351 lines
40 KiB
XML
351 lines
40 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6485232" ID-GBIF-Dataset="3dfc64e7-8c23-456d-9678-1bb0858ac470" ID-GBIF-Taxon="150645369" ID-Zenodo-Dep="6485232" checkinTime="1545431109525" checkinUser="PlaziZenodoSync" docAuthor="Jan van Tol & André Günther" docDate="2018" docId="0395A91CAC32FFCE95D3FE70FCEBF9B1" docLanguage="en" docName="10.5281zenodo.1481114_vanTol_Guenther_2018.pdf" docOrigin="Odonatologica 47 (3)" docStyle="DocumentStyle{}" docTitle="Rhinocypha togeanensis Jan van Tol & André Günther 2018, sp. nov." docType="treatment" docVersion="4" lastPageId="72" lastPageNumber="371" masterDocId="FFACD164AC71FF869443FFAFFF05FFBC" masterDocTitle="The Odonata of Sulawesi and adjacent islands. Part 8. Revision of the genus Rhinocypha Rambur, 1842 (Chlorocyphidae)" masterLastPageNumber="386" masterPageNumber="299" originalUpdateDomain="Boston" originalUpdateTime="1544520335069" originalUpdateUser="PlaziZenodoSync" pageId="67" pageNumber="366" updateTime="1650928538772" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Odonata of Sulawesi and adjacent islands. Part 8. Revision of the genus Rhinocypha Rambur, 1842 (Chlorocyphidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jan van Tol</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>André Günther</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Odonatologica</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-12-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>47</mods:number>
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||
</mods:detail>
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||
<mods:detail type="issue">
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<mods:number>3</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>299</mods:start>
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<mods:end>386</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.5281/zenodo.1481114</mods:identifier>
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<mods:identifier type="GBIF-Dataset">3dfc64e7-8c23-456d-9678-1bb0858ac470</mods:identifier>
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<mods:identifier type="Zenodo-Dep">1481114</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6485232" ID-GBIF-Taxon="150645369" ID-Zenodo-Dep="6485232" LSID="urn:lsid:plazi:treatment:0395A91CAC32FFCE95D3FE70FCEBF9B1" httpUri="http://treatment.plazi.org/id/0395A91CAC32FFCE95D3FE70FCEBF9B1" lastPageId="72" lastPageNumber="371" pageId="67" pageNumber="366">
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<subSubSection box="[400,794,479,507]" pageId="67" pageNumber="366" type="nomenclature">
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<paragraph blockId="67.[159,794,479,578]" box="[400,794,479,507]" pageId="67" pageNumber="366">
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||
<heading bold="true" box="[400,794,479,507]" centered="true" fontSize="11" level="2" pageId="67" pageNumber="366" reason="4">
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<emphasis bold="true" box="[400,794,479,507]" pageId="67" pageNumber="366">
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<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[400,695,479,507]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="366" phylum="Arthropoda" rank="species" species="togeanensis" status="sp. nov.">
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<emphasis bold="true" box="[400,695,479,507]" italics="true" pageId="67" pageNumber="366">
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Rhinocypha
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<taxonomicName box="[549,695,479,507]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="366" phylum="Arthropoda" rank="species" species="togeanensis" status="sp. nov.">togeanensis</taxonomicName>
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</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[702,794,479,507]" pageId="67" pageNumber="366" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection box="[489,706,513,541]" pageId="67" pageNumber="366" type="description">
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||
<paragraph blockId="67.[159,794,479,578]" box="[489,706,513,541]" pageId="67" pageNumber="366">
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||
<heading box="[489,706,513,541]" centered="true" fontSize="11" level="4" pageId="67" pageNumber="366" reason="4">
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(
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<figureCitation box="[495,568,513,541]" captionStart="Figure 9" captionStartId="16.[159,227,457,482]" captionTargetBox="[401,1036,249,715]" captionTargetId="figure@16.[401,1036,249,715]" captionTargetPageId="16" captionText="Figure 9. Rhinocypha togeanensis sp. nov., male. Stream Sg. Tanimpo, Batudaka, Togian Islands off Sulawesi, Indonesia. Photo: AG (24-viii- -1999)" httpUri="https://zenodo.org/record/2200395/files/figure.png" pageId="67" pageNumber="366">Figs 9</figureCitation>
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,
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<figureCitation box="[579,657,513,541]" captionStart-0="Figure 71" captionStart-1="Figures 72–76" captionStartId-0="68.[159,227,1496,1521]" captionStartId-1="70.[159,238,1373,1398]" captionTargetBox-0="[258,936,938,1457]" captionTargetBox-1="[167,1029,638,1339]" captionTargetId-0="figure@68.[258,936,938,1457]" captionTargetId-1="figure@70.[234,948,945,1021]" captionTargetPageId-0="68" captionTargetPageId-1="70" captionText-0="Figure 71. Rhinocypha togeanensis sp. nov. Male, holotype. Sulawesi Tengah, Togian Is., Batudaka, 27-viii-1999 [AG Indon 11/1999]. Photo: JvT" captionText-1="Figures 72–76. Rhinocypha togeanensis sp. nov. All from Sulawesi Tengah, Togian Is. 72 – Male, head, frontal view. Batudaka, 13viii1994 [AG 336/1994]; 73 – Male, pronotum, lef lateral view. Same locality [AG 336/1994]; 74 – Male, synthorax, lef lateral view. Same locality [AG 336/1994]; 75 – Male, abdomen, lef lateral view. Same locality [AG 336/1994]; 76 – Female, head, frontal view. Batudaka, 27 -viii-1999 [AG 13/1999]. Artwork by JvT" httpUri-0="https://zenodo.org/record/2200531/files/figure.png" httpUri-1="https://zenodo.org/record/2200535/files/figure.png" pageId="67" pageNumber="366">71–76</figureCitation>
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,
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<figureCitation box="[667,699,513,541]" captionStart="Figure 83" captionStartId="80.[159,227,1393,1418]" captionTargetBox="[162,1033,252,1349]" captionTargetId="figure@80.[159,1036,249,1352]" captionTargetPageId="80" captionText="Figure 83. Distribution of Rhinocypha flavipoda sp. nov., R. pelengensis sp. nov. and R. togeanensis sp. nov." httpUri="https://zenodo.org/record/2200551/files/figure.png" pageId="67" pageNumber="366">83</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[159,736,550,578]" pageId="67" pageNumber="366" type="reference_group">
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<paragraph blockId="67.[159,794,479,578]" box="[159,736,550,578]" pageId="67" pageNumber="366">
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<taxonomicName authorityName="Günther" authorityPageNumber="49" authorityYear="2008" box="[159,617,550,578]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="366" phylum="Arthropoda" rank="genus">
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<emphasis box="[159,283,554,578]" italics="true" pageId="67" pageNumber="366">Rhinocypha</emphasis>
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sp. B. — GÜNTHER (2008a): 49
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</taxonomicName>
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, 86, 89–94.
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</paragraph>
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</subSubSection>
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<subSubSection box="[159,360,622,648]" pageId="67" pageNumber="366" type="description">
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<paragraph blockId="67.[159,1038,622,958]" box="[159,360,622,648]" pageId="67" pageNumber="366">
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||
<heading bold="true" box="[159,360,622,648]" fontSize="11" level="3" pageId="67" pageNumber="366" reason="6">
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<emphasis bold="true" box="[159,360,622,648]" pageId="67" pageNumber="366">Material studied</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="67" pageNumber="366" type="materials_examined">
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<paragraph blockId="67.[159,1038,622,958]" pageId="67" pageNumber="366">
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<materialsCitation ID-GBIF-Occurrence="1949138277" collectingDate="1999-08-27" collectionCode="AG" collectorName="A. Günther" country="Indonesia" latitude="-0.43333334" location="To- gian Is., Batudaka, Wakai, Sg. Tanimpo" longLatPrecision="1308" longitude="121.86667" pageId="67" pageNumber="366" specimenCode="JvT 33391" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi" typeStatus="holotype">
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<emphasis bold="true" box="[159,272,660,686]" pageId="67" pageNumber="366">
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<typeStatus box="[159,272,660,686]" pageId="67" pageNumber="366">Holotype</typeStatus>
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</emphasis>
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<specimenCount box="[280,302,660,686]" pageId="67" pageNumber="366" type="male">♂</specimenCount>
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(
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<collectionCode box="[317,360,659,685]" pageId="67" pageNumber="366">AG</collectionCode>
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no. Indon 11/1999,
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<specimenCode box="[598,721,659,686]" pageId="67" pageNumber="366">JvT 33391</specimenCode>
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). »
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<collectingCountry box="[749,871,659,685]" name="Indonesia" pageId="67" pageNumber="366">Indonesia</collectingCountry>
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,
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<collectingRegion box="[882,985,659,685]" country="Indonesia" name="Sulawesi Utara" pageId="67" pageNumber="366">Sulawesi</collectingRegion>
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.
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<location LSID="urn:lsid:plazi:treatment:0395A91CAC32FFCE95D3FE70FCEBF9B1:8EE34ED1AC32FFC597A0FD3CFD5AFD6F" country="Indonesia" latitude="-0.43333334" longLatPrecision="1308" longitude="121.86667" name="To- gian Is., Batudaka, Wakai, Sg. Tanimpo" pageId="67" pageNumber="366" stateProvince="Sulawesi">To- gian Is., Batudaka, Wakai, Sg. Tanimpo</location>
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,
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<geoCoordinate box="[616,692,696,723]" direction="south" orientation="latitude" pageId="67" pageNumber="366" precision="925" value="-0.43333334">S 0°26’</geoCoordinate>
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S /
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<geoCoordinate box="[728,840,696,723]" direction="east" orientation="longitude" pageId="67" pageNumber="366" precision="925" value="121.86667">E 121°52’</geoCoordinate>
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),
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<collectingDate box="[858,986,697,723]" pageId="67" pageNumber="366" value="1999-08-27">
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<date box="[858,986,697,723]" pageId="67" pageNumber="366" value="1999-08-27">27.08.1999</date>
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</collectingDate>
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, leg.
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<collectorName box="[159,295,734,760]" pageId="67" pageNumber="366">A. Günther</collectorName>
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« (in RMNH) [with identification
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<taxonomicName authorityName="Rambur" authorityYear="1842" box="[686,822,734,760]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="366" phylum="Arthropoda" rank="genus">
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<emphasis box="[686,822,734,760]" italics="true" pageId="67" pageNumber="366">Rhinocypha</emphasis>
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</taxonomicName>
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sp. B] (
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<figureCitation box="[908,993,734,760]" captionStart="Figure 71" captionStartId="68.[159,227,1496,1521]" captionTargetBox="[258,936,938,1457]" captionTargetId="figure@68.[258,936,938,1457]" captionTargetPageId="68" captionText="Figure 71. Rhinocypha togeanensis sp. nov. Male, holotype. Sulawesi Tengah, Togian Is., Batudaka, 27-viii-1999 [AG Indon 11/1999]. Photo: JvT" httpUri="https://zenodo.org/record/2200531/files/figure.png" pageId="67" pageNumber="366">Fig. 71</figureCitation>
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||
).
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</materialsCitation>
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</paragraph>
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<paragraph blockId="67.[159,1038,622,958]" box="[159,398,771,798]" pageId="67" pageNumber="366">
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<emphasis bold="true" box="[159,279,772,798]" pageId="67" pageNumber="366">Paratypes</emphasis>
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(11♂ 6♀)
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</paragraph>
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<paragraph blockId="67.[159,1038,622,958]" box="[159,341,811,835]" pageId="67" pageNumber="366">
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<emphasis bold="true" box="[159,341,811,835]" pageId="67" pageNumber="366">Sulawesi Tengah</emphasis>
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</paragraph>
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<paragraph blockId="67.[159,1038,622,958]" pageId="67" pageNumber="366">7♂ 4♀, Togian Is., Batudaka, Wakai, Sungai Tanimpo (0°26’S, 121°52’E), 13–14- -viii-1994, leg. A. Günther & F. Randow (in AGPC); 4♂ 1♀, Togian Is., Batudaka, Wakai, Sungai Tanimpo (0°26’S, 121°52’E), 24–27-viii-1999, leg. A. Günther (in AGPC); 1♀, same data as holotype (in RMNH).</paragraph>
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</subSubSection>
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<subSubSection pageId="67" pageNumber="366" type="etymology">
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<paragraph blockId="67.[159,1037,1004,1105]" box="[159,292,1004,1030]" pageId="67" pageNumber="366">
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<heading bold="true" box="[159,292,1004,1030]" fontSize="11" level="3" pageId="67" pageNumber="366" reason="6">
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<emphasis bold="true" box="[159,292,1004,1030]" pageId="67" pageNumber="366">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="67.[159,1037,1004,1105]" pageId="67" pageNumber="366">
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<emphasis box="[159,296,1042,1067]" italics="true" pageId="67" pageNumber="366">Togeanensis</emphasis>
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, an adjective based on the type locality Kepulauan Togean (also: Togian Islands).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="68" lastPageNumber="367" pageId="67" pageNumber="366" type="diagnosis">
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<paragraph blockId="67.[159,1037,1151,1551]" box="[159,281,1151,1177]" pageId="67" pageNumber="366">
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<heading bold="true" box="[159,281,1151,1177]" fontSize="11" level="3" pageId="67" pageNumber="366" reason="6">
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<emphasis bold="true" box="[159,281,1151,1177]" pageId="67" pageNumber="366">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="67.[159,1037,1151,1551]" lastBlockId="68.[159,1037,253,728]" lastPageId="68" lastPageNumber="367" pageId="67" pageNumber="366">
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Male with head and pronotum very dark, mandibles entirely black, head with only small sub-quadrangular blue spot on gena and minute creamy spots beside lateral ocelli and on post-ocular lobe; pronotum and mesopreepisternum black; synthorax with irregular narrow blue stripe; opaque brownish black parts, with blue metallic sheen, in hind wing from Rs (some- what more distal at hind margin); abdomen dorsum black; sides with large blue markings, decreasing in size on S6–S8. Males can be distinguished from the
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<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[267,393,1449,1475]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="366" phylum="Arthropoda" rank="species" species="frontalis">
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<emphasis box="[267,393,1449,1475]" italics="true" pageId="67" pageNumber="366">
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R.
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<taxonomicName box="[298,393,1449,1475]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="67" pageNumber="366" phylum="Arthropoda" rank="species" species="frontalis">frontalis</taxonomicName>
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</emphasis>
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</taxonomicName>
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-complex by the absence of blue markings on the frons, absence of an ante-humeral stripe, and shape of band on synthorax; from
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<taxonomicName box="[159,319,1524,1550]" pageId="67" pageNumber="366">
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<emphasis box="[159,319,1524,1550]" italics="true" pageId="67" pageNumber="366">R. monochroa</emphasis>
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</taxonomicName>
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by the fully black mandibles and the absence of a ventral pro- jection along the metapleural suture; from
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<taxonomicName box="[654,812,253,279]" pageId="68" pageNumber="367">
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<emphasis box="[654,812,253,279]" italics="true" pageId="68" pageNumber="367">R. phantasma</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[868,1002,253,279]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="68" pageNumber="367" phylum="Arthropoda" rank="species" species="virgulata">
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<emphasis box="[868,1002,253,279]" italics="true" pageId="68" pageNumber="367">
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R.
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<taxonomicName box="[898,1002,253,279]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="68" pageNumber="367" phylum="Arthropoda" rank="species" species="virgulata">virgulata</taxonomicName>
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</emphasis>
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</taxonomicName>
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by a fully black dorsum of the abdomen (no pruinescence or blue rings anteri- orly on segments), from
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<taxonomicName authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[448,587,328,354]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="68" pageNumber="367" phylum="Arthropoda" rank="species" species="flavipoda">
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<emphasis box="[448,587,328,354]" italics="true" pageId="68" pageNumber="367">
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R.
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<taxonomicName box="[480,587,328,354]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="68" pageNumber="367" phylum="Arthropoda" rank="species" species="flavipoda">flavipoda</taxonomicName>
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</emphasis>
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</taxonomicName>
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by the black pronotum, and the white tibiae of mid and hind legs, and from
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<taxonomicName authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[597,756,365,391]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="68" pageNumber="367" phylum="Arthropoda" rank="species" species="pelengensis">
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<emphasis box="[597,756,365,391]" italics="true" pageId="68" pageNumber="367">
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R.
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<taxonomicName box="[629,756,365,391]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="68" pageNumber="367" phylum="Arthropoda" rank="species" species="pelengensis">pelengensis</taxonomicName>
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</emphasis>
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</taxonomicName>
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by the black mandibles, the completely black pronotum, the more extensive opaque patches on the wings and the larger blue spots on the abdomen.
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</paragraph>
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<paragraph blockId="68.[159,1037,253,728]" pageId="68" pageNumber="367">
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Females of
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<taxonomicName authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[319,483,478,503]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="68" pageNumber="367" phylum="Arthropoda" rank="species" species="togeanensis">
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<emphasis box="[319,483,478,503]" italics="true" pageId="68" pageNumber="367">
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R.
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<taxonomicName box="[353,483,478,503]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="68" pageNumber="367" phylum="Arthropoda" rank="species" species="togeanensis">togeanensis</taxonomicName>
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</emphasis>
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</taxonomicName>
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can be recognized by the absence of any pale marks on the pronotum; it is most similar to the female of
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<taxonomicName box="[867,1029,515,541]" pageId="68" pageNumber="367">
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<emphasis box="[867,1029,515,541]" italics="true" pageId="68" pageNumber="367">R. monochroa</emphasis>
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</taxonomicName>
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, with which it shares the extensive black coloration and the markings on syn- thorax and abdomen. However,
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<taxonomicName box="[543,705,589,615]" pageId="68" pageNumber="367">
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<emphasis box="[543,705,589,615]" italics="true" pageId="68" pageNumber="367">R. monochroa</emphasis>
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</taxonomicName>
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has large blue spots on the mandibles (small and creamish in
|
||
<taxonomicName authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[564,726,628,653]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="68" pageNumber="367" phylum="Arthropoda" rank="species" species="togeanensis">
|
||
<emphasis box="[564,726,628,653]" italics="true" pageId="68" pageNumber="367">
|
||
R.
|
||
<taxonomicName box="[596,726,628,653]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="68" pageNumber="367" phylum="Arthropoda" rank="species" species="togeanensis">togeanensis</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
), and usually distinct pale markings on the pronotum (may be obscured in some specimens); the transparent distal part of the fore wing is usually distinctly larger in
|
||
<taxonomicName box="[871,1029,701,727]" pageId="68" pageNumber="367">
|
||
<emphasis box="[871,1029,701,727]" italics="true" pageId="68" pageNumber="367">R. monochroa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="72" lastPageNumber="371" pageId="68" pageNumber="367" type="description">
|
||
<paragraph blockId="68.[159,1036,776,914]" box="[159,305,776,802]" pageId="68" pageNumber="367">
|
||
<heading bold="true" box="[159,305,776,802]" fontSize="11" level="3" pageId="68" pageNumber="367" reason="6">
|
||
<emphasis bold="true" box="[159,305,776,802]" pageId="68" pageNumber="367">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="68.[159,1036,776,914]" pageId="68" pageNumber="367">
|
||
<emphasis bold="true" box="[159,220,814,840]" pageId="68" pageNumber="367">Male</emphasis>
|
||
(holotype) [note: the type was previously pinned through the mid- dle of the thorax; some parts bearing diagnostic characters were damaged] (
|
||
<figureCitation box="[166,249,888,914]" captionStart="Figure 71" captionStartId="68.[159,227,1496,1521]" captionTargetBox="[258,936,938,1457]" captionTargetId="figure@68.[258,936,938,1457]" captionTargetPageId="68" captionText="Figure 71. Rhinocypha togeanensis sp. nov. Male, holotype. Sulawesi Tengah, Togian Is., Batudaka, 27-viii-1999 [AG Indon 11/1999]. Photo: JvT" httpUri="https://zenodo.org/record/2200531/files/figure.png" pageId="68" pageNumber="367">Fig. 71</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<caption httpUri="https://zenodo.org/record/2200531/files/figure.png" pageId="68" pageNumber="367" startId="68.[159,227,1496,1521]" targetBox="[258,936,938,1457]" targetPageId="68">
|
||
<paragraph blockId="68.[159,1036,1496,1550]" pageId="68" pageNumber="367">
|
||
<emphasis bold="true" box="[159,268,1496,1521]" pageId="68" pageNumber="367">Figure 71.</emphasis>
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[275,537,1496,1520]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="68" pageNumber="367" phylum="Arthropoda" rank="species" species="togeanensis" status="sp. nov.">
|
||
<emphasis box="[275,537,1496,1520]" italics="true" pageId="68" pageNumber="367">Rhinocypha togeanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[544,628,1496,1521]" pageId="68" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Male, holotype. Sulawesi Tengah, Togian Is., Batudaka, 27-viii-1999 [AG Indon 11/1999]. Photo: JvT
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="69.[159,1037,253,392]" pageId="69" pageNumber="368">
|
||
Head (
|
||
<figureCitation box="[238,325,254,280]" captionStart="Figures 72–76" captionStartId="70.[159,238,1373,1398]" captionTargetBox="[167,1029,638,1339]" captionTargetId="figure@70.[234,948,945,1021]" captionTargetPageId="70" captionText="Figures 72–76. Rhinocypha togeanensis sp. nov. All from Sulawesi Tengah, Togian Is. 72 – Male, head, frontal view. Batudaka, 13viii1994 [AG 336/1994]; 73 – Male, pronotum, lef lateral view. Same locality [AG 336/1994]; 74 – Male, synthorax, lef lateral view. Same locality [AG 336/1994]; 75 – Male, abdomen, lef lateral view. Same locality [AG 336/1994]; 76 – Female, head, frontal view. Batudaka, 27 -viii-1999 [AG 13/1999]. Artwork by JvT" httpUri="https://zenodo.org/record/2200535/files/figure.png" pageId="69" pageNumber="368">Fig. 72</figureCitation>
|
||
) — Black, anteriorly weakly lustrous, dorsally velvet black; gena with small, sub-oval blue marking, touching eyes and with short ven- tral projection; minute oblong creamy spots behind lateral ocelli, and about size of ocelli; spots on post-ocular lobe smaller, just discernable.
|
||
</paragraph>
|
||
<paragraph blockId="69.[159,1036,432,794]" pageId="69" pageNumber="368">
|
||
Thorax — Pronotum (
|
||
<figureCitation box="[413,496,432,458]" captionStart="Figures 72–76" captionStartId="70.[159,238,1373,1398]" captionTargetBox="[167,1029,638,1339]" captionTargetId="figure@70.[234,948,945,1021]" captionTargetPageId="70" captionText="Figures 72–76. Rhinocypha togeanensis sp. nov. All from Sulawesi Tengah, Togian Is. 72 – Male, head, frontal view. Batudaka, 13viii1994 [AG 336/1994]; 73 – Male, pronotum, lef lateral view. Same locality [AG 336/1994]; 74 – Male, synthorax, lef lateral view. Same locality [AG 336/1994]; 75 – Male, abdomen, lef lateral view. Same locality [AG 336/1994]; 76 – Female, head, frontal view. Batudaka, 27 -viii-1999 [AG 13/1999]. Artwork by JvT" httpUri="https://zenodo.org/record/2200535/files/figure.png" pageId="69" pageNumber="368">Fig. 73</figureCitation>
|
||
) entirely velvet black, without any trace of pale markings.
|
||
</paragraph>
|
||
<paragraph blockId="69.[159,1036,432,794]" pageId="69" pageNumber="368">
|
||
Synthorax (
|
||
<figureCitation box="[322,414,507,533]" captionStart="Figures 72–76" captionStartId="70.[159,238,1373,1398]" captionTargetBox="[167,1029,638,1339]" captionTargetId="figure@70.[234,948,945,1021]" captionTargetPageId="70" captionText="Figures 72–76. Rhinocypha togeanensis sp. nov. All from Sulawesi Tengah, Togian Is. 72 – Male, head, frontal view. Batudaka, 13viii1994 [AG 336/1994]; 73 – Male, pronotum, lef lateral view. Same locality [AG 336/1994]; 74 – Male, synthorax, lef lateral view. Same locality [AG 336/1994]; 75 – Male, abdomen, lef lateral view. Same locality [AG 336/1994]; 76 – Female, head, frontal view. Batudaka, 27 -viii-1999 [AG 13/1999]. Artwork by JvT" httpUri="https://zenodo.org/record/2200535/files/figure.png" pageId="69" pageNumber="368">Fig. 74</figureCitation>
|
||
). Mesopreepisternum glossy black. Synthorax dull black with irregular blue band from ventro-posterior corner of mesokatepisternum to ventro-posterior corner of metepimeron; anterior side on metepisternum with narrow black interruption to mesokatepisternum; dor- sal side of band irregular, ventrally running just below metathoracic spira- cle, metakatepisternum black; stripe over metepimeron bent ventrad distal to metakatepisternum, leaving a narrow black border ventrally; irregular dorsal side of stripe without dorso-posteriorly directed projection.
|
||
</paragraph>
|
||
<paragraph blockId="69.[159,1036,830,899]" pageId="69" pageNumber="368">Legs (right mid leg missing) — Black, with distal ⁵/₆ of inner sides of tibiae of mid and hind legs white.</paragraph>
|
||
<paragraph blockId="69.[159,1036,939,1115]" pageId="69" pageNumber="368">Wings — Dark brownish black with blue metallic sheen from around the nodus in fore wing, and Ax5 in hind wing to the wing tip; opaque part me- dially extended approximately to level of Ax10 in fore wing, and to fork of Rs in hind wing, but leaving the quadrangle transparent; transparent base of wings enfumed dark brown; pterostigma of fore and hind wings brown.</paragraph>
|
||
<paragraph blockId="69.[159,1037,1155,1442]" pageId="69" pageNumber="368">
|
||
Abdomen (
|
||
<figureCitation box="[292,378,1155,1182]" captionStart="Figures 72–76" captionStartId="70.[159,238,1373,1398]" captionTargetBox="[167,1029,638,1339]" captionTargetId="figure@70.[234,948,945,1021]" captionTargetPageId="70" captionText="Figures 72–76. Rhinocypha togeanensis sp. nov. All from Sulawesi Tengah, Togian Is. 72 – Male, head, frontal view. Batudaka, 13viii1994 [AG 336/1994]; 73 – Male, pronotum, lef lateral view. Same locality [AG 336/1994]; 74 – Male, synthorax, lef lateral view. Same locality [AG 336/1994]; 75 – Male, abdomen, lef lateral view. Same locality [AG 336/1994]; 76 – Female, head, frontal view. Batudaka, 27 -viii-1999 [AG 13/1999]. Artwork by JvT" httpUri="https://zenodo.org/record/2200535/files/figure.png" pageId="69" pageNumber="368">Fig. 75</figureCitation>
|
||
) — Dorsum black; sides with extensive blue markings; marking on segment 1 sub-quadrangular, ventrally somewhat irregular; that on S2 covering most of side and leaving only a small black triangle in ventro-anterior corner; markings on S3–S5 covering sides nearly completely, except the intersegmental annulae; marking on S6 ventrally over full length of segment, dorsally tapered posteriorly; markings on S7–S8 similar to S6, but increasingly smaller; S9 with just a very small pale marking at an- terior margin; S10 and anal appendages black.
|
||
</paragraph>
|
||
<paragraph blockId="69.[159,1036,1482,1547]" pageId="69" pageNumber="368">Measurements [mm] — Hind wing length 22.0, hind wing width 5.5; abdomen (including appendages) 19.0.</paragraph>
|
||
<paragraph blockId="70.[159,1036,254,504]" box="[159,625,254,280]" pageId="70" pageNumber="369">
|
||
<emphasis bold="true" box="[159,247,254,280]" pageId="70" pageNumber="369">Female</emphasis>
|
||
(AG Indon 13/1999, JvT 33392)
|
||
</paragraph>
|
||
<paragraph blockId="70.[159,1036,254,504]" pageId="70" pageNumber="369">
|
||
Head (
|
||
<figureCitation box="[237,322,291,318]" captionStart="Figures 72–76" captionStartId="70.[159,238,1373,1398]" captionTargetBox="[167,1029,638,1339]" captionTargetId="figure@70.[234,948,945,1021]" captionTargetPageId="70" captionText="Figures 72–76. Rhinocypha togeanensis sp. nov. All from Sulawesi Tengah, Togian Is. 72 – Male, head, frontal view. Batudaka, 13viii1994 [AG 336/1994]; 73 – Male, pronotum, lef lateral view. Same locality [AG 336/1994]; 74 – Male, synthorax, lef lateral view. Same locality [AG 336/1994]; 75 – Male, abdomen, lef lateral view. Same locality [AG 336/1994]; 76 – Female, head, frontal view. Batudaka, 27 -viii-1999 [AG 13/1999]. Artwork by JvT" httpUri="https://zenodo.org/record/2200535/files/figure.png" pageId="70" pageNumber="369">Fig. 76</figureCitation>
|
||
) — Black; pale parts more extensive than in the male; man- dibles medially with small, oblong creamish spot; gena with creamish, rec- tangular spot touching eye margins, ventrally somewhat extended along eye margin; narrow stripe along eye margin at level of antennae, not connected with marking on gena; dorsum of head velvet black with creamy spots near lateral ocelli and on post-ocular lobe as in male.
|
||
</paragraph>
|
||
<paragraph blockId="70.[159,760,552,579]" box="[159,760,552,579]" pageId="70" pageNumber="369">Thorax — Pronotum black without any pale marks.</paragraph>
|
||
<caption httpUri="https://zenodo.org/record/2200535/files/figure.png" pageId="70" pageNumber="369" startId="70.[159,238,1373,1398]" targetBox="[167,1029,638,1339]" targetPageId="70">
|
||
<paragraph blockId="70.[159,1037,1373,1545]" pageId="70" pageNumber="369">
|
||
<emphasis bold="true" box="[159,319,1373,1398]" pageId="70" pageNumber="369">Figures 72–76.</emphasis>
|
||
<taxonomicName authority="Jan van Tol & André Günther, 2018" authorityName="Jan van Tol & André Günther" authorityYear="2018" box="[325,587,1373,1397]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="70" pageNumber="369" phylum="Arthropoda" rank="species" species="togeanensis" status="sp. nov.">
|
||
<emphasis box="[325,587,1373,1397]" italics="true" pageId="70" pageNumber="369">Rhinocypha togeanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[594,677,1373,1398]" pageId="70" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
|
||
All from Sulawesi Tengah, Togian Is. 72 – Male, head, frontal view. Batudaka, 13viii1994 [AG 336/1994]; 73 – Male, pronotum, lef lateral view. Same locality [AG 336/1994]; 74 – Male, synthorax, lef lateral view. Same locality [AG 336/1994]; 75 – Male, abdomen, lef lateral view. Same locality [AG 336/1994]; 76 – Female, head, frontal view. Batudaka, 27 -viii-1999 [AG 13/1999]. Artwork by JvT
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="71.[159,1036,253,691]" pageId="71" pageNumber="370">Synthorax. Mesopreepisternum black. Base colour of synthorax black, semimatt with some metallic sheen; mesepisternum black without markings; mesepimeron with short and narrow cream stripe posteriorly, about the length of this stripe from hind margin; band over metepisternum and met- epimeron narrow, creamy white with some blue tones; band consists anteri- orly of small spot in ventro-posterior corner of mesokatepisternum, continues over metepisternum and metepimeron with short black interruption on suture between mesokatepisternum and metepisternum; ventral and dorsal sides of band subtly irregular, dorsally ending against hind margin halfway metepimeron, ventrally running under metathoracic spiracle, continuing posteriad well from the lower margin of metepimeron, and ending against hind margin.</paragraph>
|
||
<paragraph blockId="71.[159,525,739,765]" box="[159,525,739,765]" pageId="71" pageNumber="370">Legs — Black, somewhat shiny.</paragraph>
|
||
<paragraph blockId="71.[159,1037,813,1064]" pageId="71" pageNumber="370">Wings — Very dark. Base of fore wing transparent approximately to level of Arculus, but costal space transparent to level of nodus; tip of fore wing transparent distal to Px24, tip subtly opaque white; hind wing semi-transparent basal to Arculus, strongly enfumed dark brown, rest of wing opaque dark brown, but tip distal to distal side of pterostigma transparent and clear; pterostigma of fore and hind wings with basal half brown, anteriorly nar- rower, distal part opaque white.</paragraph>
|
||
<paragraph blockId="71.[159,1037,1112,1363]" pageId="71" pageNumber="370">Abdomen — Dark brown with relatively small pale (cream) markings: S1 with sub-oval spot against hind margin, covering approximately ¼ of segment; S2 and S3 with a short pale stripe close to lower margin of segment, about ⅓ of the segment length; small sub-triangular or sub-oval spots lat- ero-posteriorly on segment; S3–S7 with a minute oblong spot medioante- riorly on each segment; S4–S7 with a short stripe close to lower margin of tergite, length of this stripe about ⅓ of length of segment; S8–S10 black.</paragraph>
|
||
<paragraph blockId="71.[159,1036,1411,1550]" box="[159,338,1411,1437]" pageId="71" pageNumber="370">
|
||
<heading bold="true" box="[159,338,1411,1437]" fontSize="11" level="3" pageId="71" pageNumber="370" reason="6">
|
||
<emphasis bold="true" box="[159,338,1411,1437]" pageId="71" pageNumber="370">Measurements</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="71.[159,1036,1411,1550]" pageId="71" pageNumber="370">Measurements of male specimens [mm] — Hind wing length (n = 5) x= 21.5 (20.5–22.0), hind wing width (n = 5) x= 6.0 (5.5–6.0), abdomen length (in- cluding appendages) (n = 5) x= 18.5 (17.5–19.5).</paragraph>
|
||
<paragraph blockId="72.[159,1036,253,355]" pageId="72" pageNumber="371">Measurements of female specimens [mm] — Hind wing length (n = 3) x = 23.0 (22.0–24.0), hind wing width (n= 3) x=6.0 (6.0–6.5), abdomen length (including appendages) (n = 3) x= 17.5 (17.5–18.0).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="72" pageNumber="371" type="distribution">
|
||
<paragraph blockId="72.[159,1038,403,952]" box="[159,459,403,429]" pageId="72" pageNumber="371">
|
||
<heading bold="true" box="[159,459,403,429]" fontSize="11" level="3" pageId="72" pageNumber="371" reason="6">
|
||
<emphasis bold="true" box="[159,459,403,429]" pageId="72" pageNumber="371">Distribution and habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="72.[159,1038,403,952]" pageId="72" pageNumber="371">
|
||
Known from only one river on the island of Batudaka, one of the Togian Islands (
|
||
<figureCitation box="[255,340,478,504]" captionStart="Figure 83" captionStartId="80.[159,227,1393,1418]" captionTargetBox="[162,1033,252,1349]" captionTargetId="figure@80.[159,1036,249,1352]" captionTargetPageId="80" captionText="Figure 83. Distribution of Rhinocypha flavipoda sp. nov., R. pelengensis sp. nov. and R. togeanensis sp. nov." httpUri="https://zenodo.org/record/2200551/files/figure.png" pageId="72" pageNumber="371">Fig. 83</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="72.[159,1038,403,952]" pageId="72" pageNumber="371">
|
||
On Batudaka it was found only on Sungai Tanimpo south of Wakai village (
|
||
<figureCitation box="[166,254,552,578]" captionStart="Figure 14" captionStartId="17.[775,843,1168,1193]" captionTargetBox="[159,736,1125,1510]" captionTargetId="figure@17.[159,736,1125,1510]" captionTargetPageId="17" captionText="Figure 14. Type locality of Rhinocypha togeanensis sp. nov., and habitat of Libellago xanthocyana. Sg. Tanimpo, Batudaka, Togian Islands, Indonesia. This site was still in un- disturbed lowland rain forest at the time; it is now heavily deforested. Photo: AG (24-viii-1999)" httpUri="https://zenodo.org/record/2200413/files/figure.png" pageId="72" pageNumber="371">Fig. 14</figureCitation>
|
||
). In 1994 and 1999 the habitat was a fast-flowing shallow stream of about 3–4 m in width within undisturbed evergreen lowland rainforest. Important habitat requirements were apparently large broken logs stand- ing in the water. Above a small waterfall (‘Air terjun Wakai’, 0°26’10.1”S, 121°51’36.1”E) the stream was characterized by small cascades, pools and shallow, but still flowing sections with a bed of gravel. Below the water- fall, the character changed increasingly to a sandy lowland stream.
|
||
<taxonomicName authorityName="Jan van Tol & André Günther" authorityYear="2018" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="72" pageNumber="371" phylum="Arthropoda" rank="species" species="togeanensis">
|
||
<emphasis italics="true" pageId="72" pageNumber="371">
|
||
Rhinocypha
|
||
<taxonomicName box="[234,365,814,839]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="72" pageNumber="371" phylum="Arthropoda" rank="species" species="togeanensis">togeanensis</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
was absent in lower (sandy) reaches of the still fast flow- ing Sungai Tanimpo within disturbed primary forest, secondary forests and plantations.Current Google EarthTM images show that the site has now been largely deforested.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="72" pageNumber="371" type="biology_ecology">
|
||
<paragraph blockId="72.[159,1037,1000,1549]" box="[159,286,1000,1026]" pageId="72" pageNumber="371">
|
||
<heading bold="true" box="[159,286,1000,1026]" fontSize="11" level="3" pageId="72" pageNumber="371" reason="6">
|
||
<emphasis bold="true" box="[159,286,1000,1026]" pageId="72" pageNumber="371">Behaviour</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="72.[159,1037,1000,1549]" pageId="72" pageNumber="371">
|
||
Detailed observations of the reproductive behaviour were recorded by GÜNTHER (2001, 2008a) on Batudaka (Togian islands) sub »
|
||
<taxonomicName authorityName="Rambur" authorityYear="1842" box="[901,1036,1075,1101]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="72" pageNumber="371" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[901,1036,1075,1101]" italics="true" pageId="72" pageNumber="371">Rhinocypha</emphasis>
|
||
</taxonomicName>
|
||
sp. B«. The behaviour of the males was basically similar to
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[840,964,1112,1138]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="72" pageNumber="371" phylum="Arthropoda" rank="species" species="frontalis">
|
||
<emphasis box="[840,964,1112,1138]" italics="true" pageId="72" pageNumber="371">
|
||
R.
|
||
<taxonomicName box="[872,964,1112,1138]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="72" pageNumber="371" phylum="Arthropoda" rank="species" species="frontalis">frontalis</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
. They maintained small territories around driftwood and twigs lying in the water, but mostly around large logs in the stream. The agonistic display consisted mainly of threat flights with synchronous stroking wing beats. In contrast to
|
||
<taxonomicName baseAuthorityName="Selys" baseAuthorityYear="1873" box="[159,280,1261,1287]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="72" pageNumber="371" phylum="Arthropoda" rank="species" species="frontalis">
|
||
<emphasis box="[159,280,1261,1287]" italics="true" pageId="72" pageNumber="371">
|
||
R.
|
||
<taxonomicName box="[188,280,1261,1287]" class="Insecta" family="Chlorocyphidae" genus="Rhinocypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="72" pageNumber="371" phylum="Arthropoda" rank="species" species="frontalis">frontalis</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
, the flight pattern includes an additional vertical component when males alternately or simultaneously ascend and descend while facing each other. Due to the highly visible blue iridescent patches on the hind wings, pairs of fighting males were very conspicuous, their sparkling wing colours visible from some distance. During courtship males led the females into their territories.After mating, the females left the male territories.All recorded oviposition took place unguarded on the large logs well above the water surface. Aggregations of females at these oviposition sites were regularly observed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |