165 lines
21 KiB
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165 lines
21 KiB
XML
<document id="7A0BBDABF480EFB3DD474262DF638A5F" ID-DOI="10.5281/zenodo.5399742" ID-ISSN="1638-9387" ID-Zenodo-Dep="5399742" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630332575338" checkinUser="marcus" docAuthor="Kritsky, Delane C. & Boeger, Walter A." docDate="2002" docId="F15587D8DF74FF98FF57F0C5FA4BFB9A" docLanguage="en" docName="Zoosystema.2002.24.1.7-40.pdf" docOrigin="Zoosystema 24 (1)" docSource="http://dx.doi.org/10.5281/zenodo.5399767" docStyle="DocumentStyle:64A3174DE33E974D4EB6573A0A793B12.4:Zoosystema.2000-2008.journal_article.type2" docStyleId="64A3174DE33E974D4EB6573A0A793B12" docStyleName="Zoosystema.2000-2008.journal_article.type2" docStyleVersion="4" docTitle="Euryhaliotrema longibaculum Kritsky & Boeger 2002, n. comb." docType="treatment" docVersion="5" lastPageNumber="32" masterDocId="0D6CFFA0DF6DFF81FFA9F339FE64FFD8" masterDocTitle="41: New and previously described species of Dactylogyridae (Platyhelminthes) from the gills of marine and freshwater perciform fishes (Teleostei) with proposal of a new genus and a hypothesis on phylogeny" masterLastPageNumber="40" masterPageNumber="7" pageNumber="32" updateTime="1699231409364" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="6BA1FAD4678B0E044C25575DF46B18CC">
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<mods:title id="E8F11025D1FD98A3CDEE3E7E8AF607A6">41: New and previously described species of Dactylogyridae (Platyhelminthes) from the gills of marine and freshwater perciform fishes (Teleostei) with proposal of a new genus and a hypothesis on phylogeny</mods:title>
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<mods:name id="F14936E840FC3DAB704C5D9757C218FC" type="personal">
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<mods:role id="62019EC88A14AA2A5C099693D8093B24">
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<mods:roleTerm id="486D8A5A0B5D2ADD3E5BDA4E17DA8B88">Author</mods:roleTerm>
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<mods:namePart id="19970CC2525185A29796E4C30CD9F490">Kritsky, Delane C.</mods:namePart>
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<mods:affiliation id="3BBC8AA4534A75D93845ED4943962135">Department of Health and Nutrition Sciences, College of Health Professions, Campus Box 8090, Idaho State University, Pocatello, Idaho 83209 (USA) kritdela @ isu. edu</mods:affiliation>
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<mods:nameIdentifier id="0D7687255286711996DB474BFB611859" type="email">kritdela@isu.edu</mods:nameIdentifier>
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<mods:name id="54E2C72927BADCDE0A53453B98D24931" type="personal">
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<mods:namePart id="D317F4FEE168EE2E0423EF54D78668A6">Boeger, Walter A.</mods:namePart>
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<mods:affiliation id="690832304E7385821A23F530D9CCA4D7">Department of Biological Sciences, Campus Box 8007, Idaho State University, Pocatello, Idaho 83209 (USA) Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19073, Curitiba, Paraná, 81530 (Brazil) Conselho Nacional de Desenvolvimento Cientifíco e Tecnológico (CNPq) wboeger @ bio. ufpr. br.</mods:affiliation>
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<mods:nameIdentifier id="B127D88539031227BE518800EB1B72F1" type="email">wboeger@bio.ufpr.br</mods:nameIdentifier>
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<mods:title id="799602387F6D13F99C9C37377B942253">Zoosystema</mods:title>
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<mods:part id="CD859065280D2417FB98A645E0528618">
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<mods:date id="782F96DDB22B136024C5B17DCA21EB4A">2002</mods:date>
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<mods:number id="42D0DE12B0926374A26CA443BF06A10D">24</mods:number>
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<mods:number id="C391EE7995B969B78472653564C613B0">1</mods:number>
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<mods:end id="3ADF7C4A72E4960CD69869FFCD510E00">40</mods:end>
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<mods:identifier id="22CA457E564FFF92201E49FEA23B42C4" type="DOI">10.5281/zenodo.5399742</mods:identifier>
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<mods:identifier id="40D116CC58A1ECBAD2D7BD15166D8AD9" type="ISSN">1638-9387</mods:identifier>
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<treatment id="F15587D8DF74FF98FF57F0C5FA4BFB9A" ID-DOI="http://doi.org/10.5281/zenodo.5485193" ID-Zenodo-Dep="5485193" LSID="urn:lsid:plazi:treatment:F15587D8DF74FF98FF57F0C5FA4BFB9A" httpUri="http://treatment.plazi.org/id/F15587D8DF74FF98FF57F0C5FA4BFB9A" lastPageNumber="32" pageId="25" pageNumber="32">
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<subSubSection id="31E66545DF74FF98FF57F0C5FC49FBEC" pageId="25" pageNumber="32" type="nomenclature">
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<paragraph id="794336CEDF74FF98FF57F0C5FC49FBEC" blockId="25.[151,684,1020,1375]" pageId="25" pageNumber="32">
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<heading id="220B81A2DF74FF98FF57F0C5FC49FBEC" centered="true" fontSize="11" level="2" pageId="25" pageNumber="32" reason="3">
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<taxonomicName id="BEFC4D4DDF74FF98FF57F0C5FFA0FBEC" ID-CoL="3D4RJ" authority="(Zhukov, 1976)" authorityName="Kritsky & Boeger" authorityYear="2002" baseAuthorityName="Zhukov" baseAuthorityYear="1976" class="Monogenea" family="Ancyrocephalidae" genus="Euryhaliotrema" kingdom="Animalia" order="Dactylogyridea" pageId="25" pageNumber="32" phylum="Platyhelminthes" rank="species" species="longibaculum" status="comb. nov.">
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<emphasis id="4B88EADCDF74FF98FF57F0C5FC20FBCE" bold="true" box="[254,580,1020,1046]" italics="true" pageId="25" pageNumber="32">Euryhaliotrema longibaculum</emphasis>
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(
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<bibRefCitation id="1D6D4B3FDF74FF98FEB7F723FFD8FBEC" author="Zhukov" box="[286,444,1050,1076]" firstAuthor="Zhukov" pageId="25" pageNumber="32" pagination="33 - 47" refId="ref19640" refString="ZHUKOV E. V. 1976. - [New monogenean species of the genus Haliotrema Johnston and Tiegs, 1922, from the Gulf of Mexico fishes of the Fam. Lutianidae], in Fauna, systematics and phylogeny of Monogenoidea. Proceedings, Institute of Biology and Pedology, Far-East Science Centre, Academy of Sciences of the U. S. S. R., New Series 35: 33 - 47 (in Russian)." type="book chapter" year="1976">Zhukov, 1976</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="50BB57A7DF74FF98FE62F723FC49FBEC" box="[459,557,1050,1076]" pageId="25" pageNumber="32" rank="species">n. comb.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="31E66545DF74FF98FF3EF77CFC2DFB85" box="[151,585,1093,1117]" pageId="25" pageNumber="32" type="reference_group">
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<paragraph id="794336CEDF74FF98FF3EF77CFC2DFB85" blockId="25.[151,684,1020,1375]" box="[151,585,1093,1117]" pageId="25" pageNumber="32">
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<treatmentCitationGroup id="59EC11E0DF74FF98FF3EF77CFC2DFB85" box="[151,585,1093,1117]" pageId="25" pageNumber="32">
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<treatmentCitation id="F85D10DFDF74FF98FF3EF77CFC21FB85" author="ZHUKOV E. V." box="[151,581,1093,1117]" page="39" pageId="25" pageNumber="32" year="1976">
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<taxonomicName id="BEFC4D4DDF74FF98FF3EF77CFC21FB85" ID-CoL="6KYXL" authority="Zhukov, 1976: 39" authorityName="Zhukov" authorityPageNumber="39" authorityYear="1976" box="[151,581,1093,1117]" class="Monogenea" family="Ancyrocephalidae" genus="Haliotrema" kingdom="Animalia" order="Dactylogyridea" pageId="25" pageNumber="32" phylum="Platyhelminthes" rank="species" species="longibaculum">
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<emphasis id="4B88EADCDF74FF98FF3EF77CFFECFB85" box="[151,392,1093,1117]" italics="true" pageId="25" pageNumber="32">Haliotrema longibaculum</emphasis>
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<bibRefCitation id="1D6D4B3FDF74FF98FE26F77CFC21FB85" author="Zhukov" box="[399,581,1093,1117]" firstAuthor="Zhukov" page="39" pageId="25" pageNumber="32" pagination="33 - 47" refId="ref19640" refString="ZHUKOV E. V. 1976. - [New monogenean species of the genus Haliotrema Johnston and Tiegs, 1922, from the Gulf of Mexico fishes of the Fam. Lutianidae], in Fauna, systematics and phylogeny of Monogenoidea. Proceedings, Institute of Biology and Pedology, Far-East Science Centre, Academy of Sciences of the U. S. S. R., New Series 35: 33 - 47 (in Russian)." type="book chapter" year="1976">Zhukov, 1976: 39</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="31E66545DF74FF98FF3EF755FF2DFA85" pageId="25" pageNumber="32" type="materials_examined">
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<paragraph id="794336CEDF74FF98FF3EF755FF22FB60" blockId="25.[151,684,1020,1375]" pageId="25" pageNumber="32">
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MATERIAL EXAMINED. — Voucher specimen from
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<taxonomicName id="BEFC4D4DDF74FF98FF3EF7BFFF17FB46" authority=", Gulf" authorityName="Gulf" box="[151,371,1157,1182]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="32" phylum="Chordata" rank="species" species="synagris">
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<emphasis id="4B88EADCDF74FF98FF3EF7BFFF5DFB45" box="[151,313,1158,1181]" italics="true" pageId="25" pageNumber="32">Lutjanus synagris</emphasis>
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, Gulf
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</taxonomicName>
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of
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<collectingRegion id="BB38F82CDF74FF98FE3CF7BFFF99FB46" box="[405,509,1158,1182]" country="Mexico" name="Campeche" pageId="25" pageNumber="32">Campeche</collectingRegion>
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(Gulf of
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<collectingRegion id="BB38F82CDF74FF98FDF2F7BFFCC2FB46" box="[603,678,1158,1182]" country="Mexico" name="Mexico" pageId="25" pageNumber="32">Mexico</collectingRegion>
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) (
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<collectionCode id="1FEDAE0BDF74FF98FF34F799FE95FB60" box="[157,241,1184,1208]" country="USA" httpUri="http://grbio.org/cool/ppqr-es1r" name="United States National Parasite Collection" pageId="25" pageNumber="32" type="Museum">USNPC</collectionCode>
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91427).
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</paragraph>
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<paragraph id="794336CEDF74FF98FF3EF7FEFF2DFA85" blockId="25.[151,684,1020,1375]" pageId="25" pageNumber="32">
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<collectionCode id="1FEDAE0BDF74FF98FF3EF7FEFF04FB05" box="[151,352,1223,1246]" pageId="25" pageNumber="32">PREVIOUS RECORDS</collectionCode>
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. — Gills of
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<taxonomicName id="BEFC4D4DDF74FF98FE43F7FEFCC8FB38" baseAuthorityName="Cuvier" baseAuthorityYear="1828" box="[490,684,1223,1248]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="32" phylum="Chordata" rank="species" species="mahogoni">
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<emphasis id="4B88EADCDF74FF98FE43F7FEFCC8FB38" box="[490,684,1223,1248]" italics="true" pageId="25" pageNumber="32">Lutjanus mahogoni</emphasis>
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</taxonomicName>
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(type host) and
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<taxonomicName id="BEFC4D4DDF74FF98FE9BF7D8FC33FB21" box="[306,599,1249,1273]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="32" phylum="Chordata" rank="species" species="synagris">
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<emphasis id="4B88EADCDF74FF98FE9BF7D8FFB0FB20" box="[306,468,1249,1272]" italics="true" pageId="25" pageNumber="32">Lutjanus synagris</emphasis>
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(Lutjanidae)
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</taxonomicName>
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: Havana and Gulf of
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<collectingRegion id="BB38F82CDF74FF98FEB5F7C2FFEDFACB" box="[284,393,1275,1299]" country="Mexico" name="Campeche" pageId="25" pageNumber="32">Campeche</collectingRegion>
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(Gulf of
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<collectingRegion id="BB38F82CDF74FF98FE5BF7C2FC25FACB" box="[498,577,1275,1299]" country="Mexico" name="Mexico" pageId="25" pageNumber="32">Mexico</collectingRegion>
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) (
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<bibRefCitation id="1D6D4B3FDF74FF98FDF1F7C2FEB4FAF4" author="Zhukov" firstAuthor="Zhukov" pageId="25" pageNumber="32" pagination="33 - 47" refId="ref19640" refString="ZHUKOV E. V. 1976. - [New monogenean species of the genus Haliotrema Johnston and Tiegs, 1922, from the Gulf of Mexico fishes of the Fam. Lutianidae], in Fauna, systematics and phylogeny of Monogenoidea. Proceedings, Institute of Biology and Pedology, Far-East Science Centre, Academy of Sciences of the U. S. S. R., New Series 35: 33 - 47 (in Russian)." type="book chapter" year="1976">Zhukov 1976</bibRefCitation>
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). According to Timofeeva (pers. comm.), the
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<typeStatus id="A647886CDF74FF98FF3EF614FE94FA9D" box="[151,240,1325,1349]" pageId="25" pageNumber="32" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="BEFC4D4DDF74FF98FEBCF614FFDCFA9D" authorityName="Kritsky & Boeger" authorityYear="2002" baseAuthorityName="Kritsky & Boeger" baseAuthorityYear="2002" box="[277,440,1325,1349]" class="Monogenea" family="Ancyrocephalidae" genus="Haliotrema" kingdom="Animalia" order="Dactylogyridea" pageId="25" pageNumber="32" phylum="Platyhelminthes" rank="species" species="longibaculum" status="comb. nov.">
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<emphasis id="4B88EADCDF74FF98FEBCF614FFDCFA9D" box="[277,440,1325,1349]" italics="true" pageId="25" pageNumber="32">E. longibaculum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="50BB57A7DF74FF98FE69F614FC79FA9D" box="[448,541,1325,1349]" pageId="25" pageNumber="32" rank="species">n. comb.</taxonomicNameLabel>
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was collected from
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<taxonomicName id="BEFC4D4DDF74FF98FF64F67FFF20FA87" baseAuthorityName="Cuvier" baseAuthorityYear="1828" box="[205,324,1350,1375]" class="Actinopterygii" family="Lutjanidae" genus="Lutjanus" kingdom="Animalia" order="Perciformes" pageId="25" pageNumber="32" phylum="Chordata" rank="species" species="mahogoni">
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<emphasis id="4B88EADCDF74FF98FF64F67FFF20FA87" box="[205,324,1350,1375]" italics="true" pageId="25" pageNumber="32">L. mahogoni</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="31E66545DF74FF98FF3EF6BEFC7CF9DA" pageId="25" pageNumber="32" type="description">
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<paragraph id="794336CEDF74FF98FF3EF6BEFF21FA47" blockId="25.[151,684,1415,1538]" box="[151,325,1415,1441]" pageId="25" pageNumber="32">MEASUREMENTS</paragraph>
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<paragraph id="794336CEDF74FF98FF3EF691FC7CF9DA" blockId="25.[151,684,1415,1538]" pageId="25" pageNumber="32">Ventral anchor 27 (25-28; n = 2) long, base 13 (12-14; n = 2) wide; dorsal anchor 45 (43-46; n = 2) long, base 9-10 (n = 2) wide.</paragraph>
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</subSubSection>
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<subSubSection id="31E66545DF74FF98FF3EF51EFA4BFB9A" pageId="25" pageNumber="32" type="discussion">
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<paragraph id="794336CEDF74FF98FF3EF51EFE9FF9E7" blockId="25.[151,684,1575,1666]" box="[151,251,1575,1601]" pageId="25" pageNumber="32">REMARKS</paragraph>
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<paragraph id="794336CEDF74FF98FF3EF571FA4BFB9A" blockId="25.[151,684,1575,1666]" lastBlockId="25.[714,1248,167,1090]" pageId="25" pageNumber="32">
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The original description of this species as
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<taxonomicName id="BEFC4D4DDF74FF98FDCEF571FF0FF95A" authorityName="Kritsky & Boeger" authorityYear="2002" baseAuthorityName="Kritsky & Boeger" baseAuthorityYear="2002" class="Monogenea" family="Ancyrocephalidae" genus="Haliotrema" kingdom="Animalia" order="Dactylogyridea" pageId="25" pageNumber="32" phylum="Platyhelminthes" rank="species" species="longibaculum">
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<emphasis id="4B88EADCDF74FF98FDCEF571FF0FF95A" italics="true" pageId="25" pageNumber="32">Haliotrema longibaculum</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="1D6D4B3FDF74FF98FE31F551FC26F95A" author="Zhukov" box="[408,578,1640,1666]" firstAuthor="Zhukov" pageId="25" pageNumber="32" pagination="33 - 47" refId="ref19640" refString="ZHUKOV E. V. 1976. - [New monogenean species of the genus Haliotrema Johnston and Tiegs, 1922, from the Gulf of Mexico fishes of the Fam. Lutianidae], in Fauna, systematics and phylogeny of Monogenoidea. Proceedings, Institute of Biology and Pedology, Far-East Science Centre, Academy of Sciences of the U. S. S. R., New Series 35: 33 - 47 (in Russian)." type="book chapter" year="1976">Zhukov (1976)</bibRefCitation>
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indicates that the body is fusiform and the vaginal aperture is dextroventral in the body, while the direction of the coil of the copulatory organ is uncertain. Although not a part of the
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<typeStatus id="A647886CDF74FF98FC5EF23EFA41FEF9" box="[1015,1061,263,289]" pageId="25" pageNumber="32">type</typeStatus>
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series, the specimen of
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<taxonomicName id="BEFC4D4DDF74FF98FC8BF211FDB2FE9A" authorityName="Kritsky & Boeger" authorityYear="2002" baseAuthorityName="Kritsky & Boeger" baseAuthorityYear="2002" box="[802,982,296,322]" class="Monogenea" family="Ancyrocephalidae" genus="Haliotrema" kingdom="Animalia" order="Dactylogyridea" pageId="25" pageNumber="32" phylum="Platyhelminthes" rank="species" species="longibaculum">
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<emphasis id="4B88EADCDF74FF98FC8BF211FDB2FE9A" box="[802,982,296,322]" italics="true" pageId="25" pageNumber="32">H. longibaculum</emphasis>
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</taxonomicName>
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available to us was collected by Dr Zhukov and was apparently used to complement his description of the species (Timofeeva pers. comm.). The unstained specimen is severely flattened and damaged, and it was not possible to determine many diagnostic features of internal anatomy and of the sclerotized parts of the haptor and copulatory complex. Nonetheless, this species is clearly a member of
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<taxonomicName id="BEFC4D4DDF74FF98FD63F171FD0FFDBA" authority="Kritsky & Boeger, 2002" authorityName="Kritsky & Boeger" authorityYear="2002" box="[714,875,584,610]" class="Monogenea" family="Ancyrocephalidae" genus="Euryhaliotrema" kingdom="Animalia" order="Dactylogyridea" pageId="25" pageNumber="32" phylum="Platyhelminthes" rank="genus" status="gen. nov.">
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<emphasis id="4B88EADCDF74FF98FD63F171FD0FFDBA" box="[714,875,584,610]" italics="true" pageId="25" pageNumber="32">Euryhaliotrema</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="50BB57A7DF74FF98FCDDF17EFDA4FDB9" box="[884,960,583,609]" pageId="25" pageNumber="32" rank="genus">n. gen.</taxonomicNameLabel>
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as shown by the presence of a bulbous base of the copulatory organ.
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<bibRefCitation id="1D6D4B3FDF74FF98FD63F1B1FD1EFD7A" author="Zhukov" box="[714,890,648,674]" firstAuthor="Zhukov" pageId="25" pageNumber="32" pagination="33 - 47" refId="ref19640" refString="ZHUKOV E. V. 1976. - [New monogenean species of the genus Haliotrema Johnston and Tiegs, 1922, from the Gulf of Mexico fishes of the Fam. Lutianidae], in Fauna, systematics and phylogeny of Monogenoidea. Proceedings, Institute of Biology and Pedology, Far-East Science Centre, Academy of Sciences of the U. S. S. R., New Series 35: 33 - 47 (in Russian)." type="book chapter" year="1976">Zhukov (1976)</bibRefCitation>
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shows the bulbous base of the copulatory organ in his drawing of the copulatory complex but the bulb was not illustrated in his drawing of the terminal reproductive genitalia (
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<bibRefCitation id="1D6D4B3FDF74FF98FD78F031FD0FFCFA" author="Zhukov" box="[721,875,776,802]" firstAuthor="Zhukov" pageId="25" pageNumber="32" pagination="33 - 47" refId="ref19640" refString="ZHUKOV E. V. 1976. - [New monogenean species of the genus Haliotrema Johnston and Tiegs, 1922, from the Gulf of Mexico fishes of the Fam. Lutianidae], in Fauna, systematics and phylogeny of Monogenoidea. Proceedings, Institute of Biology and Pedology, Far-East Science Centre, Academy of Sciences of the U. S. S. R., New Series 35: 33 - 47 (in Russian)." type="book chapter" year="1976">Zhukov 1976</bibRefCitation>
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: fig. 6). Thus,
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<taxonomicName id="BEFC4D4DDF74FF98FBA4F031FD5DFC9A" authorityName="Kritsky & Boeger" authorityYear="2002" baseAuthorityName="Zhukov" baseAuthorityYear="1976" class="Monogenea" family="Ancyrocephalidae" genus="Euryhaliotrema" kingdom="Animalia" order="Dactylogyridea" pageId="25" pageNumber="32" phylum="Platyhelminthes" rank="species" species="longibaculum" status="comb. nov.">
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<emphasis id="4B88EADCDF74FF98FBA4F031FD5DFC9A" italics="true" pageId="25" pageNumber="32">Euryhaliotrema longibaculum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="50BB57A7DF74FF98FCEBF01EFDC3FC9A" box="[834,935,807,834]" pageId="25" pageNumber="32" rank="species">n. comb.</taxonomicNameLabel>
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is proposed. It differs from all other described species of
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<taxonomicName id="BEFC4D4DDF74FF98FB84F071FABAFCBA" authority="Kritsky & Boeger, 2002" authorityName="Kritsky & Boeger" authorityYear="2002" box="[1069,1246,840,866]" class="Monogenea" family="Ancyrocephalidae" genus="Euryhaliotrema" kingdom="Animalia" order="Dactylogyridea" pageId="25" pageNumber="32" phylum="Platyhelminthes" rank="genus" status="gen. nov.">
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<emphasis id="4B88EADCDF74FF98FB84F071FABAFCBA" box="[1069,1246,840,866]" italics="true" pageId="25" pageNumber="32">Euryhaliotrema</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="50BB57A7DF74FF98FD63F05EFD70FC59" box="[714,788,871,897]" pageId="25" pageNumber="32" rank="genus">n. gen.</taxonomicNameLabel>
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in having a comparatively broad copulatory tube comprising less than one ring. The accessory piece is articulated to the base of the copulatory organ by a straight stout articulation process. The ventral anchor possesses a narrow base with an elongate superficial root, while the deep root is short to nonexistent.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |