340 lines
32 KiB
XML
340 lines
32 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="3B1B110789ED40C81DC18F1FADE2C979" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole fitarata Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="8CF00FF0-5B19-4EA6-803E-27F62D848627" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>905</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>235</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161777004" LSID="urn:lsid:zoobank.org:act:8CF00FF0-5B19-4EA6-803E-27F62D848627" httpUri="http://treatment.plazi.org/id/3B1B110789ED40C81DC18F1FADE2C979" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/8CF00FF0-5B19-4EA6-803E-27F62D848627" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fitarata" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fitarata" status="sp. nov.">Pheidole fitarata</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">Figs 46A-F</figureCitation>
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<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84S</figureCitation>
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<figureCitation captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86S</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Antsiranana; Ampasindava,
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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<normalizedToken originalValue="d’Ambilanivy">d'Ambilanivy</normalizedToken>
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, 3.9 km 181°S Ambaliha;
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<geoCoordinate degrees="13.79861" direction="south" orientation="latitude" precision="1" value="-13.79861">-13.79861</geoCoordinate>
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,
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<geoCoordinate degrees="48.16167" direction="east" orientation="longitude" precision="1" value="48.16167">48.16167</geoCoordinate>
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; alt. 600 m; 4 Mar 2001; Fisher et al. leg.; BLF03293, CASENT0420028 (CASC).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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.
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</emphasis>
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Madagascar. •4w., 1s.; same data as for holotype; CASENT0420024-CASENT0420027, CASENT0872215-CASENT0872217 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
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: •1s.; Ambondrobe, 41.1km 175° Vohemar;
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<geoCoordinate degrees="13.71533" direction="south" orientation="latitude" precision="1" value="-13.71533">-13.71533</geoCoordinate>
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,
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<geoCoordinate degrees="50.10167" direction="east" orientation="longitude" precision="1" value="50.10167">50.10167</geoCoordinate>
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; alt. 10 m; 29 Nov 2004; B.L. Fisher leg.; CASENT0056078 (CASC). •1w., 4s.; Ampasindava,
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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<normalizedToken originalValue="d’Ambilanivy">d'Ambilanivy</normalizedToken>
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, 3.9 km 181°S Ambaliha;
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<geoCoordinate degrees="13.79861" direction="south" orientation="latitude" precision="1" value="-13.79861">-13.79861</geoCoordinate>
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,
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<geoCoordinate degrees="48.16167" direction="east" orientation="longitude" precision="1" value="48.16167">48.16167</geoCoordinate>
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; alt. 600 m; 4 Mar 2001; B.L. Fisher et al. leg.; CASENT0406700, CASENT0420217, CASENT0420226, CASENT0464109, CASENT0464294 (CASC). •2s.;
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<normalizedToken originalValue="Forêt">Foret</normalizedToken>
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<normalizedToken originalValue="d’">d'</normalizedToken>
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Andavakoera, 21.4 km 75°ENE Ambilobe; 4.6 km 356°N Betsiaka; -13.11833, 49.23; alt. 425 m; 15 Dec 2003; B.L. Fisher leg.; CASENT0044135, CASENT0044209 (CASC). •1s.; Montagne des
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<normalizedToken originalValue="Français">Francais</normalizedToken>
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, 7.2 km 142°SE Antsiranana (=Diego Suarez);
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<geoCoordinate degrees="12.32278" direction="south" orientation="latitude" precision="1" value="-12.32278">-12.32278</geoCoordinate>
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,
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<geoCoordinate degrees="49.33817" direction="east" orientation="longitude" precision="1" value="49.33817">49.33817</geoCoordinate>
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; alt. 180 m; 22 Feb 2001; B.L. Fisher et al. leg.; CASENT0461044 (CASC). •1s.; Nosy Be,
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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Naturelle
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<normalizedToken originalValue="Intégrale">Integrale</normalizedToken>
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de Lokobe, 6.3 km 112°ESE Hellville;
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<geoCoordinate degrees="13.41933" direction="south" orientation="latitude" precision="1" value="-13.41933">-13.41933</geoCoordinate>
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,
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<geoCoordinate degrees="48.33117" direction="east" orientation="longitude" precision="1" value="48.33117">48.33117</geoCoordinate>
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; alt. 30 m; 19 Mar 2001; B.L. Fisher et al. leg.; CASENT0462968 (CASC). •1s.; R.S. Manongarivo, 12.8 km 228°SW Antanambao;
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<geoCoordinate degrees="13.97667" direction="south" orientation="latitude" precision="1" value="-13.97667">-13.97667</geoCoordinate>
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,
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<geoCoordinate degrees="48.42333" direction="east" orientation="longitude" precision="1" value="48.42333">48.42333</geoCoordinate>
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; alt. 780 m; 11 Oct 1998; B.L. Fisher leg.; CASENT0198865 (CASC). •3s.;
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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<normalizedToken originalValue="d’Ambre">d'Ambre</normalizedToken>
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, 3.5 km 235°SW Sakaramy;
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<geoCoordinate degrees="12.46889" direction="south" orientation="latitude" precision="1" value="-12.46889">-12.46889</geoCoordinate>
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,
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<geoCoordinate degrees="49.24217" direction="east" orientation="longitude" precision="1" value="49.24217">49.24217</geoCoordinate>
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; alt. 325 m; 26 Jan 2001; B.L. Fisher et al. leg.; CASENT0406576, CASENT0423829, CASENT0423892 (CASC). •1w., 1s.;
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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de
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<normalizedToken originalValue="l’Ankarana">l'Ankarana</normalizedToken>
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, 13.6 km 192°SSW Anivorano Nord;
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<geoCoordinate degrees="12.86361" direction="south" orientation="latitude" precision="1" value="-12.86361">-12.86361</geoCoordinate>
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,
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<geoCoordinate degrees="49.22583" direction="east" orientation="longitude" precision="1" value="49.22583">49.22583</geoCoordinate>
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; alt. 210 m; 16 Feb 2001; B.L. Fisher et al. leg.; CASENT0420164, CASENT0420181 (CASC). •7s., 1q.;
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<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
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<normalizedToken originalValue="Spéciale">Speciale</normalizedToken>
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de
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<normalizedToken originalValue="l’Ankarana">l'Ankarana</normalizedToken>
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, 22.9 km 224°SW Anivorano Nord;
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<geoCoordinate degrees="12.90889" direction="south" orientation="latitude" precision="1" value="-12.90889">-12.90889</geoCoordinate>
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,
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<geoCoordinate degrees="49.10983" direction="east" orientation="longitude" precision="1" value="49.10983">49.10983</geoCoordinate>
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; alt. 80 m; 10 Feb 2001; B.L. Fisher et al. leg.; CASENT0420207, CASENT0439168, CASENT0439173, CASENT0439183, CASENT0439197, CASENT0439313, CASENT0439315, CASENT0008034 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
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.
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</emphasis>
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Head in full-face view elongated; sides of the head with moderately dense, long, erect pilosity; occipital lobes smooth to indistinctly foveolate, with thick, sparse, short, transverse, and sometimes arcuate rugae; inner hypostomal teeth distinct, high, closely spaced, triangular, with rounded apex directed inward; outer hypostomal teeth thinner than inner hypostomal teeth and approximately the same height, triangular.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Head foveolate, genae smooth; scape, when laid back, surpassing the posterior head margin by two-fifths of its length; promesonotum low, convex, short, with posterior declivity relatively convex; mesosoma foveolate, katepisternum with smooth notch; propodeal spines short, triangular.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 10): HL: 1.13-1.43 (1.29); HW: 0.91-1.12 (1.07); SL: 0.43-0.53 (0.49); EL: 0.12-0.17 (0.14); WL: 0.79-1.02 (0.92); PSL: 0.16-0.21 (0.18); MTL: 0.51-0.59 (0.55); PNW: 0.45-0.64 (0.56); PTW: 0.11-0.19 (0.16); PPW: 0.29-0.47 (0.41); CI: 77.9-88.4 (82.7); SI: 43.5-49.0 (46.5); PSLI: 12.7-15.3 (14.0); PPI: 34.8-42.9 (38.2); PNI: 49.1-56.7 (52.8); MTI: 48.4 -56.1 (52.0).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view longitudinal, longer than wide, anterior of eyes straight, posterior of eyes slightly convex, occipital margin of lobes straight, inclining toward centre (Fig.
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<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46B</figureCitation>
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). In lateral view sub-rectangular; ventral and dorsal faces finely convex; dorsal face finely depressed posteriorly, forming indistinct transverse depression between frons and occipital lobes; inner hypostomal teeth visible. Sides of the head with moderately dense, long, erect pilosity; whole head with dense, moderately long, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae; scrobe surface shiny, foveolate, with thick, longitudinal, short, and interrupted rugae. Occipital lobes shiny, smooth to indistinctly foveolate, with thick, sparse, short, transverse, and sometimes arcuate rugae; frons with moderately dense, thick, longitudinal and interrupted rugae, interspaces smooth and indistinctly foveolate on the posterior part, rugae weakening posteriorly and never connected with rugae on the occipital lobes; genae shiny and smooth; malar area with thick, longitudinal, moderately sparse rugae, interspaces smooth. Centre of clypeus shiny and smooth, lateral sides with longitudinal rugae; median notch present, wide, and shallow; median longitudinal carina absent; lateral longitudinal carinae absent. Scape, when laid back, not reaching the mid-length of head; pilosity suberect to erect (Fig.
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<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46B, D</figureCitation>
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). Inner hypostomal teeth distinct, high, closely spaced, triangular, with rounded apex directed inward; outer hypostomal teeth thinner than inner hypostomal teeth and approximately the same height, triangular (Fig.
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<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84S</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum relatively low and arched, posterior mesonotum relatively steep, with tubercle-like projections; promesonotal groove absent; metanotal groove indistinct; propodeal spines moderately long, triangular, with sharp apex and wide base; humeral area laterally weakly produced (Fig.
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<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46D</figureCitation>
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). Surface finely foveolate, dorsal surface of promesonotum with fading sculpture and sometimes with smooth notch; katepisternum with smooth notch; lower half of lateral sides of propodeum with additional longitudinal rugae. Pilosity sparse, very long, and erect (Fig.
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<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny; peduncle moderately long, finely foveolate, with indistinct horizontal lobes on its basal part; node smooth, relatively high, triangular, with rounded apex, in rear view node slightly convex; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46D, F</figureCitation>
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||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Shiny and finely shagreened, dorsum with fading sculpture; in dorsal view sides with moderately short, acute, and triangular projections; pilosity long to moderately long, and erect (Fig.
|
||
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46D, F</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Shiny and finely shagreened; pilosity moderately dense, long, and erect (Fig.
|
||
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46D, F</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Bright brown to brown; legs, lower frons, malar area and lower parts of lateral sides of mesosoma yellowish brown (Fig.
|
||
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46D, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1" start="Figure 46" startId="F46">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 46.</emphasis>
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fitarata" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fitarata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole fitarata</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., full-face view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
), profile (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
), and dorsal view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
) of paratype minor worker (CASENT0420025) and full-face view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
), profile (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
), and dorsal view (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
) of holotype major worker (CASENT0420028).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
|
||
Measurements (
|
||
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
||
= 6): HL: 0.45-0.47 (0.46); HW: 0.39-0.41 (0.4); SL: 0.44-0.46 (0.45); EL: 0.07-0.1 (0.08); WL: 0.52-0.56 (0.55); PSL: 0.08-0.09 (0.08); MTL: 0.33-0.36 (0.35); PNW: 0.26-0.28 (0.27); PTW: 0.06-0.06 (0.06); PPW: 0.09-0.1 (0.1); CI: 85.9-89.5 (87.7); SI: 111.0-113.6 (111.6); PSLI: 17.3-20.4 (18.5); PPI: 61.0-65.2 (62.5); PNI: 66.4-68.8 (67.8); MTI: 82.8-89.4 (86.7).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
Occipital margin straight or indistinctly concave; occipital carina absent (Fig.
|
||
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46A</figureCitation>
|
||
). Pilosity moderately dense, long, suberect. Head shiny, foveolate; genae smooth. Clypeus smooth or finely foveolate; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by two-fifths of its length; pilosity suberect to erect (Fig.
|
||
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46A, C</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
||
.
|
||
</emphasis>
|
||
In lateral view, promesonotum low, convex, short, with posterior declivity relatively convex; promesonotal groove absent; metanotal groove indistinct; propodeal spines short, triangular, with acute apex (Fig.
|
||
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46C</figureCitation>
|
||
). Sculpture foveolate; katepisternum with smooth notch. Pilosity sparse, long, and erect (Fig.
|
||
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Peduncle short and thin with ventral face slightly convex; with few short, erect setae (Fig.
|
||
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
Short, low, and convex; sometimes with few short, erect setae (Fig.
|
||
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
||
.
|
||
</emphasis>
|
||
With moderately sparse, erect pilosity (Fig.
|
||
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46C, E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Yellow (Fig.
|
||
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Pheidole fitarata sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0420025) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0420028)." figureDoi="10.3897/zookeys.905.39592.figure46" httpUri="https://binary.pensoft.net/fig/373093" pageId="0" pageNumber="1">46C, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
||
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Malagasy for mirror, in reference to smooth and shiny genae of major workers.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
||
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">The species was collected between 10-780 m in elevation, in rainforest, in tropical dry forest, and in littoral rainforest. Nests were located rotten logs.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="comments">
|
||
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fitarata" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fitarata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole fitarata</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is most similar to
|
||
<taxonomicName lsidName="P. rugofitarata" pageId="0" pageNumber="1" rank="species" species="rugofitarata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. rugofitarata</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName lsidName="P. avaratra" pageId="0" pageNumber="1" rank="species" species="avaratra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. avaratra</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fitarata" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fitarata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole fitarata</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from
|
||
<taxonomicName lsidName="P. rugofitarata" pageId="0" pageNumber="1" rank="species" species="rugofitarata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. rugofitarata</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in less distinct and interrupted longitudinal rugae on frons and antennal scrobes, lack of connection between rugae on frons and occipital lobes, smooth genae, and never lobe-like outer hypostomal teeth; from
|
||
<taxonomicName lsidName="P. avaratra" pageId="0" pageNumber="1" rank="species" species="avaratra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. avaratra</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in presence of longitudinal, thick, and interrupted rugae on frons and antennal scrobes, frons with rugae weakening posteriorly and never connected with rugae on the occipital lobes, genae smooth, and smaller and directed inward inner hypostomal teeth.
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole fitarata" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fitarata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole fitarata</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from
|
||
<taxonomicName lsidName="P. avaratra" pageId="0" pageNumber="1" rank="species" species="avaratra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. avaratra</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in longer and lower promesonotum and katepisternum with smooth notch.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |