taxonomicName box="[779,879,1462,1487]" pageId="37" pageNumber="336" rank="subSpecies" subSpecies="nov">ssp. nov</taxonomicName>
.
</taxonomicNameLabel>
Male, head, dorsal view. Locality as in Fig. 34. Artwork by JvT
</paragraph>
</caption>
<subSubSection pageId="38" pageNumber="337" type="distribution">
<paragraph blockId="38.[159,1036,253,392]" box="[159,459,253,280]" pageId="38" pageNumber="337">
<heading bold="true" box="[159,459,253,280]" fontSize="11" level="3" pageId="38" pageNumber="337" reason="6">
<emphasis bold="true" box="[159,459,253,280]" pageId="38" pageNumber="337">Distribution and habitat</emphasis>
</heading>
</paragraph>
<paragraph blockId="38.[159,1036,253,392]" pageId="38" pageNumber="337">
South-western Sulawesi. The limited number of observations suggests that this species occurs above
<emphasis box="[464,489,329,354]" italics="true" pageId="38" pageNumber="337">ca</emphasis>
400 m a.s.l. However, we also have records of the sympatric
<taxonomicName box="[325,486,365,391]" pageId="38" pageNumber="337">
<emphasis box="[325,486,365,391]" italics="true" pageId="38" pageNumber="337">R. monochroa</emphasis>
</taxonomicName>
from altitudes up to 1 0 0 0 m.
</paragraph>
</subSubSection>
<subSubSection lastPageId="39" lastPageNumber="338" pageId="38" pageNumber="337" type="discussion">
<paragraph blockId="38.[159,1036,440,728]" box="[159,267,440,466]" pageId="38" pageNumber="337">
<heading bold="true" box="[159,267,440,466]" fontSize="11" level="3" pageId="38" pageNumber="337" reason="6">
<emphasis bold="true" box="[159,267,440,466]" pageId="38" pageNumber="337">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="38.[159,1036,440,728]" pageId="38" pageNumber="337">We identified this taxon from a series in RMNH, collected in 1949 (the year 1948 on some labels is incorrect) at Borong Rapoa, a village on the flank of the Mt. Lompobatang, east of Makassar, by A. Diakonoff. Other specimens in RMNH included a female with label »C. S. Celebes, 670 m / S. Bone, Bu- ludua / vi.1935 / C. Veen« (in RMNH, JvT 22891), and one female from the same site, collected by L. Coomans de Ruiter on 27-iv-1941 (JvT 22890), which both agree with a female from the Borong Rapoa series. The Buludua</paragraph>
<caption httpUri="https://zenodo.org/record/2200471/files/figure.png" pageId="38" pageNumber="337" targetBox="[469,1036,782,1539]" targetPageId="38">
<paragraph blockId="38.[159,432,1170,1547]" pageId="38" pageNumber="337">
<emphasis bold="true" box="[159,337,1170,1195]" pageId="38" pageNumber="337">Figures 3637.</emphasis>
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="38" pageNumber="337" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis italics="true" pageId="38" pageNumber="337">Rhinocypha frontalis</emphasis>
</taxonomicName>
subspecies. Wings, female. 36 — Subspecies
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="38" pageNumber="337" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis italics="true" pageId="38" pageNumber="337">frontalis</emphasis>
</taxonomicName>
. Sulawesi Utara, surroundings Tondano, 09-vi-1940 [JvT 22866]; 37 — subspecies
<taxonomicName authorityName="Jan van Tol &amp; André Günther" authorityYear="2018" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="38" pageNumber="337" phylum="Arthropoda" rank="subSpecies" species="frontalis" subSpecies="sulselensis">
<emphasis italics="true" pageId="38" pageNumber="337">sulselensis</emphasis>
</taxonomicName>
. Sulawesi Selatan, Boeloedoea, vi-1935 [JvT 22891]. Both with same magnification. Photos: JvT
</paragraph>
</caption>
<paragraph blockId="39.[159,1037,253,914]" pageId="39" pageNumber="338">
site presumably is at a distance of about 10 km WNW of Bulu Bulu (
<emphasis box="[1010,1035,254,279]" italics="true" pageId="39" pageNumber="338">ca</emphasis>
4°5132”S, 120°0943”E; Buludua is just another spelling of the same place), since there is no place of more than 600 m altitude closer to that village. Recently, we have received two specimens collected about 20 km east of Padaelo (about 4°30S, 119°47E) at
<emphasis box="[578,603,404,429]" italics="true" pageId="39" pageNumber="338">ca</emphasis>
700 m a.s.l., agreeing with the specimens mentioned above.
</paragraph>
<paragraph blockId="39.[159,1037,253,914]" pageId="39" pageNumber="338">The status of these populations needs to be studied more in detail. We de- scribe these specimens as a separate subspecies, since we are unaware of any populations with characters intermediate between those from northern and those from southwest Sulawesi.</paragraph>
<paragraph blockId="39.[159,1037,253,914]" pageId="39" pageNumber="338">Fortunately, we were able to compare molecular characters (CO1) of these specimens with those of specimens collected near Manado (northern pe- ninsula, near the type locality). The results are equivocal. Nearly all specimens from the Minahassa agree to a large extend in molecular characters, and the specimens from SW Sulawesi differ in these characters. However, one specimen from the Minahassa is sister to the clade of the other Mina- hassa specimens plus the South-western population. These results are too preliminary to draw firm conclusions.</paragraph>
</subSubSection>
</treatment>
</document>