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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1126.90233" ID-Pensoft-Pub="1313-2970-1126-55" ID-Pensoft-UUID="22EA20E1A9255729AAA61C4099DDD686" ID-ZooBank="0ABE7C8DBD9C44ED89D3CACB78D12AB9" ModsDocID="1313-2970-1126-55" checkinTime="1667357868610" checkinUser="pensoft" docAuthor="Kundrata, Robin, Hoffmannova, Johana, Hinson, Kevin R., Keller, Oliver &amp; Packova, Gabriela" docDate="2022" docId="EAF9466D67F3565B8A0624E3B422EDC4" docLanguage="en" docName="ZooKeys 1126: 55-130" docOrigin="ZooKeys 1126" docPubDate="2022-11-01" docSource="http://dx.doi.org/10.3897/zookeys.1126.90233" docTitle="Haplocladon Gorham 1883" docType="treatment" docVersion="1" id="22EA20E1A9255729AAA61C4099DDD686" lastPageNumber="55" masterDocId="22EA20E1A9255729AAA61C4099DDD686" masterDocTitle="Rhagophthalmidae Olivier, 1907 (Coleoptera, Elateroidea): described genera and species, current problems, and prospects for the bioluminescent and paedomorphic beetle lineage" masterLastPageNumber="130" masterPageNumber="55" pageNumber="55" updateTime="1667357868610" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Rhagophthalmidae Olivier, 1907 (Coleoptera, Elateroidea): described genera and species, current problems, and prospects for the bioluminescent and paedomorphic beetle lineage</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kundrata, Robin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9397-1030</mods:nameIdentifier>
<mods:affiliation>Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, 77900, Olomouc, Czech Republic</mods:affiliation>
<mods:nameIdentifier type="email">robin.kundrata@upol.cz</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hoffmannova, Johana</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0216-6031</mods:nameIdentifier>
<mods:affiliation>Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, 77900, Olomouc, Czech Republic</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hinson, Kevin R.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3111-4513</mods:nameIdentifier>
<mods:affiliation>EpiLogic GmbH Agrarbiologische Forschung und Beratung, Hohenbachernstr. 19 - 21, 85354, Freising, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Keller, Oliver</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5067-3316</mods:nameIdentifier>
<mods:affiliation>Florida State Collection of Arthropods, Florida Department of Agriculture and Consumer Services, P. O. Box 147100, Gainesville, FL, 32614 - 7100, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Packova, Gabriela</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7949-619X</mods:nameIdentifier>
<mods:affiliation>Department of Zoology, Faculty of Science, Palacky University, 17. listopadu 50, 77900, Olomouc, Czech Republic</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1126</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>55</mods:start>
<mods:end>130</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1126.90233</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1126.90233</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1126-55</mods:identifier>
<mods:identifier type="ZooBank">0ABE7C8DBD9C44ED89D3CACB78D12AB9</mods:identifier>
<mods:identifier type="Pensoft-UUID">22EA20E1A9255729AAA61C4099DDD686</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:EAF9466D67F3565B8A0624E3B422EDC4" httpUri="http://treatment.plazi.org/id/EAF9466D67F3565B8A0624E3B422EDC4" lastPageNumber="55" pageId="0" pageNumber="55">
<subSubSection pageId="0" pageNumber="55" type="nomenclature">
<paragraph pageId="0" pageNumber="55">
Genus
<taxonomicName LSID="EAF9466D-67F3-565B-8A06-24E3B422EDC4" authority="Gorham, 1883" authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Haplocladon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">Haplocladon Gorham, 1883</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="reference_group">
<paragraph pageId="0" pageNumber="55">
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Haplocladon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">Haplocladon</taxonomicName>
Gorham, 1883b: 249 [as a subgenus of
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Diplocladon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">Diplocladon</taxonomicName>
Gorham, 1883]. Gender: neuter. Type species.
<taxonomicName authorityName="Kundrata" authorityYear="2022" class="Insecta" family="Rhagophthalmidae" genus="Haplocladon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haplocladon gorhami" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="gorhami">Haplocladon gorhami</taxonomicName>
Kundrata, 2022, nom. nov. [replacement name for
<taxonomicName authorityName="Kundrata &amp; Hoffmannova &amp; Hinson &amp; Keller &amp; Packova" authorityYear="2022" class="Insecta" family="Rhagophthalmidae" genus="Diplocladon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocladon hasseltii" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="hasseltii">Diplocladon hasseltii</taxonomicName>
Gorham, 1883b]; by monotypy.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="literature">
<paragraph pageId="0" pageNumber="55">Literature.</paragraph>
<paragraph pageId="0" pageNumber="55">
<bibRefCitation author="Gorham, HS" journalOrPublisher="Notes from the Leyden Museum" pageId="0" pageNumber="55" pagination="249 - 250" refId="B87" refString="Gorham, HS, 1883b. A new subgenus of the coleopterous family Drilidae. Notes from the Leyden Museum 5: 249 - 250" title="A new subgenus of the coleopterous family Drilidae." volume="5" year="1883 b">Gorham (1883b</bibRefCitation>
: 249): original description;
<bibRefCitation author="Gorham, HS" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="55" pagination="293 - 330" refId="B91" refString="Gorham, HS, 1895. List of the Coleoptera in the collection of H. E. Andrewes, Esq., from India and Burma, with descriptions of new species and notes. Families: Malacodermata, Erotylidae, Endomychidae. Annales de la Societe Entomologique de Belgique 39: 293 - 330" title="List of the Coleoptera in the collection of H. E. Andrewes, Esq., from India and Burma, with descriptions of new species and notes. Families: Malacodermata, Erotylidae, Endomychidae." volume="39" year="1895">Gorham (1895</bibRefCitation>
: 310): remark;
<bibRefCitation author="Gorham, HS" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="55" pagination="323 - 330" refId="B92" refString="Gorham, HS, 1903. On Coleoptera collected in India by MM. H.-E. and H.-L. Andrewes. Families Malacodermata, Erotylidae, Endomychidae and Coccinellidae. Annales de la Societe Entomologique de Belgique 47: 323 - 330" title="On Coleoptera collected in India by MM. H. - E. and H. - L. Andrewes. Families Malacodermata, Erotylidae, Endomychidae and Coccinellidae." volume="47" year="1903">Gorham (1903</bibRefCitation>
: 330): species description;
<bibRefCitation DOI="https://doi.org/10.1007/978-94-011-9697-0_1" author="Olivier, E" editor="Junk, W" journalOrPublisher="W. Junk, Berlin, Germany" pageId="0" pageNumber="55" pagination="1 - 10" refId="B220" refString="Olivier, E, 1910. Rhagophthalmidae, Drilidae. In: Junk, W, Schenkling, S, Eds., Coleopterorum Catalogus, pars 10. W. Junk, Berlin, Germany: 1 - 10, DOI: https://doi.org/10.1007/978-94-011-9697-0_1" title="Rhagophthalmidae, Drilidae." url="https://doi.org/10.1007/978-94-011-9697-0_1" volumeTitle="Coleopterorum Catalogus, pars 10." year="1910">Olivier (1910</bibRefCitation>
: 8): catalogue;
<bibRefCitation author="Wittmer, W" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="55" refId="B294" refString="Wittmer, W, 1944. Catalogue des Drilidae E. Oliv. (Coleoptera-Malacodermata). Revista de la Sociedad Entomologica Argentina XII: 203-221." title="Catalogue des Drilidae E. Oliv. (Coleoptera-Malacodermata). Revista de la Sociedad Entomologica Argentina XII: 203 - 221." year="1944">Wittmer (1944</bibRefCitation>
: 211): catalogue;
<bibRefCitation author="Crowson, RA" journalOrPublisher=", Ltd, London" pageId="0" pageNumber="55" refId="B54" refString="Crowson, RA, 1955. The natural classification of the families of Coleoptera. Nathaniel Lloyd &amp; Co., Ltd, London" title="The natural classification of the families of Coleoptera. Nathaniel Lloyd &amp; Co." year="1955">Crowson (1955</bibRefCitation>
: 68): remark;
<bibRefCitation author="Paulus, HF" journalOrPublisher="Senckenbergiana Biologica" pageId="0" pageNumber="55" pagination="37 - 54" refId="B225" refString="Paulus, HF, 1972. Die systematische und phylogenetische Stellung der Karumiidae, mit einer beschreibung von Escalerina serraticornis n.sp. aus S-Persien. Senckenbergiana Biologica 53: 37 - 54" title="Die systematische und phylogenetische Stellung der Karumiidae, mit einer beschreibung von Escalerina serraticornis n. sp. aus S-Persien." volume="53" year="1972">Paulus (1972</bibRefCitation>
: 49): remark;
<bibRefCitation DOI="https://doi.org/10.3157/0013-872X(2008)119[109:AGBSCR]2.0.CO;2" author="Li, X" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="55" refId="B165" refString="Li, X, Liang, X, 2008. A gigantic bioluminescent starworm (Coleoptera: Rhagophthalmidae) from Northwest Yunnan, China. Entomological News 119(2): 109-112. https://doi.org/10.3157/0013-872X(2008)119[109:AGBSCR]2.0.CO;2" title="A gigantic bioluminescent starworm (Coleoptera: Rhagophthalmidae) from Northwest Yunnan, China. Entomological News 119 (2): 109 - 112." url="https://doi.org/10.3157/0013-872X(2008)119[109:AGBSCR]2.0.CO;2" year="2008">Li and Liang (2008</bibRefCitation>
: 109): remark [as
<taxonomicName family="Rhagophthalmidae" kingdom="Animalia" lsidName="D. haplocladon" order="Coleoptera" pageId="0" pageNumber="55" rank="species" species="haplocladon">
<emphasis italics="true" pageId="0" pageNumber="55">D. haplocladon</emphasis>
</taxonomicName>
[sic!]]. In addition to the aforementioned literature, this genus was included in a PhD thesis by
<bibRefCitation author="Jeng, M-L" journalOrPublisher="PhD Thesis, University of Kansas, Lawrence, Kansas, USA" pageId="0" pageNumber="55" refId="B121" refString="Jeng, M-L, 2008. Comprehensive phylogenetics, systematics, and evolution of neoteny of Lampyridae (Insecta: Coleoptera). PhD Thesis, University of Kansas, Lawrence, Kansas, USA" title="Comprehensive phylogenetics, systematics, and evolution of neoteny of Lampyridae (Insecta: Coleoptera)." year="2008">Jeng (2008)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="55" type="remarks">
<paragraph pageId="0" pageNumber="55">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="55">
<bibRefCitation author="Gorham, HS" journalOrPublisher="Notes from the Leyden Museum" pageId="0" pageNumber="55" pagination="249 - 250" refId="B87" refString="Gorham, HS, 1883b. A new subgenus of the coleopterous family Drilidae. Notes from the Leyden Museum 5: 249 - 250" title="A new subgenus of the coleopterous family Drilidae." volume="5" year="1883 b">Gorham (1883b)</bibRefCitation>
originally described
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Haplocladon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Haplocladon</emphasis>
</taxonomicName>
as a subgenus of
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Diplocladon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Diplocladon</emphasis>
</taxonomicName>
but later treated it as a separate genus (
<bibRefCitation author="Gorham, HS" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="55" pagination="293 - 330" refId="B91" refString="Gorham, HS, 1895. List of the Coleoptera in the collection of H. E. Andrewes, Esq., from India and Burma, with descriptions of new species and notes. Families: Malacodermata, Erotylidae, Endomychidae. Annales de la Societe Entomologique de Belgique 39: 293 - 330" title="List of the Coleoptera in the collection of H. E. Andrewes, Esq., from India and Burma, with descriptions of new species and notes. Families: Malacodermata, Erotylidae, Endomychidae." volume="39" year="1895">Gorham 1895</bibRefCitation>
,
<bibRefCitation author="Gorham, HS" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="55" pagination="323 - 330" refId="B92" refString="Gorham, HS, 1903. On Coleoptera collected in India by MM. H.-E. and H.-L. Andrewes. Families Malacodermata, Erotylidae, Endomychidae and Coccinellidae. Annales de la Societe Entomologique de Belgique 47: 323 - 330" title="On Coleoptera collected in India by MM. H. - E. and H. - L. Andrewes. Families Malacodermata, Erotylidae, Endomychidae and Coccinellidae." volume="47" year="1903">1903</bibRefCitation>
). Unfortunately, he named type species of both
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Diplocladon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Diplocladon</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Haplocladon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Haplocladon</emphasis>
</taxonomicName>
as &quot;
<emphasis italics="true" pageId="0" pageNumber="55">hasseltii</emphasis>
&quot; (
<bibRefCitation author="Gorham, HS" journalOrPublisher="Notes from the Leyden Museum" pageId="0" pageNumber="55" pagination="5 - 6" refId="B86" refString="Gorham, HS, 1883a. Description of a new genus and species of the coleopterous family Drilidae. Notes from the Leyden Museum 5: 5 - 6" title="Description of a new genus and species of the coleopterous family Drilidae." volume="5" year="1883 a">Gorham 1883a</bibRefCitation>
,
<bibRefCitation author="Gorham, HS" journalOrPublisher="Notes from the Leyden Museum" pageId="0" pageNumber="55" pagination="249 - 250" refId="B87" refString="Gorham, HS, 1883b. A new subgenus of the coleopterous family Drilidae. Notes from the Leyden Museum 5: 249 - 250" title="A new subgenus of the coleopterous family Drilidae." volume="5" year="1883 b">b</bibRefCitation>
), which probably confused some subsequent authors who treated
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Haplocladon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Haplocladon</emphasis>
</taxonomicName>
as a synonym of
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Diplocladon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Diplocladon</emphasis>
</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1007/978-94-011-9697-0_1" author="Olivier, E" editor="Junk, W" journalOrPublisher="W. Junk, Berlin, Germany" pageId="0" pageNumber="55" pagination="1 - 10" refId="B220" refString="Olivier, E, 1910. Rhagophthalmidae, Drilidae. In: Junk, W, Schenkling, S, Eds., Coleopterorum Catalogus, pars 10. W. Junk, Berlin, Germany: 1 - 10, DOI: https://doi.org/10.1007/978-94-011-9697-0_1" title="Rhagophthalmidae, Drilidae." url="https://doi.org/10.1007/978-94-011-9697-0_1" volumeTitle="Coleopterorum Catalogus, pars 10." year="1910">Olivier 1910</bibRefCitation>
;
<bibRefCitation author="Wittmer, W" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="55" refId="B294" refString="Wittmer, W, 1944. Catalogue des Drilidae E. Oliv. (Coleoptera-Malacodermata). Revista de la Sociedad Entomologica Argentina XII: 203-221." title="Catalogue des Drilidae E. Oliv. (Coleoptera-Malacodermata). Revista de la Sociedad Entomologica Argentina XII: 203 - 221." year="1944">Wittmer 1944</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3157/0013-872X(2008)119[109:AGBSCR]2.0.CO;2" author="Li, X" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="55" refId="B165" refString="Li, X, Liang, X, 2008. A gigantic bioluminescent starworm (Coleoptera: Rhagophthalmidae) from Northwest Yunnan, China. Entomological News 119(2): 109-112. https://doi.org/10.3157/0013-872X(2008)119[109:AGBSCR]2.0.CO;2" title="A gigantic bioluminescent starworm (Coleoptera: Rhagophthalmidae) from Northwest Yunnan, China. Entomological News 119 (2): 109 - 112." url="https://doi.org/10.3157/0013-872X(2008)119[109:AGBSCR]2.0.CO;2" year="2008">Li and Liang 2008</bibRefCitation>
).
<bibRefCitation author="Crowson, RA" journalOrPublisher=", Ltd, London" pageId="0" pageNumber="55" refId="B54" refString="Crowson, RA, 1955. The natural classification of the families of Coleoptera. Nathaniel Lloyd &amp; Co., Ltd, London" title="The natural classification of the families of Coleoptera. Nathaniel Lloyd &amp; Co." year="1955">Crowson (1955)</bibRefCitation>
and
<bibRefCitation author="Paulus, HF" journalOrPublisher="Senckenbergiana Biologica" pageId="0" pageNumber="55" pagination="37 - 54" refId="B225" refString="Paulus, HF, 1972. Die systematische und phylogenetische Stellung der Karumiidae, mit einer beschreibung von Escalerina serraticornis n.sp. aus S-Persien. Senckenbergiana Biologica 53: 37 - 54" title="Die systematische und phylogenetische Stellung der Karumiidae, mit einer beschreibung von Escalerina serraticornis n. sp. aus S-Persien." volume="53" year="1972">Paulus (1972)</bibRefCitation>
again considered
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Haplocladon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Haplocladon</emphasis>
</taxonomicName>
a separate genus. Since
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Haplocladon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Haplocladon</emphasis>
</taxonomicName>
differs at first sight from
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Diplocladon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Diplocladon</emphasis>
</taxonomicName>
by the unipectinate antennae (versus bipectinate in
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Diplocladon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Diplocladon</emphasis>
</taxonomicName>
), we prefer to keep
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Haplocladon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Haplocladon</emphasis>
</taxonomicName>
at a generic level. Because
<taxonomicName authorityName="Kundrata &amp; Hoffmannova &amp; Hinson &amp; Keller &amp; Packova" authorityYear="2022" class="Insecta" family="Rhagophthalmidae" genus="Diplocladon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocladon hasseltii" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="hasseltii">
<emphasis italics="true" pageId="0" pageNumber="55">Diplocladon hasseltii</emphasis>
</taxonomicName>
Gorham, 1883a and
<taxonomicName authorityName="Kundrata &amp; Hoffmannova &amp; Hinson &amp; Keller &amp; Packova" authorityYear="2022" class="Insecta" family="Rhagophthalmidae" genus="Diplocladon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocladon hasseltii" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="hasseltii">
<emphasis italics="true" pageId="0" pageNumber="55">Diplocladon hasseltii</emphasis>
</taxonomicName>
Gorham, 1883b (described in subgenus
<taxonomicName authorityName="Kundrata &amp; Hoffmannova &amp; Hinson &amp; Keller &amp; Packova" authorityYear="2022" class="Insecta" family="Rhagophthalmidae" genus="Diplocladon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocladon (Haplocladon)" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="subGenus" subGenus="Haplocladon">Diplocladon Haplocladon</taxonomicName>
) are primary homonyms, the latter junior name is permanently invalid (Art. 57.2 of the Code;
<bibRefCitation author="Hunt, T" journalOrPublisher="Science" pageId="0" pageNumber="55" refId="B117" refString="ICZN [International Commission on Zoological Nomenclature] (1999) International Code of Zoological Nomenclature. Fourth Edition.The International Trust for Zoological Nomenclature, London, 305 pp." year="2007">ICZN 1999</bibRefCitation>
) and should be replaced by a new name (see below). Currently,
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Haplocladon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Haplocladon</emphasis>
</taxonomicName>
contains two species, one from Indonesia and one from southern India. Based on the available figures, specimens reported by
<bibRefCitation author="Haneda, Y" journalOrPublisher="Shin Konchu" pageId="0" pageNumber="55" pagination="2 - 5" refId="B99" refString="Haneda, Y, 1950. Star-worm in Singapore. Shin Konchu 3: 2 - 5" title="Star-worm in Singapore." volume="3" year="1950">Haneda (1950)</bibRefCitation>
from Singapore and identified as
<taxonomicName authorityName="Kundrata &amp; Hoffmannova &amp; Hinson &amp; Keller &amp; Packova" authorityYear="2022" class="Insecta" family="Rhagophthalmidae" genus="Diplocladon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplocladon hasseltii" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="species" species="hasseltii">
<emphasis italics="true" pageId="0" pageNumber="55">Diplocladon hasseltii</emphasis>
</taxonomicName>
, which were later mentioned by other authors (e.g.,
<bibRefCitation author="Harvey, EN" editor="Harvey, EN" journalOrPublisher="Academic Press, New York" pageId="0" pageNumber="55" pagination="389 - 471" refId="B102" refString="Harvey, EN, 1952. Coleoptera. In: Harvey, EN, Ed., Bioluminescence. Academic Press, New York: 389 - 471" title="Coleoptera." volumeTitle="Bioluminescence." year="1952">Harvey 1952</bibRefCitation>
;
<bibRefCitation author="Haneda, Y" editor="Johnson, FH" journalOrPublisher="AAAS, Washington DC" pageId="0" pageNumber="55" pagination="335 - 385" refId="B100" refString="Haneda, Y, 1955. Luminous organisms of Japan and the Far East. In: Johnson, FH, Ed., The luminescense of biological systems. AAAS, Washington DC: 335 - 385" title="Luminous organisms of Japan and the Far East." volumeTitle="The luminescense of biological systems." year="1955">Haneda 1955</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1146/annurev.en.16.010171.000525" author="Lloyd, JE" journalOrPublisher="Annual Review of Entomology" pageId="0" pageNumber="55" pagination="97 - 122" refId="B175" refString="Lloyd, JE, 1971. Bioluminescent communication in insects. Annual Review of Entomology 16 (1): 97 - 122, DOI: https://doi.org/10.1146/annurev.en.16.010171.000525" title="Bioluminescent communication in insects." url="https://doi.org/10.1146/annurev.en.16.010171.000525" volume="16" year="1971">Lloyd 1971</bibRefCitation>
;
<bibRefCitation author="Lloyd, JE" editor="Herring, PJ" journalOrPublisher="Academic Press, London" pageId="0" pageNumber="55" pagination="241 - 272" refId="B176" refString="Lloyd, JE, 1978. Insect Bioluminescence. Chap. 8. In: Herring, PJ, Ed., Bioluminescence in action. Academic Press, London: 241 - 272" title="Insect Bioluminescence. Chap. 8." volumeTitle="Bioluminescence in action." year="1978">Lloyd 1978</bibRefCitation>
;
<bibRefCitation author="Crowson, RA" journalOrPublisher="Academic Press, London" pageId="0" pageNumber="55" refId="B56" refString="Crowson, RA, 1981. The Biology of the Coleoptera. Academic Press, London" title="The Biology of the Coleoptera." year="1981">Crowson 1981</bibRefCitation>
;
<bibRefCitation author="Haneda, Y" journalOrPublisher="Kōseisha Kōseikaku, Tokyo, Japan" pageId="0" pageNumber="55" refId="B101" refString="Haneda, Y, 1985. Luminous organisms. Kōseisha Kōseikaku, Tokyo, Japan" title="Luminous organisms." year="1985">Haneda 1985</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1515/9783110911213.135" author="Kawashima, I" editor="Leschen, RAB" journalOrPublisher="In: Kristensen NP, Beutel RG (Eds) Handbook of Zoology, Arthropoda: Insecta. Walter de Gruyter GmbH and Co KG, Berlin / New York" pageId="0" pageNumber="55" pagination="135 - 140" refId="B133" refString="Kawashima, I, Lawrence, JF, Branham, MA, 2010. Rhagophthalmidae Olivier, 1907. In: Leschen, RAB, Beutel, RG, Lawrence, JF, Eds., Coleoptera, Beetles; Volume 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen NP, Beutel RG (Eds) Handbook of Zoology, Arthropoda: Insecta. Walter de Gruyter GmbH and Co KG, Berlin/New York: 135 - 140, DOI: https://doi.org/10.1515/9783110911213.135" title="Rhagophthalmidae Olivier, 1907." url="https://doi.org/10.1515/9783110911213.135" volumeTitle="Coleoptera, Beetles; Volume 2: Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim)." year="2010">Kawashima et al. 2010</bibRefCitation>
), are probably members of
<taxonomicName authorityName="Gorham" authorityYear="1883" class="Insecta" family="Rhagophthalmidae" genus="Haplocladon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Haplocladon" order="Coleoptera" pageId="0" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="55">Haplocladon</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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